treatments-xml/data/03/B8/59/03B8593794445F7D3EAFFF143CA2A8EF.xml

<document id="1D0008FC29118AEFB2A3A6B668272D6B" ID-CLB-Dataset="9440" ID-DOI="10.11646/zootaxa.4885.4.4" ID-GBIF-Dataset="e5646187-d56d-4e50-9cb9-9104a026b833" ID-ISSN="1175-5326" ID-Zenodo-Dep="4296928" ID-ZooBank="4D338244-A880-4E31-AABB-FAFC06AF4E3F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606683977728" checkinUser="plazi" docAuthor="Kaplin, Vladimir &amp; Vargovitsh, Robert S." docDate="2020" docId="03B8593794445F7D3EAFFF143CA2A8EF" docLanguage="en" docName="zootaxa.4885.4.4.pdf" docOrigin="Zootaxa 4885 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coryphophthalmus troglophilus Kaplin &amp; Vargovitsh 2020, sp. nov." docType="treatment" docVersion="9" lastPageNumber="536" masterDocId="FF81214F94465F7A3E38FF833E7BA83A" masterDocTitle="New species of bristletails of the family Machilidae (Microcoryphia) from caves in Abkhazia and Ukraine" masterLastPageNumber="540" masterPageNumber="530" pageNumber="532" updateTime="1698846852124" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3F7D51684612782A58E90C651DE349C4">New species of bristletails of the family Machilidae (Microcoryphia) from caves in Abkhazia and Ukraine</mods:title>
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<mods:namePart id="22FA6D7AEAD9F4A132CBFE642DE4EE2B">Vargovitsh, Robert S.</mods:namePart>
<mods:affiliation id="9FEBD4FE66781EBCADBD1FC6D90B8577">Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, 15, B. Khmelnitsky Str., Kyiv, 01030 Ukraine. arete @ izan. kiev. ua; https: // orcid. org / 0000 - 0002 - 4921 - 0098</mods:affiliation>
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<treatment id="03B8593794445F7D3EAFFF143CA2A8EF" ID-DOI="http://doi.org/10.5281/zenodo.4324163" ID-GBIF-Taxon="170676424" ID-Zenodo-Dep="4324163" LSID="urn:lsid:plazi:treatment:03B8593794445F7D3EAFFF143CA2A8EF" httpUri="http://treatment.plazi.org/id/03B8593794445F7D3EAFFF143CA2A8EF" lastPageId="7" lastPageNumber="536" pageId="2" pageNumber="532">
<subSubSection id="C30BBBAA94445F783EAFFF143C9FA88B" box="[151,740,151,178]" pageId="2" pageNumber="532" type="nomenclature">
<paragraph id="8BAEE82194445F783EAFFF143C9FA88B" blockId="2.[151,740,151,213]" box="[151,740,151,178]" pageId="2" pageNumber="532">
<heading id="D0E65F4D94445F783EAFFF143C9FA88B" bold="true" box="[151,740,151,178]" fontSize="11" level="1" pageId="2" pageNumber="532" reason="1">
<emphasis id="B965343394445F783EAFFF143C9FA88B" bold="true" box="[151,740,151,178]" pageId="2" pageNumber="532">
<taxonomicName id="4C1193A294445F783EAFFF143C01A88B" authority="Kaplin" authorityName="Kaplin &amp; Vargovitsh" authorityYear="2020" box="[151,634,151,178]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="532" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov.">
<emphasis id="B965343394445F783EAFFF143C60A888" bold="true" box="[151,539,151,178]" italics="true" pageId="2" pageNumber="532">Coryphophthalmus troglophilus</emphasis>
Kaplin
</taxonomicName>
, 
<taxonomicNameLabel id="A256894894445F783CBFFF143C9FA88B" box="[647,740,151,177]" pageId="2" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30BBBAA94445F783EAFFF383F75A8EF" box="[151,270,187,213]" pageId="2" pageNumber="532" type="description">
<paragraph id="8BAEE82194445F783EAFFF383F75A8EF" blockId="2.[151,740,151,213]" box="[151,270,187,213]" pageId="2" pageNumber="532">
<figureCitation id="132AF4A494445F783EAFFF383F75A8EF" box="[151,270,187,213]" captionStart-0="FIGURE 2" captionStart-1="FIGURES 3–9" captionStart-2="FIGURES 10–14" captionStartId-0="2.[151,250,1875,1899]" captionStartId-1="3.[151,264,1943,1967]" captionStartId-2="4.[151,264,1943,1967]" captionTargetBox-0="[151,1436,633,1852]" captionTargetBox-1="[218,1368,194,1910]" captionTargetBox-2="[254,1334,192,1907]" captionTargetId-0="figure-185@2.[151,1436,633,1852]" captionTargetId-1="figure-17@3.[209,1378,181,1919]" captionTargetId-2="figure-17@4.[243,1343,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 2. Coryphophthalmus troglophilus sp. nov. A—holotype, male, in situ; B—paratypes, females, in situ (photos R. S. Vargovitsh, 20.09.2018)" captionText-1="FIGURES 3–9. Coryphophthalmus troglophilus sp. nov. 3–5, 7–9, holotype male; 6, paratype female. 3) Eyes and paired ocelli, front view. 4) Maxillary palp. 5) Labial palp, with part of labium. 6, 7) Distal part of mandible. 8) Tarsus and tibia of hind leg. 9) Pretarsa of hind leg. Scale bars: 0.1 mm." captionText-2="FIGURES 10–14. Coryphophthalmus troglophilus sp. nov. 10–12, paratype female; 13, 14, holotype, male. 10) Urocoxite IX and anterior gonapophyses. 11) Distal divisions of anterior gonapophyse. 12) Distal divisions of posterior gonapophyse. 13) Urocoxite IX and paramere. 14) Penis. Scale bars: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4296932" figureDoi-1="http://doi.org/10.5281/zenodo.4296934" figureDoi-2="http://doi.org/10.5281/zenodo.4296936" httpUri-0="https://zenodo.org/record/4296932/files/figure.png" httpUri-1="https://zenodo.org/record/4296934/files/figure.png" httpUri-2="https://zenodo.org/record/4296936/files/figure.png" pageId="2" pageNumber="532">Figs. 2–14</figureCitation>
</paragraph>
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<subSubSection id="C30BBBAA94445F783EAFFE803AC0A9B3" pageId="2" pageNumber="532" type="diagnosis">
<paragraph id="8BAEE82194445F783EAFFE803AC0A9B3" blockId="2.[151,1436,259,610]" pageId="2" pageNumber="532">
<emphasis id="B965343394445F783EAFFE803F69A927" bold="true" box="[151,274,259,285]" pageId="2" pageNumber="532">Diagnosis.</emphasis>
<taxonomicName id="4C1193A294445F783F21FE803CFBA927" authority="Kaplin &amp; Vargovitsh, 2020" authorityName="Kaplin &amp; Vargovitsh" authorityYear="2020" box="[281,640,259,285]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="532" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov.">
<emphasis id="B965343394445F783F21FE803CFBA927" box="[281,640,259,285]" italics="true" pageId="2" pageNumber="532">Coryphophthalmus troglophilus</emphasis>
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<taxonomicNameLabel id="A256894894445F783CB0FE803C98A927" box="[648,739,259,285]" pageId="2" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
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, with 2 + 2 eversible vesicles on urocoxites II–IV, belongs to the subgenus 
<taxonomicName id="4C1193A294445F783F17FEA43C7DA97B" authorityName="Verhoeff" authorityYear="1910" box="[303,518,295,321]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="532" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B965343394445F783F17FEA43C7DA97B" box="[303,518,295,321]" italics="true" pageId="2" pageNumber="532">Coryphophthalmus</emphasis>
</taxonomicName>
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, a group of species with long thin chaetae on the legs, clypeus and maxillary palpomeres 2‒4, but lacking on male labial palp; without needle-like macrochaetae on legs. Only one species, 
<taxonomicName id="4C1193A294445F783BBBFECF3FBBA9B3" authority="(Kaplin, 2017)" baseAuthorityName="Kaplin" baseAuthorityYear="2017" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="532" phylum="Arthropoda" rank="species" species="abchasicus">
<emphasis id="B965343394445F783BBBFECF3F68A9B3" italics="true" pageId="2" pageNumber="532">C. abchasicus</emphasis>
(
<bibRefCitation id="EF8095D094445F783F1AFEEC3FC3A9B3" author="Kaplin, V. G." box="[290,440,367,393]" pageId="2" pageNumber="532" pagination="207 - 229" refId="ref4714" refString="Kaplin, V. G. (2017) New species of the bristletail family Machilidae (Insecta, Microcoryphia) from Abkhazia. Entomological Review, 97 (2), 207 - 229. https: // doi. org / 10.1134 / S 0013873817020075" type="journal article" year="2017">Kaplin, 2017</bibRefCitation>
)
</taxonomicName>
is affiliated to this group other than the new species (
<bibRefCitation id="EF8095D094445F783A2FFEEC3AD4A9B3" author="Kaplin, V. G." box="[1047,1199,367,393]" pageId="2" pageNumber="532" pagination="207 - 229" refId="ref4714" refString="Kaplin, V. G. (2017) New species of the bristletail family Machilidae (Insecta, Microcoryphia) from Abkhazia. Entomological Review, 97 (2), 207 - 229. https: // doi. org / 10.1134 / S 0013873817020075" type="journal article" year="2017">Kaplin, 2017</bibRefCitation>
).
</paragraph>
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<subSubSection id="C30BBBAA94445F7C3EFFFE103C14ADB9" lastPageId="6" lastPageNumber="536" pageId="2" pageNumber="532" type="description">
<paragraph id="8BAEE82194445F783EFFFE103D03AA58" blockId="2.[151,1436,259,610]" pageId="2" pageNumber="532">
<emphasis id="B965343394445F783EFFFE103F22A997" bold="true" box="[199,345,403,429]" pageId="2" pageNumber="532">Description.</emphasis>
Body length of males 7.0– 
<quantity id="4CE945C494445F783CBDFE103CA0A994" box="[645,731,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="2" pageNumber="532" unit="mm" value="7.3">7.3 mm</quantity>
, females 
<quantity id="4CE945C494445F783D7CFE103DB6A994" box="[836,973,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="7.699999999999999" metricValueMax="8.2" metricValueMin="7.2" pageId="2" pageNumber="532" unit="mm" value="7.699999999999999" valueMax="8.2" valueMin="7.2">7.2–8.2 mm</quantity>
; width of males and females 
<quantity id="4CE945C494445F783B37FE103BE3A994" box="[1295,1432,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="2" pageNumber="532" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8–2.1 mm</quantity>
; antennae length of males 
<quantity id="4CE945C494445F783F84FE343C3DA9E8" box="[444,582,439,466]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" metricValueMax="8.7" metricValueMin="8.5" pageId="2" pageNumber="532" unit="mm" value="8.6" valueMax="8.7" valueMin="8.5">8.5–8.7 mm</quantity>
, female 
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(slightly broken); cercus length of males 
<quantity id="4CE945C494445F783AE6FE343B43A9E8" box="[1246,1336,439,466]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="532" unit="mm" value="3.8">3.8 mm</quantity>
, females 
<quantity id="4CE945C494445F783EAFFE583F5FA9CC" box="[151,292,475,502]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.2" metricValueMin="3.6" pageId="2" pageNumber="532" unit="mm" value="3.9" valueMax="4.2" valueMin="3.6">3.6–4.2 mm</quantity>
; total eyes width of males 
<quantity id="4CE945C494445F783C64FE583D7FA9CC" box="[604,772,475,502]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" metricValueMax="8.8" metricValueMin="8.6" pageId="2" pageNumber="532" unit="mm" value="0.87" valueMax="0.88" valueMin="0.86">0.86–0.88 mm</quantity>
, females 0.86–1.00 mm; eye length of males and females 0.43–0.46 and 
<quantity id="4CE945C494445F783F7AFE7C3F92AA20" box="[322,489,511,538]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="2" pageNumber="532" unit="mm" value="0.45999999999999996" valueMax="0.48" valueMin="0.44">0.44–0.48 mm</quantity>
, respectively; paired ocelli width of males 
<quantity id="4CE945C494445F783DE2FE7C3AFAAA20" box="[986,1153,511,538]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="2" pageNumber="532" unit="mm" value="0.265" valueMax="0.27" valueMin="0.26">0.26–0.27 mm</quantity>
, females 
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; paired ocellus length of males and females 0.15–0.16 and 
<quantity id="4CE945C494445F783D11FDA03DB4AA04" box="[809,975,547,574]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="2" pageNumber="532" unit="mm" value="0.135" valueMax="0.14" valueMin="0.13">0.13–0.14 mm</quantity>
, respectively; coxal styli length of males 0.65–0.70, females 
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; ovipositor length 
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.
</paragraph>
<caption id="DF6EB8A994445F783EAFF8D03FF7AFAA" ID-DOI="http://doi.org/10.5281/zenodo.4296932" ID-Zenodo-Dep="4296932" httpUri="https://zenodo.org/record/4296932/files/figure.png" pageId="2" pageNumber="532" startId="2.[151,250,1875,1899]" targetBox="[151,1436,633,1852]" targetPageId="2">
<paragraph id="8BAEE82194445F783EAFF8D03FF7AFAA" blockId="2.[151,1436,1875,1936]" pageId="2" pageNumber="532">
<emphasis id="B965343394445F783EAFF8D03F6FAF51" bold="true" box="[151,276,1875,1899]" pageId="2" pageNumber="532">FIGURE 2.</emphasis>
<taxonomicName id="4C1193A294445F783F23F8D03C24AF51" authority="Kaplin &amp; Vargovitsh, 2020" authorityName="Kaplin &amp; Vargovitsh" authorityYear="2020" box="[283,607,1875,1899]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="532" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov.">
<emphasis id="B965343394445F783F23F8D03C24AF51" box="[283,607,1875,1899]" italics="true" pageId="2" pageNumber="532">Coryphophthalmus troglophilus</emphasis>
</taxonomicName>
<emphasis id="B965343394445F783C5EF8D73CCCAF56" bold="true" box="[614,695,1876,1900]" pageId="2" pageNumber="532">
<taxonomicNameLabel id="A256894894445F783C5EF8D73CCCAF56" box="[614,695,1876,1900]" pageId="2" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A—holotype, male, 
<emphasis id="B965343394445F783DB6F8D73DB6AF51" box="[910,973,1876,1899]" italics="true" pageId="2" pageNumber="532">in situ</emphasis>
; B—paratypes, females, 
<emphasis id="B965343394445F783AE8F8D73B6BAF51" box="[1232,1296,1876,1899]" italics="true" pageId="2" pageNumber="532">in situ</emphasis>
(photos R. S. Vargovitsh, 20.09.2018)
</paragraph>
</caption>
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<paragraph id="8BAEE82194455F793EAFF8143C35AFCD" blockId="3.[151,1437,1943,2040]" pageId="3" pageNumber="533">
<emphasis id="B965343394455F793EAFF8143F41AF95" bold="true" box="[151,314,1943,1967]" pageId="3" pageNumber="533">FIGURES 3–9.</emphasis>
<taxonomicName id="4C1193A294455F793F07F81B3CFAAF8A" authority="Kaplin &amp; Vargovitsh, 2020" authorityName="Kaplin &amp; Vargovitsh" authorityYear="2020" box="[319,641,1944,1968]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov.">
<emphasis id="B965343394455F793F07F81B3CFAAF8A" box="[319,641,1944,1968]" italics="true" pageId="3" pageNumber="533">Coryphophthalmus troglophilus</emphasis>
</taxonomicName>
<emphasis id="B965343394455F793CBEF81B3CAEAF8A" bold="true" box="[646,725,1944,1968]" pageId="3" pageNumber="533">
<taxonomicNameLabel id="A256894894455F793CBEF81B3CAEAF8A" box="[646,725,1944,1968]" pageId="3" pageNumber="533" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
3–5, 7–9, holotype male; 6, paratype female. 3) Eyes and paired ocelli, front view. 4) Maxillary palp. 5) Labial palp, with part of labium. 6, 7) Distal part of mandible. 8) Tarsus and tibia of hind leg. 9) Pretarsa of hind leg. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="DF6EB8A994425F7E3EAFF8143C94AFCD" ID-DOI="http://doi.org/10.5281/zenodo.4296936" ID-Zenodo-Dep="4296936" httpUri="https://zenodo.org/record/4296936/files/figure.png" pageId="4" pageNumber="534" startId="4.[151,264,1943,1967]" targetBox="[254,1334,192,1907]" targetPageId="4">
<paragraph id="8BAEE82194425F7E3EAFF8143C94AFCD" blockId="4.[151,1437,1943,2040]" pageId="4" pageNumber="534">
<emphasis id="B965343394425F7E3EAFF8143F2EAF95" bold="true" box="[151,341,1943,1967]" pageId="4" pageNumber="534">FIGURES 10–14.</emphasis>
<taxonomicName id="4C1193A294425F7E3F63F81B3CE4AF8A" authority="Kaplin &amp; Vargovitsh, 2020" authorityName="Kaplin &amp; Vargovitsh" authorityYear="2020" box="[347,671,1944,1968]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov.">
<emphasis id="B965343394425F7E3F63F81B3CE4AF8A" box="[347,671,1944,1968]" italics="true" pageId="4" pageNumber="534">Coryphophthalmus troglophilus</emphasis>
</taxonomicName>
<emphasis id="B965343394425F7E3C9EF81B3C8CAF8A" bold="true" box="[678,759,1944,1968]" pageId="4" pageNumber="534">
<taxonomicNameLabel id="A256894894425F7E3C9EF81B3C8CAF8A" box="[678,759,1944,1968]" pageId="4" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
10–12, paratype female; 13, 14, holotype, male. 10) Urocoxite IX and anterior gonapophyses. 11) Distal divisions of anterior gonapophyse. 12) Distal divisions of posterior gonapophyse. 13) Urocoxite IX and paramere. 14) Penis. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BAEE82194435F7F3EFFFF143D77A9EB" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="535">
General body color (in ethanol) whitish or light yellow, almost without hypodermal pigment. Antennal base, frons, gena, lateral sides of clypeus, basal parts of mandibles, thoracic sterna with brown-violet hypodermal pigment of light to medium intensity. Scale color on surface of body grey, dark grey, almost white and black, light brown or brown, spotted on the upper side of body (
<figureCitation id="132AF4A494435F7F3C4BFE803CC2A927" box="[627,697,259,285]" captionStart="FIGURE 2" captionStartId="2.[151,250,1875,1899]" captionTargetBox="[151,1436,633,1852]" captionTargetId="figure-185@2.[151,1436,633,1852]" captionTargetPageId="2" captionText="FIGURE 2. Coryphophthalmus troglophilus sp. nov. A—holotype, male, in situ; B—paratypes, females, in situ (photos R. S. Vargovitsh, 20.09.2018)" figureDoi="http://doi.org/10.5281/zenodo.4296932" httpUri="https://zenodo.org/record/4296932/files/figure.png" pageId="5" pageNumber="535">Fig. 2</figureCitation>
). Antennae of males and females slightly longer than body. Distal chains of male and female flagella divided into 8–12 annuli. Аpical, 3rd, 5th and 7th annuli of distal chains with one or two basiconic sensilla of D form; each annulus of chains also with one “rosette-shaped” sensillum. Clypeus and labrum of male with long thin chaetae. Cerci of males and females about 0.50–0.55 times body length, divided into 22–23 annuli, with apically bifurcated spikes. Two distal annuli of cerci without lateral hyaline spines. Remaining cercal annuli with 1–5 inner lateral spines in both sexes.
</paragraph>
<paragraph id="8BAEE82194435F7F3EFFFE583D8AAABF" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="535">
Compound eyes round, green or dark green (in ethanol). Length to width ratio of compound eye 1.00– 
<quantity id="4CE945C494435F7F3B77FE583BE7A9CF" box="[1359,1436,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="2.6162" pageId="5" pageNumber="535" unit="in" value="1.03">1.03 in</quantity>
both sexes; ratio of line of contact to length of male and female eyes 0.50–0.53 and 0.42–0.46, respectively. Paired ocelli submedian, pyriform, dark brown with narrow white borders. Ratios of width to length of male and female ocelli 1.7 and 1.8, respectively. Ratio of distance between inner and outer margins of ocelli to total width of compound eyes 0.16–0.18 and 0.64–0.66 eye in both sexes, respectively (
<figureCitation id="132AF4A494435F7F3DA7FDE83D98AABF" box="[927,995,619,645]" captionStart="FIGURES 3–9" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[218,1368,194,1910]" captionTargetId="figure-17@3.[209,1378,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–9. Coryphophthalmus troglophilus sp. nov. 3–5, 7–9, holotype male; 6, paratype female. 3) Eyes and paired ocelli, front view. 4) Maxillary palp. 5) Labial palp, with part of labium. 6, 7) Distal part of mandible. 8) Tarsus and tibia of hind leg. 9) Pretarsa of hind leg. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4296934" httpUri="https://zenodo.org/record/4296934/files/figure.png" pageId="5" pageNumber="535">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BAEE82194435F7F3EFFFD0C3CFDABBB" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="535">
Apical maxillary palpomere 0.55–0.65 times (male) and 0.72–0.75 times (female) that of the preceding one, ratio of length of 5th and 4th palpomeres 
<quantity id="4CE945C494435F7F3C77FD303CF5AAF7" box="[591,654,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="3.302" pageId="5" pageNumber="535" unit="in" value="1.3">1.3 in</quantity>
male and 
<quantity id="4CE945C494435F7F3D38FD303D3BAAF7" box="[768,832,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="3.5559999999999996" pageId="5" pageNumber="535" unit="in" value="1.4">1.4 in</quantity>
female. Dorsal surface of 7th, 6th and 5th palpomeres of maxillary palp with 10–11, 13–14 and 3–4 hyaline spines in male, and 12–14, 14–16, and 4–7 spines in female, respectively. Underside of palpomeres 2–4 of male maxillary palp with relatively numerous and long thin chaetae, missing on the dorsal surface of male labial palpomeres (
<figureCitation id="132AF4A494435F7F3D2DFC9C3D21AB03" box="[789,858,799,825]" captionStart="FIGURES 3–9" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[218,1368,194,1910]" captionTargetId="figure-17@3.[209,1378,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–9. Coryphophthalmus troglophilus sp. nov. 3–5, 7–9, holotype male; 6, paratype female. 3) Eyes and paired ocelli, front view. 4) Maxillary palp. 5) Labial palp, with part of labium. 6, 7) Distal part of mandible. 8) Tarsus and tibia of hind leg. 9) Pretarsa of hind leg. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4296934" httpUri="https://zenodo.org/record/4296934/files/figure.png" pageId="5" pageNumber="535">Fig. 4</figureCitation>
). Apical labial palpomere triangularly oval, 2.3–2.5 times (male) and 2.6–2.7 times (female) longer than wide, with 30–35 sensorial cones (
<figureCitation id="132AF4A494435F7F3A61FCC03AE7AB67" box="[1113,1180,834,861]" captionStart="FIGURES 3–9" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[218,1368,194,1910]" captionTargetId="figure-17@3.[209,1378,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–9. Coryphophthalmus troglophilus sp. nov. 3–5, 7–9, holotype male; 6, paratype female. 3) Eyes and paired ocelli, front view. 4) Maxillary palp. 5) Labial palp, with part of labium. 6, 7) Distal part of mandible. 8) Tarsus and tibia of hind leg. 9) Pretarsa of hind leg. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4296934" httpUri="https://zenodo.org/record/4296934/files/figure.png" pageId="5" pageNumber="535">Fig. 5</figureCitation>
). Mandibles with three teeth in female and four in male (
<figureCitation id="132AF4A494435F7F3C34FCE43C0CABBB" box="[524,631,871,897]" captionStart="FIGURES 3–9" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[218,1368,194,1910]" captionTargetId="figure-17@3.[209,1378,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–9. Coryphophthalmus troglophilus sp. nov. 3–5, 7–9, holotype male; 6, paratype female. 3) Eyes and paired ocelli, front view. 4) Maxillary palp. 5) Labial palp, with part of labium. 6, 7) Distal part of mandible. 8) Tarsus and tibia of hind leg. 9) Pretarsa of hind leg. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4296934" httpUri="https://zenodo.org/record/4296934/files/figure.png" pageId="5" pageNumber="535">Figs. 6, 7</figureCitation>
).
</paragraph>
<paragraph id="8BAEE82194435F7F3EFFFC083FBEAC9B" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="535">
Fore and middle femora and tibiae of female widened. Hind tarsus 1.2–1.3 times longer than fore and middle tarsi in both sexes. Fore tibia longer than middle tibia 1.3 times in female and 1.2 times in male. Hind tibia longer than middle tibia, 1.5 times in female and 1.3 times in male. Fore femur of male without sensory field. Ratios of length to width of femur, tibia and tarsus given in 
<tableCitation id="C693DD9A94435F7F3CFBFC743D6DAC2B" box="[707,790,1015,1041]" captionStart="TABLE 1" captionStartId="5.[151,239,1232,1256]" captionTargetPageId="5" captionText="TABLE 1. Ratios of length to width of main leg segments of Coryphophthalmus troglophilus sp. nov." httpUri="http://table.plazi.org/id/DF6EB8A994435F7F3EAFFB533AA3ACD3" pageId="5" pageNumber="535" tableUuid="DF6EB8A994435F7F3EAFFB533AA3ACD3">Table 1</tableCitation>
. Ratio of length of apical hind tarsomere to total length of hind tarsus 
<quantity id="4CE945C494435F7F3F21FB983F1CAC0F" box="[281,359,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="8.128" pageId="5" pageNumber="535" unit="in" value="0.32">0.32 in</quantity>
male and 
<quantity id="4CE945C494435F7F3FE4FB993C51AC0F" box="[476,554,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="8.889999999999999" pageId="5" pageNumber="535" unit="in" value="0.35">0.35 in</quantity>
female (
<figureCitation id="132AF4A494435F7F3CB5FB983CAAAC0F" box="[653,721,1051,1077]" captionStart="FIGURES 3–9" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[218,1368,194,1910]" captionTargetId="figure-17@3.[209,1378,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–9. Coryphophthalmus troglophilus sp. nov. 3–5, 7–9, holotype male; 6, paratype female. 3) Eyes and paired ocelli, front view. 4) Maxillary palp. 5) Labial palp, with part of labium. 6, 7) Distal part of mandible. 8) Tarsus and tibia of hind leg. 9) Pretarsa of hind leg. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4296934" httpUri="https://zenodo.org/record/4296934/files/figure.png" pageId="5" pageNumber="535">Fig. 8</figureCitation>
). Trochanter and femur of male with long, thin chaetae. Ventral surface of femora, tibiae and tarsi without spine-like macrochaetae in both sexes. All tibiae with 1–2 lateral, relatively long, colorless, thickened macrochaetae. Pretarsa with two well-developed claws and arolium with flexible pad between them (
<figureCitation id="132AF4A494435F7F3F4AFB043FCCAC9B" box="[370,439,1159,1185]" captionStart="FIGURES 3–9" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[218,1368,194,1910]" captionTargetId="figure-17@3.[209,1378,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–9. Coryphophthalmus troglophilus sp. nov. 3–5, 7–9, holotype male; 6, paratype female. 3) Eyes and paired ocelli, front view. 4) Maxillary palp. 5) Labial palp, with part of labium. 6, 7) Distal part of mandible. 8) Tarsus and tibia of hind leg. 9) Pretarsa of hind leg. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4296934" httpUri="https://zenodo.org/record/4296934/files/figure.png" pageId="5" pageNumber="535">Fig. 9</figureCitation>
).
</paragraph>
<caption id="DF6EB8A994435F7F3EAFFB533AA3ACD3" ID-Table-UUID="DF6EB8A994435F7F3EAFFB533AA3ACD3" box="[151,1240,1232,1257]" httpUri="http://table.plazi.org/id/DF6EB8A994435F7F3EAFFB533AA3ACD3" pageId="5" pageNumber="535" targetBox="[166,1231,1273,1699]" targetIsTable="true" targetPageId="5">
<paragraph id="8BAEE82194435F7F3EAFFB533AA3ACD3" blockId="5.[151,1240,1232,1257]" box="[151,1240,1232,1257]" pageId="5" pageNumber="535">
<emphasis id="B965343394435F7F3EAFFB533F71ACD2" bold="true" box="[151,266,1232,1256]" pageId="5" pageNumber="535">TABLE 1.</emphasis>
Ratios of length to width of main leg segments of 
<taxonomicName id="4C1193A294435F7F3D10FB533A07ACD2" authority="Kaplin &amp; Vargovitsh, 2020" authorityName="Kaplin &amp; Vargovitsh" authorityYear="2020" box="[808,1148,1232,1256]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="5" pageNumber="535" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov.">
<emphasis id="B965343394435F7F3D10FB533A07ACD2" box="[808,1148,1232,1256]" italics="true" pageId="5" pageNumber="535">Coryphophthalmus troglophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A256894894435F7F3ABBFB523AA3ACD3" box="[1155,1240,1233,1257]" pageId="5" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BAEE82194435F7F3E9EFB7A3AB4AE99" pageId="5" pageNumber="535">
<table id="F9111A819443A0853E9EFB7A3AB4AE99" box="[166,1231,1273,1699]" gridcols="4" gridrows="11" pageId="5" pageNumber="535">
<tr id="3521EA639443A0853E9EFB7A3AB4AD2B" box="[166,1231,1273,1297]" gridrow="0" pageId="5" pageNumber="535" rowspan-1="1">
<th id="76F0831F9443A0853E9EFB7A3E84AD2B" box="[166,255,1273,1297]" gridcol="0" gridrow="0" pageId="5" pageNumber="535">Segment</th>
<th id="76F0831F9443A0853CF9FB7A3AB4AD2B" box="[705,1231,1273,1297]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="5" pageNumber="535">Sex</th>
</tr>
<tr id="3521EA639443A0853E9EFAA23AB4AD03" box="[166,1231,1313,1337]" gridrow="1" pageId="5" pageNumber="535" rowspan-0="1" rowspan-1="1">
<td id="76F0831F9443A0853CF9FAA23D5DAD03" box="[705,806,1313,1337]" gridcol="2" gridrow="1" pageId="5" pageNumber="535">Female</td>
<td id="76F0831F9443A0853A52FAA23AB4AD03" box="[1130,1231,1313,1337]" gridcol="3" gridrow="1" pageId="5" pageNumber="535">Male</td>
</tr>
<tr id="3521EA639443A0853E9EFACA3AB4AD5B" box="[166,1231,1353,1377]" gridrow="2" pageId="5" pageNumber="535">
<th id="76F0831F9443A0853E9EFACA3E84AD5B" box="[166,255,1353,1377]" gridcol="0" gridrow="2" pageId="5" pageNumber="535">Tarsus</th>
<td id="76F0831F9443A0853FB0FACA3FB5AD5B" box="[392,462,1353,1377]" gridcol="1" gridrow="2" pageId="5" pageNumber="535">fore</td>
<td id="76F0831F9443A0853CF9FACA3D5DAD5B" box="[705,806,1353,1377]" gridcol="2" gridrow="2" pageId="5" pageNumber="535">4.70</td>
<td id="76F0831F9443A0853A52FACA3AB4AD5B" box="[1130,1231,1353,1377]" gridcol="3" gridrow="2" pageId="5" pageNumber="535">4.82–5.02</td>
</tr>
<tr id="3521EA639443A0853E9EFAF23AB4ADB3" box="[166,1231,1393,1417]" gridrow="3" pageId="5" pageNumber="535" rowspan-0="1">
<td id="76F0831F9443A0853FB0FAF23FB5ADB3" box="[392,462,1393,1417]" gridcol="1" gridrow="3" pageId="5" pageNumber="535">middle</td>
<td id="76F0831F9443A0853CF9FAF23D5DADB3" box="[705,806,1393,1417]" gridcol="2" gridrow="3" pageId="5" pageNumber="535">4.46–4.57</td>
<td id="76F0831F9443A0853A52FAF23AB4ADB3" box="[1130,1231,1393,1417]" gridcol="3" gridrow="3" pageId="5" pageNumber="535">4.84–5.05</td>
</tr>
<tr id="3521EA639443A0853E9EFA1A3AB4AD88" box="[166,1231,1433,1458]" gridrow="4" pageId="5" pageNumber="535" rowspan-0="1">
<td id="76F0831F9443A0853FB0FA1A3FB5AD88" box="[392,462,1433,1458]" gridcol="1" gridrow="4" pageId="5" pageNumber="535">hind</td>
<td id="76F0831F9443A0853CF9FA1A3D5DAD88" box="[705,806,1433,1458]" gridcol="2" gridrow="4" pageId="5" pageNumber="535">5.72–5.75</td>
<td id="76F0831F9443A0853A52FA1A3AB4AD88" box="[1130,1231,1433,1458]" gridcol="3" gridrow="4" pageId="5" pageNumber="535">5.72–6.15</td>
</tr>
<tr id="3521EA639443A0853E9EFA413AB4ADE0" box="[166,1231,1474,1498]" gridrow="5" pageId="5" pageNumber="535">
<th id="76F0831F9443A0853E9EFA413E84ADE0" box="[166,255,1474,1498]" gridcol="0" gridrow="5" pageId="5" pageNumber="535">Tibia</th>
<td id="76F0831F9443A0853FB0FA413FB5ADE0" box="[392,462,1474,1498]" gridcol="1" gridrow="5" pageId="5" pageNumber="535">fore</td>
<td id="76F0831F9443A0853CF9FA413D5DADE0" box="[705,806,1474,1498]" gridcol="2" gridrow="5" pageId="5" pageNumber="535">2.18</td>
<td id="76F0831F9443A0853A52FA413AB4ADE0" box="[1130,1231,1474,1498]" gridcol="3" gridrow="5" pageId="5" pageNumber="535">2.47–2.53</td>
</tr>
<tr id="3521EA639443A0853E9EFA693AB4AE38" box="[166,1231,1514,1538]" gridrow="6" pageId="5" pageNumber="535" rowspan-0="1">
<td id="76F0831F9443A0853FB0FA693FB5AE38" box="[392,462,1514,1538]" gridcol="1" gridrow="6" pageId="5" pageNumber="535">middle</td>
<td id="76F0831F9443A0853CF9FA693D5DAE38" box="[705,806,1514,1538]" gridcol="2" gridrow="6" pageId="5" pageNumber="535">1.89–1.92</td>
<td id="76F0831F9443A0853A52FA693AB4AE38" box="[1130,1231,1514,1538]" gridcol="3" gridrow="6" pageId="5" pageNumber="535">2.41–2.55</td>
</tr>
<tr id="3521EA639443A0853E9EF9913AB4AE10" box="[166,1231,1554,1578]" gridrow="7" pageId="5" pageNumber="535" rowspan-0="1">
<td id="76F0831F9443A0853FB0F9913FB5AE10" box="[392,462,1554,1578]" gridcol="1" gridrow="7" pageId="5" pageNumber="535">hind</td>
<td id="76F0831F9443A0853CF9F9913D5DAE10" box="[705,806,1554,1578]" gridcol="2" gridrow="7" pageId="5" pageNumber="535">3.34–3.36</td>
<td id="76F0831F9443A0853A52F9913AB4AE10" box="[1130,1231,1554,1578]" gridcol="3" gridrow="7" pageId="5" pageNumber="535">3.19–3.30</td>
</tr>
<tr id="3521EA639443A0853E9EF9B83AB4AE69" box="[166,1231,1595,1619]" gridrow="8" pageId="5" pageNumber="535">
<th id="76F0831F9443A0853E9EF9B83E84AE69" box="[166,255,1595,1619]" gridcol="0" gridrow="8" pageId="5" pageNumber="535">Femur</th>
<td id="76F0831F9443A0853FB0F9B83FB5AE69" box="[392,462,1595,1619]" gridcol="1" gridrow="8" pageId="5" pageNumber="535">fore</td>
<td id="76F0831F9443A0853CF9F9B83D5DAE69" box="[705,806,1595,1619]" gridcol="2" gridrow="8" pageId="5" pageNumber="535">2.10</td>
<td id="76F0831F9443A0853A52F9B83AB4AE69" box="[1130,1231,1595,1619]" gridcol="3" gridrow="8" pageId="5" pageNumber="535">2.23–2.24</td>
</tr>
<tr id="3521EA639443A0853E9EF9E03AB4AE41" box="[166,1231,1635,1659]" gridrow="9" pageId="5" pageNumber="535" rowspan-0="1">
<td id="76F0831F9443A0853FB0F9E03FB5AE41" box="[392,462,1635,1659]" gridcol="1" gridrow="9" pageId="5" pageNumber="535">middle</td>
<td id="76F0831F9443A0853CF9F9E03D5DAE41" box="[705,806,1635,1659]" gridcol="2" gridrow="9" pageId="5" pageNumber="535">2.18</td>
<td id="76F0831F9443A0853A52F9E03AB4AE41" box="[1130,1231,1635,1659]" gridcol="3" gridrow="9" pageId="5" pageNumber="535">2.46–2.54</td>
</tr>
<tr id="3521EA639443A0853E9EF9083AB4AE99" box="[166,1231,1675,1699]" gridrow="10" pageId="5" pageNumber="535" rowspan-0="1">
<td id="76F0831F9443A0853FB0F9083FB5AE99" box="[392,462,1675,1699]" gridcol="1" gridrow="10" pageId="5" pageNumber="535">hind</td>
<td id="76F0831F9443A0853CF9F9083D5DAE99" box="[705,806,1675,1699]" gridcol="2" gridrow="10" pageId="5" pageNumber="535">2.59–2.62</td>
<td id="76F0831F9443A0853A52F9083AB4AE99" box="[1130,1231,1675,1699]" gridcol="3" gridrow="10" pageId="5" pageNumber="535">2.63–2.65</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAEE82194435F7F3EFFF9503BF2AF0F" blockId="5.[151,1436,1747,1953]" pageId="5" pageNumber="535">
Urocoxites I and V–VII with 1 + 1, II–IV with 2 + 2 eversible vesicles. Posterior angle of urosternites II–VI 74–80°, VII about 85° in both sexes. Length ratios of urosternites, urocoxites and urostyli given in 
<tableCitation id="C693DD9A94435F7F3AC6F9743B2EAF2B" box="[1278,1365,1783,1809]" captionStart="TABLE 2" captionStartId="6.[151,239,151,175]" captionTargetPageId="6" captionText="TABLE 2. Length ratios of urosternites and urocoxites of Coryphophthalmus troglophilus sp. nov." httpUri="http://table.plazi.org/id/DF6EB8A994405F7C3EAFFF143ACDA895" pageId="5" pageNumber="535" tableUuid="DF6EB8A994405F7C3EAFFF143ACDA895">Table 2</tableCitation>
. Inner posterior lobes of female urocoxites VII protruding. Ratio of length to total width of protruding lobes about 0.34.
</paragraph>
<paragraph id="8BAEE82194435F7F3EFFF8BC3B0BAF9B" blockId="5.[151,1436,1747,1953]" pageId="5" pageNumber="535">
Thoracic tergites, urotergites I–IV, all urosternites, urocoxites I–V without macrochaetae in both sexes. Distribution of sublateral spines on urocoxites and urotergites given in 
<tableCitation id="C693DD9A94435F7F3D5FF8E03DC0AF47" box="[871,955,1891,1917]" captionStart="TABLE 3" captionStartId="6.[151,239,663,687]" captionTargetPageId="6" captionText="TABLE 3. Distribution of sublateral spines on urotergites and urocoxites of Coryphophthalmus troglophilus sp. nov." httpUri="http://table.plazi.org/id/DF6EB8A994405F7C3EAFFD143B0CAA95" pageId="5" pageNumber="535" tableUuid="DF6EB8A994405F7C3EAFFD143B0CAA95">Table 3</tableCitation>
. Urocoxites IX with 2–3 + 2–3 outer spines in both sexes and with 6–8 + 6–8 inner sublateral spines in male and 7–11 + 7–11 ones in female (
<figureCitation id="132AF4A494435F7F3AD9F8043B18AF9B" box="[1249,1379,1927,1953]" captionStart="FIGURES 10–14" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[254,1334,192,1907]" captionTargetId="figure-17@4.[243,1343,181,1919]" captionTargetPageId="4" captionText="FIGURES 10–14. Coryphophthalmus troglophilus sp. nov. 10–12, paratype female; 13, 14, holotype, male. 10) Urocoxite IX and anterior gonapophyses. 11) Distal divisions of anterior gonapophyse. 12) Distal divisions of posterior gonapophyse. 13) Urocoxite IX and paramere. 14) Penis. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4296936" httpUri="https://zenodo.org/record/4296936/files/figure.png" pageId="5" pageNumber="535">Figs 10, 13</figureCitation>
).
</paragraph>
<caption id="DF6EB8A994405F7C3EAFFF143ACDA895" ID-Table-UUID="DF6EB8A994405F7C3EAFFF143ACDA895" box="[151,1206,151,175]" httpUri="http://table.plazi.org/id/DF6EB8A994405F7C3EAFFF143ACDA895" pageId="6" pageNumber="536" targetBox="[166,1369,191,614]" targetIsTable="true" targetPageId="6">
<paragraph id="8BAEE82194405F7C3EAFFF143ACDA895" blockId="6.[151,1206,151,175]" box="[151,1206,151,175]" pageId="6" pageNumber="536">
<emphasis id="B965343394405F7C3EAFFF143F71A895" bold="true" box="[151,266,151,175]" pageId="6" pageNumber="536">TABLE 2.</emphasis>
Length ratios of urosternites and urocoxites of 
<taxonomicName id="4C1193A294405F7C3D3DFF143A22A895" authority="Kaplin &amp; Vargovitsh, 2020" authorityName="Kaplin &amp; Vargovitsh" authorityYear="2020" box="[773,1113,151,175]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov.">
<emphasis id="B965343394405F7C3D3DFF143A22A895" box="[773,1113,151,175]" italics="true" pageId="6" pageNumber="536">Coryphophthalmus troglophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A256894894405F7C3A58FF143ACDA895" box="[1120,1206,151,175]" pageId="6" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BAEE82194405F7C3E9EFF3C3B2CAA5F" pageId="6" pageNumber="536">
<table id="F9111A819440A0853E9EFF3C3B22AA5C" box="[166,1369,191,614]" gridcols="7" gridrows="10" pageId="6" pageNumber="536">
<tr id="3521EA639440A0853E9EFF3C3B22A8C6" box="[166,1369,191,252]" gridrow="0" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFF3C3F63A8C6" box="[166,280,191,252]" gridcol="0" gridrow="0" pageId="6" pageNumber="536">Abdominal segment</th>
<th id="76F0831F9440A0853FB0FF3C3C00A8C6" box="[392,635,191,252]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="6" pageNumber="536">Urosternite:urocoxite</th>
<th id="76F0831F9440A0853CD8FF3C3A70A8C6" box="[736,1035,191,252]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="6" pageNumber="536">Urostyli (not including apical spines):urocoxites</th>
<th id="76F0831F9440A0853A5FFF3C3B22A8C6" box="[1127,1369,191,252]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="6" pageNumber="536">Apical spines:styli</th>
</tr>
<tr id="3521EA639440A0853E9EFE8F3B22A91E" box="[166,1369,268,292]" gridrow="1" pageId="6" pageNumber="536" rowspan-0="1">
<td id="76F0831F9440A0853FB0FE8F3FB6A91E" box="[392,461,268,292]" gridcol="1" gridrow="1" pageId="6" pageNumber="536">female</td>
<td id="76F0831F9440A0853C72FE8F3C00A91E" box="[586,635,268,292]" gridcol="2" gridrow="1" pageId="6" pageNumber="536">male</td>
<td id="76F0831F9440A0853CD8FE8F3D46A91E" box="[736,829,268,292]" gridcol="3" gridrow="1" pageId="6" pageNumber="536">female</td>
<td id="76F0831F9440A0853D81FE8F3A70A91E" box="[953,1035,268,292]" gridcol="4" gridrow="1" pageId="6" pageNumber="536">male</td>
<td id="76F0831F9440A0853A5FFE8F3AD7A91E" box="[1127,1196,268,292]" gridcol="5" gridrow="1" pageId="6" pageNumber="536">female</td>
<td id="76F0831F9440A0853B10FE8F3B22A91E" box="[1320,1369,268,292]" gridcol="6" gridrow="1" pageId="6" pageNumber="536">male</td>
</tr>
<tr id="3521EA639440A0853E9EFEB73B22A976" box="[166,1369,308,332]" gridrow="2" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFEB73F63A976" box="[166,280,308,332]" gridcol="0" gridrow="2" pageId="6" pageNumber="536">II</th>
<td id="76F0831F9440A0853FB0FEB73FB6A976" box="[392,461,308,332]" gridcol="1" gridrow="2" pageId="6" pageNumber="536">0.62</td>
<td id="76F0831F9440A0853C72FEB73C00A976" box="[586,635,308,332]" gridcol="2" gridrow="2" pageId="6" pageNumber="536">0.64</td>
<td id="76F0831F9440A0853CD8FEB73D46A976" box="[736,829,308,332]" gridcol="3" gridrow="2" pageId="6" pageNumber="536">0.56</td>
<td id="76F0831F9440A0853D81FEB73A70A976" box="[953,1035,308,332]" gridcol="4" gridrow="2" pageId="6" pageNumber="536">0.57</td>
<td id="76F0831F9440A0853A5FFEB73AD7A976" box="[1127,1196,308,332]" gridcol="5" gridrow="2" pageId="6" pageNumber="536">0.58</td>
<td id="76F0831F9440A0853B10FEB73B22A976" box="[1320,1369,308,332]" gridcol="6" gridrow="2" pageId="6" pageNumber="536">0.63</td>
</tr>
<tr id="3521EA639440A0853E9EFEDF3B22A94E" box="[166,1369,348,372]" gridrow="3" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFEDF3F63A94E" box="[166,280,348,372]" gridcol="0" gridrow="3" pageId="6" pageNumber="536">III</th>
<td id="76F0831F9440A0853FB0FEDF3FB6A94E" box="[392,461,348,372]" gridcol="1" gridrow="3" pageId="6" pageNumber="536">0.66</td>
<td id="76F0831F9440A0853C72FEDF3C00A94E" box="[586,635,348,372]" gridcol="2" gridrow="3" pageId="6" pageNumber="536">0.68</td>
<td id="76F0831F9440A0853CD8FEDF3D46A94E" box="[736,829,348,372]" gridcol="3" gridrow="3" pageId="6" pageNumber="536">0.58</td>
<td id="76F0831F9440A0853D81FEDF3A70A94E" box="[953,1035,348,372]" gridcol="4" gridrow="3" pageId="6" pageNumber="536">0.61</td>
<td id="76F0831F9440A0853A5FFEDF3AD7A94E" box="[1127,1196,348,372]" gridcol="5" gridrow="3" pageId="6" pageNumber="536">0.56</td>
<td id="76F0831F9440A0853B10FEDF3B22A94E" box="[1320,1369,348,372]" gridcol="6" gridrow="3" pageId="6" pageNumber="536">0.58</td>
</tr>
<tr id="3521EA639440A0853E9EFE073B22A9A6" box="[166,1369,388,412]" gridrow="4" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFE073F63A9A6" box="[166,280,388,412]" gridcol="0" gridrow="4" pageId="6" pageNumber="536">IV</th>
<td id="76F0831F9440A0853FB0FE073FB6A9A6" box="[392,461,388,412]" gridcol="1" gridrow="4" pageId="6" pageNumber="536">0.65</td>
<td id="76F0831F9440A0853C72FE073C00A9A6" box="[586,635,388,412]" gridcol="2" gridrow="4" pageId="6" pageNumber="536">0.66</td>
<td id="76F0831F9440A0853CD8FE073D46A9A6" box="[736,829,388,412]" gridcol="3" gridrow="4" pageId="6" pageNumber="536">0.53</td>
<td id="76F0831F9440A0853D81FE073A70A9A6" box="[953,1035,388,412]" gridcol="4" gridrow="4" pageId="6" pageNumber="536">0.58</td>
<td id="76F0831F9440A0853A5FFE073AD7A9A6" box="[1127,1196,388,412]" gridcol="5" gridrow="4" pageId="6" pageNumber="536">0.60</td>
<td id="76F0831F9440A0853B10FE073B22A9A6" box="[1320,1369,388,412]" gridcol="6" gridrow="4" pageId="6" pageNumber="536">0.64</td>
</tr>
<tr id="3521EA639440A0853E9EFE2F3B22A9FE" box="[166,1369,428,452]" gridrow="5" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFE2F3F63A9FE" box="[166,280,428,452]" gridcol="0" gridrow="5" pageId="6" pageNumber="536">V</th>
<td id="76F0831F9440A0853FB0FE2F3FB6A9FE" box="[392,461,428,452]" gridcol="1" gridrow="5" pageId="6" pageNumber="536">0.64</td>
<td id="76F0831F9440A0853C72FE2F3C00A9FE" box="[586,635,428,452]" gridcol="2" gridrow="5" pageId="6" pageNumber="536">0.65</td>
<td id="76F0831F9440A0853CD8FE2F3D46A9FE" box="[736,829,428,452]" gridcol="3" gridrow="5" pageId="6" pageNumber="536">0.53</td>
<td id="76F0831F9440A0853D81FE2F3A70A9FE" box="[953,1035,428,452]" gridcol="4" gridrow="5" pageId="6" pageNumber="536">0.57</td>
<td id="76F0831F9440A0853A5FFE2F3AD7A9FE" box="[1127,1196,428,452]" gridcol="5" gridrow="5" pageId="6" pageNumber="536">0.60</td>
<td id="76F0831F9440A0853B10FE2F3B22A9FE" box="[1320,1369,428,452]" gridcol="6" gridrow="5" pageId="6" pageNumber="536">0.65</td>
</tr>
<tr id="3521EA639440A0853E9EFE563B22A9D7" box="[166,1369,469,493]" gridrow="6" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFE563F63A9D7" box="[166,280,469,493]" gridcol="0" gridrow="6" pageId="6" pageNumber="536">VI</th>
<td id="76F0831F9440A0853FB0FE563FB6A9D7" box="[392,461,469,493]" gridcol="1" gridrow="6" pageId="6" pageNumber="536">0.60</td>
<td id="76F0831F9440A0853C72FE563C00A9D7" box="[586,635,469,493]" gridcol="2" gridrow="6" pageId="6" pageNumber="536">0.62</td>
<td id="76F0831F9440A0853CD8FE563D46A9D7" box="[736,829,469,493]" gridcol="3" gridrow="6" pageId="6" pageNumber="536">0.54</td>
<td id="76F0831F9440A0853D81FE563A70A9D7" box="[953,1035,469,493]" gridcol="4" gridrow="6" pageId="6" pageNumber="536">0.58</td>
<td id="76F0831F9440A0853A5FFE563AD7A9D7" box="[1127,1196,469,493]" gridcol="5" gridrow="6" pageId="6" pageNumber="536">0.63</td>
<td id="76F0831F9440A0853B10FE563B22A9D7" box="[1320,1369,469,493]" gridcol="6" gridrow="6" pageId="6" pageNumber="536">0.63</td>
</tr>
<tr id="3521EA639440A0853E9EFE7E3B22AA2F" box="[166,1369,509,533]" gridrow="7" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFE7E3F63AA2F" box="[166,280,509,533]" gridcol="0" gridrow="7" pageId="6" pageNumber="536">VII</th>
<td id="76F0831F9440A0853FB0FE7E3FB6AA2F" box="[392,461,509,533]" gridcol="1" gridrow="7" pageId="6" pageNumber="536">0.49</td>
<td id="76F0831F9440A0853C72FE7E3C00AA2F" box="[586,635,509,533]" gridcol="2" gridrow="7" pageId="6" pageNumber="536">0.61</td>
<td id="76F0831F9440A0853CD8FE7E3D46AA2F" box="[736,829,509,533]" gridcol="3" gridrow="7" pageId="6" pageNumber="536">0.52</td>
<td id="76F0831F9440A0853D81FE7E3A70AA2F" box="[953,1035,509,533]" gridcol="4" gridrow="7" pageId="6" pageNumber="536">0.50</td>
<td id="76F0831F9440A0853A5FFE7E3AD7AA2F" box="[1127,1196,509,533]" gridcol="5" gridrow="7" pageId="6" pageNumber="536">0.65</td>
<td id="76F0831F9440A0853B10FE7E3B22AA2F" box="[1320,1369,509,533]" gridcol="6" gridrow="7" pageId="6" pageNumber="536">0.59</td>
</tr>
<tr id="3521EA639440A0853E9EFDA63B22AA07" box="[166,1369,549,573]" gridrow="8" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFDA63F63AA07" box="[166,280,549,573]" gridcol="0" gridrow="8" pageId="6" pageNumber="536">VIII</th>
<td id="76F0831F9440A0853FB0FDA63FB6AA07" box="[392,461,549,573]" gridcol="1" gridrow="8" pageId="6" pageNumber="536">–</td>
<td id="76F0831F9440A0853C72FDA63C00AA07" box="[586,635,549,573]" gridcol="2" gridrow="8" pageId="6" pageNumber="536">0.39</td>
<td id="76F0831F9440A0853CD8FDA63D46AA07" box="[736,829,549,573]" gridcol="3" gridrow="8" pageId="6" pageNumber="536">0.77</td>
<td id="76F0831F9440A0853D81FDA63A70AA07" box="[953,1035,549,573]" gridcol="4" gridrow="8" pageId="6" pageNumber="536">0.64</td>
<td id="76F0831F9440A0853A5FFDA63AD7AA07" box="[1127,1196,549,573]" gridcol="5" gridrow="8" pageId="6" pageNumber="536">0.52</td>
<td id="76F0831F9440A0853B10FDA63B22AA07" box="[1320,1369,549,573]" gridcol="6" gridrow="8" pageId="6" pageNumber="536">0.57</td>
</tr>
<tr id="3521EA639440A0853E9EFDCE3B22AA5C" box="[166,1369,589,614]" gridrow="9" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFDCE3F63AA5C" box="[166,280,589,614]" gridcol="0" gridrow="9" pageId="6" pageNumber="536">IX</th>
<td id="76F0831F9440A0853FB0FDCE3FB6AA5C" box="[392,461,589,614]" gridcol="1" gridrow="9" pageId="6" pageNumber="536">–</td>
<td id="76F0831F9440A0853C72FDCE3C00AA5C" box="[586,635,589,614]" gridcol="2" gridrow="9" pageId="6" pageNumber="536">–</td>
<td id="76F0831F9440A0853CD8FDCE3D46AA5C" box="[736,829,589,614]" gridcol="3" gridrow="9" pageId="6" pageNumber="536">0.66</td>
<td id="76F0831F9440A0853D81FDCE3A70AA5C" box="[953,1035,589,614]" gridcol="4" gridrow="9" pageId="6" pageNumber="536">0.82</td>
<td id="76F0831F9440A0853A5FFDCE3AD7AA5C" box="[1127,1196,589,614]" gridcol="5" gridrow="9" pageId="6" pageNumber="536">0.35</td>
<td id="76F0831F9440A0853B10FDCE3B22AA5C" box="[1320,1369,589,614]" gridcol="6" gridrow="9" pageId="6" pageNumber="536">0.51</td>
</tr>
</table>
</paragraph>
<caption id="DF6EB8A994405F7C3EAFFD143B0CAA95" ID-Table-UUID="DF6EB8A994405F7C3EAFFD143B0CAA95" box="[151,1399,663,687]" httpUri="http://table.plazi.org/id/DF6EB8A994405F7C3EAFFD143B0CAA95" pageId="6" pageNumber="536" targetBox="[166,1384,703,1049]" targetIsTable="true" targetPageId="6">
<paragraph id="8BAEE82194405F7C3EAFFD143B0CAA95" blockId="6.[151,1399,663,687]" box="[151,1399,663,687]" pageId="6" pageNumber="536">
<emphasis id="B965343394405F7C3EAFFD143F71AA95" bold="true" box="[151,266,663,687]" pageId="6" pageNumber="536">TABLE 3.</emphasis>
Distribution of sublateral spines on urotergites and urocoxites of 
<taxonomicName id="4C1193A294405F7C3DFEFD143B60AA95" authority="Kaplin &amp; Vargovitsh, 2020" authorityName="Kaplin &amp; Vargovitsh" authorityYear="2020" box="[966,1307,663,687]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov.">
<emphasis id="B965343394405F7C3DFEFD143B60AA95" box="[966,1307,663,687]" italics="true" pageId="6" pageNumber="536">Coryphophthalmus troglophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A256894894405F7C3B1AFD143B0CAA95" box="[1314,1399,663,687]" pageId="6" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BAEE82194405F7C3E9EFD3C3B79AC23" pageId="6" pageNumber="536">
<table id="F9111A819440A0853E9EFD3C3B13AC23" box="[166,1384,703,1049]" gridcols="5" gridrows="9" pageId="6" pageNumber="536">
<tr id="3521EA639440A0853E9EFD3C3B13AAE2" box="[166,1384,703,728]" gridrow="0" pageId="6" pageNumber="536" rowspan-2="1">
<th id="76F0831F9440A0853E9EFD3C3F09AAE2" box="[166,370,703,728]" gridcol="0" gridrow="0" pageId="6" pageNumber="536">Abdominal segment</th>
<th id="76F0831F9440A0853FE2FD3C3CDAAAE2" box="[474,673,703,728]" gridcol="1" gridrow="0" pageId="6" pageNumber="536">Urotergites</th>
<th id="76F0831F9440A0853D55FD3C3B13AAE2" box="[877,1384,703,728]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="6" pageNumber="536">Urocoxites</th>
</tr>
<tr id="3521EA639440A0853E9EFD643B13AAC5" box="[166,1384,743,767]" gridrow="1" pageId="6" pageNumber="536" rowspan-0="1">
<td id="76F0831F9440A0853FE2FD643CDAAAC5" box="[474,673,743,767]" gridcol="1" gridrow="1" pageId="6" pageNumber="536">male</td>
<td id="76F0831F9440A0853CFAFD643D7CAAC5" box="[706,775,743,767]" gridcol="2" gridrow="1" pageId="6" pageNumber="536">female</td>
<td id="76F0831F9440A0853D55FD643A55AAC5" box="[877,1070,743,767]" gridcol="3" gridrow="1" pageId="6" pageNumber="536">male</td>
<td id="76F0831F9440A0853AA8FD643B13AAC5" box="[1168,1384,743,767]" gridcol="4" gridrow="1" pageId="6" pageNumber="536">female</td>
</tr>
<tr id="3521EA639440A0853E9EFC933B13AB12" box="[166,1384,784,808]" gridrow="2" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFC933F09AB12" box="[166,370,784,808]" gridcol="0" gridrow="2" pageId="6" pageNumber="536">I–IV</th>
<td id="76F0831F9440A0853FE2FC933CDAAB12" box="[474,673,784,808]" gridcol="1" gridrow="2" pageId="6" pageNumber="536">0</td>
<td id="76F0831F9440A0853CFAFC933D7CAB12" box="[706,775,784,808]" gridcol="2" gridrow="2" pageId="6" pageNumber="536">0</td>
<td id="76F0831F9440A0853D55FC933A55AB12" box="[877,1070,784,808]" gridcol="3" gridrow="2" pageId="6" pageNumber="536">0</td>
<td id="76F0831F9440A0853AA8FC933B13AB12" box="[1168,1384,784,808]" gridcol="4" gridrow="2" pageId="6" pageNumber="536">0</td>
</tr>
<tr id="3521EA639440A0853E9EFCBB3B13AB6A" box="[166,1384,824,848]" gridrow="3" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFCBB3F09AB6A" box="[166,370,824,848]" gridcol="0" gridrow="3" pageId="6" pageNumber="536">V</th>
<td id="76F0831F9440A0853FE2FCBB3CDAAB6A" box="[474,673,824,848]" gridcol="1" gridrow="3" pageId="6" pageNumber="536">1–2 + 1–2</td>
<td id="76F0831F9440A0853CFAFCBB3D7CAB6A" box="[706,775,824,848]" gridcol="2" gridrow="3" pageId="6" pageNumber="536">0</td>
<td id="76F0831F9440A0853D55FCBB3A55AB6A" box="[877,1070,824,848]" gridcol="3" gridrow="3" pageId="6" pageNumber="536">0</td>
<td id="76F0831F9440A0853AA8FCBB3B13AB6A" box="[1168,1384,824,848]" gridcol="4" gridrow="3" pageId="6" pageNumber="536">0</td>
</tr>
<tr id="3521EA639440A0853E9EFCE33B13AB42" box="[166,1384,864,888]" gridrow="4" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFCE33F09AB42" box="[166,370,864,888]" gridcol="0" gridrow="4" pageId="6" pageNumber="536">VI</th>
<td id="76F0831F9440A0853FE2FCE33CDAAB42" box="[474,673,864,888]" gridcol="1" gridrow="4" pageId="6" pageNumber="536">3 + 3</td>
<td id="76F0831F9440A0853CFAFCE33D7CAB42" box="[706,775,864,888]" gridcol="2" gridrow="4" pageId="6" pageNumber="536">2 + 2</td>
<td id="76F0831F9440A0853D55FCE33A55AB42" box="[877,1070,864,888]" gridcol="3" gridrow="4" pageId="6" pageNumber="536">0</td>
<td id="76F0831F9440A0853AA8FCE33B13AB42" box="[1168,1384,864,888]" gridcol="4" gridrow="4" pageId="6" pageNumber="536">0</td>
</tr>
<tr id="3521EA639440A0853E9EFC0B3B13AB9A" box="[166,1384,904,928]" gridrow="5" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFC0B3F09AB9A" box="[166,370,904,928]" gridcol="0" gridrow="5" pageId="6" pageNumber="536">VII</th>
<td id="76F0831F9440A0853FE2FC0B3CDAAB9A" box="[474,673,904,928]" gridcol="1" gridrow="5" pageId="6" pageNumber="536">2 + 2</td>
<td id="76F0831F9440A0853CFAFC0B3D7CAB9A" box="[706,775,904,928]" gridcol="2" gridrow="5" pageId="6" pageNumber="536">2 + 2</td>
<td id="76F0831F9440A0853D55FC0B3A55AB9A" box="[877,1070,904,928]" gridcol="3" gridrow="5" pageId="6" pageNumber="536">2–3 + 2–3</td>
<td id="76F0831F9440A0853AA8FC0B3B13AB9A" box="[1168,1384,904,928]" gridcol="4" gridrow="5" pageId="6" pageNumber="536">3–4 + 3–4</td>
</tr>
<tr id="3521EA639440A0853E9EFC323B13ABF3" box="[166,1384,945,969]" gridrow="6" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFC323F09ABF3" box="[166,370,945,969]" gridcol="0" gridrow="6" pageId="6" pageNumber="536">VIII</th>
<td id="76F0831F9440A0853FE2FC323CDAABF3" box="[474,673,945,969]" gridcol="1" gridrow="6" pageId="6" pageNumber="536">2 + 2</td>
<td id="76F0831F9440A0853CFAFC323D7CABF3" box="[706,775,945,969]" gridcol="2" gridrow="6" pageId="6" pageNumber="536">3 + 3</td>
<td id="76F0831F9440A0853D55FC323A55ABF3" box="[877,1070,945,969]" gridcol="3" gridrow="6" pageId="6" pageNumber="536">3 + 3</td>
<td id="76F0831F9440A0853AA8FC323B13ABF3" box="[1168,1384,945,969]" gridcol="4" gridrow="6" pageId="6" pageNumber="536">4 + 4</td>
</tr>
<tr id="3521EA639440A0853E9EFC5A3B13ABCB" box="[166,1384,985,1009]" gridrow="7" pageId="6" pageNumber="536">
<th id="76F0831F9440A0853E9EFC5A3F09ABCB" box="[166,370,985,1009]" gridcol="0" gridrow="7" pageId="6" pageNumber="536">IX</th>
<td id="76F0831F9440A0853FE2FC5A3CDAABCB" box="[474,673,985,1009]" gridcol="1" gridrow="7" pageId="6" pageNumber="536">4 + 4</td>
<td id="76F0831F9440A0853CFAFC5A3D7CABCB" box="[706,775,985,1009]" gridcol="2" gridrow="7" pageId="6" pageNumber="536">2 + 2</td>
<td id="76F0831F9440A0853D55FC5A3A55ABCB" box="[877,1070,985,1009]" gridcol="3" gridrow="7" pageId="6" pageNumber="536">2–3/6–8 + 6–8/2–3</td>
<td id="76F0831F9440A0853AA8FC5A3B13ABCB" box="[1168,1384,985,1009]" gridcol="4" gridrow="7" pageId="6" pageNumber="536">2–3/7–11 + 7–11/2–3</td>
</tr>
<tr id="3521EA639440A0853E9EFB823B13AC23" box="[166,1384,1025,1049]" gridrow="8" pageId="6" pageNumber="536" rowspan-3="1">
<th id="76F0831F9440A0853E9EFB823F09AC23" box="[166,370,1025,1049]" gridcol="0" gridrow="8" pageId="6" pageNumber="536">X</th>
<td id="76F0831F9440A0853FE2FB823CDAAC23" box="[474,673,1025,1049]" gridcol="1" gridrow="8" pageId="6" pageNumber="536">4 + 4</td>
<td id="76F0831F9440A0853CFAFB823D7CAC23" box="[706,775,1025,1049]" gridcol="2" gridrow="8" pageId="6" pageNumber="536">5 + 5</td>
<td id="76F0831F9440A0853AA8FB823B13AC23" box="[1168,1384,1025,1049]" gridcol="4" gridrow="8" pageId="6" pageNumber="536">–</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAEE82194405F7C3EFFFBCB3C09ACC9" blockId="6.[151,1437,1096,2023]" pageId="6" pageNumber="536">
Ovipositor slender, elongate, visibly surpassing apex of styli IX (
<figureCitation id="132AF4A494405F7C3DA2FBCB3D96AC59" box="[922,1005,1096,1123]" captionStart="FIGURES 10–14" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[254,1334,192,1907]" captionTargetId="figure-17@4.[243,1343,181,1919]" captionTargetPageId="4" captionText="FIGURES 10–14. Coryphophthalmus troglophilus sp. nov. 10–12, paratype female; 13, 14, holotype, male. 10) Urocoxite IX and anterior gonapophyses. 11) Distal divisions of anterior gonapophyse. 12) Distal divisions of posterior gonapophyse. 13) Urocoxite IX and paramere. 14) Penis. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4296936" httpUri="https://zenodo.org/record/4296936/files/figure.png" pageId="6" pageNumber="536">Fig. 10</figureCitation>
). Anterior and posterior gonapophyses with approximately 38–40 or 41–43 divisions, respectively. One or two basal divisions of anterior gonapophyses and 20–21 basal divisions of posterior gonapophyses glabrous. Distal spines of gonapophyses as long as 3–4 apical divisions combined. Apical divisions of anterior and posterior gonapophyses with 8–9 and 5–8 chaetae, respectively (not including apical spines) (
<figureCitation id="132AF4A494405F7C3FDCFB5B3C1EACC9" box="[484,613,1240,1267]" captionStart="FIGURES 10–14" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[254,1334,192,1907]" captionTargetId="figure-17@4.[243,1343,181,1919]" captionTargetPageId="4" captionText="FIGURES 10–14. Coryphophthalmus troglophilus sp. nov. 10–12, paratype female; 13, 14, holotype, male. 10) Urocoxite IX and anterior gonapophyses. 11) Distal divisions of anterior gonapophyse. 12) Distal divisions of posterior gonapophyse. 13) Urocoxite IX and paramere. 14) Penis. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4296936" httpUri="https://zenodo.org/record/4296936/files/figure.png" pageId="6" pageNumber="536">Figs 11, 12</figureCitation>
).
</paragraph>
<paragraph id="8BAEE82194405F7C3EFFFB7F3C14ADB9" blockId="6.[151,1437,1096,2023]" pageId="6" pageNumber="536">
Male genitalia with one pair of parameres on urite IX. Parameres with 1 + 5 divisions (
<figureCitation id="132AF4A494405F7C3A82FB7F3B75AD2D" box="[1210,1294,1276,1303]" captionStart="FIGURES 10–14" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[254,1334,192,1907]" captionTargetId="figure-17@4.[243,1343,181,1919]" captionTargetPageId="4" captionText="FIGURES 10–14. Coryphophthalmus troglophilus sp. nov. 10–12, paratype female; 13, 14, holotype, male. 10) Urocoxite IX and anterior gonapophyses. 11) Distal divisions of anterior gonapophyse. 12) Distal divisions of posterior gonapophyse. 13) Urocoxite IX and paramere. 14) Penis. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4296936" httpUri="https://zenodo.org/record/4296936/files/figure.png" pageId="6" pageNumber="536">Fig. 12</figureCitation>
), surpassing apex of penis. Penis and parameres clearly not attaining level of apex of urocoxites IX, ratio of the distance between apexes of penis and of urocoxites IX to width of distal division of penis 5.0–6.5. Basal division of penis 1.3–1.4 times longer than distal division (
<figureCitation id="132AF4A494405F7C3C36FAEB3C1AADB9" box="[526,609,1384,1411]" captionStart="FIGURES 10–14" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[254,1334,192,1907]" captionTargetId="figure-17@4.[243,1343,181,1919]" captionTargetPageId="4" captionText="FIGURES 10–14. Coryphophthalmus troglophilus sp. nov. 10–12, paratype female; 13, 14, holotype, male. 10) Urocoxite IX and anterior gonapophyses. 11) Distal divisions of anterior gonapophyse. 12) Distal divisions of posterior gonapophyse. 13) Urocoxite IX and paramere. 14) Penis. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4296936" httpUri="https://zenodo.org/record/4296936/files/figure.png" pageId="6" pageNumber="536">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30BBBAA94405F7C3EFFFA0F3D9CAF08" pageId="6" pageNumber="536" type="materials_examined">
<paragraph id="8BAEE82194405F7C3EFFFA0F3F12AE29" blockId="6.[151,1437,1096,2023]" pageId="6" pageNumber="536">
<emphasis id="B965343394405F7C3EFFFA0F3FCAAD9C" bold="true" box="[199,433,1420,1446]" pageId="6" pageNumber="536">Material examined.</emphasis>
<materialsCitation id="3B79E27C94405F7C3F80FA0F3F66ADD5" ID-GBIF-Occurrence="2988979302" collectingDate="2018-09-20" collectionCode="ZIN, R" collectorName="W Caucasus &amp; S. Vargovitsh" county="Kotsha Cave" elevation="440" latitude="42.933334" location="Abkhazia" longLatPrecision="1146" longitude="41.516666" municipality="Aymara" pageId="6" pageNumber="536" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AA568394405F7C3F80FA0F3C5EAD9C" box="[440,549,1420,1446]" pageId="6" pageNumber="536" type="holotype">Holotype</typeStatus>
: male (on slide) (
<collectionCode id="ED0070E494405F7C3CD0FA0E3D67AD9D" box="[744,796,1421,1447]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="536" type="Museum">ZIN</collectionCode>
); 
<location id="8ECEBEFA94405F7C3D17FA0F3DE6AD9C" LSID="urn:lsid:plazi:treatment:03B8593794445F7D3EAFFF143CA2A8EF:8ECEBEFA94405F7C3D17FA0F3DE6AD9C" box="[815,925,1420,1446]" county="Kotsha Cave" latitude="42.933334" longLatPrecision="1146" longitude="41.516666" municipality="Aymara" name="Abkhazia" pageId="6" pageNumber="536">Abkhazia</location>
, 
<location id="8ECEBEFA94405F7C3D91FA0F3AEBAD9C" LSID="urn:lsid:plazi:treatment:03B8593794445F7D3EAFFF143CA2A8EF:8ECEBEFA94405F7C3D91FA0F3AEBAD9C" box="[937,1168,1420,1446]" county="Kotsha Cave" latitude="42.933334" longLatPrecision="1146" longitude="41.516666" municipality="Aymara" name="Ochamchira District" pageId="6" pageNumber="536">Ochamchira District</location>
, 
<location id="8ECEBEFA94405F7C3AA4FA0E3AAEAD9D" LSID="urn:lsid:plazi:treatment:03B8593794445F7D3EAFFF143CA2A8EF:8ECEBEFA94405F7C3AA4FA0E3AAEAD9D" box="[1180,1237,1421,1447]" county="Kotsha Cave" latitude="42.933334" longLatPrecision="1146" longitude="41.516666" municipality="Aymara" name="Otap" pageId="6" pageNumber="536">Otap</location>
, 
<collectingMunicipality id="6BCA725B94405F7C3AE7FA0E3B41AD9D" box="[1247,1338,1421,1447]" pageId="6" pageNumber="536">Aymara</collectingMunicipality>
, 
<collectorName id="26E48DF794405F7C3B7CFA0E3EADADF1" pageId="6" pageNumber="536">W Caucasus</collectorName>
, 
<geoCoordinate id="EE258EE694405F7C3EDDFA313F3EADF7" box="[229,325,1458,1485]" degrees="42" direction="north" minutes="56" orientation="latitude" pageId="6" pageNumber="536" precision="925" value="42.933334">42°56′N</geoCoordinate>
, 
<geoCoordinate id="EE258EE694405F7C3F6BFA313FCBADF7" box="[339,432,1458,1485]" degrees="41" direction="east" minutes="31" orientation="longitude" pageId="6" pageNumber="536" precision="925" value="41.516666">41°31′E</geoCoordinate>
, elev. 
<quantity id="4CE945C494405F7C3FC5FA323C32ADF1" box="[509,585,1457,1483]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="536" unit="m" value="440.0">
<elevation id="003C0F1294405F7C3FC5FA323C32ADF1" box="[509,585,1457,1483]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="536" unit="m" value="440.0">440 m</elevation>
</quantity>
, 
<collectingCounty id="62CF90AD94405F7C3C6FFA333C96ADF1" box="[599,749,1456,1483]" pageId="6" pageNumber="536">Kotsha Cave</collectingCounty>
, ˗20 to ˗ 
<elevation id="003C0F1294405F7C3D60FA333D9BADF0" box="[856,992,1456,1483]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="536" unit="m" value="25.0">
<quantity id="4CE945C494405F7C3D60FA333DEEADF1" box="[856,917,1456,1483]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="536" unit="m" value="25.0">25 m</quantity>
depth
</elevation>
, on wet stoneflows, 
<date id="FFAFCEE194405F7C3AE9FA323B35ADF1" box="[1233,1358,1457,1483]" pageId="6" pageNumber="536" value="2018-09-20">
<collectingDate id="EFEB370994405F7C3AE9FA323B35ADF1" box="[1233,1358,1457,1483]" pageId="6" pageNumber="536" value="2018-09-20">20.09.2018</collectingDate>
</date>
(
<collectionCode id="ED0070E494405F7C3B59FA323B0CADF1" box="[1377,1399,1457,1483]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="536">R</collectionCode>
. 
<collectorName id="26E48DF794405F7C3BBEFA323F6EADD5" pageId="6" pageNumber="536">S. Vargovitsh</collectorName>
)
</materialsCitation>
. 
<materialsCitation id="3B79E27C94405F7C3F12FA563F1EAE29" ID-GBIF-Occurrence="2988979301" collectingDate="2018-09-20" collectionCode="R, ZIN" collectorName="S. Vargovitsh" county="Kotsha Cave" elevation="440" latitude="42.933334" location="Abkhazia" longLatPrecision="1146" longitude="41.516666" municipality="Aymara" pageId="6" pageNumber="536" specimenCount="9" specimenCount-female="7" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="54AA568394405F7C3F12FA563FE0ADD5" box="[298,411,1493,1519]" pageId="6" pageNumber="536" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D1723A894405F7C3F90FA563C7FADD4" box="[424,516,1492,1519]" pageId="6" pageNumber="536" type="male">2 males</specimenCount>
, 
<specimenCount id="9D1723A894405F7C3C2AFA563CFAADD4" box="[530,641,1492,1519]" pageId="6" pageNumber="536" type="female">7 females</specimenCount>
(one male and two females on slides); same locality and data (
<collectionCode id="ED0070E494405F7C3B5AFA563B03ADD5" box="[1378,1400,1493,1519]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="536">R</collectionCode>
. 
<collectorName id="26E48DF794405F7C3BBEFA563F69AE29" pageId="6" pageNumber="536">S. Vargovitsh</collectorName>
) (
<collectionCode id="ED0070E494405F7C3F10FA7A3F27AE29" box="[296,348,1529,1555]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="536" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAEE82194405F7C3EFFF99F3D9CAF08" blockId="6.[151,1437,1096,2023]" pageId="6" pageNumber="536">
<emphasis id="B965343394405F7C3EFFF99F3F4EAE0D" bold="true" box="[199,309,1564,1591]" pageId="6" pageNumber="536">Habitats.</emphasis>
All specimens of 
<taxonomicName id="4C1193A294405F7C3C3BF99F3D11AE0C" authority="Kaplin &amp; Vargovitsh, 2020" authorityName="Kaplin &amp; Vargovitsh" authorityYear="2020" box="[515,874,1564,1590]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov.">
<emphasis id="B965343394405F7C3C3BF99F3D11AE0C" box="[515,874,1564,1590]" italics="true" pageId="6" pageNumber="536">Coryphophthalmus troglophilus</emphasis>
</taxonomicName>
<emphasis id="B965343394405F7C3D4AF99E3DB7AE0D" bold="true" box="[882,972,1565,1591]" pageId="6" pageNumber="536">
<taxonomicNameLabel id="A256894894405F7C3D4AF99E3DB7AE0D" box="[882,972,1565,1591]" pageId="6" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were collected on walls, stalagmites and among stones in a cave at depths of ˗20 to ˗ 
<specimenCount id="9D1723A894405F7C3C43F9C33CC0AE61" box="[635,699,1600,1627]" pageId="6" pageNumber="536" type="generic">
<quantity id="4CE945C494405F7C3C43F9C33CCCAE61" box="[635,695,1600,1627]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="536" unit="m" value="25.0">25 m</quantity>
.
</specimenCount>
The cave is narrow, almost all vertical, with a total depth of ˗ 
<quantity id="4CE945C494405F7C3B5BF9C23BE7AE61" box="[1379,1436,1601,1627]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="536" unit="m" value="33.0">33 m</quantity>
(
<figureCitation id="132AF4A494405F7C3E98F9E73E97AE45" box="[160,236,1636,1663]" captionStart="FIGURES 10–14" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[254,1334,192,1907]" captionTargetId="figure-17@4.[243,1343,181,1919]" captionTargetPageId="4" captionText="FIGURES 10–14. Coryphophthalmus troglophilus sp. nov. 10–12, paratype female; 13, 14, holotype, male. 10) Urocoxite IX and anterior gonapophyses. 11) Distal divisions of anterior gonapophyse. 12) Distal divisions of posterior gonapophyse. 13) Urocoxite IX and paramere. 14) Penis. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4296936" httpUri="https://zenodo.org/record/4296936/files/figure.png" pageId="6" pageNumber="536">Fig.14</figureCitation>
). The arthropod fauna in the cave is quite rich. The main adaptations of this new species to life in conditions of low light on the rocky surface substrate include green or dark green eyes; weak body pigmentation; pretarsa with well-developed arolium and claws; relatively long cerci, tarsi and tibiae of the hind legs; apical needles of the urostyli; large sublateral spines on urocoxites IX, especially in the male; reduced number of apical sensorial cones on the third labial palpomere and sensilla on the apical part of labium; and camouflaged by the mottled color of scales on the upper side of the body. These characteristics place it as a troglophile.
</paragraph>
</subSubSection>
<subSubSection id="C30BBBAA94405F7C3EFFF8BE3AABAF6C" box="[199,1232,1852,1879]" pageId="6" pageNumber="536" type="etymology">
<paragraph id="8BAEE82194405F7C3EFFF8BE3AABAF6C" blockId="6.[151,1437,1096,2023]" box="[199,1232,1852,1879]" pageId="6" pageNumber="536">
<emphasis id="B965343394405F7C3EFFF8BE3F35AF6D" bold="true" box="[199,334,1853,1879]" pageId="6" pageNumber="536">Etymology.</emphasis>
The species name 
<taxonomicName id="4C1193A294405F7C3C1AF8BF3CD1AF6C" authorityName="Kaplin &amp; Vargovitsh" authorityYear="2020" box="[546,682,1852,1878]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="troglophilus">
<emphasis id="B965343394405F7C3C1AF8BF3CD1AF6C" box="[546,682,1852,1878]" italics="true" pageId="6" pageNumber="536">troglophilus</emphasis>
</taxonomicName>
is Latin for “troglo = cave; philus = preference”.
</paragraph>
</subSubSection>
<subSubSection id="C30BBBAA94405F7D3EFFF8E33CA2A8EF" lastPageId="7" lastPageNumber="537" pageId="6" pageNumber="536" type="discussion">
<paragraph id="8BAEE82194405F7D3EFFF8E33CA2A8EF" blockId="6.[151,1437,1096,2023]" lastBlockId="7.[151,1436,151,213]" lastPageId="7" lastPageNumber="537" pageId="6" pageNumber="536">
<emphasis id="B965343394405F7C3EFFF8E33F37AF40" bold="true" box="[199,332,1888,1914]" pageId="6" pageNumber="536">Discussion.</emphasis>
<taxonomicName id="4C1193A294405F7C3F6EF8E33CC4AF40" authority="Kaplin &amp; Vargovitsh, 2020" authorityName="Kaplin &amp; Vargovitsh" authorityYear="2020" box="[342,703,1888,1914]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov.">
<emphasis id="B965343394405F7C3F6EF8E33CC4AF40" box="[342,703,1888,1914]" italics="true" pageId="6" pageNumber="536">Coryphophthalmus troglophilus</emphasis>
</taxonomicName>
<emphasis id="B965343394405F7C3CF1F8E23D5DAF41" bold="true" box="[713,806,1889,1915]" pageId="6" pageNumber="536">
<taxonomicNameLabel id="A256894894405F7C3CF1F8E23D5DAF41" box="[713,806,1889,1915]" pageId="6" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles 
<taxonomicName id="4C1193A294405F7C3D92F8E23A31AF40" baseAuthorityName="Kaplin" baseAuthorityYear="2017" box="[938,1098,1888,1915]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="abchasicus">
<emphasis id="B965343394405F7C3D92F8E23A31AF40" box="[938,1098,1888,1915]" italics="true" pageId="6" pageNumber="536">C. abchasicus</emphasis>
</taxonomicName>
in the presence of numerous long thin chaetae on the undersurface of the male maxillary palp, absence of such chaetae on the upper surface of the male labial palp and absence of spine-like chaetae on the undersurface of the tarsus. The main differences between 
<taxonomicName id="4C1193A294405F7C3EAFF84E3F3BAFDC" authorityName="Kaplin &amp; Vargovitsh" authorityYear="2020" box="[151,320,1996,2023]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="troglophilus">
<emphasis id="B965343394405F7C3EAFF84E3F3BAFDC" box="[151,320,1996,2023]" italics="true" pageId="6" pageNumber="536">C. troglophilus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1193A294405F7C3F4FF84E3C6EAFDC" baseAuthorityName="Kaplin" baseAuthorityYear="2017" box="[375,533,1996,2023]" class="Insecta" family="Machilidae" genus="Coryphophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="abchasicus">
<emphasis id="B965343394405F7C3F4FF84E3C6EAFDC" box="[375,533,1996,2023]" italics="true" pageId="6" pageNumber="536">C. abchasicus</emphasis>
</taxonomicName>
consist in the color of compound eyes, the line of eye contact, lengths of cercus and apical spines of the urostyli, ratio of length to width of the apical palpomere of the labial palp and number of divisions of the ovipositor and parameres (
<tableCitation id="C693DD9A94415F7D3C4CFF383CB7A8EF" box="[628,716,187,213]" captionStart="TABLE 4" captionStartId="7.[151,239,260,284]" captionTargetPageId="7" captionText="TABLE 4. Main morphological differences of Coryphophthalmus troglophilus sp. nov. and C. abchasicus (Kaplin, 2017)." httpUri="http://table.plazi.org/id/DF6EB8A994415F7D3EAFFE873EA6A97B" pageId="7" pageNumber="537" tableUuid="DF6EB8A994415F7D3EAFFE873EA6A97B">Table 4</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>