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<document id="E22A3CACD6209C9CF7879D35D59BE500" ID-DOI="10.11646/zootaxa.5370.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10147814" ID-ZooBank="48903495-7C6C-46E4-9B1B-D6A2F2781873" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1700202702953" checkinUser="plazi" docAuthor="Harzhauser, Mathias &amp; Landau, Bernard" docDate="2023" docId="03BEE17BFFF3C056FF0EFF657CE231ED" docLanguage="en" docName="zootaxa.5370.1.1.pdf" docOrigin="Zootaxa 5370 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5370.1.1/52270" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Mathilda margaritula O. Semper 1865" docType="treatment" docVersion="8" lastPageNumber="51" masterDocId="FF879903FFC1C064FF99FFF27C023738" masterDocTitle="The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition" masterLastPageNumber="74" masterPageNumber="1" pageNumber="51" updateTime="1706539845549" updateUser="ExternalLinkService">
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<mods:title id="C5B2DEF2E30BE83CCD59EF7CC2B2F43D">The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition</mods:title>
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<mods:affiliation id="3DEC33EFAA4E7B02A1123A34095D249D">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;</mods:affiliation>
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<mods:affiliation id="FE07A34D7B9C6439B066E92C923EACCF">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655; &amp; Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal. bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494 &amp; Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;</mods:affiliation>
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<taxonomicName id="4C172BEEFFF3C056FF0EFF657E4F3789" ID-CoL="BXSW9" authority="Semper, 1865" authorityName="O. Semper" authorityYear="1865" box="[151,589,151,178]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="margaritula">
<emphasis id="B9638C7FFFF3C056FF0EFF657E4F3789" bold="true" box="[151,589,151,178]" pageId="50" pageNumber="51">
<emphasis id="B9638C7FFFF3C056FF0EFF657D9C378A" bold="true" box="[151,414,151,178]" italics="true" pageId="50" pageNumber="51">Mathilda margaritula</emphasis>
Semper, 1865
</emphasis>
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<figureCitation id="132C4CE8FFF3C056FF0EFF497D1D37EE" box="[151,287,187,214]" captionStart="FIGURE 25" captionStartId="49.[151,250,1686,1710]" captionTargetBox="[213,1375,181,1662]" captionTargetId="figure-25@49.[213,1375,181,1662]" captionTargetPageId="49" captionText="FIGURE 25 A1A2. Mathilda gloriosa Boettger, 1907, holotype, SMF 372691, Coşteiu de Sus (Romania). B1B2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549a, Lăpugiu de Sus (Romania). C1C2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1876/0011/0098, Lăpugiu des Sus (Romania). D. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549b, Lăpugiu de Sus (Romania). E. Mathilda margaritula Semper, 1865, lecto- type, NHMW 1868/0001/0549, Lăpugiu de Sus (Romania). F1F2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549c, Lăpugiu de Sus (Romania). G. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549d, Lăpugiu de Sus (Romania). H1H2. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039a, Sedlec (Czech Republic). I1I2. Mathilda margaritula Semper, 1865, NHMW 1866/0001/0912, Niederleis (Austria). J. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039b, Sedlec (Czech Republic). K. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039c, Sedlec (Czech Republic)." figureDoi="http://doi.org/10.5281/zenodo.10147866" httpUri="https://zenodo.org/record/10147866/files/figure.png" pageId="50" pageNumber="51">Figs 25BK</figureCitation>
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<paragraph id="8BA8506DFFF3C056FF0EFEF27F67362F" blockId="50.[151,1436,255,525]" box="[151,869,255,280]" pageId="50" pageNumber="51">
<emphasis id="B9638C7FFFF3C056FF0EFEF27CA6362F" box="[151,164,256,279]" italics="true" pageId="50" pageNumber="51">*</emphasis>
<taxonomicName id="4C172BEEFFF3C056FF33FEF27DD83620" authority="Semper" authorityName="O. Semper" authorityYear="1865" box="[170,474,256,280]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="margaritula">
<emphasis id="B9638C7FFFF3C056FF33FEF27D853620" box="[170,391,256,280]" italics="true" pageId="50" pageNumber="51">Mathilda margaritula</emphasis>
Semper
</taxonomicName>
—Semper 1865: 343, pl. 13, figs 55a.
</paragraph>
<paragraph id="8BA8506DFFF3C056FF0EFEEC7ED2360E" blockId="50.[151,1436,255,525]" box="[151,720,286,311]" pageId="50" pageNumber="51">
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<taxonomicName id="4C172BEEFFF3C056FF0EFEEC7DE5360E" ID-CoL="BXSW9" authority="O. Semper" authorityName="O. Semper" authorityYear="1865" box="[151,487,286,310]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="margaritula">
<emphasis id="B9638C7FFFF3C056FF0EFEEC7D76360E" box="[151,372,286,310]" italics="true" pageId="50" pageNumber="51">Mathilda margaritula</emphasis>
O. Semper
</taxonomicName>
—de
<treatmentCitation id="0AB6767CFFF3C056FD87FEED7ECE360E" author="Boury, E. de" box="[542,716,286,311]" page="118" pageId="50" pageNumber="51" year="1883">
<bibRefCitation id="EF862D9CFFF3C056FD87FEED7ECE360E" author="Boury, E. de" box="[542,716,286,311]" pageId="50" pageNumber="51" pagination="110 - 153" refId="ref49657" refString="Boury, E. de (1883) Description d'especes nouvelles de Mathilda du basin de Paris. Journal de Conchyliologie, 31, 110 - 153. [https: // www. biodiversitylibrary. org / page / 15857125]" type="journal article" year="1883">Boury 1883: 118</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BA8506DFFF3C056FF0EFECF7EBF366D" blockId="50.[151,1436,255,525]" box="[151,701,317,341]" pageId="50" pageNumber="51">
<treatmentCitationGroup id="AB077743FFF3C056FF0EFECF7EBF366D" box="[151,701,317,341]" pageId="50" pageNumber="51">
<taxonomicName id="4C172BEEFFF3C056FF0EFECF7DD5366D" ID-CoL="BXSW9" authority="O. Semp." authorityName="O. Semper" authorityYear="1865" box="[151,471,317,341]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="margaritula">
<emphasis id="B9638C7FFFF3C056FF0EFECF7D76366D" box="[151,372,317,341]" italics="true" pageId="50" pageNumber="51">Mathilda margaritula</emphasis>
O. Semp.
</taxonomicName>
<treatmentCitation id="0AB6767CFFF3C056FE68FECF7EBB366D" author="Boettger, O." box="[497,697,317,341]" page="154" pageId="50" pageNumber="51" year="1902">
<bibRefCitation id="EF862D9CFFF3C056FE68FECF7EBB366D" author="Boettger, O." box="[497,697,317,341]" pageId="50" pageNumber="51" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger 1902: 154</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BA8506DFFF3C056FF0EFEAE7E8F364B" blockId="50.[151,1436,255,525]" box="[151,653,347,372]" pageId="50" pageNumber="51">
<taxonomicName id="4C172BEEFFF3C056FF0EFEAE7DBA364C" authority="Semp." authorityName="O. Semper" authorityYear="1865" box="[151,440,348,372]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="margaritula">
<emphasis id="B9638C7FFFF3C056FF0EFEAE7D76364C" box="[151,372,348,372]" italics="true" pageId="50" pageNumber="51">Mathilda margaritula</emphasis>
Semp.
</taxonomicName>
—Vetters 1910: 155.
</paragraph>
<paragraph id="8BA8506DFFF3C056FF0EFE887F1136AA" blockId="50.[151,1436,255,525]" box="[151,787,378,402]" pageId="50" pageNumber="51">
<taxonomicName id="4C172BEEFFF3C056FF0EFE887E4636AA" authority="Semp." authorityName="O. Semper" authorityYear="1865" box="[151,580,378,402]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF3C056FF0EFE887CF636AA" box="[151,244,378,402]" italics="true" pageId="50" pageNumber="51">Mathilda</emphasis>
(
<emphasis id="B9638C7FFFF3C056FE9AFE897D1A36AA" box="[259,280,379,402]" italics="true" pageId="50" pageNumber="51">M</emphasis>
[
<emphasis id="B9638C7FFFF3C056FEB9FE887D6A36AA" box="[288,360,378,402]" italics="true" pageId="50" pageNumber="51">athilda</emphasis>
].)
<emphasis id="B9638C7FFFF3C056FE1FFE887E0236AA" box="[390,512,378,402]" italics="true" pageId="50" pageNumber="51">margaritula</emphasis>
Semp.
</taxonomicName>
—Sieber 1958: 139.
</paragraph>
<paragraph id="8BA8506DFFF3C056FF0EFE6B7F0B3689" blockId="50.[151,1436,255,525]" box="[151,777,409,433]" pageId="50" pageNumber="51">
<taxonomicName id="4C172BEEFFF3C056FF0EFE6B7E393689" authority="Semper" authorityName="O. Semper" authorityYear="1865" box="[151,571,409,433]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="margaritula" subGenus="Mathilda">
<emphasis id="B9638C7FFFF3C056FF0EFE6B7CF63689" box="[151,244,409,433]" italics="true" pageId="50" pageNumber="51">Mathilda</emphasis>
(
<emphasis id="B9638C7FFFF3C056FE9AFE6B7D623689" box="[259,352,409,433]" italics="true" pageId="50" pageNumber="51">Mathilda</emphasis>
)
<emphasis id="B9638C7FFFF3C056FEF7FE6B7DEA3689" box="[366,488,409,433]" italics="true" pageId="50" pageNumber="51">margaritula</emphasis>
Semper
</taxonomicName>
—Sieber 1960: 265.
</paragraph>
<paragraph id="8BA8506DFFF3C056FF0EFE4A7DDB36D6" blockId="50.[151,1436,255,525]" pageId="50" pageNumber="51">
<taxonomicName id="4C172BEEFFF3C056FF0EFE4A7EEE36E8" authority="Semper" authorityName="Semper" baseAuthorityName="Borson" baseAuthorityYear="1821" box="[151,748,440,464]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="granosa">
<emphasis id="B9638C7FFFF3C056FF0EFE4A7D5236F7" box="[151,336,440,464]" italics="true" pageId="50" pageNumber="51">Mathilda granosa</emphasis>
(Borsoni [sic]) var.
<emphasis id="B9638C7FFFF3C056FDBFFE4A7E9536E8" box="[550,663,440,464]" italics="true" pageId="50" pageNumber="51">gemmulata</emphasis>
Semper
</taxonomicName>
—Kecskemétiné K̂rmendy 1962: 87, pl. 10, fig. 4 [
<emphasis id="B9638C7FFFF3C056FA88FE4A793536F7" box="[1297,1335,440,463]" italics="true" pageId="50" pageNumber="51">non</emphasis>
<taxonomicName id="4C172BEEFFF3C056FAA6FE4A7DCD36D6" authority="Semper, 1865" authorityName="Semper" authorityYear="1865" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="gemmulata">
<emphasis id="B9638C7FFFF3C056FAA6FE4A7D3A36D6" italics="true" pageId="50" pageNumber="51">Mathilda gemmulata</emphasis>
Semper, 1865
</taxonomicName>
].
</paragraph>
<paragraph id="8BA8506DFFF3C056FF0EFE0778A23535" blockId="50.[151,1436,255,525]" box="[151,1184,501,525]" pageId="50" pageNumber="51">
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<taxonomicName id="4C172BEEFFF3C056FF0EFE077DD63535" ID-CoL="BXTLZ" authority="Semper, 1865" authorityName="O. Semper" authorityYear="1865" box="[151,468,501,525]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="monilis">
<emphasis id="B9638C7FFFF3C056FF0EFE077D473535" box="[151,325,501,525]" italics="true" pageId="50" pageNumber="51">Mathilda monilis</emphasis>
Semper, 1865
</taxonomicName>
<treatmentCitation id="0AB6767CFFF3C056FE76FE077E9F3535" author="Baluk, W." box="[495,669,501,525]" page="196" pageId="50" pageNumber="51" year="2006">
<bibRefCitation id="EF862D9CFFF3C056FE76FE077E9F3535" author="Baluk, W." box="[495,669,501,525]" pageId="50" pageNumber="51" pagination="177 - 220" refId="ref48759" refString="Baluk, W. (2006) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part V Addenda et corrigenda ad Prosobranchia. Acta Geologica Polonica, 56, 177 - 220. [https: // geojournals. pgi. gov. pl / agp / article / view / 9847 / 8381]" type="journal article" year="2006">Bałuk 2006: 196</bibRefCitation>
</treatmentCitation>
: pl. 7, fig. 3 [
<emphasis id="B9638C7FFFF3C056FCBCFE047F493535" box="[805,843,502,525]" italics="true" pageId="50" pageNumber="51">non</emphasis>
<taxonomicName id="4C172BEEFFF3C056FCC8FE0778943535" authority="Semper, 1865" authorityName="O. Semper" authorityYear="1865" box="[849,1174,501,525]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="monilis">
<emphasis id="B9638C7FFFF3C056FCC8FE077FFD3535" box="[849,1023,501,525]" italics="true" pageId="50" pageNumber="51">Mathilda monilis</emphasis>
Semper, 1865
</taxonomicName>
].
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<emphasis id="B9638C7FFFF3C056FF0EFDC97D45356D" bold="true" box="[151,327,571,597]" pageId="50" pageNumber="51">Type material.</emphasis>
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<typeStatus id="54ACEECFFFF3C056FEC9FDCE7DC2356E" box="[336,448,572,598]" pageId="50" pageNumber="51" type="lectotype">Lectotype</typeStatus>
(designated herein):
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/0001/0549, SL:
<quantity id="4CEFFD88FFF3C056FB96FDC9786B356E" box="[1039,1129,571,598]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="50" pageNumber="51" unit="mm" value="4.8">4.8 mm</quantity>
, MD:
<quantity id="4CEFFD88FFF3C056FB2DFDC9790C356E" box="[1204,1294,571,598]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="50" pageNumber="51" unit="mm" value="1.7">1.7 mm</quantity>
,
<location id="8EC806B6FFF3C056FA85FDC97CBD3541" LSID="urn:lsid:plazi:treatment:03BEE17BFFF3C056FF0EFF657CE231ED:8EC806B6FFF3C056FA85FDC97CBD3541" country="Romania" name="Lapugiu de Sus" pageId="50" pageNumber="51">
Lăpugiu
<collectorName id="26E235BBFFF3C056FA1BFDC97CBD3541" pageId="50" pageNumber="51">de Sus</collectorName>
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(
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<figureCitation id="132C4CE8FFF3C056FEC9FDAD7DB43542" box="[336,438,607,634]" captionStart="FIGURE 25" captionStartId="49.[151,250,1686,1710]" captionTargetBox="[213,1375,181,1662]" captionTargetId="figure-25@49.[213,1375,181,1662]" captionTargetPageId="49" captionText="FIGURE 25 A1A2. Mathilda gloriosa Boettger, 1907, holotype, SMF 372691, Coşteiu de Sus (Romania). B1B2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549a, Lăpugiu de Sus (Romania). C1C2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1876/0011/0098, Lăpugiu des Sus (Romania). D. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549b, Lăpugiu de Sus (Romania). E. Mathilda margaritula Semper, 1865, lecto- type, NHMW 1868/0001/0549, Lăpugiu de Sus (Romania). F1F2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549c, Lăpugiu de Sus (Romania). G. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549d, Lăpugiu de Sus (Romania). H1H2. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039a, Sedlec (Czech Republic). I1I2. Mathilda margaritula Semper, 1865, NHMW 1866/0001/0912, Niederleis (Austria). J. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039b, Sedlec (Czech Republic). K. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039c, Sedlec (Czech Republic)." figureDoi="http://doi.org/10.5281/zenodo.10147866" httpUri="https://zenodo.org/record/10147866/files/figure.png" pageId="50" pageNumber="51">Fig. 25E</figureCitation>
</materialsCitation>
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<materialsCitation id="3B7F5A30FFF3C056FE5BFDAD7DB7359C" collectionCode="NHMW" country="Romania" location="Lapugiu de Sus" pageId="50" pageNumber="51" specimenCode="NHMW 1868" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="54ACEECFFFF3C056FE5BFDAD7E683541" box="[450,618,607,633]" pageId="50" pageNumber="51" type="paralectotype">Paralectotypes</typeStatus>
:
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/0001/0549a, SL: 4.0 mm, MD:
<quantity id="4CEFFD88FFF3C056FB1AFDAD78DF3542" box="[1155,1245,607,634]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="50" pageNumber="51" unit="mm" value="1.7">1.7 mm</quantity>
,
<location id="8EC806B6FFF3C056FB73FDAD799E3541" LSID="urn:lsid:plazi:treatment:03BEE17BFFF3C056FF0EFF657CE231ED:8EC806B6FFF3C056FB73FDAD799E3541" box="[1258,1436,607,633]" country="Romania" name="Lapugiu de Sus" pageId="50" pageNumber="51">Lăpugiu de Sus</location>
(
<collectingCountry id="F30010FDFFF3C056FF38FD717D0935A5" box="[161,267,643,669]" name="Romania" pageId="50" pageNumber="51">Romania</collectingCountry>
),
<figureCitation id="132C4CE8FFF3C056FE87FD717D8635A6" box="[286,388,643,670]" captionStart="FIGURE 25" captionStartId="49.[151,250,1686,1710]" captionTargetBox="[213,1375,181,1662]" captionTargetId="figure-25@49.[213,1375,181,1662]" captionTargetPageId="49" captionText="FIGURE 25 A1A2. Mathilda gloriosa Boettger, 1907, holotype, SMF 372691, Coşteiu de Sus (Romania). B1B2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549a, Lăpugiu de Sus (Romania). C1C2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1876/0011/0098, Lăpugiu des Sus (Romania). D. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549b, Lăpugiu de Sus (Romania). E. Mathilda margaritula Semper, 1865, lecto- type, NHMW 1868/0001/0549, Lăpugiu de Sus (Romania). F1F2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549c, Lăpugiu de Sus (Romania). G. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549d, Lăpugiu de Sus (Romania). H1H2. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039a, Sedlec (Czech Republic). I1I2. Mathilda margaritula Semper, 1865, NHMW 1866/0001/0912, Niederleis (Austria). J. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039b, Sedlec (Czech Republic). K. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039c, Sedlec (Czech Republic)." figureDoi="http://doi.org/10.5281/zenodo.10147866" httpUri="https://zenodo.org/record/10147866/files/figure.png" pageId="50" pageNumber="51">Figs 25B</figureCitation>
<figureCitation id="132C4CE8FFF3C056FE1DFD647DAF35A6" box="[388,429,644,676]" captionStart="FIGURE 1" captionStartId="5.[151,250,1522,1546]" captionTargetBox="[232,1366,234,1455]" captionTargetId="figure-58@5.[232,1366,234,1455]" captionTargetPageId="5" captionText="FIGURE 1. Descriptive terms used in descriptions of Architectonicidae (modified from Bieler 1993)." figureDoi="http://doi.org/10.5281/zenodo.10147816" httpUri="https://zenodo.org/record/10147816/files/figure.png" pageId="50" pageNumber="51">
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B
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<figureCitation id="132C4CE8FFF3C056FE34FD647DB7359C" box="[429,437,662,676]" captionStart="FIGURE 2" captionStartId="6.[151,250,1086,1110]" captionTargetBox="[236,1353,159,1066]" captionTargetId="figure-113@6.[236,1353,159,1066]" captionTargetPageId="6" captionText="FIGURE 2. Whorl cross-sections of Architectonicidae genera. All modified after Bieler (1993) except for Simplexollata, Basisulcata and Ammotectonica. Solatisonax? refers to Solatisonax? kostejana (Boettger, 1907), Bold genera are represented in the Paratethys Sea. SSC: subsutural cord, MC: mid cord(s), UPC: upper peripheral cord, LPC: lower peripheral cord, IPC: infraperipheral cord; UC: umbilical crenae; PUC: proxumbilical cord." figureDoi="http://doi.org/10.5281/zenodo.10147818" httpUri="https://zenodo.org/record/10147818/files/figure.png" pageId="50" pageNumber="51">
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<materialsCitation id="3B7F5A30FFF3C056FE5DFD767CF535F0" collectionCode="NHMW" country="Romania" location="Lapugiu des Sus" pageId="50" pageNumber="51" specimenCode="NHMW 1876" specimenCount="1" typeStatus="paralectotype">
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/0011/0098, SL:
<quantity id="4CEFFD88FFF3C056FC85FD717F7735A6" box="[796,885,643,670]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="50" pageNumber="51" unit="mm" value="3.7">3.7 mm</quantity>
, MD:
<quantity id="4CEFFD88FFF3C056FC27FD71781535A6" box="[958,1047,643,670]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="50" pageNumber="51" unit="mm" value="1.5">1.5 mm</quantity>
,
<location id="8EC806B6FFF3C056FBBAFD7178DF35A5" LSID="urn:lsid:plazi:treatment:03BEE17BFFF3C056FF0EFF657CE231ED:8EC806B6FFF3C056FBBAFD7178DF35A5" box="[1059,1245,643,669]" country="Romania" name="Lapugiu des Sus" pageId="50" pageNumber="51">Lăpugiu des Sus</location>
(
<collectingCountry id="F30010FDFFF3C056FB74FD71795A35A5" box="[1261,1368,643,669]" name="Romania" pageId="50" pageNumber="51">Romania</collectingCountry>
),
<figureCitation id="132C4CE8FFF3C056FAF5FD717CC435FA" captionStart="FIGURE 25" captionStartId="49.[151,250,1686,1710]" captionTargetBox="[213,1375,181,1662]" captionTargetId="figure-25@49.[213,1375,181,1662]" captionTargetPageId="49" captionText="FIGURE 25 A1A2. Mathilda gloriosa Boettger, 1907, holotype, SMF 372691, Coşteiu de Sus (Romania). B1B2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549a, Lăpugiu de Sus (Romania). C1C2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1876/0011/0098, Lăpugiu des Sus (Romania). D. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549b, Lăpugiu de Sus (Romania). E. Mathilda margaritula Semper, 1865, lecto- type, NHMW 1868/0001/0549, Lăpugiu de Sus (Romania). F1F2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549c, Lăpugiu de Sus (Romania). G. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549d, Lăpugiu de Sus (Romania). H1H2. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039a, Sedlec (Czech Republic). I1I2. Mathilda margaritula Semper, 1865, NHMW 1866/0001/0912, Niederleis (Austria). J. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039b, Sedlec (Czech Republic). K. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039c, Sedlec (Czech Republic)." figureDoi="http://doi.org/10.5281/zenodo.10147866" httpUri="https://zenodo.org/record/10147866/files/figure.png" pageId="50" pageNumber="51">Figs 25C</figureCitation>
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C
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<figureCitation id="132C4CE8FFF3C056FF76FD487CF535F0" box="[239,247,698,712]" captionStart="FIGURE 2" captionStartId="6.[151,250,1086,1110]" captionTargetBox="[236,1353,159,1066]" captionTargetId="figure-113@6.[236,1353,159,1066]" captionTargetPageId="6" captionText="FIGURE 2. Whorl cross-sections of Architectonicidae genera. All modified after Bieler (1993) except for Simplexollata, Basisulcata and Ammotectonica. Solatisonax? refers to Solatisonax? kostejana (Boettger, 1907), Bold genera are represented in the Paratethys Sea. SSC: subsutural cord, MC: mid cord(s), UPC: upper peripheral cord, LPC: lower peripheral cord, IPC: infraperipheral cord; UC: umbilical crenae; PUC: proxumbilical cord." figureDoi="http://doi.org/10.5281/zenodo.10147818" httpUri="https://zenodo.org/record/10147818/files/figure.png" pageId="50" pageNumber="51">
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<materialsCitation id="3B7F5A30FFF3C056FE9EFD5A793635FA" box="[263,1332,679,706]" collectionCode="NHMW" country="Romania" location="Lapugiu de Sus" pageId="50" pageNumber="51" specimenCode="NHMW 1868" specimenCount="1" typeStatus="paralectotype">
<specimenCode id="DBB1F816FFF3C056FE9EFD5A7DA135FA" box="[263,419,679,706]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="50" pageNumber="51" type="Museum">NHMW 1868</specimenCode>
/0001/0549b, SL:
<quantity id="4CEFFD88FFF3C056FDECFD557ECD35FA" box="[629,719,679,706]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="50" pageNumber="51" unit="mm" value="4.8">4.8 mm</quantity>
, MD:
<quantity id="4CEFFD88FFF3C056FC85FD557F7535FA" box="[796,887,679,706]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="50" pageNumber="51" unit="mm" value="1.7">1.7 mm</quantity>
,
<location id="8EC806B6FFF3C056FC1CFD55783A35F9" LSID="urn:lsid:plazi:treatment:03BEE17BFFF3C056FF0EFF657CE231ED:8EC806B6FFF3C056FC1CFD55783A35F9" box="[901,1080,679,705]" country="Romania" name="Lapugiu de Sus" pageId="50" pageNumber="51">Lăpugiu de Sus</location>
(
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<figureCitation id="132C4CE8FFF3C056FB53FD55793635FA" box="[1226,1332,679,706]" captionStart="FIGURE 25" captionStartId="49.[151,250,1686,1710]" captionTargetBox="[213,1375,181,1662]" captionTargetId="figure-25@49.[213,1375,181,1662]" captionTargetPageId="49" captionText="FIGURE 25 A1A2. Mathilda gloriosa Boettger, 1907, holotype, SMF 372691, Coşteiu de Sus (Romania). B1B2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549a, Lăpugiu de Sus (Romania). C1C2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1876/0011/0098, Lăpugiu des Sus (Romania). D. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549b, Lăpugiu de Sus (Romania). E. Mathilda margaritula Semper, 1865, lecto- type, NHMW 1868/0001/0549, Lăpugiu de Sus (Romania). F1F2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549c, Lăpugiu de Sus (Romania). G. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549d, Lăpugiu de Sus (Romania). H1H2. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039a, Sedlec (Czech Republic). I1I2. Mathilda margaritula Semper, 1865, NHMW 1866/0001/0912, Niederleis (Austria). J. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039b, Sedlec (Czech Republic). K. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039c, Sedlec (Czech Republic)." figureDoi="http://doi.org/10.5281/zenodo.10147866" httpUri="https://zenodo.org/record/10147866/files/figure.png" pageId="50" pageNumber="51">Fig. 25D</figureCitation>
</materialsCitation>
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<materialsCitation id="3B7F5A30FFF3C056FAD8FD5A786235D4" collectionCode="NHMW" country="Romania" location="Lapugiu de Sus" pageId="50" pageNumber="51" specimenCode="NHMW 1868" specimenCount="1" typeStatus="paralectotype">
<specimenCode id="DBB1F816FFF3C056FAD8FD5A7CCC35DE" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="50" pageNumber="51" type="Museum">NHMW 1868</specimenCode>
/0001/0549c, SL: mm, 4.8 MD:
<quantity id="4CEFFD88FFF3C056FDACFD397E8F35DE" box="[565,653,715,742]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="50" pageNumber="51" unit="mm" value="1.8">1.8 mm</quantity>
,
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(
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<figureCitation id="132C4CE8FFF3C056FC49FD39783135DE" box="[976,1075,715,742]" captionStart="FIGURE 25" captionStartId="49.[151,250,1686,1710]" captionTargetBox="[213,1375,181,1662]" captionTargetId="figure-25@49.[213,1375,181,1662]" captionTargetPageId="49" captionText="FIGURE 25 A1A2. Mathilda gloriosa Boettger, 1907, holotype, SMF 372691, Coşteiu de Sus (Romania). B1B2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549a, Lăpugiu de Sus (Romania). C1C2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1876/0011/0098, Lăpugiu des Sus (Romania). D. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549b, Lăpugiu de Sus (Romania). E. Mathilda margaritula Semper, 1865, lecto- type, NHMW 1868/0001/0549, Lăpugiu de Sus (Romania). F1F2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549c, Lăpugiu de Sus (Romania). G. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549d, Lăpugiu de Sus (Romania). H1H2. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039a, Sedlec (Czech Republic). I1I2. Mathilda margaritula Semper, 1865, NHMW 1866/0001/0912, Niederleis (Austria). J. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039b, Sedlec (Czech Republic). K. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039c, Sedlec (Czech Republic)." figureDoi="http://doi.org/10.5281/zenodo.10147866" httpUri="https://zenodo.org/record/10147866/files/figure.png" pageId="50" pageNumber="51">Figs 25F</figureCitation>
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<figureCitation id="132C4CE8FFF3C056FBC1FD2C786235D4" box="[1112,1120,734,748]" captionStart="FIGURE 2" captionStartId="6.[151,250,1086,1110]" captionTargetBox="[236,1353,159,1066]" captionTargetId="figure-113@6.[236,1353,159,1066]" captionTargetPageId="6" captionText="FIGURE 2. Whorl cross-sections of Architectonicidae genera. All modified after Bieler (1993) except for Simplexollata, Basisulcata and Ammotectonica. Solatisonax? refers to Solatisonax? kostejana (Boettger, 1907), Bold genera are represented in the Paratethys Sea. SSC: subsutural cord, MC: mid cord(s), UPC: upper peripheral cord, LPC: lower peripheral cord, IPC: infraperipheral cord; UC: umbilical crenae; PUC: proxumbilical cord." figureDoi="http://doi.org/10.5281/zenodo.10147818" httpUri="https://zenodo.org/record/10147818/files/figure.png" pageId="50" pageNumber="51">
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<materialsCitation id="3B7F5A30FFF3C056FBF7FD3E7F6E3432" collectionCode="NHMW" country="Romania" location="Lapugiu de Sus" pageId="50" pageNumber="51" specimenCode="NHMW 1868" specimenCount="1" typeStatus="paralectotype">
<specimenCode id="DBB1F816FFF3C056FBF7FD3E790535DE" box="[1134,1287,715,742]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="50" pageNumber="51" type="Museum">NHMW 1868</specimenCode>
/0001/0549d, SL:
<quantity id="4CEFFD88FFF3C056FF5EFD1D7D1D3432" box="[199,287,751,778]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="50" pageNumber="51" unit="mm" value="4.8">4.8 mm</quantity>
, MD:
<quantity id="4CEFFD88FFF3C056FEFFFD1D7DBC3432" box="[358,446,751,778]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="50" pageNumber="51" unit="mm" value="1.8">1.8 mm</quantity>
,
<location id="8EC806B6FFF3C056FE53FD1D7E753431" LSID="urn:lsid:plazi:treatment:03BEE17BFFF3C056FF0EFF657CE231ED:8EC806B6FFF3C056FE53FD1D7E753431" box="[458,631,751,777]" country="Romania" name="Lapugiu de Sus" pageId="50" pageNumber="51">Lăpugiu de Sus</location>
(
<collectingCountry id="F30010FDFFF3C056FD1EFD1D7EF03431" box="[647,754,751,777]" name="Romania" pageId="50" pageNumber="51">Romania</collectingCountry>
),
<figureCitation id="132C4CE8FFF3C056FC9CFD1D7F6E3432" box="[773,876,751,778]" captionStart="FIGURE 25" captionStartId="49.[151,250,1686,1710]" captionTargetBox="[213,1375,181,1662]" captionTargetId="figure-25@49.[213,1375,181,1662]" captionTargetPageId="49" captionText="FIGURE 25 A1A2. Mathilda gloriosa Boettger, 1907, holotype, SMF 372691, Coşteiu de Sus (Romania). B1B2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549a, Lăpugiu de Sus (Romania). C1C2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1876/0011/0098, Lăpugiu des Sus (Romania). D. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549b, Lăpugiu de Sus (Romania). E. Mathilda margaritula Semper, 1865, lecto- type, NHMW 1868/0001/0549, Lăpugiu de Sus (Romania). F1F2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549c, Lăpugiu de Sus (Romania). G. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549d, Lăpugiu de Sus (Romania). H1H2. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039a, Sedlec (Czech Republic). I1I2. Mathilda margaritula Semper, 1865, NHMW 1866/0001/0912, Niederleis (Austria). J. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039b, Sedlec (Czech Republic). K. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039c, Sedlec (Czech Republic)." figureDoi="http://doi.org/10.5281/zenodo.10147866" httpUri="https://zenodo.org/record/10147866/files/figure.png" pageId="50" pageNumber="51">Fig. 25G</figureCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA8506DFFF3C056FF5EFCE178C734A2" blockId="50.[151,1437,571,1749]" pageId="50" pageNumber="51">
<emphasis id="B9638C7FFFF3C056FF5EFCE17DBB3415" bold="true" box="[199,441,787,813]" pageId="50" pageNumber="51">Illustrated material.</emphasis>
<collectionCode id="ED06C8A8FFF3C056FE58FCE67E1E3416" box="[449,540,788,814]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="50" pageNumber="51" type="Museum">NHMW</collectionCode>
1858/0047/0039a, SL:
<quantity id="4CEFFD88FFF3C056FCBEFCE17F823416" box="[807,896,787,814]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="50" pageNumber="51" unit="mm" value="4.3">4.3 mm</quantity>
, MD:
<quantity id="4CEFFD88FFF3C056FC50FCE178203416" box="[969,1058,787,814]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="50" pageNumber="51" unit="mm" value="1.6">1.6 mm</quantity>
, Sedlec (= Porzteich) (
<collectingCountry id="F30010FDFFF3C056FABDFCE17CE33469" name="Czech Republic" pageId="50" pageNumber="51">Czech Republic</collectingCountry>
),
<figureCitation id="132C4CE8FFF3C056FF61FCC57D60346A" box="[248,354,823,850]" captionStart="FIGURE 25" captionStartId="49.[151,250,1686,1710]" captionTargetBox="[213,1375,181,1662]" captionTargetId="figure-25@49.[213,1375,181,1662]" captionTargetPageId="49" captionText="FIGURE 25 A1A2. Mathilda gloriosa Boettger, 1907, holotype, SMF 372691, Coşteiu de Sus (Romania). B1B2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549a, Lăpugiu de Sus (Romania). C1C2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1876/0011/0098, Lăpugiu des Sus (Romania). D. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549b, Lăpugiu de Sus (Romania). E. Mathilda margaritula Semper, 1865, lecto- type, NHMW 1868/0001/0549, Lăpugiu de Sus (Romania). F1F2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549c, Lăpugiu de Sus (Romania). G. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549d, Lăpugiu de Sus (Romania). H1H2. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039a, Sedlec (Czech Republic). I1I2. Mathilda margaritula Semper, 1865, NHMW 1866/0001/0912, Niederleis (Austria). J. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039b, Sedlec (Czech Republic). K. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039c, Sedlec (Czech Republic)." figureDoi="http://doi.org/10.5281/zenodo.10147866" httpUri="https://zenodo.org/record/10147866/files/figure.png" pageId="50" pageNumber="51">Figs 25H</figureCitation>
<figureCitation id="132C4CE8FFF3C056FEFBFCB87D8E346A" box="[354,396,824,856]" captionStart="FIGURE 1" captionStartId="5.[151,250,1522,1546]" captionTargetBox="[232,1366,234,1455]" captionTargetId="figure-58@5.[232,1366,234,1455]" captionTargetPageId="5" captionText="FIGURE 1. Descriptive terms used in descriptions of Architectonicidae (modified from Bieler 1993)." figureDoi="http://doi.org/10.5281/zenodo.10147816" httpUri="https://zenodo.org/record/10147816/files/figure.png" pageId="50" pageNumber="51">
<subScript id="17935228FFF3C056FEFBFCB87D683460" attach="left" box="[354,362,842,856]" fontSize="6" pageId="50" pageNumber="51">1</subScript>
H
</figureCitation>
<figureCitation id="132C4CE8FFF3C056FE14FCB87D973460" box="[397,405,842,856]" captionStart="FIGURE 2" captionStartId="6.[151,250,1086,1110]" captionTargetBox="[236,1353,159,1066]" captionTargetId="figure-113@6.[236,1353,159,1066]" captionTargetPageId="6" captionText="FIGURE 2. Whorl cross-sections of Architectonicidae genera. All modified after Bieler (1993) except for Simplexollata, Basisulcata and Ammotectonica. Solatisonax? refers to Solatisonax? kostejana (Boettger, 1907), Bold genera are represented in the Paratethys Sea. SSC: subsutural cord, MC: mid cord(s), UPC: upper peripheral cord, LPC: lower peripheral cord, IPC: infraperipheral cord; UC: umbilical crenae; PUC: proxumbilical cord." figureDoi="http://doi.org/10.5281/zenodo.10147818" httpUri="https://zenodo.org/record/10147818/files/figure.png" pageId="50" pageNumber="51">
<subScript id="17935228FFF3C056FE14FCB87D973460" attach="right" box="[397,405,842,856]" fontSize="6" pageId="50" pageNumber="51">2</subScript>
</figureCitation>
.
<collectionCode id="ED06C8A8FFF3C056FE3FFCCA7E03346A" box="[422,513,824,850]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="50" pageNumber="51" type="Museum">NHMW</collectionCode>
1866/0001/0912, SL:
<quantity id="4CEFFD88FFF3C056FC9EFCC57F60346A" box="[775,866,823,850]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="50" pageNumber="51" unit="mm" value="4.3">4.3 mm</quantity>
, MD:
<quantity id="4CEFFD88FFF3C056FC29FCC57809346A" box="[944,1035,823,850]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="50" pageNumber="51" unit="mm" value="1.7">1.7 mm</quantity>
, Niederleis (
<collectingCountry id="F30010FDFFF3C056FB06FCC578F53469" box="[1183,1271,823,849]" name="Austria" pageId="50" pageNumber="51">Austria</collectingCountry>
),
<figureCitation id="132C4CE8FFF3C056FA94FCC5796F346A" box="[1293,1389,823,850]" captionStart="FIGURE 25" captionStartId="49.[151,250,1686,1710]" captionTargetBox="[213,1375,181,1662]" captionTargetId="figure-25@49.[213,1375,181,1662]" captionTargetPageId="49" captionText="FIGURE 25 A1A2. Mathilda gloriosa Boettger, 1907, holotype, SMF 372691, Coşteiu de Sus (Romania). B1B2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549a, Lăpugiu de Sus (Romania). C1C2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1876/0011/0098, Lăpugiu des Sus (Romania). D. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549b, Lăpugiu de Sus (Romania). E. Mathilda margaritula Semper, 1865, lecto- type, NHMW 1868/0001/0549, Lăpugiu de Sus (Romania). F1F2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549c, Lăpugiu de Sus (Romania). G. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549d, Lăpugiu de Sus (Romania). H1H2. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039a, Sedlec (Czech Republic). I1I2. Mathilda margaritula Semper, 1865, NHMW 1866/0001/0912, Niederleis (Austria). J. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039b, Sedlec (Czech Republic). K. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039c, Sedlec (Czech Republic)." figureDoi="http://doi.org/10.5281/zenodo.10147866" httpUri="https://zenodo.org/record/10147866/files/figure.png" pageId="50" pageNumber="51">Figs 25I</figureCitation>
<figureCitation id="132C4CE8FFF3C056FAF4FCB8798F346A" box="[1389,1421,824,856]" captionStart="FIGURE 1" captionStartId="5.[151,250,1522,1546]" captionTargetBox="[232,1366,234,1455]" captionTargetId="figure-58@5.[232,1366,234,1455]" captionTargetPageId="5" captionText="FIGURE 1. Descriptive terms used in descriptions of Architectonicidae (modified from Bieler 1993)." figureDoi="http://doi.org/10.5281/zenodo.10147816" httpUri="https://zenodo.org/record/10147816/files/figure.png" pageId="50" pageNumber="51">
<subScript id="17935228FFF3C056FAF4FCB879773460" attach="left" box="[1389,1397,842,856]" fontSize="6" pageId="50" pageNumber="51">1</subScript>
I
</figureCitation>
<figureCitation id="132C4CE8FFF3C056FA15FCB879963460" box="[1420,1428,842,856]" captionStart="FIGURE 2" captionStartId="6.[151,250,1086,1110]" captionTargetBox="[236,1353,159,1066]" captionTargetId="figure-113@6.[236,1353,159,1066]" captionTargetPageId="6" captionText="FIGURE 2. Whorl cross-sections of Architectonicidae genera. All modified after Bieler (1993) except for Simplexollata, Basisulcata and Ammotectonica. Solatisonax? refers to Solatisonax? kostejana (Boettger, 1907), Bold genera are represented in the Paratethys Sea. SSC: subsutural cord, MC: mid cord(s), UPC: upper peripheral cord, LPC: lower peripheral cord, IPC: infraperipheral cord; UC: umbilical crenae; PUC: proxumbilical cord." figureDoi="http://doi.org/10.5281/zenodo.10147818" httpUri="https://zenodo.org/record/10147818/files/figure.png" pageId="50" pageNumber="51">
<subScript id="17935228FFF3C056FA15FCB879963460" attach="left" box="[1420,1428,842,856]" fontSize="6" pageId="50" pageNumber="51">2</subScript>
</figureCitation>
;
<collectionCode id="ED06C8A8FFF3C056FF0EFCAE7CF0344E" box="[151,242,860,886]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="50" pageNumber="51" type="Museum">NHMW</collectionCode>
1858/0047/0039b, SL:
<quantity id="4CEFFD88FFF3C056FD9BFCA97E5E344E" box="[514,604,859,886]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="50" pageNumber="51" unit="mm" value="4.2">4.2 mm</quantity>
, MD:
<quantity id="4CEFFD88FFF3C056FD3EFCA97F02344E" box="[679,768,859,886]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="50" pageNumber="51" unit="mm" value="1.5">1.5 mm</quantity>
, Sedlec (= Porzteich) (
<collectingCountry id="F30010FDFFF3C056FB9FFCA978C3344D" box="[1030,1217,859,885]" name="Czech Republic" pageId="50" pageNumber="51">Czech Republic</collectingCountry>
),
<figureCitation id="132C4CE8FFF3C056FB4CFCA97937344E" box="[1237,1333,859,886]" captionStart="FIGURE 25" captionStartId="49.[151,250,1686,1710]" captionTargetBox="[213,1375,181,1662]" captionTargetId="figure-25@49.[213,1375,181,1662]" captionTargetPageId="49" captionText="FIGURE 25 A1A2. Mathilda gloriosa Boettger, 1907, holotype, SMF 372691, Coşteiu de Sus (Romania). B1B2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549a, Lăpugiu de Sus (Romania). C1C2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1876/0011/0098, Lăpugiu des Sus (Romania). D. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549b, Lăpugiu de Sus (Romania). E. Mathilda margaritula Semper, 1865, lecto- type, NHMW 1868/0001/0549, Lăpugiu de Sus (Romania). F1F2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549c, Lăpugiu de Sus (Romania). G. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549d, Lăpugiu de Sus (Romania). H1H2. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039a, Sedlec (Czech Republic). I1I2. Mathilda margaritula Semper, 1865, NHMW 1866/0001/0912, Niederleis (Austria). J. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039b, Sedlec (Czech Republic). K. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039c, Sedlec (Czech Republic)." figureDoi="http://doi.org/10.5281/zenodo.10147866" httpUri="https://zenodo.org/record/10147866/files/figure.png" pageId="50" pageNumber="51">Fig. 25J</figureCitation>
.
<collectionCode id="ED06C8A8FFF3C056FAD8FCAE799E344E" box="[1345,1436,860,886]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="50" pageNumber="51" type="Museum">NHMW</collectionCode>
1858/0047/0039c, SL:
<quantity id="4CEFFD88FFF3C056FE00FC8D7DF334A2" box="[409,497,895,922]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="50" pageNumber="51" unit="mm" value="2.6">2.6 mm</quantity>
, MD:
<quantity id="4CEFFD88FFF3C056FDA1FC8D7E9234A2" box="[568,656,895,922]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="50" pageNumber="51" unit="mm" value="1.2">1.2 mm</quantity>
, Sedlec (= Porzteich) (
<collectingCountry id="F30010FDFFF3C056FC16FC8D784A34A1" box="[911,1096,895,921]" name="Czech Republic" pageId="50" pageNumber="51">Czech Republic</collectingCountry>
),
<figureCitation id="132C4CE8FFF3C056FBC3FC8D78C334A2" box="[1114,1217,895,922]" captionStart="FIGURE 25" captionStartId="49.[151,250,1686,1710]" captionTargetBox="[213,1375,181,1662]" captionTargetId="figure-25@49.[213,1375,181,1662]" captionTargetPageId="49" captionText="FIGURE 25 A1A2. Mathilda gloriosa Boettger, 1907, holotype, SMF 372691, Coşteiu de Sus (Romania). B1B2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549a, Lăpugiu de Sus (Romania). C1C2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1876/0011/0098, Lăpugiu des Sus (Romania). D. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549b, Lăpugiu de Sus (Romania). E. Mathilda margaritula Semper, 1865, lecto- type, NHMW 1868/0001/0549, Lăpugiu de Sus (Romania). F1F2. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549c, Lăpugiu de Sus (Romania). G. Mathilda margaritula Semper, 1865, paralectotype, NHMW 1868/0001/0549d, Lăpugiu de Sus (Romania). H1H2. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039a, Sedlec (Czech Republic). I1I2. Mathilda margaritula Semper, 1865, NHMW 1866/0001/0912, Niederleis (Austria). J. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039b, Sedlec (Czech Republic). K. Mathilda margaritula Semper, 1865, NHMW 1858/0047/0039c, Sedlec (Czech Republic)." figureDoi="http://doi.org/10.5281/zenodo.10147866" httpUri="https://zenodo.org/record/10147866/files/figure.png" pageId="50" pageNumber="51">Fig. 25K</figureCitation>
.
</paragraph>
<paragraph id="8BA8506DFFF3C056FF5EFC517F8233E5" blockId="50.[151,1437,571,1749]" pageId="50" pageNumber="51">
<emphasis id="B9638C7FFFF3C056FF5EFC517DB83485" bold="true" box="[199,442,931,957]" pageId="50" pageNumber="51">Revised description.</emphasis>
Small, slender, robust turritelliform shell; apical angle ~20°. Protoconch multispiral, of about 2.5 smooth whorls, heterostrophic, nucleus just exposed, tilted at angle of about 120°. Teleoconch of six subcylindrical whorls separated by deeply impressed, suture. Sculpture of three strongly raised primary spiral cords: S1 slightly weaker, recessed; S2 forming periphery; S3 placed at some distance above suture, equal in strength to S2. Forth spiral cord appears just above suture on third whorl largely covered by subsequent whorl. Axial sculpture of about 25 raised axial lamellae forming marked, cancellate sculpture with prominent rounded tubercles developed at intersections. Base strongly angulated, almost flat, sharply delimited by double peribasal cord, with weak periumbilical cord. Aperture small, subcircular. Outer lip simple, crenulated by edge of spiral cords. Columella nearly straight. Columellar callus forming narrow rim. No parietal callus.
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFF3C056FF5EFB157CE23119" pageId="50" pageNumber="51" type="discussion">
<paragraph id="8BA8506DFFF3C056FF5EFB157D70328E" blockId="50.[151,1437,571,1749]" pageId="50" pageNumber="51">
<emphasis id="B9638C7FFFF3C056FF5EFB157D4E3239" bold="true" box="[199,332,1255,1281]" pageId="50" pageNumber="51">Discussion.</emphasis>
<taxonomicName id="4C172BEEFFF3C056FECDFB157ED03239" authority="(Borson, 1821)" baseAuthorityName="Borson" baseAuthorityYear="1821" box="[340,722,1255,1281]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="granosa">
<emphasis id="B9638C7FFFF3C056FECDFB157E1C3239" box="[340,542,1255,1281]" italics="true" pageId="50" pageNumber="51">Mathilda granosa</emphasis>
(
<bibRefCitation id="EF862D9CFFF3C056FDB7FB157EC83239" author="Borson, S." box="[558,714,1255,1281]" pageId="50" pageNumber="51" pagination="297 - 364" refId="ref49541" refString="Borson, S. (1821) Continuazione del saggio di orittografia Piemontese. Memorie della reale accademia delle scienze di Torino, Series 1, 26, 297 - 364, pls. 5 - 6. [https: // www. biodiversitylibrary. org / page / 7653833]" type="journal article" year="1821">Borson, 1821</bibRefCitation>
)
</taxonomicName>
, from the Burdigalian of the Colli Torinesi (
<collectingCountry id="F30010FDFFF3C056FB5EFB1578FD3239" box="[1223,1279,1255,1281]" name="Italy" pageId="50" pageNumber="51">Italy</collectingCountry>
), differs in its weaker subsutural cord and the keel-like mid-cord (see Pavia 1976: pl. 1, figs 2a2b;
<bibRefCitation id="EF862D9CFFF3C056FBFBFAF97976321D" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[1122,1396,1291,1318]" pageId="50" pageNumber="51" pagination="3 - 576" refId="ref56228" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576." type="journal article" year="2013">
Landau
<emphasis id="B9638C7FFFF3C056FB27FAFE78F2321D" box="[1214,1264,1291,1317]" italics="true" pageId="50" pageNumber="51">et al</emphasis>
. 2013: 303
</bibRefCitation>
, pl. 52, fig. 4) [note that
<taxonomicName id="4C172BEEFFF3C056FE19FADD7F773271" authority="sensu Landau et al. (2011" authorityName="Landau" authorityYear="2011" box="[384,885,1327,1354]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="granosa">
<emphasis id="B9638C7FFFF3C056FE19FADD7E4B3271" box="[384,585,1327,1353]" italics="true" pageId="50" pageNumber="51">Mathilda granosa</emphasis>
sensu
<bibRefCitation id="EF862D9CFFF3C056FD0FFADD7F773271" author="Landau, B. &amp; Da Silva, C. M. &amp; Mayoral, E." box="[662,885,1327,1353]" pageId="50" pageNumber="51" refId="ref56098" refString="Landau, B., Da Silva, C. M. &amp; Mayoral, E. (2011) The Lower Pliocene gastropods of the Huelva Sands Formation, Guadalquivir Basin, Southwestern Spain. Palaeofocus, 4, 155 pp." type="book" year="2011">
Landau
<emphasis id="B9638C7FFFF3C056FD68FAC27F283271" box="[753,810,1327,1353]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
(2011
</bibRefCitation>
</taxonomicName>
: pl. 22, fig. 3) is
<taxonomicName id="4C172BEEFFF3C056FBADFADD7D0A3255" authority="(Brocchi, 1814)" baseAuthorityName="Brocchi" baseAuthorityYear="1814" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="quadricarinata">
<emphasis id="B9638C7FFFF3C056FBADFADD794E3271" box="[1076,1356,1327,1353]" italics="true" pageId="50" pageNumber="51">Mathilda quadricarinata</emphasis>
(
<bibRefCitation id="EF862D9CFFF3C056FAC2FADD7D023255" author="Brocchi, G." pageId="50" pageNumber="51" refId="ref49699" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Stamperia reale, Milano, 2 + 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up]" type="book" year="1814">Brocchi, 1814</bibRefCitation>
)
</taxonomicName>
according to Sacchetti
<emphasis id="B9638C7FFFF3C056FD94FAA67E473255" box="[525,581,1363,1389]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
(2023)].The extant to Pliocene Mediterranean
<taxonomicName id="4C172BEEFFF3C056FBC8FAA17CCD32AA" authority="Semper, 1865" authorityName="Semper" authorityYear="1865" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="gemmulata">
<emphasis id="B9638C7FFFF3C056FBC8FAA179383255" box="[1105,1338,1363,1389]" italics="true" pageId="50" pageNumber="51">Mathilda gemmulata</emphasis>
Semper, 1865
</taxonomicName>
differs in its more slender shell, the more incised suture, the more pronounced mid-whorl angulation and the wider sutural ramp.
</paragraph>
<paragraph id="8BA8506DFFF3C056FF5EFA4D7CE23119" blockId="50.[151,1437,571,1749]" pageId="50" pageNumber="51">
<emphasis id="B9638C7FFFF3C056FF5EFA4D7DAA32E1" bold="true" box="[199,424,1471,1497]" pageId="50" pageNumber="51">Paleoenvironment.</emphasis>
Offshore clays in middle to outer neritic settings. At the Niederleis section, sediment and biota from coastal and lagoonal environments not exceeding
<quantity id="4CEFFD88FFF3C056FCCDFA117F9232C6" box="[852,912,1507,1534]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="m" value="30.0">30 m</quantity>
water depth were documented (
<bibRefCitation id="EF862D9CFFF3C056FA98FA117CD63119" author="Mandic, O. &amp; Harzhauser, M. &amp; Spezzaferri, S. &amp; Zuschin, M." pageId="50" pageNumber="51" pagination="193 - 206" refId="ref56609" refString="Mandic, O., Harzhauser, M., Spezzaferri, S. &amp; Zuschin, M. (2002) The paleoenvironment of an early Middle Miocene Paratethys sequence in NE Austria with special emphasis on paleoecology of mollusks and foraminifera. Geobios, Memoirs Special, 24, 193 - 206. https: // doi. org / 10.1016 / S 0016 - 6995 (02) 00059 - 1" type="journal article" year="2002">
Mandic
<emphasis id="B9638C7FFFF3C056FAFBFA16799F32C5" box="[1378,1437,1507,1533]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
2002
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFF3C056FF5EF9D97CE231ED" pageId="50" pageNumber="51" type="distribution">
<paragraph id="8BA8506DFFF3C056FF5EF9D97952317D" blockId="50.[151,1437,571,1749]" box="[199,1360,1579,1605]" pageId="50" pageNumber="51">
<emphasis id="B9638C7FFFF3C056FF5EF9D97D63317D" bold="true" box="[199,353,1579,1605]" pageId="50" pageNumber="51">Distribution.</emphasis>
Only known from the Langhian equivalent of the Badenian in the Central Paratethys Sea.
</paragraph>
<paragraph id="8BA8506DFFF3C056FF5EF9BD7CE231ED" blockId="50.[151,1437,571,1749]" pageId="50" pageNumber="51">
<emphasis id="B9638C7FFFF3C056FF5EF9BD7DE83151" bold="true" box="[199,490,1615,1641]" pageId="50" pageNumber="51">Central Paratethys Sea.</emphasis>
Badenian (Middle Miocene):
<emphasis id="B9638C7FFFF3C056FCD2F9A278073151" box="[843,1029,1616,1641]" italics="true" pageId="50" pageNumber="51">Korytnica Basin</emphasis>
: Korytnica (
<collectingCountry id="F30010FDFFF3C056FB00F9BD78E83151" box="[1177,1258,1615,1641]" name="Poland" pageId="50" pageNumber="51">Poland</collectingCountry>
) (
<bibRefCitation id="EF862D9CFFF3C056FA9DF9BD79923151" author="Baluk, W." box="[1284,1424,1615,1641]" pageId="50" pageNumber="51" pagination="177 - 220" refId="ref48759" refString="Baluk, W. (2006) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part V Addenda et corrigenda ad Prosobranchia. Acta Geologica Polonica, 56, 177 - 220. [https: // geojournals. pgi. gov. pl / agp / article / view / 9847 / 8381]" type="journal article" year="2006">Bałuk 2006</bibRefCitation>
);
<emphasis id="B9638C7FFFF3C056FF0EF9867D2931B5" box="[151,299,1652,1677]" italics="true" pageId="50" pageNumber="51">
<collectingRegion id="49D39E8FFFF3C056FF0EF9867CE631B5" box="[151,228,1652,1677]" country="Austria" name="Wien" pageId="50" pageNumber="51">Vienna</collectingRegion>
Basin
</emphasis>
: Niederleis (
<collectingCountry id="F30010FDFFF3C056FE23F9817E1031B5" box="[442,530,1651,1677]" name="Austria" pageId="50" pageNumber="51">Austria</collectingCountry>
), Sedlec (= Porzteich) (
<collectingCountry id="F30010FDFFF3C056FC8EF9817FCE31B5" box="[791,972,1651,1677]" name="Czech Republic" pageId="50" pageNumber="51">Czech Republic</collectingCountry>
) (
<emphasis id="B9638C7FFFF3C056FC7AF981784531B5" box="[995,1095,1651,1677]" italics="true" pageId="50" pageNumber="51">hoc opus</emphasis>
);
<emphasis id="B9638C7FFFF3C056FBC6F986792231B5" box="[1119,1312,1652,1677]" italics="true" pageId="50" pageNumber="51">Pannonian Basin</emphasis>
: Várpalota (
<collectingCountry id="F30010FDFFF3C056FF38F9657D073189" box="[161,261,1687,1713]" name="Hungary" pageId="50" pageNumber="51">Hungary</collectingCountry>
) (Kecskemétiné K̂rmendy 1962);
<emphasis id="B9638C7FFFF3C056FD09F96A7F1A3189" box="[656,792,1688,1714]" italics="true" pageId="50" pageNumber="51">Făget Basin</emphasis>
: Coşteiu de Sus, Lăpugiu de Sus (Semper 1865;
<bibRefCitation id="EF862D9CFFF3C056FAA5F96A7CD131ED" author="Boettger, O." pageId="50" pageNumber="51" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger 1902</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>