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<document id="21241897C873FD4098CBA7A9FA44E2A1" ID-DOI="10.11646/zootaxa.5370.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10147814" ID-ZooBank="48903495-7C6C-46E4-9B1B-D6A2F2781873" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1700202702953" checkinUser="plazi" docAuthor="Harzhauser, Mathias &amp; Landau, Bernard" docDate="2023" docId="03BEE17BFFDBC07EFF0EFE287E4E3344" docLanguage="en" docName="zootaxa.5370.1.1.pdf" docOrigin="Zootaxa 5370 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5370.1.1/52270" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Heliacus d'Orbigny 1842" docType="treatment" docVersion="1" lastPageNumber="27" masterDocId="FF879903FFC1C064FF99FFF27C023738" masterDocTitle="The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition" masterLastPageNumber="74" masterPageNumber="1" pageNumber="27" updateTime="1706539087537" updateUser="carolina">
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<mods:title id="F4382B080FD8EC3707F86BE382919E8D">The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition</mods:title>
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<mods:namePart id="453D4344BA0185FFBD119AEACAB570B7">Harzhauser, Mathias</mods:namePart>
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<mods:affiliation id="A5DC16842E50AFCE9911D3C5ACC6B962">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;</mods:affiliation>
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<mods:namePart id="5B9BF0D61F031E75153345D9BC764672">Landau, Bernard</mods:namePart>
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<mods:affiliation id="57390A2162095965D2320A506FE16C8A">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655; &amp; Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal. bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494 &amp; Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;</mods:affiliation>
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<subSubSection id="C30D03E6FFDBC07EFF0EFE287E2F36CC" box="[151,557,474,501]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BA8506DFFDBC07EFF0EFE287E2F36CC" blockId="26.[151,557,474,501]" box="[151,557,474,501]" pageId="26" pageNumber="27">
<emphasis id="B9638C7FFFDBC07EFF0EFE287E2F36CC" bold="true" box="[151,557,474,501]" pageId="26" pageNumber="27">
Genus
<taxonomicName id="4C172BEEFFDBC07EFF77FE287E2F36CC" ID-CoL="8V5JL" authority="d'Orbigny, 1842" authorityName="d'Orbigny" authorityYear="1842" box="[238,557,474,501]" class="Gastropoda" family="Architectonicidae" genus="Heliacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFDBC07EFF77FE287D5836CD" bold="true" box="[238,346,474,501]" italics="true" pageId="26" pageNumber="27">Heliacus</emphasis>
dOrbigny, 1842
</taxonomicName>
</emphasis>
</paragraph>
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<subSubSection id="C30D03E6FFDBC07EFF0EFDD07D2E3558" pageId="26" pageNumber="27" type="type_taxon">
<paragraph id="8BA8506DFFDBC07EFF0EFDD07D2E3558" blockId="26.[151,1437,546,1148]" pageId="26" pageNumber="27">
<emphasis id="B9638C7FFFDBC07EFF0EFDD07D313504" bold="true" box="[151,307,546,572]" pageId="26" pageNumber="27">
<typeStatus id="54ACEECFFFDBC07EFF0EFDD07CD03504" box="[151,210,546,572]" pageId="26" pageNumber="27">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C172BEEFFDBC07EFEA2FDD07EBC3504" ID-CoL="8V9FD" authority="Deshayes, 1830" authorityName="Deshayes" authorityYear="1830" box="[315,702,546,572]" class="Gastropoda" family="Architectonicidae" genus="Solarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="herberti">
<emphasis id="B9638C7FFFDBC07EFEA2FDD07E063504" box="[315,516,546,572]" italics="true" pageId="26" pageNumber="27">Solarium herberti</emphasis>
<bibRefCitation id="EF862D9CFFDBC07EFD95FDD07EBC3504" author="Deshayes, G. P." box="[524,702,546,572]" pageId="26" pageNumber="27" refId="ref51768" refString="Deshayes, G. P. (1830) Encyclopedie Methodique. Histoire naturelle des vers. 2 (1). Agasse, Paris, 256 pp. [https: // www. biodiversitylibrary. org / item / 105145 # page / 7 / mode / 1 up]" type="book" year="1830">Deshayes, 1830</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="4C172BEEFFDBC07EFD7EFDD078883504" ID-CoL="8V863" authority="(Gmelin, 1791)" baseAuthorityName="Gmelin" baseAuthorityYear="1791" box="[743,1162,546,572]" class="Gastropoda" family="Architectonicidae" genus="Heliacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="cylindricus">
<emphasis id="B9638C7FFFDBC07EFD7EFDD07FCD3504" box="[743,975,546,572]" italics="true" pageId="26" pageNumber="27">Heliacus cylindricus</emphasis>
(
<bibRefCitation id="EF862D9CFFDBC07EFC46FDD078833504" author="Gmelin, J. F." box="[991,1153,546,572]" pageId="26" pageNumber="27" pagination="3021 - 3910" refId="ref52952" refString="Gmelin, J. F. (1791) Vermes. In: Gmelin, J. F. (Ed.), Caroli a Linnaei systema naturae per regna tria naturae. Tome 1 (6). 13 th Edition. G. E. Beer, Lipsiae [Leipzig], pp. 3021 - 3910. [http: // www. biodiversitylibrary. org / item / 83098 # 5]" type="book chapter" year="1791">Gmelin, 1791</bibRefCitation>
)
</taxonomicName>
]; by monotypy. Present-day, Atlantic.
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFDBC07EFF5EFD987F2B330C" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8BA8506DFFDBC07EFF5EFD98798735C8" blockId="26.[151,1437,546,1148]" pageId="26" pageNumber="27">
<emphasis id="B9638C7FFFDBC07EFF5EFD987DAB35BC" bold="true" box="[199,425,618,644]" pageId="26" pageNumber="27">Original diagnosis.</emphasis>
<emphasis id="B9638C7FFFDBC07EFE25FD997CC135F4" italics="true" pageId="26" pageNumber="27">
Testa orbiculato-conica, trochiformi, transversim costata, longitudinaliter
<taxonomicName id="4C172BEEFFDBC07EFA99FD99794E35BC" authorityName="I. Lea" authorityYear="1833" box="[1280,1356,619,644]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="striata">striata</taxonomicName>
, nigrofusca; spira elevata, anfractibus quinis subconvexis; umbilico angusto, crenato, intus tricostate; apertura subrotunda”
</emphasis>
[Shell orbicular-conical, trochiform, transversely ribbed, longitudinally striated, black-brown; a raised spiral, subconvex with bends; umbilicus narrow, crenate, tricostate inside; aperture sub-rounded] (dOrbigny 1842: 68).
</paragraph>
<paragraph id="8BA8506DFFDBC07EFF5EFD087F2B330C" blockId="26.[151,1437,546,1148]" pageId="26" pageNumber="27">
<emphasis id="B9638C7FFFDBC07EFF5EFD087DB8342C" bold="true" box="[199,442,762,788]" pageId="26" pageNumber="27">Revised description.</emphasis>
<emphasis id="B9638C7FFFDBC07EFE56FD087DE7330C" italics="true" pageId="26" pageNumber="27">
Teleoconch: small to medium-sized (usually
<quantity id="4CEFFD88FFDBC07EFC53FD087833342C" box="[970,1073,762,788]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="13.0" metricValueMin="6.0" pageId="26" pageNumber="27" unit="mm" value="9.5" valueMax="13.0" valueMin="6.0">6-13 mm</quantity>
, rarely over
<quantity id="4CEFFD88FFDBC07EFB5CFD087917342C" box="[1221,1301,762,788]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="mm" value="20.0">20 mm</quantity>
}, depressed to tall roundly cone-shaped, with narrow to very wide umbilicus (ca. 8-40% of shell diameter); whorls weakly to distinctly bulging; sculpture: spiral ribs with relatively large nodules on entire surface; number of ribs constant, only between lower peripheral and infraperipheral ribs occasionally with 1 ± strong additional rib; apical side: subsutural rib, single midrib (or two narrow ones) and upper peripheral rib usually equally developed, rarely upper peripheral rib somewhat stronger; periphery formed by lower peripheral rib and only slightly weaker infraperipheral rib (occasionally with additional rib between them); upper point of whorl attachment at or below lower peripheral rib; base usually with 5 spiral ribs, increasing in width towards umbilicus; umbilical wall usually with 1-2 strong, narrow spiral ribs
</emphasis>
” (
<bibRefCitation id="EF862D9CFFDBC07EFE66FBE87EBC330C" author="Bieler, R." box="[511,702,1050,1076]" pageId="26" pageNumber="27" pagination="1 - 376" refId="ref49146" refString="Bieler, R. (1993) Architectonicidae of the Indo-Pacific (Mollusca, Gastropoda). Abhandlungen des naturwissenschaftlichen Vereins in Hamburg, Neue Folge, 30, 1 - 376." type="journal article" year="1993">Bieler 1993: 184</bibRefCitation>
) (
<figureCitation id="132C4CE8FFDBC07EFD4FFBE87F19330C" box="[726,795,1050,1076]" captionStart="FIGURE 2" captionStartId="6.[151,250,1086,1110]" captionTargetBox="[236,1353,159,1066]" captionTargetId="figure-113@6.[236,1353,159,1066]" captionTargetPageId="6" captionText="FIGURE 2. Whorl cross-sections of Architectonicidae genera. All modified after Bieler (1993) except for Simplexollata, Basisulcata and Ammotectonica. Solatisonax? refers to Solatisonax? kostejana (Boettger, 1907), Bold genera are represented in the Paratethys Sea. SSC: subsutural cord, MC: mid cord(s), UPC: upper peripheral cord, LPC: lower peripheral cord, IPC: infraperipheral cord; UC: umbilical crenae; PUC: proxumbilical cord." figureDoi="http://doi.org/10.5281/zenodo.10147818" httpUri="https://zenodo.org/record/10147818/files/figure.png" pageId="26" pageNumber="27">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFDBC07EFF5EFBCC7E4E3344" pageId="26" pageNumber="27" type="description">
<paragraph id="8BA8506DFFDBC07EFF5EFBCC7E4E3344" blockId="26.[151,1437,546,1148]" pageId="26" pageNumber="27">
<emphasis id="B9638C7FFFDBC07EFF5EFBCC7D4B3360" bold="true" box="[199,329,1086,1112]" pageId="26" pageNumber="27">Synonyms.</emphasis>
<emphasis id="B9638C7FFFDBC07EFEC9FBCD7DA33360" box="[336,417,1087,1112]" italics="true" pageId="26" pageNumber="27">Torinia</emphasis>
J. E.
<bibRefCitation id="EF862D9CFFDBC07EFE79FBCC7E5C3360" author="Gray, J. E." box="[480,606,1086,1112]" pageId="26" pageNumber="27" pagination="48 - 92" refId="ref53236" refString="Gray, J. E. (1842) [Mollusca]. In: Synopsis of the contents of the British Museum. 44 th Edition. G. Woodfall and Son, London., pp. 48 - 92. https: // www. biodiversitylibrary. org / page / 55295773" type="book chapter" year="1842">Gray, 1842</bibRefCitation>
,
<typeStatus id="54ACEECFFFDBC07EFDF0FBCD7E9B3361" box="[617,665,1087,1113]" pageId="26" pageNumber="27">type</typeStatus>
species
<taxonomicName id="4C172BEEFFDBC07EFD61FBCC788D3360" authority="Dillwyn, 1817" authorityName="Dillwyn" authorityYear="1817" box="[760,1167,1086,1112]" class="Gastropoda" family="Trochidae" genus="Trochus" kingdom="Animalia" order="Trochida" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="cylindraceus">
<emphasis id="B9638C7FFFDBC07EFD61FBCC7FE53360" box="[760,999,1086,1112]" italics="true" pageId="26" pageNumber="27">Trochus cylindraceus</emphasis>
<bibRefCitation id="EF862D9CFFDBC07EFC74FBCC788D3360" author="Dillwyn, L. W." box="[1005,1167,1086,1112]" pageId="26" pageNumber="27" refId="ref51963" refString="Dillwyn, L. W. (1817) n. k. In: A descriptive catalogue of Recent shells, arranged according to the Linnaean method; with particular attention to the synonymy. Vol. 2. J. &amp; A. Arch, London, pp. 581 - 1092, index. [https: // www. biodiversitylibrary. org / item / 41315 # page / 9 / mode / 1 up]" type="url" year="1817">Dillwyn, 1817</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="4C172BEEFFDBC07EFB2FFBCC7D4B3344" ID-CoL="8V863" authority="(Gmelin, 1791)" baseAuthorityName="Gmelin" baseAuthorityYear="1791" class="Gastropoda" family="Architectonicidae" genus="Heliacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="cylindricus">
<emphasis id="B9638C7FFFDBC07EFB2FFBCC799E3360" box="[1206,1436,1086,1112]" italics="true" pageId="26" pageNumber="27">Heliacus cylindricus</emphasis>
(
<bibRefCitation id="EF862D9CFFDBC07EFF06FB907D433344" author="Gmelin, J. F." box="[159,321,1122,1148]" pageId="26" pageNumber="27" pagination="3021 - 3910" refId="ref52952" refString="Gmelin, J. F. (1791) Vermes. In: Gmelin, J. F. (Ed.), Caroli a Linnaei systema naturae per regna tria naturae. Tome 1 (6). 13 th Edition. G. E. Beer, Lipsiae [Leipzig], pp. 3021 - 3910. [http: // www. biodiversitylibrary. org / item / 83098 # 5]" type="book chapter" year="1791">Gmelin, 1791</bibRefCitation>
)
</taxonomicName>
]. Present-day, Atlantic.
</paragraph>
</subSubSection>
</treatment>
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