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<document id="002C27BF4E97ACD562F35C117D4A6162" ID-DOI="10.11646/zootaxa.5370.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10147814" ID-ZooBank="48903495-7C6C-46E4-9B1B-D6A2F2781873" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1700202702953" checkinUser="plazi" docAuthor="Harzhauser, Mathias &amp; Landau, Bernard" docDate="2023" docId="03BEE17BFFD4C071FF0EFE447DCE30CE" docLanguage="en" docName="zootaxa.5370.1.1.pdf" docOrigin="Zootaxa 5370 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5370.1.1/52270" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Solatisonax transversa Harzhauser &amp; Landau 2023, nov. sp." docType="treatment" docVersion="8" lastPageNumber="22" masterDocId="FF879903FFC1C064FF99FFF27C023738" masterDocTitle="The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition" masterLastPageNumber="74" masterPageNumber="1" pageNumber="22" updateTime="1706539087537" updateUser="carolina">
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<mods:title id="301E457C76E2ED96959AA3AB117C950F">The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition</mods:title>
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<mods:namePart id="D4B4D204CA0ED186B57A5E46A0E57818">Harzhauser, Mathias</mods:namePart>
<mods:nameIdentifier id="729611D0DC7D0F5569276F46BFD59D44" type="ORCID">0000-0002-4471-6655</mods:nameIdentifier>
<mods:affiliation id="FD615EFFB419D7E5B9D2A96C5B994B1F">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;</mods:affiliation>
<mods:nameIdentifier id="7B2E54A88020FF52AB15A736A75F0F0D" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:namePart id="FF2784F42DACCC1860CDD43033908D22">Landau, Bernard</mods:namePart>
<mods:nameIdentifier id="4638252D342E98ABB2121C2BDE0BEFC5" type="ORCID">0000-0002-4471-6655</mods:nameIdentifier>
<mods:affiliation id="5637D51043629EE13F51A6A9855877C6">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655; &amp; Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal. bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494 &amp; Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;</mods:affiliation>
<mods:nameIdentifier id="675385FE25AE374FCC57BEA865EFB478" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:title id="91D7E594B59B1F1D49235446F5523BFA">Zootaxa</mods:title>
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<mods:part id="5B956792B75FD7B9C79168BBB73CB20F">
<mods:date id="0C48843027FEADE12BEB6DF6408D06D2">2023</mods:date>
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<mods:number id="F984A7E18D66926919788BE6587347CA">2023-11-14</mods:number>
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<mods:number id="A7EA64D6E48A3A7513A7FDDC86169995">5370</mods:number>
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<treatment id="03BEE17BFFD4C071FF0EFE447DCE30CE" ID-DOI="http://doi.org/10.5281/zenodo.10356891" ID-Zenodo-Dep="10356891" LSID="urn:lsid:plazi:treatment:03BEE17BFFD4C071FF0EFE447DCE30CE" httpUri="http://treatment.plazi.org/id/03BEE17BFFD4C071FF0EFE447DCE30CE" lastPageNumber="22" pageId="21" pageNumber="22">
<subSubSection id="C30D03E6FFD4C071FF0EFE447E1C36E9" box="[151,542,438,465]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BA8506DFFD4C071FF0EFE447E1C36E9" blockId="21.[151,542,438,507]" box="[151,542,438,465]" pageId="21" pageNumber="22">
<heading id="D0E0E701FFD4C071FF0EFE447E1C36E9" bold="true" box="[151,542,438,465]" fontSize="11" level="2" pageId="21" pageNumber="22" reason="4">
<emphasis id="B9638C7FFFD4C071FF0EFE447E1C36E9" bold="true" box="[151,542,438,465]" pageId="21" pageNumber="22">
<taxonomicName id="4C172BEEFFD4C071FF0EFE447DBA36E9" ID-CoL="BXSSJ" authority="Harzhauser &amp; Landau, 2023" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[151,440,438,465]" class="Gastropoda" family="Architectonicidae" genus="Solatisonax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Heterostropha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="transversa" status="sp. nov.">
<emphasis id="B9638C7FFFD4C071FF0EFE447D2736E9" bold="true" box="[151,293,438,465]" italics="true" pageId="21" pageNumber="22">Solatisonax</emphasis>
?
<emphasis id="B9638C7FFFD4C071FEA2FE457DBA36E9" bold="true" box="[315,440,439,465]" italics="true" pageId="21" pageNumber="22">transversa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2503104FFD4C071FE59FE457E1C36E9" box="[448,542,439,465]" pageId="21" pageNumber="22" rank="species">nov. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFD4C071FF0EFE297D5536C3" box="[151,343,475,507]" pageId="21" pageNumber="22" type="description">
<paragraph id="8BA8506DFFD4C071FF0EFE297D5536C3" blockId="21.[151,542,438,507]" box="[151,343,475,507]" pageId="21" pageNumber="22">
<figureCitation id="132C4CE8FFD4C071FF0EFE297CE836CD" box="[151,234,475,501]" captionStart="FIGURE 4" captionStartId="9.[151,250,1590,1614]" captionTargetBox="[151,1436,151,1566]" captionTargetId="figure-233@9.[151,1436,151,1569]" captionTargetPageId="9" captionText="FIGURE 4 Protoconchs of Paratethyan Architectonicidae. A. Ammotectonica gregorovae nov. sp., paratype, NHMW 1862/0029/0078c. B. Simplexollata anticollata nov. sp., paratype, NHMW 1854/0035/0331, Lăpugiu de Sus (Romania). C. Simplexollata carocollata (Lamarck, 1822), NHMW 1850/0018/0115, Léognan (France), Burdigalian. D. Simplexollata simplex (Bronn, 1831), NHMW 2013/0078/0109, Baden (Austria).E. Nipteraxis deformatusnov. sp., paratype, NHMW 2010/0004/0456a, Bad Vöslau (Austria). F. Nipteraxis exmoniliferus (Sacco, 1892), NHMW 1907/0323/1959, M̂llersdorf (Austria). G. Solatectonica kostejana (Boettger, 1907), NHMW 2012/0229/0049b, Traiskirchen (Austria). H. Nipteraxis monilifer (Bronn, 1831), NHMW 1846/0037/0427, M̂llersdorf (Austria). I. Solatisonax tavianii nov. sp., NHMW2016/0266/0002, SL: 8.0 mm, MD: 15.0 mm, Lăpugiu des Sus (Romania). J. Solatisonax transversa nov. sp., holotype, NHMW 2016/0177/0341, Lăpugiu de Sus (Roma- nia). K. Granosolarium semilaevis (Sacco, 1892), NHMW 1846/0037/0428d, M̂llersdorf (Austria). L. Heliacus globosus nov. sp., holotype NHMW 1907/0328/1959a, M̂llersdorf (Austria). M. Heliacus miser (Dujardin, 1837), NHMW 2023/0267/0001, Steinebrunn (Austria). N. Pseudotorinia grasemanni nov. sp., paratype, NHMW 2023/0267/0010, Steinebrunn (Austria). O. Pseudotorinia marthae (Boettger, 1902), NHMW 2012/0229/0049, Traiskirchen (Austria). All figures same scale." figureDoi="http://doi.org/10.5281/zenodo.10147820" httpUri="https://zenodo.org/record/10147820/files/figure.png" pageId="21" pageNumber="22">Figs 4J</figureCitation>
,
<figureCitation id="132C4CE8FFD4C071FF6CFE297D2636CD" box="[245,292,475,501]" captionStart="FIGURE 11" captionStartId="22.[151,250,1209,1233]" captionTargetBox="[151,1436,181,1185]" captionTargetId="figure-30@22.[151,1436,181,1185]" captionTargetPageId="22" captionText="FIGURE 11 A1–A3. Solatisonax transversa nov. sp., holotype, NHMW 2016/0177/0341, Lăpugiu de Sus (Romania). B1–B3. Heliacus globosus nov. sp., holotype NHMW 1907/0328/1959a, M̂llersdorf (Austria). C1–C3. Nipteraxis pozsgayae (Kovács Leél-Őssy &amp; Vicián, 2023), holotype, PAL 2022.52, Ottnangian, Parádfürdö, Ilona Valley (Hungary)." figureDoi="http://doi.org/10.5281/zenodo.10147834" httpUri="https://zenodo.org/record/10147834/files/figure.png" pageId="21" pageNumber="22">11A</figureCitation>
<figureCitation id="132C4CE8FFD4C071FEBCFE1F7D4D36CD" box="[293,335,475,507]" captionStart="FIGURE 1" captionStartId="5.[151,250,1522,1546]" captionTargetBox="[232,1366,234,1455]" captionTargetId="figure-58@5.[232,1366,234,1455]" captionTargetPageId="5" captionText="FIGURE 1. Descriptive terms used in descriptions of Architectonicidae (modified from Bieler 1993)." figureDoi="http://doi.org/10.5281/zenodo.10147816" httpUri="https://zenodo.org/record/10147816/files/figure.png" pageId="21" pageNumber="22">
<subScript id="17935228FFD4C071FEBCFE1F7D2F36C3" attach="right" box="[293,301,493,507]" fontSize="6" pageId="21" pageNumber="22">1</subScript>
–A
</figureCitation>
<figureCitation id="132C4CE8FFD4C071FED6FE1F7D5536C3" box="[335,343,493,507]" captionStart="FIGURE 3" captionStartId="6.[151,250,2002,2026]" captionTargetId="figure-99@6.[337,1250,1242,1978]" captionTargetPageId="6" captionText="FIGURE 3. Descriptive terms used in descriptions of Mathildidae." figureDoi="http://doi.org/10.5281/zenodo.10307672" httpUri="https://zenodo.org/record/10307672/files/figure.png" pageId="21" pageNumber="22">
<subScript id="17935228FFD4C071FED6FE1F7D5536C3" attach="left" box="[335,343,493,507]" fontSize="6" pageId="21" pageNumber="22">3</subScript>
</figureCitation>
</paragraph>
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<subSubSection id="C30D03E6FFD4C071FF0EFDED7D10356E" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="8BA8506DFFD4C071FF0EFDED7D10356E" blockId="21.[151,1437,543,598]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB077743FFD4C071FF0EFDED7D10356E" pageId="21" pageNumber="22">
<taxonomicName id="4C172BEEFFD4C071FF0EFDED7DF7350F" ID-CoL="94LC4" authority="(Lam.)" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[151,501,543,567]" class="Gastropoda" family="Architectonicidae" genus="Architectonica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="carocollata">
<emphasis id="B9638C7FFFD4C071FF0EFDED7DAA350F" box="[151,424,543,567]" italics="true" pageId="21" pageNumber="22">Architectonica carocollata</emphasis>
(Lam.)
</taxonomicName>
juv. an n. sp.—
<treatmentCitation id="0AB6767CFFD4C071FD05FDED7F39350F" author="Baldi, T." box="[668,827,543,567]" page="60" pageId="21" pageNumber="22" year="1960">
<bibRefCitation id="EF862D9CFFD4C071FD05FDED7F39350F" author="Baldi, T." box="[668,827,543,567]" pageId="21" pageNumber="22" pagination="51 - 99" refId="ref48411" refString="Baldi, T. (1960) Tortonische Molluskenfauna von &quot; Badener Tegelfazies' aus Szokolya, Nordungarn. Annales historico-naturales Musei nationalis hungarici, pars Mineralogica et Palaeontologica, 52, 51 - 99. [http: // publication. nhmus. hu / pdf / annHNHM / Annals _ HNHM _ 1960 _ Vol _ 52 _ 51. pdf]" type="journal article" year="1960">Báldi 1960: 60</bibRefCitation>
</treatmentCitation>
, pl. 1, figs 8a–b [
<emphasis id="B9638C7FFFD4C071FC6CFDD27819350F" box="[1013,1051,544,567]" italics="true" pageId="21" pageNumber="22">non</emphasis>
<taxonomicName id="4C172BEEFFD4C071FBBAFDED7D0A356E" authority="(Lamarck, 1822)" baseAuthorityName="Lamarck" baseAuthorityYear="1822" class="Gastropoda" family="Mathildidae" genus="Simplexollata" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="carocollata">
<emphasis id="B9638C7FFFD4C071FBBAFDED7929350F" box="[1059,1323,543,567]" italics="true" pageId="21" pageNumber="22">Simplexollata carocollata</emphasis>
(
<bibRefCitation id="EF862D9CFFD4C071FAA3FDED7CFD356E" author="Lamarck, J. P. B. A. de" pageId="21" pageNumber="22" refId="ref55913" refString="Lamarck, J. P. B. A. de Monet de (1822) Histoire naturelle des animaux sans vertebres ... precedee d'une introduction offrant la determination des caracteres essentiels de l'animal, sa distinction du vegetal et des autres corps naturels, enfin, l'exposition des principes fondamentaux de la zoologie. Lamarck, Paris, 711 pp. [https: // www. biodiversitylibrary. org / item / 47433 # page / 9 / mode / 1 up]" type="book" year="1822">Lamarck, 1822</bibRefCitation>
)
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFD4C071FF0EFD767F57344E" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8BA8506DFFD4C071FF0EFD767D2535FB" blockId="21.[151,1437,644,2038]" pageId="21" pageNumber="22">
<emphasis id="B9638C7FFFD4C071FF0EFD767D4435A6" bold="true" box="[151,326,644,670]" pageId="21" pageNumber="22">Type material.</emphasis>
<materialsCitation id="3B7F5A30FFD4C071FED7FD767D2535FB" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="21" pageNumber="22" specimenCode="NHMW 2016" specimenCount="1" typeStatus="holotype">
<typeStatus id="54ACEECFFFD4C071FED7FD767DB935A6" box="[334,443,644,670]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBB1F816FFD4C071FE5EFD777E6135A6" box="[455,611,644,671]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="22" type="Museum">NHMW 2016</specimenCode>
/0177/0341, SL:
<quantity id="4CEFFD88FFD4C071FCBBFD767F7935A7" box="[802,891,644,671]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="21" pageNumber="22" unit="mm" value="1.8">1.8 mm</quantity>
, MD:
<quantity id="4CEFFD88FFD4C071FC5CFD76781D35A7" box="[965,1055,644,671]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="21" pageNumber="22" unit="mm" value="6.7">6.7 mm</quantity>
,
<location id="8EC806B6FFD4C071FBB5FD7678DE35A6" LSID="urn:lsid:plazi:treatment:03BEE17BFFD4C071FF0EFE447DCE30CE:8EC806B6FFD4C071FBB5FD7678DE35A6" box="[1068,1244,644,670]" country="Romania" name="Lapugiu de Sus" pageId="21" pageNumber="22">
Lăpugiu
<collectorName id="26E235BBFFD4C071FB08FD7678DE35A6" box="[1169,1244,644,670]" pageId="21" pageNumber="22">de Sus</collectorName>
</location>
(
<collectingCountry id="F30010FDFFD4C071FB74FD76795A35A6" box="[1261,1368,644,670]" name="Romania" pageId="21" pageNumber="22">Romania</collectingCountry>
),
<figureCitation id="132C4CE8FFD4C071FAF5FD767CB035FA" captionStart="FIGURE 4" captionStartId="9.[151,250,1590,1614]" captionTargetBox="[151,1436,151,1566]" captionTargetId="figure-233@9.[151,1436,151,1569]" captionTargetPageId="9" captionText="FIGURE 4 Protoconchs of Paratethyan Architectonicidae. A. Ammotectonica gregorovae nov. sp., paratype, NHMW 1862/0029/0078c. B. Simplexollata anticollata nov. sp., paratype, NHMW 1854/0035/0331, Lăpugiu de Sus (Romania). C. Simplexollata carocollata (Lamarck, 1822), NHMW 1850/0018/0115, Léognan (France), Burdigalian. D. Simplexollata simplex (Bronn, 1831), NHMW 2013/0078/0109, Baden (Austria).E. Nipteraxis deformatusnov. sp., paratype, NHMW 2010/0004/0456a, Bad Vöslau (Austria). F. Nipteraxis exmoniliferus (Sacco, 1892), NHMW 1907/0323/1959, M̂llersdorf (Austria). G. Solatectonica kostejana (Boettger, 1907), NHMW 2012/0229/0049b, Traiskirchen (Austria). H. Nipteraxis monilifer (Bronn, 1831), NHMW 1846/0037/0427, M̂llersdorf (Austria). I. Solatisonax tavianii nov. sp., NHMW2016/0266/0002, SL: 8.0 mm, MD: 15.0 mm, Lăpugiu des Sus (Romania). J. Solatisonax transversa nov. sp., holotype, NHMW 2016/0177/0341, Lăpugiu de Sus (Roma- nia). K. Granosolarium semilaevis (Sacco, 1892), NHMW 1846/0037/0428d, M̂llersdorf (Austria). L. Heliacus globosus nov. sp., holotype NHMW 1907/0328/1959a, M̂llersdorf (Austria). M. Heliacus miser (Dujardin, 1837), NHMW 2023/0267/0001, Steinebrunn (Austria). N. Pseudotorinia grasemanni nov. sp., paratype, NHMW 2023/0267/0010, Steinebrunn (Austria). O. Pseudotorinia marthae (Boettger, 1902), NHMW 2012/0229/0049, Traiskirchen (Austria). All figures same scale." figureDoi="http://doi.org/10.5281/zenodo.10147820" httpUri="https://zenodo.org/record/10147820/files/figure.png" pageId="21" pageNumber="22">Figs 4J</figureCitation>
,
<figureCitation id="132C4CE8FFD4C071FF27FD5A7CEF35FA" box="[190,237,680,706]" captionStart="FIGURE 11" captionStartId="22.[151,250,1209,1233]" captionTargetBox="[151,1436,181,1185]" captionTargetId="figure-30@22.[151,1436,181,1185]" captionTargetPageId="22" captionText="FIGURE 11 A1–A3. Solatisonax transversa nov. sp., holotype, NHMW 2016/0177/0341, Lăpugiu de Sus (Romania). B1–B3. Heliacus globosus nov. sp., holotype NHMW 1907/0328/1959a, M̂llersdorf (Austria). C1–C3. Nipteraxis pozsgayae (Kovács Leél-Őssy &amp; Vicián, 2023), holotype, PAL 2022.52, Ottnangian, Parádfürdö, Ilona Valley (Hungary)." figureDoi="http://doi.org/10.5281/zenodo.10147834" httpUri="https://zenodo.org/record/10147834/files/figure.png" pageId="21" pageNumber="22">11A</figureCitation>
<figureCitation id="132C4CE8FFD4C071FF74FD487D1A35FA" box="[237,280,680,712]" captionStart="FIGURE 1" captionStartId="5.[151,250,1522,1546]" captionTargetBox="[232,1366,234,1455]" captionTargetId="figure-58@5.[232,1366,234,1455]" captionTargetPageId="5" captionText="FIGURE 1. Descriptive terms used in descriptions of Architectonicidae (modified from Bieler 1993)." figureDoi="http://doi.org/10.5281/zenodo.10147816" httpUri="https://zenodo.org/record/10147816/files/figure.png" pageId="21" pageNumber="22">
<subScript id="17935228FFD4C071FF74FD487CF735F0" attach="left" box="[237,245,698,712]" fontSize="6" pageId="21" pageNumber="22">1</subScript>
–A
</figureCitation>
<figureCitation id="132C4CE8FFD4C071FE81FD487D2235F0" box="[280,288,698,712]" captionStart="FIGURE 3" captionStartId="6.[151,250,2002,2026]" captionTargetId="figure-99@6.[337,1250,1242,1978]" captionTargetPageId="6" captionText="FIGURE 3. Descriptive terms used in descriptions of Mathildidae." figureDoi="http://doi.org/10.5281/zenodo.10307672" httpUri="https://zenodo.org/record/10307672/files/figure.png" pageId="21" pageNumber="22">
<subScript id="17935228FFD4C071FE81FD487D2235F0" attach="left" box="[280,288,698,712]" fontSize="6" pageId="21" pageNumber="22">3</subScript>
</figureCitation>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA8506DFFD4C071FF5EFD3E7EC23432" blockId="21.[151,1437,644,2038]" pageId="21" pageNumber="22">
<emphasis id="B9638C7FFFD4C071FF5EFD3E7DBB35DE" bold="true" box="[199,441,716,742]" pageId="21" pageNumber="22">Additional material.</emphasis>
<materialsCitation id="3B7F5A30FFD4C071FE59FD3E7EBE3432" country="Hungary" location="Szokolya" municipality="Hungarian Natural History Museum" pageId="21" pageNumber="22" specimenCount="1">
<specimenCount id="9D119BE4FFD4C071FE59FD3E7E1235DF" box="[448,528,716,743]" count="1" pageId="21" pageNumber="22" type="generic">1 spec.</specimenCount>
,
<location id="8EC806B6FFD4C071FD83FD3E7E8035DE" LSID="urn:lsid:plazi:treatment:03BEE17BFFD4C071FF0EFE447DCE30CE:8EC806B6FFD4C071FD83FD3E7E8035DE" box="[538,642,716,742]" country="Hungary" municipality="Hungarian Natural History Museum" name="Szokolya" pageId="21" pageNumber="22">Szokolya</location>
(
<collectingCountry id="F30010FDFFD4C071FD0BFD3E7EFA35DE" box="[658,760,716,742]" name="Hungary" pageId="21" pageNumber="22">Hungary</collectingCountry>
), SL:
<quantity id="4CEFFD88FFD4C071FCA3FD3E7F9035DF" box="[826,914,716,743]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="21" pageNumber="22" unit="mm" value="3.6">3.6 mm</quantity>
, MD: 7.0 mm,
<collectingMunicipality id="6BCCCA17FFD4C071FBA3FD3E7CD73433" pageId="21" pageNumber="22">Hungarian Natural History Museum</collectingMunicipality>
, illustrated in
<bibRefCitation id="EF862D9CFFD4C071FEEDFD027DFF3432" author="Baldi, T." box="[372,509,752,778]" pageId="21" pageNumber="22" pagination="51 - 99" refId="ref48411" refString="Baldi, T. (1960) Tortonische Molluskenfauna von &quot; Badener Tegelfazies' aus Szokolya, Nordungarn. Annales historico-naturales Musei nationalis hungarici, pars Mineralogica et Palaeontologica, 52, 51 - 99. [http: // publication. nhmus. hu / pdf / annHNHM / Annals _ HNHM _ 1960 _ Vol _ 52 _ 51. pdf]" type="journal article" year="1960">Báldi (1960</bibRefCitation>
: pl. 1, figs 8a–b)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA8506DFFD4C071FF5EFCE67EA33416" blockId="21.[151,1437,644,2038]" box="[199,673,788,814]" pageId="21" pageNumber="22">
<materialsCitation id="3B7F5A30FFD4C071FF5EFCE67E9F3416" box="[199,669,788,814]" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="21" pageNumber="22" specimenCount="1" typeStatus="holotype">
<emphasis id="B9638C7FFFD4C071FF5EFCE67D643416" bold="true" box="[199,358,788,814]" pageId="21" pageNumber="22">
<typeStatus id="54ACEECFFFD4C071FF5EFCE67D003416" box="[199,258,788,814]" pageId="21" pageNumber="22">Type</typeStatus>
locality.
</emphasis>
<location id="8EC806B6FFD4C071FEF5FCE67E183416" LSID="urn:lsid:plazi:treatment:03BEE17BFFD4C071FF0EFE447DCE30CE:8EC806B6FFD4C071FEF5FCE67E183416" box="[364,538,788,814]" country="Romania" name="Lapugiu de Sus" pageId="21" pageNumber="22">
Lăpugiu
<collectorName id="26E235BBFFD4C071FE48FCE67E183416" box="[465,538,788,814]" pageId="21" pageNumber="22">de Sus</collectorName>
</location>
(
<collectingCountry id="F30010FDFFD4C071FDB0FCE67E973416" box="[553,661,788,814]" name="Romania" pageId="21" pageNumber="22">Romania</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA8506DFFD4C071FF5EFCCA7EFE346A" blockId="21.[151,1437,644,2038]" box="[199,764,824,850]" pageId="21" pageNumber="22">
<emphasis id="B9638C7FFFD4C071FF5EFCCA7D6D346A" bold="true" box="[199,367,824,850]" pageId="21" pageNumber="22">
<typeStatus id="54ACEECFFFD4C071FF5EFCCA7D00346A" box="[199,258,824,850]" pageId="21" pageNumber="22">Type</typeStatus>
stratum.
</emphasis>
Silt and clay of the Dej Formation.
</paragraph>
<paragraph id="8BA8506DFFD4C071FF5EFCAE7F57344E" blockId="21.[151,1437,644,2038]" box="[199,853,860,886]" pageId="21" pageNumber="22">
<emphasis id="B9638C7FFFD4C071FF5EFCAE7CFF344E" bold="true" box="[199,253,860,886]" pageId="21" pageNumber="22">Age.</emphasis>
Middle Miocene, early/middle Badenian (Langhian).
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFD4C071FF5EFC72788A34A2" box="[199,1160,896,923]" pageId="21" pageNumber="22" type="etymology">
<paragraph id="8BA8506DFFD4C071FF5EFC72788A34A2" blockId="21.[151,1437,644,2038]" box="[199,1160,896,923]" pageId="21" pageNumber="22">
<emphasis id="B9638C7FFFD4C071FF5EFC727D4C34A2" bold="true" box="[199,334,896,922]" pageId="21" pageNumber="22">Etymology.</emphasis>
From Latin
<emphasis id="B9638C7FFFD4C071FE42FC737E5E34A2" box="[475,604,897,922]" italics="true" pageId="21" pageNumber="22">transversus</emphasis>
(= oblique), referring to the prosocline axial ribs.
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFD4C071FF5EFC56798034DA" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8BA8506DFFD4C071FF5EFC56798034DA" blockId="21.[151,1437,644,2038]" pageId="21" pageNumber="22">
<emphasis id="B9638C7FFFD4C071FF5EFC567D403486" bold="true" box="[199,322,932,958]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
Small, moderately elevated lentiform shell with low conical spire, characterized by prosocline axial ribs between beaded SSC and UPC and wide basal field with reduced spiral sculpture and prominent axial folds.
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFD4C071FF5EFC1E7D1832AA" pageId="21" pageNumber="22" type="description">
<paragraph id="8BA8506DFFD4C071FF5EFC1E7D1832AA" blockId="21.[151,1437,644,2038]" pageId="21" pageNumber="22">
<emphasis id="B9638C7FFFD4C071FF5EFC1E7D5B333E" bold="true" box="[199,345,1004,1030]" pageId="21" pageNumber="22">Description.</emphasis>
Small, moderately elevated lentiform shell with low conical spire and angulated periphery. Protoconch, heterostrophic, medium-sized of &lt;1 visible whorls (diameter
<quantity id="4CEFFD88FFD4C071FC0BFBE27FE93312" box="[914,1003,1040,1066]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="21" pageNumber="22" unit="mm" value="1.1">1.1 mm</quantity>
). Teleoconch of about 3.75 whorls. Teleoconch whorls nearly flat. Early teleoconch whorls with beaded SSC, beaded UPC and prominent, nearly smooth LPC. No MCs; instead, mid-whorl with densely spaced, prominent, prosocline axial ribs. SSC fading on second teleoconch whorl. Beads on UPC fade on third teleoconch whorl. Periphery formed by blunt keel; whorls attached slightly below keel. IPC forming distinctly weaker angulation below keel. Base with nearly smooth LPC and IPC, three weaker spiral cords intercalated. Basal field lacking spiral sculpture except for few weak spiral threads close to IPC. Basal field with long, narrow, prominent axial folds strengthening towards umbilicus; partly bifurcating towards IPC. No PUC. UC tubercular, overhanging umbilicus, weakly delimited from basal field by narrow shallow groove. Umbilicus narrowly funnel-shaped, moderately wide (~28% of total diameter). Umbilical wall with prominent prosocline growth lines. Aperture subquadratic with moderate peripheral angulation and distinct, shallow CG, no PG.
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFD4C071FF5EFA6E7D3730B2" pageId="21" pageNumber="22" type="discussion">
<paragraph id="8BA8506DFFD4C071FF5EFA6E7DA1307A" blockId="21.[151,1437,644,2038]" pageId="21" pageNumber="22">
<emphasis id="B9638C7FFFD4C071FF5EFA6E7D4E328E" bold="true" box="[199,332,1436,1462]" pageId="21" pageNumber="22">Discussion.</emphasis>
The placement in
<taxonomicName id="4C172BEEFFD4C071FDBFFA6E7EAB328E" authorityName="Iredale" authorityYear="1931" box="[550,681,1436,1462]" class="Gastropoda" family="Architectonicidae" genus="Solatisonax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFD4C071FDBFFA6E7EAB328E" box="[550,681,1436,1462]" italics="true" pageId="21" pageNumber="22">Solatisonax</emphasis>
</taxonomicName>
is tentative. Axial ribs occur in several extant species, such as
<taxonomicName id="4C172BEEFFD4C071FA1EFA6E7DAB32E2" authority="(Smith, 1891)" baseAuthorityName="Smith" baseAuthorityYear="1891" class="Gastropoda" family="Architectonicidae" genus="Solarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="atkinsoni">
<emphasis id="B9638C7FFFD4C071FA1EFA6E7CFC32E2" italics="true" pageId="21" pageNumber="22">S. atkinsoni</emphasis>
(Smith, 1891)
</taxonomicName>
,
<taxonomicName id="4C172BEEFFD4C071FE20FA327F2032E3" authority="(Thiele, 1925)" baseAuthorityName="Thiele" baseAuthorityYear="1925" box="[441,802,1472,1499]" class="Gastropoda" family="Architectonicidae" genus="Solatisonax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="acutecarinata">
<emphasis id="B9638C7FFFD4C071FE20FA327E7632E2" box="[441,628,1472,1498]" italics="true" pageId="21" pageNumber="22">S. acutecarinata</emphasis>
(Thiele, 1925)
</taxonomicName>
and
<taxonomicName id="4C172BEEFFD4C071FCC6FA32790932E2" authority="(von Martens, 1904)" baseAuthorityName="von Martens" baseAuthorityYear="1904" box="[863,1291,1472,1499]" class="Gastropoda" family="Architectonicidae" genus="Solatisonax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="supraradiata">
<emphasis id="B9638C7FFFD4C071FCC6FA32781332E2" box="[863,1041,1472,1498]" italics="true" pageId="21" pageNumber="22">S. supraradiata</emphasis>
(von
<bibRefCitation id="EF862D9CFFD4C071FBC3FA33790132E2" author="Martens, E. von" box="[1114,1283,1472,1499]" pageId="21" pageNumber="22" pagination="1 - 146" refId="ref56732" refString="Martens, E. von (1904) Die beschalten Gastropoden der deutschen Tiefsee-Expedition, 1898 - 1899. A. Systematisch-geographischer Theil. Wissenschaftliche Ergebnisse der deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899, 7 (A), 1 - 146, pls. 1 - 5. [https: // www. biodiversitylibrary. org / item / 43022 # page / 7 / mode / 1 up]" type="journal article" year="1904">Martens, 1904</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C172BEEFFD4C071FA83FA327D2B32C6" baseAuthorityName="von Martens" baseAuthorityYear="1904" class="Gastropoda" family="Architectonicidae" genus="Solatisonax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="supraradiata">
<emphasis id="B9638C7FFFD4C071FA83FA327D2B32C6" italics="true" pageId="21" pageNumber="22">Solatisonax supraradiata</emphasis>
</taxonomicName>
is especially similar to
<taxonomicName id="4C172BEEFFD4C071FDAAFA167EC732C6" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[563,709,1508,1534]" class="Gastropoda" family="Architectonicidae" genus="Solatisonax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="transversa">
<emphasis id="B9638C7FFFD4C071FDAAFA167EC732C6" box="[563,709,1508,1534]" italics="true" pageId="21" pageNumber="22">S. transversa</emphasis>
</taxonomicName>
concerning the wide basal field with axial ribs and has a comparably prominent IPC (see
<bibRefCitation id="EF862D9CFFD4C071FE2AF9FA7E3A311A" author="Bieler, R." box="[435,568,1544,1570]" pageId="21" pageNumber="22" pagination="1 - 376" refId="ref49146" refString="Bieler, R. (1993) Architectonicidae of the Indo-Pacific (Mollusca, Gastropoda). Abhandlungen des naturwissenschaftlichen Vereins in Hamburg, Neue Folge, 30, 1 - 376." type="journal article" year="1993">Bieler 1993</bibRefCitation>
; fig. 139). The Pliocene Mediterraean
<taxonomicName id="4C172BEEFFD4C071FC46F9FA7986311A" authority="(Chirli, 2013)" baseAuthorityName="Chirli" baseAuthorityYear="2013" box="[991,1412,1544,1570]" class="Gastropoda" family="Architectonicidae" genus="Solatisonax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="pliohumilis">
<emphasis id="B9638C7FFFD4C071FC46F9FA78E4311A" box="[991,1254,1544,1570]" italics="true" pageId="21" pageNumber="22">Solatisonax pliohumilis</emphasis>
(
<bibRefCitation id="EF862D9CFFD4C071FB6AF9FA797F311A" author="Chirli, C." box="[1267,1405,1544,1570]" pageId="21" pageNumber="22" refId="ref50127" refString="Chirli, C. (2013) Malacofauna pliocenica Toscana. Vol. 9. Gastropoda fine Scaphopoda. Chirli, C., Firenze, 169 pp., 24 pls." type="book" year="2013">Chirli, 2013</bibRefCitation>
)
</taxonomicName>
is superficially similar but lacks the angulation by the IPC, has less prominent axial sculpture, and has a narrower umbilicus (see
<bibRefCitation id="EF862D9CFFD4C071FE82F9A27D9C3152" author="Chirli, C." box="[283,414,1616,1642]" pageId="21" pageNumber="22" refId="ref50127" refString="Chirli, C. (2013) Malacofauna pliocenica Toscana. Vol. 9. Gastropoda fine Scaphopoda. Chirli, C., Firenze, 169 pp., 24 pls." type="book" year="2013">Chirli 2013</bibRefCitation>
: pl. 5, figs 1–5).
<taxonomicName id="4C172BEEFFD4C071FDCEF9A278FE3152" authority="Cossmann &amp; Peyrot, 1919" authorityName="Cossmann &amp; Peyrot" authorityYear="1919" box="[599,1276,1616,1643]" class="Gastropoda" family="Architectonicidae" genus="Solarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="peyreirense" subGenus="Nipteraxis">
<emphasis id="B9638C7FFFD4C071FDCEF9A27EBC3152" box="[599,702,1616,1642]" italics="true" pageId="21" pageNumber="22">Solarium</emphasis>
(
<emphasis id="B9638C7FFFD4C071FD54F9A37F403152" box="[717,834,1617,1642]" italics="true" pageId="21" pageNumber="22">Nipteraxis</emphasis>
)
<emphasis id="B9638C7FFFD4C071FCC8F9A37FD03152" box="[849,978,1617,1642]" italics="true" pageId="21" pageNumber="22">peyreirense</emphasis>
<bibRefCitation id="EF862D9CFFD4C071FC4EF9A278FE3152" author="Cossmann, M. &amp; Peyrot, A." box="[983,1276,1616,1643]" pageId="21" pageNumber="22" pagination="213 - 491" refId="ref50539" refString="Cossmann, M. &amp; Peyrot, A. (1919) Conchologie neogenique de l'Aquitaine. Actes de la societe Linneenne de Bordeaux, 70, 213 - 491. [https: // www. biodiversitylibrary. org / item / 103369 # page / 520 / mode / 1 up]" type="journal article" year="1919">Cossmann &amp; Peyrot, 1919</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EF862D9CFFD4C071FA93F9A27D5931B6" author="Cossmann, M. &amp; Peyrot, A." pageId="21" pageNumber="22" pagination="213 - 491" refId="ref50539" refString="Cossmann, M. &amp; Peyrot, A. (1919) Conchologie neogenique de l'Aquitaine. Actes de la societe Linneenne de Bordeaux, 70, 213 - 491. [https: // www. biodiversitylibrary. org / item / 103369 # page / 520 / mode / 1 up]" type="journal article" year="1919">Cossmann &amp; Peyrot 1919: 462</bibRefCitation>
, pl. 15, figs 45–47), from the Chattian of Peyrehorade (
<collectingCountry id="F30010FDFFD4C071FC51F986781831B6" box="[968,1050,1652,1678]" name="France" pageId="21" pageNumber="22">France</collectingCountry>
), is reminiscent of the Paratethyan species in its sculpture on the dorsal side but differs distinctly in its much larger and coarser UC and fewer blunter axial folds on the basal field. Placement in the genus
<taxonomicName id="4C172BEEFFD4C071FD6AF94F782831EE" authority="Cossmann, 1916" authorityName="Cossmann" authorityYear="1916" box="[755,1066,1724,1750]" class="Gastropoda" family="Architectonicidae" genus="Nipteraxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="21" pageNumber="22" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFD4C071FD6AF94F7F6A31EE" box="[755,872,1725,1750]" italics="true" pageId="21" pageNumber="22">Nipteraxis</emphasis>
<bibRefCitation id="EF862D9CFFD4C071FCF6F94E782831EE" author="Cossmann M." box="[879,1066,1724,1750]" pageId="21" pageNumber="22" refId="ref50491" refString="Cossmann M. (1916) Essais de paleoconchologie comparee. Dixieme livraison. Paris, published by the author. 292 pp., 12 pls. [&quot; 1915 &quot;; 1916, https: // www. biodiversitylibrary. org / item / 111980]" type="book" year="1916">Cossmann, 1916</bibRefCitation>
</taxonomicName>
[
<typeStatus id="54ACEECFFFD4C071FBA0F94F786E31EF" box="[1081,1132,1725,1751]" pageId="21" pageNumber="22">type</typeStatus>
species
<taxonomicName id="4C172BEEFFD4C071FB55F94E7D4131C2" authority="Lamarck, 1804" authorityName="Lamarck" authorityYear="1804" class="Gastropoda" family="Architectonicidae" genus="Solarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="plicatum">
<emphasis id="B9638C7FFFD4C071FB55F94E799E31EE" box="[1228,1436,1724,1750]" italics="true" pageId="21" pageNumber="22">Solarium plicatum</emphasis>
<bibRefCitation id="EF862D9CFFD4C071FF0EF9127D4131C2" author="Lamarck, J. - B." box="[151,323,1760,1786]" pageId="21" pageNumber="22" pagination="163 - 170" refId="ref55809" refString="Lamarck, J. - B. (1804) Memoires sur les fossiles des environs de Paris (suite 2). Annales du museum d'histoire naturelle, 3, 163 - 170 + 266 - 274 + 343 - 352 + 436 - 441. [https: // biodiversitylibrary. org / page / 14762943]" type="journal article" year="1804">Lamarck, 1804</bibRefCitation>
</taxonomicName>
; Eocene,
<collectingCountry id="F30010FDFFD4C071FE36F9137E0331C3" box="[431,513,1761,1787]" name="France" pageId="21" pageNumber="22">France</collectingCountry>
], as proposed by
<bibRefCitation id="EF862D9CFFD4C071FD58F9127FF731C2" author="Cossmann, M. &amp; Peyrot, A." box="[705,1013,1760,1787]" pageId="21" pageNumber="22" pagination="213 - 491" refId="ref50539" refString="Cossmann, M. &amp; Peyrot, A. (1919) Conchologie neogenique de l'Aquitaine. Actes de la societe Linneenne de Bordeaux, 70, 213 - 491. [https: // www. biodiversitylibrary. org / item / 103369 # page / 520 / mode / 1 up]" type="journal article" year="1919">Cossmann &amp; Peyrot (1919)</bibRefCitation>
, is rejected herein for ‘
<taxonomicName id="4C172BEEFFD4C071FB6EF9127CF33026" authorityName="Cossmann &amp; Peyrot" authorityYear="1919" class="Gastropoda" family="Architectonicidae" genus="Solarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="peyreirense">
<emphasis id="B9638C7FFFD4C071FB6EF912795C31C2" box="[1271,1374,1760,1786]" italics="true" pageId="21" pageNumber="22">Solarium</emphasis>
’
<emphasis id="B9638C7FFFD4C071FAF5F9137CF33026" italics="true" pageId="21" pageNumber="22">peyreirense</emphasis>
</taxonomicName>
and for
<taxonomicName id="4C172BEEFFD4C071FEC9F8F67E523026" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[336,592,1796,1822]" class="Gastropoda" family="Architectonicidae" genus="Solatisonax" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="transversa">
<emphasis id="B9638C7FFFD4C071FEC9F8F67E523026" box="[336,592,1796,1822]" italics="true" pageId="21" pageNumber="22">Solatisonax transversa</emphasis>
</taxonomicName>
, as
<taxonomicName id="4C172BEEFFD4C071FDE4F8F77F483026" box="[637,842,1796,1822]" pageId="21" pageNumber="22">
<emphasis id="B9638C7FFFD4C071FDE4F8F77EF03026" box="[637,754,1797,1822]" italics="true" pageId="21" pageNumber="22">Nipteraxis</emphasis>
species
</taxonomicName>
have a wider umbilicus and prominent beaded spiral cords on the basal field.
</paragraph>
<paragraph id="8BA8506DFFD4C071FF5EF8BE7D3730B2" blockId="21.[151,1437,644,2038]" pageId="21" pageNumber="22">
<emphasis id="B9638C7FFFD4C071FF5EF8BE7D65305E" bold="true" box="[199,359,1868,1894]" pageId="21" pageNumber="22">Paleoecology.</emphasis>
Unknown; probably outer neritic environments, which are typical for many
<taxonomicName id="4C172BEEFFD4C071FB5BF8BE799E305E" box="[1218,1436,1868,1894]" pageId="21" pageNumber="22">
<emphasis id="B9638C7FFFD4C071FB5BF8BE7947305E" box="[1218,1349,1868,1894]" italics="true" pageId="21" pageNumber="22">Solatisonax</emphasis>
species
</taxonomicName>
(
<bibRefCitation id="EF862D9CFFD4C071FF06F8827D2A30B2" author="Bieler, R." box="[159,296,1904,1930]" pageId="21" pageNumber="22" pagination="1 - 376" refId="ref49146" refString="Bieler, R. (1993) Architectonicidae of the Indo-Pacific (Mollusca, Gastropoda). Abhandlungen des naturwissenschaftlichen Vereins in Hamburg, Neue Folge, 30, 1 - 376." type="journal article" year="1993">Bieler 1993</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFD4C071FF5EF8667DCE30CE" pageId="21" pageNumber="22" type="distribution">
<paragraph id="8BA8506DFFD4C071FF5EF86679523096" blockId="21.[151,1437,644,2038]" box="[199,1360,1940,1966]" pageId="21" pageNumber="22">
<emphasis id="B9638C7FFFD4C071FF5EF8667D633096" bold="true" box="[199,353,1940,1966]" pageId="21" pageNumber="22">Distribution.</emphasis>
Only known from the Langhian equivalent of the Badenian in the Central Paratethys Sea.
</paragraph>
<paragraph id="8BA8506DFFD4C071FF5EF84A7DCE30CE" blockId="21.[151,1437,644,2038]" pageId="21" pageNumber="22">
<emphasis id="B9638C7FFFD4C071FF5EF84A7DE530EA" bold="true" box="[199,487,1976,2002]" pageId="21" pageNumber="22">Central Paratethys Sea.</emphasis>
Badenian (Middle Miocene):
<emphasis id="B9638C7FFFD4C071FCD8F84B780730EA" box="[833,1029,1977,2002]" italics="true" pageId="21" pageNumber="22">Pannonian Basin</emphasis>
: Szokolya (
<collectingCountry id="F30010FDFFD4C071FB16F84A78F730EA" box="[1167,1269,1976,2002]" name="Hungary" pageId="21" pageNumber="22">Hungary</collectingCountry>
),
<emphasis id="B9638C7FFFD4C071FA92F84A799730EA" box="[1291,1429,1976,2002]" italics="true" pageId="21" pageNumber="22">Făget Basin</emphasis>
: Lăpugiu de Sus (
<collectingCountry id="F30010FDFFD4C071FECDF82E7DC230CE" box="[340,448,2012,2038]" name="Romania" pageId="21" pageNumber="22">Romania</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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