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<mods:title id="98C2EBC476023A975F88E5589868D755">The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition</mods:title>
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<mods:affiliation id="3B2B5B772B25D256DF7698B38E4441B4">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655; &amp; Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal. bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494 &amp; Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;</mods:affiliation>
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<heading id="D0E0E701FFE8C04DFF0EFA597E5B32FD" bold="true" box="[151,601,1451,1478]" fontSize="11" level="2" pageId="41" pageNumber="42" reason="4">
<taxonomicName id="4C172BEEFFE8C04DFF0EFA597E5B32FD" ID-CoL="BXTCR" authority="(Boettger, 1902)" authorityName="" baseAuthorityName="Boettger" baseAuthorityYear="1902" box="[151,601,1451,1478]" class="Gastropoda" family="Architectonicidae" genus="Spirolaxis" kingdom="Animalia" order="Heterostropha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="cornicula">
<emphasis id="B9638C7FFFE8C04DFF0EFA597E5B32FD" bold="true" box="[151,601,1451,1478]" pageId="41" pageNumber="42">
<emphasis id="B9638C7FFFE8C04DFF0EFA597D8F32FE" bold="true" box="[151,397,1451,1478]" italics="true" pageId="41" pageNumber="42">Spirolaxis cornicula</emphasis>
(
<bibRefCitation id="EF862D9CFFE8C04DFE07FA5E7E5332FD" author="Boettger, O." box="[414,593,1451,1478]" pageId="41" pageNumber="42" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger, 1902</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C30D03E6FFE8C04DFF0EFA227D1D32D2" box="[151,287,1488,1514]" pageId="41" pageNumber="42" type="description">
<paragraph id="8BA8506DFFE8C04DFF0EFA227D1D32D2" blockId="41.[151,601,1451,1514]" box="[151,287,1488,1514]" pageId="41" pageNumber="42">
<figureCitation id="132C4CE8FFE8C04DFF0EFA227D1D32D2" box="[151,287,1488,1514]" captionStart="FIGURE 21" captionStartId="42.[151,250,1833,1857]" captionTargetBox="[275,1333,257,1797]" captionTargetId="figure-30@42.[275,1333,257,1797]" captionTargetPageId="42" captionText="FIGURE 21 A1A3. Spirolaxis cornicula (Boettger, 1902), lectotype, SMF 372697, Coşteiu de Sus (Romania). B1B3. Spirolaxis cornicula (Boettger, 1902), holotype of Discohelix quinquangularis Boettger, 1902, SMF 372694, Lăpugiu de Sus (Romania). C1C2. Spirolaxis cornicula (Boettger, 1902), NHMW 2016/0177/0342, Lăpugiu des Sus (Romania)." figureDoi="http://doi.org/10.5281/zenodo.10147854" httpUri="https://zenodo.org/record/10147854/files/figure.png" pageId="41" pageNumber="42">Figs 21AC</figureCitation>
</paragraph>
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<paragraph id="8BA8506DFFE8C04DFF0EF9FD7F62311F" blockId="41.[151,1186,1551,1943]" box="[151,864,1551,1575]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="AB077743FFE8C04DFF0EF9FD7F62311F" box="[151,864,1551,1575]" pageId="41" pageNumber="42">
<emphasis id="B9638C7FFFE8C04DFF0EF9FD7CA6311E" box="[151,164,1551,1574]" italics="true" pageId="41" pageNumber="42">*</emphasis>
<taxonomicName id="4C172BEEFFE8C04DFF33F9FD7E42311F" authority="Harzhauser &amp; Landau, 2023" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[170,576,1551,1575]" class="Gastropoda" family="Discohelicidae" genus="Discohelix" kingdom="Animalia" order="Seguenziida" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="corniculum" status="sp. nov." subGenus="Pseudomalaxis">
<emphasis id="B9638C7FFFE8C04DFF33F9FD7D14311F" box="[170,278,1551,1575]" italics="true" pageId="41" pageNumber="42">Discohelix</emphasis>
(
<emphasis id="B9638C7FFFE8C04DFEBCF9FD7DBC311F" box="[293,446,1551,1575]" italics="true" pageId="41" pageNumber="42">Pseudomalaxis</emphasis>
)
<emphasis id="B9638C7FFFE8C04DFE54F9FD7E42311F" box="[461,576,1551,1575]" italics="true" pageId="41" pageNumber="42">corniculum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2503104FFE8C04DFDDEF9FD7E78311F" box="[583,634,1551,1575]" pageId="41" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
<treatmentCitation id="0AB6767CFFE8C04DFD0DF9FD7F5E311F" author="Boettger, O." box="[660,860,1551,1575]" page="116" pageId="41" pageNumber="42" year="1902">
<bibRefCitation id="EF862D9CFFE8C04DFD0DF9FD7F5E311F" author="Boettger, O." box="[660,860,1551,1575]" pageId="41" pageNumber="42" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger 1902: 116</bibRefCitation>
</treatmentCitation>
.
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<paragraph id="8BA8506DFFE8C04DFF0EF9DF7ED3317D" blockId="41.[151,1186,1551,1943]" box="[151,721,1581,1606]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="AB077743FFE8C04DFF0EF9DF7ED3317D" box="[151,721,1581,1606]" pageId="41" pageNumber="42">
<taxonomicName id="4C172BEEFFE8C04DFF0EF9DF7DB0317D" authority="Harzhauser &amp; Landau, 2023" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[151,434,1581,1605]" class="Gastropoda" family="Discohelicidae" genus="Discohelix" kingdom="Animalia" order="Seguenziida" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="quinquangularis" status="sp. nov.">
<emphasis id="B9638C7FFFE8C04DFF0EF9DF7DB0317D" box="[151,434,1581,1605]" italics="true" pageId="41" pageNumber="42">Discohelix quinquangularis</emphasis>
</taxonomicName>
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<treatmentCitation id="0AB6767CFFE8C04DFD9DF9DC7ECF317D" author="Boettger, O." box="[516,717,1581,1606]" page="116" pageId="41" pageNumber="42" year="1902">
<bibRefCitation id="EF862D9CFFE8C04DFD9DF9DC7ECF317D" author="Boettger, O." box="[516,717,1581,1606]" pageId="41" pageNumber="42" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger 1902: 116</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BA8506DFFE8C04DFF0EF9BE7F6D315C" blockId="41.[151,1186,1551,1943]" box="[151,879,1612,1637]" pageId="41" pageNumber="42">
<taxonomicName id="4C172BEEFFE8C04DFF0EF9BE7E89315D" authority="Boettger" authorityName="Boettger" authorityYear="1902" box="[151,651,1612,1637]" class="Gastropoda" family="Discohelicidae" genus="Discohelix" kingdom="Animalia" order="Seguenziida" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="corniculum" subGenus="Pseudomalaxis">
<emphasis id="B9638C7FFFE8C04DFF0EF9BE7D01315C" box="[151,259,1612,1636]" italics="true" pageId="41" pageNumber="42">Discohelix</emphasis>
(
<emphasis id="B9638C7FFFE8C04DFE8BF9BE7DA9315C" box="[274,427,1612,1636]" italics="true" pageId="41" pageNumber="42">Pseudomalaxis</emphasis>
)
<emphasis id="B9638C7FFFE8C04DFE23F9BE7E2F315C" box="[442,557,1612,1636]" italics="true" pageId="41" pageNumber="42">corniculum</emphasis>
Boettger
</taxonomicName>
—Boettger 1907: 138.
</paragraph>
<paragraph id="8BA8506DFFE8C04DFF0EF9997FCC31BB" blockId="41.[151,1186,1551,1943]" box="[151,974,1643,1667]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="AB077743FFE8C04DFF0EF9997FCC31BB" box="[151,974,1643,1667]" pageId="41" pageNumber="42">
<taxonomicName id="4C172BEEFFE8C04DFF0EF9997E1531BB" ID-CoL="BXTJV" authority="(Boettger)" baseAuthorityName="O. Boettger" baseAuthorityYear="1902" box="[151,535,1643,1667]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="corniculum">
<emphasis id="B9638C7FFFE8C04DFF0EF9997DA831BB" box="[151,426,1643,1667]" italics="true" pageId="41" pageNumber="42">Pseudomalaxis corniculum</emphasis>
(Boettger)
</taxonomicName>
<treatmentCitation id="0AB6767CFFE8C04DFDA8F9997F0831BB" author="Cossmann M." box="[561,778,1643,1667]" page="143" pageId="41" pageNumber="42" year="1916">
<bibRefCitation id="EF862D9CFFE8C04DFDA8F9997F0831BB" author="Cossmann M." box="[561,778,1643,1667]" pageId="41" pageNumber="42" refId="ref50491" refString="Cossmann M. (1916) Essais de paleoconchologie comparee. Dixieme livraison. Paris, published by the author. 292 pp., 12 pls. [&quot; 1915 &quot;; 1916, https: // www. biodiversitylibrary. org / item / 111980]" type="book" year="1916">Cossmann 1916: 143</bibRefCitation>
</treatmentCitation>
, pl. 12, figs 1921.
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</paragraph>
<paragraph id="8BA8506DFFE8C04DFF0EF97B7F8F319A" blockId="41.[151,1186,1551,1943]" box="[151,909,1673,1698]" pageId="41" pageNumber="42">
<taxonomicName id="4C172BEEFFE8C04DFF0EF97B7E153199" authority="(Boettger)" baseAuthorityName="O. Boettger" baseAuthorityYear="1902" box="[151,535,1673,1697]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="corniculum">
<emphasis id="B9638C7FFFE8C04DFF0EF97B7DA83199" box="[151,426,1673,1697]" italics="true" pageId="41" pageNumber="42">Pseudomalaxis corniculum</emphasis>
(Boettger)
</taxonomicName>
—Zilch 1934: 218, pl. 7, figs 1920.
</paragraph>
<paragraph id="8BA8506DFFE8C04DFF0EF95A7FC231F8" blockId="41.[151,1186,1551,1943]" box="[151,960,1704,1728]" pageId="41" pageNumber="42">
<taxonomicName id="4C172BEEFFE8C04DFF0EF95A7E4931F8" authority="(Boettger)" baseAuthorityName="O. Boettger" baseAuthorityYear="1902" box="[151,587,1704,1728]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="quinquangularis">
<emphasis id="B9638C7FFFE8C04DFF0EF95A7DDD31F8" box="[151,479,1704,1728]" italics="true" pageId="41" pageNumber="42">Pseudomalaxis quinquangularis</emphasis>
(Boettger)
</taxonomicName>
—Zilch 1934: 219, pl. 7, figs 21ab.
</paragraph>
<paragraph id="8BA8506DFFE8C04DFF0EF93578A031E7" blockId="41.[151,1186,1551,1943]" box="[151,1186,1735,1759]" pageId="41" pageNumber="42">
<taxonomicName id="4C172BEEFFE8C04DFF0EF9357E8831E7" authority="Boettger" authorityName="Boettger" box="[151,650,1735,1759]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="corniculum" subGenus="Discosolix">
<emphasis id="B9638C7FFFE8C04DFF0EF9357D3231E7" box="[151,304,1735,1759]" italics="true" pageId="41" pageNumber="42">Pseudomalaxis</emphasis>
(
<emphasis id="B9638C7FFFE8C04DFEA6F9357DA831E7" box="[319,426,1735,1759]" italics="true" pageId="41" pageNumber="42">Discosolix</emphasis>
)
<emphasis id="B9638C7FFFE8C04DFE20F9357E2E31E7" box="[441,556,1735,1759]" italics="true" pageId="41" pageNumber="42">corniculum</emphasis>
Boettger
</taxonomicName>
<bibRefCitation id="EF862D9CFFE8C04DFD3AF9357F4431E7" author="Korobkov, I. A." box="[675,838,1735,1759]" pageId="41" pageNumber="42" refId="ref55459" refString="Korobkov, I. A. (1955) Spravochnik i metodicheskoe rukovodstvo po tretichnym mollyuskam. Bryukhonogie. Gostoptekhizdat, Leningrad. 795 pp. [Koрobkov, I. A. (1955) СpрavocнiK i мitodiciсKoi рyKovodсtvo po tрiticным мoЛЛюсKaм. Bрюkoнogii. GoсtoptikiZDat, ЛiнiнgрaD, 795 pp.]" type="book" year="1955">Korobkov 1955</bibRefCitation>
: plate captions, pl. 13, figs 2628.
</paragraph>
<paragraph id="8BA8506DFFE8C04DFF0EF9177F7131C6" blockId="41.[151,1186,1551,1943]" box="[151,883,1765,1790]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="AB077743FFE8C04DFF0EF9177F7131C6" box="[151,883,1765,1790]" pageId="41" pageNumber="42">
<taxonomicName id="4C172BEEFFE8C04DFF0EF9177E0A31C5" ID-CoL="BXTJV" authority="Boettger" authorityName="Boettger" box="[151,520,1765,1789]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="corniculum">
<emphasis id="B9638C7FFFE8C04DFF0EF9177DA831C5" box="[151,426,1765,1789]" italics="true" pageId="41" pageNumber="42">Pseudomalaxis corniculum</emphasis>
Boettger
</taxonomicName>
<treatmentCitation id="0AB6767CFFE8C04DFDBBF9177F6D31C6" author="Csepreghy-Meznerics, I." box="[546,879,1765,1790]" page="431" pageId="41" pageNumber="42" year="1956">
<bibRefCitation id="EF862D9CFFE8C04DFDBBF9177F6D31C6" author="Csepreghy-Meznerics, I." box="[546,879,1765,1790]" pageId="41" pageNumber="42" pagination="361 - 477" refId="ref50934" refString="Csepreghy-Meznerics, I. (1956) Die Molluskenfauna von Szob und Letkes. A magyar allami f ˆ ldtani intezet evk ˆ nyve, 45, 361 - 477. [https: // epa. oszk. hu / 03200 / 03274 / 00094 / pdf / EPA 03274 _ mafi _ evkonyv _ 1956 _ 02 _ 363 - 442. pdf and https: // epa. oszk. hu / 03200 / 03274 / 00094 / pdf / EPA 03274 _ mafi _ evkonyv _ 1956 _ 02 _ 450 - 477. pdf]" type="journal article" year="1956">Csepreghy-Meznerics 1956: 431</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA8506DFFE8C04DFF0EF8F67FB03024" blockId="41.[151,1186,1551,1943]" box="[151,946,1796,1821]" pageId="41" pageNumber="42">
<taxonomicName id="4C172BEEFFE8C04DFF0EF8F67E413024" authority="Boettger, 1901" authorityName="Boettger" authorityYear="1901" box="[151,579,1796,1821]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="corniculum">
<emphasis id="B9638C7FFFE8C04DFF0EF8F67DA83024" box="[151,426,1796,1820]" italics="true" pageId="41" pageNumber="42">Pseudomalaxis corniculum</emphasis>
Boettger, 1901
</taxonomicName>
—Strausz 1966: 119, pl. 57, fig. 20.
</paragraph>
<paragraph id="8BA8506DFFE8C04DFF0EF8D178363003" blockId="41.[151,1186,1551,1943]" box="[151,1076,1827,1851]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="AB077743FFE8C04DFF0EF8D178363003" box="[151,1076,1827,1851]" pageId="41" pageNumber="42">
<taxonomicName id="4C172BEEFFE8C04DFF0EF8D17EE93003" ID-CoL="BXTJV" authority="(O. Boettger 1902)" baseAuthorityName="O. Boettger" baseAuthorityYear="1902" box="[151,747,1827,1851]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="corniculum" subGenus="Spirolaxis">
<emphasis id="B9638C7FFFE8C04DFF0EF8D17D323003" box="[151,304,1827,1851]" italics="true" pageId="41" pageNumber="42">Pseudomalaxis</emphasis>
(
<emphasis id="B9638C7FFFE8C04DFEA6F8D17DA73003" box="[319,421,1827,1851]" italics="true" pageId="41" pageNumber="42">Spirolaxis</emphasis>
)
<emphasis id="B9638C7FFFE8C04DFE2DF8D17E253003" box="[436,551,1827,1851]" italics="true" pageId="41" pageNumber="42">corniculum</emphasis>
(O.
<bibRefCitation id="EF862D9CFFE8C04DFDCCF8D17EE73003" author="Boettger, O." box="[597,741,1827,1851]" pageId="41" pageNumber="42" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger 1902</bibRefCitation>
)
</taxonomicName>
<treatmentCitation id="0AB6767CFFE8C04DFC9FF8D17FA53002" author="Bieler, R." box="[774,935,1827,1851]" page="77" pageId="41" pageNumber="42" year="1984">
<bibRefCitation id="EF862D9CFFE8C04DFC9FF8D17FA53002" author="Bieler, R." box="[774,935,1827,1851]" pageId="41" pageNumber="42" pagination="53 - 103" refId="ref48969" refString="Bieler, R. (1984) Die Gattungen der Architectonicidae (Gastropoda: &quot; Heterogastropoda &quot;). Allgemeines und Teil 1: Pseudomalaxis. Archiv f ¸ r Molluskenkunde, 115, 53 - 103." type="journal article" year="1984">Bieler 1984: 77</bibRefCitation>
</treatmentCitation>
, pl. 3, fig. 21.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA8506DFFE8C04DFF0EF8B3786A3062" blockId="41.[151,1186,1551,1943]" box="[151,1128,1857,1882]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="AB077743FFE8C04DFF0EF8B3786A3062" box="[151,1128,1857,1882]" pageId="41" pageNumber="42">
<taxonomicName id="4C172BEEFFE8C04DFF0EF8B37F1D3061" authority="(O. Boettger 1902)" baseAuthorityName="O. Boettger" baseAuthorityYear="1902" box="[151,799,1857,1882]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="quinquangularis" subGenus="Spirolaxis">
<emphasis id="B9638C7FFFE8C04DFF0EF8B37D323061" box="[151,304,1857,1881]" italics="true" pageId="41" pageNumber="42">Pseudomalaxis</emphasis>
(
<emphasis id="B9638C7FFFE8C04DFEA6F8B37DA73061" box="[319,421,1857,1881]" italics="true" pageId="41" pageNumber="42">Spirolaxis</emphasis>
)
<emphasis id="B9638C7FFFE8C04DFE2DF8B37E5E3061" box="[436,604,1857,1881]" italics="true" pageId="41" pageNumber="42">quinquangularis</emphasis>
(O.
<bibRefCitation id="EF862D9CFFE8C04DFD10F8B07F1B3061" author="Boettger, O." box="[649,793,1857,1882]" pageId="41" pageNumber="42" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger 1902</bibRefCitation>
)
</taxonomicName>
<treatmentCitation id="0AB6767CFFE8C04DFCA3F8B37FD93061" author="Bieler, R." box="[826,987,1857,1882]" page="77" pageId="41" pageNumber="42" year="1984">
<bibRefCitation id="EF862D9CFFE8C04DFCA3F8B37FD93061" author="Bieler, R." box="[826,987,1857,1882]" pageId="41" pageNumber="42" pagination="53 - 103" refId="ref48969" refString="Bieler, R. (1984) Die Gattungen der Architectonicidae (Gastropoda: &quot; Heterogastropoda &quot;). Allgemeines und Teil 1: Pseudomalaxis. Archiv f ¸ r Molluskenkunde, 115, 53 - 103." type="journal article" year="1984">Bieler 1984: 77</bibRefCitation>
</treatmentCitation>
, pl. 3, fig. 22.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA8506DFFE8C04DFF0EF8927F353040" blockId="41.[151,1186,1551,1943]" box="[151,823,1888,1913]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="AB077743FFE8C04DFF0EF8927F353040" box="[151,823,1888,1913]" pageId="41" pageNumber="42">
<taxonomicName id="4C172BEEFFE8C04DFF0EF8927E2A3040" ID-CoL="7P335" authorityName="Dunker" authorityYear="1847" baseAuthorityName="O. Boettger" baseAuthorityYear="1902" box="[151,552,1888,1913]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="corniculum">
<emphasis id="B9638C7FFFE8C04DFF0EF8927D083040" box="[151,266,1888,1912]" italics="true" pageId="41" pageNumber="42">corniculum</emphasis>
(
<emphasis id="B9638C7FFFE8C04DFE80F8927D873040" box="[281,389,1888,1912]" italics="true" pageId="41" pageNumber="42">Discohelix</emphasis>
)
<bibRefCitation id="EF862D9CFFE8C04DFE0DF8937E2A3040" author="Boettger, O." box="[404,552,1888,1913]" pageId="41" pageNumber="42" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger, 1902</bibRefCitation>
</taxonomicName>
<treatmentCitation id="0AB6767CFFE8C04DFDDAF8927F303040" author="Bieler, R. &amp; Petit, R. E." box="[579,818,1888,1912]" page="30" pageId="41" pageNumber="42" year="2005">
<bibRefCitation id="EF862D9CFFE8C04DFDDAF8927F303040" author="Bieler, R. &amp; Petit, R. E." box="[579,818,1888,1912]" pageId="41" pageNumber="42" pagination="1 - 119" refId="ref49237" refString="Bieler, R. &amp; Petit, R. E. (2005) Catalogue of Recent and fossil taxa of the family Architectonicidae Gray, 1850 (Mollusca: Gastropoda). Zootaxa, 1101 (1), 1 - 119. https: // doi. org / 10.11646 / zootaxa. 1101.1.1" type="journal article" year="2005">Bieler &amp; Petit 2005: 30</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA8506DFFE8C04DFF0EF88D7F6930AF" blockId="41.[151,1186,1551,1943]" box="[151,875,1919,1943]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="AB077743FFE8C04DFF0EF88D7F6930AF" box="[151,875,1919,1943]" pageId="41" pageNumber="42">
<taxonomicName id="4C172BEEFFE8C04DFF0EF88D7DBB30AF" ID-CoL="7P335" authorityName="Dunker" authorityYear="1847" baseAuthorityName="O. Boettger" baseAuthorityYear="1902" box="[151,441,1919,1943]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="quinquangularis">
<emphasis id="B9638C7FFFE8C04DFF0EF88D7D3D30AF" box="[151,319,1919,1943]" italics="true" pageId="41" pageNumber="42">quinquangularis</emphasis>
(
<emphasis id="B9638C7FFFE8C04DFED4F88D7DBB30AF" box="[333,441,1919,1943]" italics="true" pageId="41" pageNumber="42">Discohelix</emphasis>
</taxonomicName>
)
<bibRefCitation id="EF862D9CFFE8C04DFE50F88D7E5E30AF" author="Boettger, O." box="[457,604,1919,1943]" pageId="41" pageNumber="42" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger, 1902</bibRefCitation>
<treatmentCitation id="0AB6767CFFE8C04DFDEEF88D7F6530AF" author="Bieler, R. &amp; Petit, R. E." box="[631,871,1919,1943]" page="60" pageId="41" pageNumber="42" year="2005">
<bibRefCitation id="EF862D9CFFE8C04DFDEEF88D7F6530AF" author="Bieler, R. &amp; Petit, R. E." box="[631,871,1919,1943]" pageId="41" pageNumber="42" pagination="1 - 119" refId="ref49237" refString="Bieler, R. &amp; Petit, R. E. (2005) Catalogue of Recent and fossil taxa of the family Architectonicidae Gray, 1850 (Mollusca: Gastropoda). Zootaxa, 1101 (1), 1 - 119. https: // doi. org / 10.11646 / zootaxa. 1101.1.1" type="journal article" year="2005">Bieler &amp; Petit 2005: 60</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA8506DFFE8C04DFF0EF84F7D0B30CB" blockId="41.[151,1436,1980,2035]" pageId="41" pageNumber="42">
<emphasis id="B9638C7FFFE8C04DFF0EF84F7CBF30EC" box="[151,189,1981,2004]" italics="true" pageId="41" pageNumber="42">non</emphasis>
<taxonomicName id="4C172BEEFFE8C04DFF5AF84E799E30EC" authority="(Boettger)" authorityName="Jaume &amp; Borro" authorityYear="1946" baseAuthorityName="Boettger" box="[195,1436,1980,2005]" class="Gastropoda" family="Discohelicidae" genus="Discohelix" kingdom="Animalia" order="Seguenziida" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="corniculum" subGenus="Pseudomalaxis">
<emphasis id="B9638C7FFFE8C04DFF5AF84E7D2D30EC" box="[195,303,1980,2004]" italics="true" pageId="41" pageNumber="42">Discohelix</emphasis>
(
<emphasis id="B9638C7FFFE8C04DFEA4F84E7DD430EC" box="[317,470,1980,2004]" italics="true" pageId="41" pageNumber="42">Pseudomalaxis</emphasis>
)
<emphasis id="B9638C7FFFE8C04DFE7CF84E7E5A30EC" box="[485,600,1980,2004]" italics="true" pageId="41" pageNumber="42">corniculum</emphasis>
(Boettger)—Hornung 1920: 85, pl. 2, fig. 15 [=
<emphasis id="B9638C7FFFE8C04DFBD9F84E78F730EC" box="[1088,1269,1980,2004]" italics="true" pageId="41" pageNumber="42">Spirolaxis clenchi</emphasis>
<bibRefCitation id="EF862D9CFFE8C04DFB62F84F799E30EC" author="Jaume, M. L. &amp; Borro, P." box="[1275,1436,1980,2004]" pageId="41" pageNumber="42" pagination="13 - 22" refId="ref55028" refString="Jaume, M. L. &amp; Borro, P. (1946) Novedades en moluscos marinos cubanos. Revista de la sociedad malaceologica &quot; Carlos de la Torre &quot;, 4 (1), 13 - 22." type="journal article" year="1946">Jaume &amp; Borro,</bibRefCitation>
</taxonomicName>
1946].
</paragraph>
</subSubSection>
<caption id="DF6800E5FFEBC04EFF0EF8DB789A30B1" ID-DOI="http://doi.org/10.5281/zenodo.10147854" ID-Zenodo-Dep="10147854" httpUri="https://zenodo.org/record/10147854/files/figure.png" pageId="42" pageNumber="43" startId="42.[151,250,1833,1857]" targetBox="[275,1333,257,1797]" targetPageId="42" targetType="figure">
<paragraph id="8BA8506DFFEBC04EFF0EF8DB799F305D" blockId="42.[151,1437,1833,1935]" pageId="42" pageNumber="43">
<emphasis id="B9638C7FFFEBC04EFF0EF8DB7D603079" bold="true" box="[151,354,1833,1863]" pageId="42" pageNumber="43">
FIGURE 21 A
<subScript id="17935228FFEBC04EFEB6F8CB7D34307F" attach="right" box="[303,310,1849,1863]" fontSize="6" pageId="42" pageNumber="43">1</subScript>
A
<subScript id="17935228FFEBC04EFECCF8CB7D5E307F" attach="left" box="[341,348,1849,1863]" fontSize="6" pageId="42" pageNumber="43">3</subScript>
.
</emphasis>
<taxonomicName id="4C172BEEFFEBC04EFEF1F8DB7EE33079" authority="(Boettger, 1902)" authorityName="" baseAuthorityName="Boettger" baseAuthorityYear="1902" box="[360,737,1833,1857]" class="Gastropoda" family="Architectonicidae" genus="Spirolaxis" kingdom="Animalia" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="cornicula">
<emphasis id="B9638C7FFFEBC04EFEF1F8DB7E363079" box="[360,564,1833,1857]" italics="true" pageId="42" pageNumber="43">Spirolaxis cornicula</emphasis>
(
<bibRefCitation id="EF862D9CFFEBC04EFDD8F8DB7ED83079" author="Boettger, O." box="[577,730,1833,1857]" pageId="42" pageNumber="43" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger, 1902</bibRefCitation>
)
</taxonomicName>
, lectotype, SMF 372697, Coşteiu de Sus (Romania).
<emphasis id="B9638C7FFFEBC04EFB6FF8DB793B3079" bold="true" box="[1270,1337,1833,1863]" pageId="42" pageNumber="43">
B
<subScript id="17935228FFEBC04EFA9EF8CB790C307F" attach="left" box="[1287,1294,1849,1863]" fontSize="6" pageId="42" pageNumber="43">1</subScript>
B
<subScript id="17935228FFEBC04EFAB5F8CB7931307F" attach="left" box="[1324,1331,1849,1863]" fontSize="6" pageId="42" pageNumber="43">3</subScript>
.
</emphasis>
<taxonomicName id="4C172BEEFFEBC04EFAA6F8DB7DB8305D" authority="(Boettger, 1902)" authorityName="" baseAuthorityName="Boettger" baseAuthorityYear="1902" class="Gastropoda" family="Architectonicidae" genus="Spirolaxis" kingdom="Animalia" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="cornicula">
<emphasis id="B9638C7FFFEBC04EFAA6F8DB7D0C305D" italics="true" pageId="42" pageNumber="43">Spirolaxis cornicula</emphasis>
(
<bibRefCitation id="EF862D9CFFEBC04EFE83F8BF7DB6305D" author="Boettger, O." box="[282,436,1869,1893]" pageId="42" pageNumber="43" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger, 1902</bibRefCitation>
)
</taxonomicName>
, holotype of
<taxonomicName id="4C172BEEFFEBC04EFDA5F8BF7FF3305D" authority="Boettger, 1902" authorityName="Boettger" authorityYear="1902" box="[572,1009,1869,1894]" class="Gastropoda" family="Discohelicidae" genus="Discohelix" kingdom="Animalia" order="Seguenziida" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="quinquangularis">
<emphasis id="B9638C7FFFEBC04EFDA5F8BF7F54305D" box="[572,854,1869,1893]" italics="true" pageId="42" pageNumber="43">Discohelix quinquangularis</emphasis>
<bibRefCitation id="EF862D9CFFEBC04EFCC3F8BC7FF3305D" author="Boettger, O." box="[858,1009,1869,1894]" pageId="42" pageNumber="43" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger, 1902</bibRefCitation>
</taxonomicName>
, SMF 372694, Lăpugiu de Sus (Romania).
</paragraph>
<paragraph id="8BA8506DFFEBC04EFF0EF883789A30B1" blockId="42.[151,1437,1833,1935]" box="[151,1176,1905,1935]" pageId="42" pageNumber="43">
<emphasis id="B9638C7FFFEBC04EFF0EF8837CDF30B1" bold="true" box="[151,221,1905,1935]" pageId="42" pageNumber="43">
C
<subScript id="17935228FFEBC04EFF30F8737CB230B7" attach="left" box="[169,176,1921,1935]" fontSize="6" pageId="42" pageNumber="43">1</subScript>
C
<subScript id="17935228FFEBC04EFF49F8737CD530B7" attach="right" box="[208,215,1921,1935]" fontSize="6" pageId="42" pageNumber="43">2</subScript>
.
</emphasis>
<taxonomicName id="4C172BEEFFEBC04EFF7DF8837E6230B1" authority="(Boettger, 1902)" authorityName="" baseAuthorityName="Boettger" baseAuthorityYear="1902" box="[228,608,1905,1929]" class="Gastropoda" family="Architectonicidae" genus="Spirolaxis" kingdom="Animalia" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="cornicula">
<emphasis id="B9638C7FFFEBC04EFF7DF8837DB330B1" box="[228,433,1905,1929]" italics="true" pageId="42" pageNumber="43">Spirolaxis cornicula</emphasis>
(
<bibRefCitation id="EF862D9CFFEBC04EFE27F8837E5B30B1" author="Boettger, O." box="[446,601,1905,1929]" pageId="42" pageNumber="43" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger, 1902</bibRefCitation>
)
</taxonomicName>
, NHMW 2016/0177/0342, Lăpugiu des Sus (Romania).
</paragraph>
</caption>
<subSubSection id="C30D03E6FFEAC04FFF0EFF657DD8342D" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8BA8506DFFEAC04FFF0EFF657D6E37C1" blockId="43.[151,1437,150,1689]" pageId="43" pageNumber="44">
<emphasis id="B9638C7FFFEAC04FFF0EFF657D443788" bold="true" box="[151,326,150,177]" pageId="43" pageNumber="44">Type material.</emphasis>
<materialsCitation id="3B7F5A30FFEAC04FFED4FF657D6E37C1" collectionCode="SMF" collectorName="de Sus" country="Germany" location="Costeiu de Sus" pageId="43" pageNumber="44" specimenCode="SMF 372697" specimenCount="1" typeStatus="lectotype">
<typeStatus id="54ACEECFFFEAC04FFED4FF657DBF3789" box="[333,445,151,177]" pageId="43" pageNumber="44" type="lectotype">Lectotype</typeStatus>
(designated by Zilch 1934: 218):
<specimenCode id="DBB1F816FFEAC04FFCA7FF657FD03789" box="[830,978,151,177]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="43" pageNumber="44">SMF 372697</specimenCode>
(XII 2373a), SL:
<quantity id="4CEFFD88FFEAC04FFB07FF6578F43789" box="[1182,1270,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="43" pageNumber="44" unit="mm" value="0.7">0.7 mm</quantity>
, MD:
<quantity id="4CEFFD88FFEAC04FFAA6FF65799A3789" box="[1343,1432,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="43" pageNumber="44" unit="mm" value="2.8">2.8 mm</quantity>
, Senckenberg Museum, Frankfurt/Main, Germany,
<location id="8EC806B6FFEAC04FFD4BFF497F7837ED" LSID="urn:lsid:plazi:treatment:03BEE17BFFE8C04FFF0EFA5978A231A1:8EC806B6FFEAC04FFD4BFF497F7837ED" box="[722,890,187,213]" country="Germany" name="Costeiu de Sus" pageId="43" pageNumber="44">
Coşteiu
<collectorName id="26E235BBFFEAC04FFCA9FF497F7837ED" box="[816,890,187,213]" pageId="43" pageNumber="44">de Sus</collectorName>
</location>
(
<collectingCountry id="F30010FDFFEAC04FFC13FF497FF737ED" box="[906,1013,187,213]" name="Romania" pageId="43" pageNumber="44">Romania</collectingCountry>
), illustrated in Zilch (1934: pl. 7, fig. 19),
<figureCitation id="132C4CE8FFEAC04FFF52FF2D7D3037C1" box="[203,306,223,249]" captionStart="FIGURE 21" captionStartId="42.[151,250,1833,1857]" captionTargetBox="[275,1333,257,1797]" captionTargetId="figure-30@42.[275,1333,257,1797]" captionTargetPageId="42" captionText="FIGURE 21 A1A3. Spirolaxis cornicula (Boettger, 1902), lectotype, SMF 372697, Coşteiu de Sus (Romania). B1B3. Spirolaxis cornicula (Boettger, 1902), holotype of Discohelix quinquangularis Boettger, 1902, SMF 372694, Lăpugiu de Sus (Romania). C1C2. Spirolaxis cornicula (Boettger, 1902), NHMW 2016/0177/0342, Lăpugiu des Sus (Romania)." figureDoi="http://doi.org/10.5281/zenodo.10147854" httpUri="https://zenodo.org/record/10147854/files/figure.png" pageId="43" pageNumber="44">Figs 21A</figureCitation>
<figureCitation id="132C4CE8FFEAC04FFEABFF037D5F37C1" box="[306,349,223,255]" captionStart="FIGURE 1" captionStartId="5.[151,250,1522,1546]" captionTargetBox="[232,1366,234,1455]" captionTargetId="figure-58@5.[232,1366,234,1455]" captionTargetPageId="5" captionText="FIGURE 1. Descriptive terms used in descriptions of Architectonicidae (modified from Bieler 1993)." figureDoi="http://doi.org/10.5281/zenodo.10147816" httpUri="https://zenodo.org/record/10147816/files/figure.png" pageId="43" pageNumber="44">
<subScript id="17935228FFEAC04FFEABFF037D3837C7" attach="left" box="[306,314,241,255]" fontSize="6" pageId="43" pageNumber="44">1</subScript>
A
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<figureCitation id="132C4CE8FFEAC04FFEC5FF037D6637C7" box="[348,356,241,255]" captionStart="FIGURE 3" captionStartId="6.[151,250,2002,2026]" captionTargetId="figure-99@6.[337,1250,1242,1978]" captionTargetPageId="6" captionText="FIGURE 3. Descriptive terms used in descriptions of Mathildidae." figureDoi="http://doi.org/10.5281/zenodo.10307672" httpUri="https://zenodo.org/record/10307672/files/figure.png" pageId="43" pageNumber="44">
<subScript id="17935228FFEAC04FFEC5FF037D6637C7" attach="left" box="[348,356,241,255]" fontSize="6" pageId="43" pageNumber="44">3</subScript>
</figureCitation>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA8506DFFEAC04FFF5EFEF17DB936B1" blockId="43.[151,1437,150,1689]" pageId="43" pageNumber="44">
<emphasis id="B9638C7FFFEAC04FFF5EFEF17DB93624" bold="true" box="[199,443,258,285]" pageId="43" pageNumber="44">Illustrated material:</emphasis>
<materialsCitation id="3B7F5A30FFEAC04FFE5BFEF17E70365D" collectionCode="SMF, NHMW" collectorName="de Sus" country="Germany" location="Lapugiu de Sus" municipality="Lapugiu des Sus" pageId="43" pageNumber="44" specimenCode="SMF 372694, NHMW 2016" specimenCount="1" typeStatus="holotype">
<typeStatus id="54ACEECFFFEAC04FFE5BFEF17E283625" box="[450,554,259,285]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C172BEEFFEAC04FFDC8FEF1783B3625" authority="Boettger, 1902" authorityName="Boettger" authorityYear="1902" box="[593,1081,259,285]" class="Gastropoda" family="Discohelicidae" genus="Discohelix" kingdom="Animalia" order="Seguenziida" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="quinquangularis">
<emphasis id="B9638C7FFFEAC04FFDC8FEF17F8B3625" box="[593,905,259,285]" italics="true" pageId="43" pageNumber="44">Discohelix quinquangularis</emphasis>
<bibRefCitation id="EF862D9CFFEAC04FFC08FEF1783B3625" author="Boettger, O." box="[913,1081,259,285]" pageId="43" pageNumber="44" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger, 1902</bibRefCitation>
</taxonomicName>
,
<specimenCode id="DBB1F816FFEAC04FFBDCFEF178DA3625" box="[1093,1240,259,285]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="43" pageNumber="44">SMF 372694</specimenCode>
(XII 2374a), SL:
<quantity id="4CEFFD88FFEAC04FFF0EFED47CED3679" box="[151,239,294,321]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="43" pageNumber="44" unit="mm" value="0.5">0.5 mm</quantity>
, MD:
<quantity id="4CEFFD88FFEAC04FFEA1FED57D923679" box="[312,400,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="43" pageNumber="44" unit="mm" value="1.7">1.7 mm</quantity>
, Senckenberg Museum, Frankfurt/Main, Germany,
<location id="8EC806B6FFEAC04FFC4AFED578833679" LSID="urn:lsid:plazi:treatment:03BEE17BFFE8C04FFF0EFA5978A231A1:8EC806B6FFEAC04FFC4AFED578833679" box="[979,1153,295,321]" country="Germany" municipality="Lapugiu des Sus" name="Lapugiu de Sus" pageId="43" pageNumber="44">
Lăpugiu
<collectorName id="26E235BBFFEAC04FFBA1FED578833679" box="[1080,1153,295,321]" pageId="43" pageNumber="44">de Sus</collectorName>
</location>
(
<collectingCountry id="F30010FDFFEAC04FFB08FED578FE3679" box="[1169,1276,295,321]" name="Romania" pageId="43" pageNumber="44">Romania</collectingCountry>
), illustrated in Zilch (1934: pl. 7, fig. 21),
<figureCitation id="132C4CE8FFEAC04FFE4BFEB97E38365D" box="[466,570,331,357]" captionStart="FIGURE 21" captionStartId="42.[151,250,1833,1857]" captionTargetBox="[275,1333,257,1797]" captionTargetId="figure-30@42.[275,1333,257,1797]" captionTargetPageId="42" captionText="FIGURE 21 A1A3. Spirolaxis cornicula (Boettger, 1902), lectotype, SMF 372697, Coşteiu de Sus (Romania). B1B3. Spirolaxis cornicula (Boettger, 1902), holotype of Discohelix quinquangularis Boettger, 1902, SMF 372694, Lăpugiu de Sus (Romania). C1C2. Spirolaxis cornicula (Boettger, 1902), NHMW 2016/0177/0342, Lăpugiu des Sus (Romania)." figureDoi="http://doi.org/10.5281/zenodo.10147854" httpUri="https://zenodo.org/record/10147854/files/figure.png" pageId="43" pageNumber="44">Figs 21B</figureCitation>
<figureCitation id="132C4CE8FFEAC04FFDA3FEAF7E60365D" box="[570,610,331,363]" captionStart="FIGURE 1" captionStartId="5.[151,250,1522,1546]" captionTargetBox="[232,1366,234,1455]" captionTargetId="figure-58@5.[232,1366,234,1455]" captionTargetPageId="5" captionText="FIGURE 1. Descriptive terms used in descriptions of Architectonicidae (modified from Bieler 1993)." figureDoi="http://doi.org/10.5281/zenodo.10147816" httpUri="https://zenodo.org/record/10147816/files/figure.png" pageId="43" pageNumber="44">
<subScript id="17935228FFEAC04FFDA3FEAF7E403653" attach="left" box="[570,578,349,363]" fontSize="6" pageId="43" pageNumber="44">1</subScript>
B
</figureCitation>
<figureCitation id="132C4CE8FFEAC04FFDFAFEAF7E693653" box="[611,619,349,363]" captionStart="FIGURE 3" captionStartId="6.[151,250,2002,2026]" captionTargetId="figure-99@6.[337,1250,1242,1978]" captionTargetPageId="6" captionText="FIGURE 3. Descriptive terms used in descriptions of Mathildidae." figureDoi="http://doi.org/10.5281/zenodo.10307672" httpUri="https://zenodo.org/record/10307672/files/figure.png" pageId="43" pageNumber="44">
<subScript id="17935228FFEAC04FFDFAFEAF7E693653" attach="right" box="[611,619,349,363]" fontSize="6" pageId="43" pageNumber="44">3</subScript>
</figureCitation>
.
</materialsCitation>
<materialsCitation id="3B7F5A30FFEAC04FFDE3FEB97DB936B1" collectionCode="SMF, NHMW" collectorName="de Sus" country="Germany" location="Lapugiu de Sus" municipality="Lapugiu des Sus" pageId="43" pageNumber="44" specimenCode="SMF 372694, NHMW 2016" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB1F816FFEAC04FFDE3FEB97F14365D" box="[634,790,331,357]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="43" pageNumber="44" type="Museum">NHMW 2016</specimenCode>
/0177/0342, SL:
<quantity id="4CEFFD88FFEAC04FFC4FFEB9782D365D" box="[982,1071,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="43" pageNumber="44" unit="mm" value="1.2">1.2 mm</quantity>
, MD:
<quantity id="4CEFFD88FFEAC04FFBE3FEB978D1365D" box="[1146,1235,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="43" pageNumber="44" unit="mm" value="4.4">4.4 mm</quantity>
,
<collectingMunicipality id="6BCCCA17FFEAC04FFB79FEB9799E365D" box="[1248,1436,331,357]" pageId="43" pageNumber="44">Lăpugiu des Sus</collectingMunicipality>
(
<collectingCountry id="F30010FDFFEAC04FFF38FE9D7D0936B1" box="[161,267,367,393]" name="Romania" pageId="43" pageNumber="44">Romania</collectingCountry>
),
<figureCitation id="132C4CE8FFEAC04FFE84FE9D7D8136B1" box="[285,387,367,393]" captionStart="FIGURE 21" captionStartId="42.[151,250,1833,1857]" captionTargetBox="[275,1333,257,1797]" captionTargetId="figure-30@42.[275,1333,257,1797]" captionTargetPageId="42" captionText="FIGURE 21 A1A3. Spirolaxis cornicula (Boettger, 1902), lectotype, SMF 372697, Coşteiu de Sus (Romania). B1B3. Spirolaxis cornicula (Boettger, 1902), holotype of Discohelix quinquangularis Boettger, 1902, SMF 372694, Lăpugiu de Sus (Romania). C1C2. Spirolaxis cornicula (Boettger, 1902), NHMW 2016/0177/0342, Lăpugiu des Sus (Romania)." figureDoi="http://doi.org/10.5281/zenodo.10147854" httpUri="https://zenodo.org/record/10147854/files/figure.png" pageId="43" pageNumber="44">Figs 21C</figureCitation>
<figureCitation id="132C4CE8FFEAC04FFE1AFE737DAE36B1" box="[387,428,367,399]" captionStart="FIGURE 1" captionStartId="5.[151,250,1522,1546]" captionTargetBox="[232,1366,234,1455]" captionTargetId="figure-58@5.[232,1366,234,1455]" captionTargetPageId="5" captionText="FIGURE 1. Descriptive terms used in descriptions of Architectonicidae (modified from Bieler 1993)." figureDoi="http://doi.org/10.5281/zenodo.10147816" httpUri="https://zenodo.org/record/10147816/files/figure.png" pageId="43" pageNumber="44">
<subScript id="17935228FFEAC04FFE1AFE737D8936B7" attach="left" box="[387,395,385,399]" fontSize="6" pageId="43" pageNumber="44">1</subScript>
C
</figureCitation>
<figureCitation id="132C4CE8FFEAC04FFE35FE737DB636B7" box="[428,436,385,399]" captionStart="FIGURE 2" captionStartId="6.[151,250,1086,1110]" captionTargetBox="[236,1353,159,1066]" captionTargetId="figure-113@6.[236,1353,159,1066]" captionTargetPageId="6" captionText="FIGURE 2. Whorl cross-sections of Architectonicidae genera. All modified after Bieler (1993) except for Simplexollata, Basisulcata and Ammotectonica. Solatisonax? refers to Solatisonax? kostejana (Boettger, 1907), Bold genera are represented in the Paratethys Sea. SSC: subsutural cord, MC: mid cord(s), UPC: upper peripheral cord, LPC: lower peripheral cord, IPC: infraperipheral cord; UC: umbilical crenae; PUC: proxumbilical cord." figureDoi="http://doi.org/10.5281/zenodo.10147818" httpUri="https://zenodo.org/record/10147818/files/figure.png" pageId="43" pageNumber="44">
<subScript id="17935228FFEAC04FFE35FE737DB636B7" attach="left" box="[428,436,385,399]" fontSize="6" pageId="43" pageNumber="44">2</subScript>
</figureCitation>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA8506DFFEAC04FFF5EFE607F0E36E9" blockId="43.[151,1437,150,1689]" pageId="43" pageNumber="44">
<emphasis id="B9638C7FFFEAC04FFF5EFE607DBF3694" bold="true" box="[199,445,402,428]" pageId="43" pageNumber="44">Additional material:</emphasis>
<materialsCitation id="3B7F5A30FFEAC04FFE5FFE617F0A36E9" collectorName="de Sus" country="Germany" location="Costeiu de Sus" pageId="43" pageNumber="44" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="54ACEECFFFEAC04FFE5FFE617E5C3695" box="[454,606,403,429]" pageId="43" pageNumber="44" type="paralectotype">Paralectotype</typeStatus>
XII 2373c, Senckenberg Museum, Frankfurt/Main, Germany,
<location id="8EC806B6FFEAC04FFABDFE617CBD36E9" LSID="urn:lsid:plazi:treatment:03BEE17BFFE8C04FFF0EFA5978A231A1:8EC806B6FFEAC04FFABDFE617CBD36E9" country="Germany" name="Costeiu de Sus" pageId="43" pageNumber="44">
Coşteiu
<collectorName id="26E235BBFFEAC04FFA1BFE617CBD36E9" pageId="43" pageNumber="44">de Sus</collectorName>
</location>
(
<collectingCountry id="F30010FDFFEAC04FFF49FE457D3836E9" box="[208,314,439,465]" name="Romania" pageId="43" pageNumber="44">Romania</collectingCountry>
), illustrated in Zilch (1934: pl. 7, fig. 20)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA8506DFFEAC04FFF5EFE287DD8342D" blockId="43.[151,1437,150,1689]" pageId="43" pageNumber="44">
<emphasis id="B9638C7FFFEAC04FFF5EFE287DBE36CC" bold="true" box="[199,444,474,500]" pageId="43" pageNumber="44">Revised description.</emphasis>
Small, delicate, glossy, translucent, open-coiled, almost planispiral shell. Protoconch small, of 2.25 whorls, planispiral (diameter
<quantity id="4CEFFD88FFEAC04FFD0BFE0C7EFF3521" box="[658,765,510,537]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="43" pageNumber="44" unit="mm" value="0.75">0.75 mm</quantity>
). Smooth with distinct protoconch lip. Teleoconch of 1.24 whorls. Dorsal side with faint growth lines. SSC reduced to smooth, narrow spiral thread. First half teleoconch whorl with weak central angulation on dorsum; later whorl profile weakly convex on dorsal side. Periphery with two finely crenulated keels (upper one corresponding to LPC). Nearly straight sided between peripheral keels. Lower keel (on basal area sensu
<bibRefCitation id="EF862D9CFFEAC04FFE22FD7D7E7E3591" author="Bieler, R." box="[443,636,655,681]" pageId="43" pageNumber="44" pagination="1 - 376" refId="ref49146" refString="Bieler, R. (1993) Architectonicidae of the Indo-Pacific (Mollusca, Gastropoda). Abhandlungen des naturwissenschaftlichen Vereins in Hamburg, Neue Folge, 30, 1 - 376." type="journal article" year="1993">Bieler 1993: 322</bibRefCitation>
) strengthens with ontogeny. Base weakly convex with delicate growth lines. Columellar side of whorls convex resulting in subcircular aperture with two weak peripheral angulations. UC reduced to faintly crenulated spiral thread. Apertural cross-section changing from sub-circular towards markedly rectangular during ontogeny.
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFEAC04FFF5EFCEC792A3115" pageId="43" pageNumber="44" type="discussion">
<paragraph id="8BA8506DFFEAC04FFF5EFCEC7CC834D5" blockId="43.[151,1437,150,1689]" pageId="43" pageNumber="44">
<emphasis id="B9638C7FFFEAC04FFF5EFCEC7D4E3400" bold="true" box="[199,332,798,824]" pageId="43" pageNumber="44">Discussion.</emphasis>
<bibRefCitation id="EF862D9CFFEAC04FFECFFCED7E093401" author="Boettger, O." box="[342,523,799,825]" pageId="43" pageNumber="44" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger (1902)</bibRefCitation>
separated two open-coiled species based in the slight angulation on the dorsal side of
<taxonomicName id="4C172BEEFFEAC04FFF6DFCB17EE23465" authority="Boettger, 1902" authorityName="Boettger" authorityYear="1902" box="[244,736,835,861]" class="Gastropoda" family="Discohelicidae" genus="Discohelix" kingdom="Animalia" order="Seguenziida" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="quinquangularis">
<emphasis id="B9638C7FFFEAC04FFF6DFCB17D6E3465" box="[244,364,835,861]" italics="true" pageId="43" pageNumber="44">Discohelix</emphasis>
<emphasis id="B9638C7FFFEAC04FFEE3FCB17E313465" box="[378,563,835,861]" italics="true" pageId="43" pageNumber="44">quinquangularis</emphasis>
<bibRefCitation id="EF862D9CFFEAC04FFDA3FCB17EE23465" author="Boettger, O." box="[570,736,835,861]" pageId="43" pageNumber="44" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger, 1902</bibRefCitation>
</taxonomicName>
and based on the narrower coiling and the weaker crenulation of the keels. Placing both
<typeStatus id="54ACEECFFFEAC04FFE26FC957DED34B9" box="[447,495,871,897]" pageId="43" pageNumber="44">type</typeStatus>
specimens at the same scale shows, that
<emphasis id="B9638C7FFFEAC04FFC55FC957FE234B8" box="[972,992,871,896]" italics="true" pageId="43" pageNumber="44">D</emphasis>
.
<taxonomicName id="4C172BEEFFEAC04FFC60FC9578B034B9" authorityName="Boettger" authorityYear="1902" box="[1017,1202,871,897]" class="Gastropoda" family="Discohelicidae" genus="Discohelix" kingdom="Animalia" order="Seguenziida" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="quinquangularis">
<emphasis id="B9638C7FFFEAC04FFC60FC9578B034B9" box="[1017,1202,871,897]" italics="true" pageId="43" pageNumber="44">quinquangularis</emphasis>
</taxonomicName>
fully corresponds to
<taxonomicName id="4C172BEEFFEAC04FFF0EFC797D7B349D" authorityName="" baseAuthorityName="Boettger" baseAuthorityYear="1902" box="[151,377,907,933]" class="Gastropoda" family="Architectonicidae" genus="Spirolaxis" kingdom="Animalia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="cornicula">
<emphasis id="B9638C7FFFEAC04FFF0EFC797D7B349D" box="[151,377,907,933]" italics="true" pageId="43" pageNumber="44">Spirolaxis cornicula</emphasis>
</taxonomicName>
at the corresponding growth stage. In our opinion, the differences discussed by
<bibRefCitation id="EF862D9CFFEAC04FFB72FC79799E349D" author="Boettger, O." box="[1259,1436,907,933]" pageId="43" pageNumber="44" pagination="1 - 200" refId="ref49293" refString="Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des siebenb ¸ rgischen Vereins f ¸ r Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]" type="journal article" year="1902">Boettger (1902)</bibRefCitation>
are related only to different ontogenetic stages. As first revisers we chose
<taxonomicName id="4C172BEEFFEAC04FFC54FC5D78AD34F1" authorityName="" baseAuthorityName="Boettger" baseAuthorityYear="1902" box="[973,1199,943,969]" class="Gastropoda" family="Architectonicidae" genus="Spirolaxis" kingdom="Animalia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="cornicula">
<emphasis id="B9638C7FFFEAC04FFC54FC5D78AD34F1" box="[973,1199,943,969]" italics="true" pageId="43" pageNumber="44">Spirolaxis cornicula</emphasis>
</taxonomicName>
as name for this species.
</paragraph>
<paragraph id="8BA8506DFFEAC04FFF5EFC057E1732DD" blockId="43.[151,1437,150,1689]" pageId="43" pageNumber="44">
Two extant species with an Amphiatlantic distribution have been described:
<taxonomicName id="4C172BEEFFEAC04FFB84FC05799E3329" authority="(Monterosato, 1890)" baseAuthorityName="Monterosato" baseAuthorityYear="1890" box="[1053,1436,1015,1041]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" kingdom="Animalia" order="Littorinimorpha" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="centrifuga">
<emphasis id="B9638C7FFFEAC04FFB84FC0578AD3329" box="[1053,1199,1015,1041]" italics="true" pageId="43" pageNumber="44">A. centrifuga</emphasis>
(Monterosato, 1890)
</taxonomicName>
and
<taxonomicName id="4C172BEEFFEAC04FFF5EFBE97E32330D" authority="Jaume &amp; Borro, 1946" authorityName="Jaume &amp; Borro" authorityYear="1946" box="[199,560,1051,1077]" class="Gastropoda" family="Architectonicidae" genus="Spirolaxis" kingdom="Animalia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="clenchi">
<emphasis id="B9638C7FFFEAC04FFF5EFBE97D35330D" box="[199,311,1051,1077]" italics="true" pageId="43" pageNumber="44">A. clenchi</emphasis>
<bibRefCitation id="EF862D9CFFEAC04FFEA4FBE97E32330D" author="Jaume, M. L. &amp; Borro, P." box="[317,560,1051,1077]" pageId="43" pageNumber="44" pagination="13 - 22" refId="ref55028" refString="Jaume, M. L. &amp; Borro, P. (1946) Novedades en moluscos marinos cubanos. Revista de la sociedad malaceologica &quot; Carlos de la Torre &quot;, 4 (1), 13 - 22." type="journal article" year="1946">Jaume &amp; Borro, 1946</bibRefCitation>
</taxonomicName>
(
<emphasis id="B9638C7FFFEAC04FFDD9FBE97E6A330C" box="[576,616,1051,1076]" italics="true" pageId="43" pageNumber="44">e.g.</emphasis>
,
<bibRefCitation id="EF862D9CFFEAC04FFDEFFBE97EFC330D" author="Bieler, R." box="[630,766,1051,1077]" pageId="43" pageNumber="44" pagination="53 - 103" refId="ref48969" refString="Bieler, R. (1984) Die Gattungen der Architectonicidae (Gastropoda: &quot; Heterogastropoda &quot;). Allgemeines und Teil 1: Pseudomalaxis. Archiv f ¸ r Molluskenkunde, 115, 53 - 103." type="journal article" year="1984">Bieler 1984</bibRefCitation>
: pl. 3, figs 1920; Negri &amp; Corselli 2016: figs 16oq). Both have been recorded from the Mediterranean Pliocene. Differences between the two species center around the degree of disjunction of the teleoconch whorls and whether the peripheral keels are bifid or not. It is still unclear whether there is one or two species present in the Atlanto-Mediterranean Pliocene to present day. In any case, the Paratethyan specimens are like
<taxonomicName id="4C172BEEFFEAC04FFEF6FB597DDE33FD" authorityName="Jaume &amp; Borro" authorityYear="1946" box="[367,476,1195,1221]" class="Gastropoda" family="Architectonicidae" genus="Spirolaxis" kingdom="Animalia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="clenchi">
<emphasis id="B9638C7FFFEAC04FFEF6FB597DDE33FD" box="[367,476,1195,1221]" italics="true" pageId="43" pageNumber="44">S. clenchi</emphasis>
</taxonomicName>
in having simple as opposed to the bifid peripheral carinae seen in
<taxonomicName id="4C172BEEFFEAC04FFB4FFB59796733FD" baseAuthorityName="Monterosato" baseAuthorityYear="1890" box="[1238,1381,1195,1221]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" kingdom="Animalia" order="Littorinimorpha" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="centrifuga">
<emphasis id="B9638C7FFFEAC04FFB4FFB59796733FD" box="[1238,1381,1195,1221]" italics="true" pageId="43" pageNumber="44">S. centrifuga</emphasis>
</taxonomicName>
, and like
<taxonomicName id="4C172BEEFFEAC04FFF50FB3D7D5833D1" baseAuthorityName="Monterosato" baseAuthorityYear="1890" box="[201,346,1231,1257]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" kingdom="Animalia" order="Littorinimorpha" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="centrifuga">
<emphasis id="B9638C7FFFEAC04FFF50FB3D7D5833D1" box="[201,346,1231,1257]" italics="true" pageId="43" pageNumber="44">S. centrifuga</emphasis>
</taxonomicName>
in having finely crenulated peripheral carinae and not coarse as in
<taxonomicName id="4C172BEEFFEAC04FFBCEFB3D78C433D1" authorityName="Jaume &amp; Borro" authorityYear="1946" box="[1111,1222,1231,1257]" class="Gastropoda" family="Architectonicidae" genus="Spirolaxis" kingdom="Animalia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="clenchi">
<emphasis id="B9638C7FFFEAC04FFBCEFB3D78C433D1" box="[1111,1222,1231,1257]" italics="true" pageId="43" pageNumber="44">S. clenchi</emphasis>
</taxonomicName>
. The teleoconch is widely uncoiled, whereas in both extant species coiling is tighter. However, specimens ascribed to
<taxonomicName id="4C172BEEFFEAC04FFB77FB0179593235" authorityName="Jaume &amp; Borro" authorityYear="1946" box="[1262,1371,1267,1293]" class="Gastropoda" family="Architectonicidae" genus="Spirolaxis" kingdom="Animalia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="clenchi">
<emphasis id="B9638C7FFFEAC04FFB77FB0179593235" box="[1262,1371,1267,1293]" italics="true" pageId="43" pageNumber="44">S. clenchi</emphasis>
</taxonomicName>
illustrate a considerable variability in disjunction from the earliest whorls (own data B.L.). The Paratethyan specimens are very similar to one of the specimens from Estepona, but considerably smaller. We consider the Paratethyan specimens to represent a separate species, but this requires confirmation based on a larger number of specimens and a better understanding on the present-day species. The extant Indo-West Pacific
<taxonomicName id="4C172BEEFFEAC04FFB97FA71799E32A5" authority="Bieler, 1993" authorityName="Bieler" authorityYear="1993" box="[1038,1436,1411,1437]" class="Gastropoda" family="Architectonicidae" genus="Spirolaxis" kingdom="Animalia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="cornuarietis">
<emphasis id="B9638C7FFFEAC04FFB97FA71790C32A4" box="[1038,1294,1411,1437]" italics="true" pageId="43" pageNumber="44">Spirolaxis cornuarietis</emphasis>
<bibRefCitation id="EF862D9CFFEAC04FFA8DFA71799E32A5" author="Bieler, R." box="[1300,1436,1411,1437]" pageId="43" pageNumber="44" pagination="1 - 376" refId="ref49146" refString="Bieler, R. (1993) Architectonicidae of the Indo-Pacific (Mollusca, Gastropoda). Abhandlungen des naturwissenschaftlichen Vereins in Hamburg, Neue Folge, 30, 1 - 376." type="journal article" year="1993">Bieler, 1993</bibRefCitation>
</taxonomicName>
is another comparable species but differs in its prominent beads and additional beaded threads on the umbilical wall (
<bibRefCitation id="EF862D9CFFEAC04FFF44FA397D6432DD" author="Bieler, R." box="[221,358,1483,1509]" pageId="43" pageNumber="44" pagination="1 - 376" refId="ref49146" refString="Bieler, R. (1993) Architectonicidae of the Indo-Pacific (Mollusca, Gastropoda). Abhandlungen des naturwissenschaftlichen Vereins in Hamburg, Neue Folge, 30, 1 - 376." type="journal article" year="1993">Bieler 1993</bibRefCitation>
: figs 276268).
</paragraph>
<paragraph id="8BA8506DFFEAC04FFF5EFA1C792A3115" blockId="43.[151,1437,150,1689]" pageId="43" pageNumber="44">
<emphasis id="B9638C7FFFEAC04FFF5EFA1C7DAA3130" bold="true" box="[199,424,1518,1544]" pageId="43" pageNumber="44">Paleoenvironment.</emphasis>
Extant
<taxonomicName id="4C172BEEFFEAC04FFD9FFA1D7ED13131" box="[518,723,1519,1545]" pageId="43" pageNumber="44">
<emphasis id="B9638C7FFFEAC04FFD9FFA1D7E753131" box="[518,631,1519,1545]" italics="true" pageId="43" pageNumber="44">Spirolaxis</emphasis>
species
</taxonomicName>
occur from few tens of meters water depth down to bathyal depths (
<bibRefCitation id="EF862D9CFFEAC04FFF76F9E17D7A3115" author="Bieler, R." box="[239,376,1555,1581]" pageId="43" pageNumber="44" pagination="1 - 376" refId="ref49146" refString="Bieler, R. (1993) Architectonicidae of the Indo-Pacific (Mollusca, Gastropoda). Abhandlungen des naturwissenschaftlichen Vereins in Hamburg, Neue Folge, 30, 1 - 376." type="journal article" year="1993">Bieler 1993</bibRefCitation>
).
<taxonomicName id="4C172BEEFFEAC04FFE12F9E17E6C3115" authorityName="" baseAuthorityName="Boettger" baseAuthorityYear="1902" box="[395,622,1555,1581]" class="Gastropoda" family="Architectonicidae" genus="Spirolaxis" kingdom="Animalia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="cornicula">
<emphasis id="B9638C7FFFEAC04FFE12F9E17E6C3115" box="[395,622,1555,1581]" italics="true" pageId="43" pageNumber="44">Spirolaxis cornicula</emphasis>
</taxonomicName>
seems to have preferred middle to outer neritic environments.
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFEAC04FFF5EF9C478A231A1" pageId="43" pageNumber="44" type="distribution">
<paragraph id="8BA8506DFFEAC04FFF5EF9C479693169" blockId="43.[151,1437,150,1689]" box="[199,1387,1590,1617]" pageId="43" pageNumber="44">
<emphasis id="B9638C7FFFEAC04FFF5EF9C47D633168" bold="true" box="[199,353,1590,1616]" pageId="43" pageNumber="44">Distribution.</emphasis>
Known so far only from the middle Badenian (late Langhian) of the Central Paratethys Sea.
</paragraph>
<paragraph id="8BA8506DFFEAC04FFF5EF9A878A231A1" blockId="43.[151,1437,150,1689]" pageId="43" pageNumber="44">
<emphasis id="B9638C7FFFEAC04FFF5EF9A87DE2314C" bold="true" box="[199,480,1626,1653]" pageId="43" pageNumber="44">Central Paratethys Sea.</emphasis>
Badenian (Middle Miocene):
<emphasis id="B9638C7FFFEAC04FFCB7F9A97FEC314C" box="[814,1006,1627,1652]" italics="true" pageId="43" pageNumber="44">Pannonian Basin</emphasis>
: Szob (
<collectingCountry id="F30010FDFFEAC04FFBDAF9A978A5314D" box="[1091,1191,1627,1653]" name="Hungary" pageId="43" pageNumber="44">Hungary</collectingCountry>
) (
<bibRefCitation id="EF862D9CFFEAC04FFB22F9A97CFB31A1" author="Csepreghy-Meznerics, I." pageId="43" pageNumber="44" pagination="361 - 477" refId="ref50934" refString="Csepreghy-Meznerics, I. (1956) Die Molluskenfauna von Szob und Letkes. A magyar allami f ˆ ldtani intezet evk ˆ nyve, 45, 361 - 477. [https: // epa. oszk. hu / 03200 / 03274 / 00094 / pdf / EPA 03274 _ mafi _ evkonyv _ 1956 _ 02 _ 363 - 442. pdf and https: // epa. oszk. hu / 03200 / 03274 / 00094 / pdf / EPA 03274 _ mafi _ evkonyv _ 1956 _ 02 _ 450 - 477. pdf]" type="journal article" year="1956">Csepreghy-Meznerics 1956</bibRefCitation>
);
<emphasis id="B9638C7FFFEAC04FFE94F98D7D9431A0" box="[269,406,1663,1689]" italics="true" pageId="43" pageNumber="44">Făget Basin</emphasis>
: Coşteiu de Sus, Lăpugiu de Sus (
<collectingCountry id="F30010FDFFEAC04FFC8CF98D7F7C31A1" box="[789,894,1663,1689]" name="Romania" pageId="43" pageNumber="44">Romania</collectingCountry>
) (
<collectingCountry id="F30010FDFFEAC04FFC0DF98D7FFF31A1" box="[916,1021,1663,1689]" name="Romania" pageId="43" pageNumber="44">Romania</collectingCountry>
) (Zilch 1934).
</paragraph>
</subSubSection>
</treatment>
</document>