196 lines
23 KiB
XML
196 lines
23 KiB
XML
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<mods:title id="FFE5B266F1F6114417D02A798024F4B0">Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
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<mods:namePart id="8779257D07C40A8F88433C099B9C490C">Ketwetsuriya, Chatchalerm</mods:namePart>
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<mods:namePart id="6AC9304542F852975D8FAE92940325C8">Karapunar, Baran</mods:namePart>
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<mods:namePart id="ED3E7DC8AAB2DB175334F64CF0C18C4D">Charoentitirat, Thasinee</mods:namePart>
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<mods:namePart id="032C8AA160D6BEDB43FD519F9006B9A5">Nützel, Al- Exander</mods:namePart>
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<mods:title id="E1258A51F1F6845814FF7192CB5AB13D">Zootaxa</mods:title>
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<mods:date id="F1F2D6C8BC5B617563DE8F077499B431">2020</mods:date>
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<mods:number id="40595D8CE23B5956A58C433FA92050AB">2020-04-17</mods:number>
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<mods:number id="3741DC98A50C7B1C955E6955482314CA">4766</mods:number>
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<treatment id="03B587AB4F201569FF5178E7FDEFFBB7" ID-DOI="http://doi.org/10.5281/zenodo.3803851" ID-GBIF-Taxon="163737156" ID-Zenodo-Dep="3803851" LSID="urn:lsid:plazi:treatment:03B587AB4F201569FF5178E7FDEFFBB7" httpUri="http://treatment.plazi.org/id/03B587AB4F201569FF5178E7FDEFFBB7" lastPageNumber="24" pageId="23" pageNumber="24">
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<subSubSection id="C30665364F201569FF5178E7FD7AFF36" box="[151,689,153,180]" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="8BA336BD4F201569FF5178E7FD7AFF36" blockId="23.[151,689,153,215]" box="[151,689,153,180]" pageId="23" pageNumber="24">
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<heading id="D0EB81D14F201569FF5178E7FD7AFF36" bold="true" box="[151,689,153,180]" fontSize="10" level="2" pageId="23" pageNumber="24" reason="2">
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<emphasis id="B968EAAF4F201569FF5178E7FD7AFF36" bold="true" box="[151,689,153,180]" pageId="23" pageNumber="24">
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<taxonomicName id="4C1C4D3E4F201569FF5178E7FEF9FF31" ID-CoL="7NQ4L" authorityName="Meek & Worthen" authorityYear="1866" box="[151,306,153,180]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F201569FF5178E7FEF9FF31" bold="true" box="[151,306,153,180]" italics="true" pageId="23" pageNumber="24">Anomphalus</emphasis>
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</taxonomicName>
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?
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<emphasis id="B968EAAF4F201569FE8E78E7FE6CFF31" bold="true" box="[328,423,153,180]" italics="true" pageId="23" pageNumber="24">blancus</emphasis>
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<bibRefCitation id="EF8D4B4C4F201569FE6878E4FD7AFF36" author="Kues & Batten" box="[430,689,153,180]" firstAuthor="Kues" pageId="23" pageNumber="24" pagination="1 - 95" refId="ref25729" refString="Kues, B. S. & Batten, R. L. (2001) Middle Pennsylvanian gastropods from the Flechado Formation, north-central New Mexico. Journal of Paleontology, 75 (Supplement 1), 1 - 95. https: // doi. org / 10.1666 / 0022 - 3360 (2001) 75 [1: MPGFTF] 2.0. CO; 2" type="journal article" year="2001">Kues & Batten, 2001</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F201569FF5178C3FEE3FF52" box="[151,296,189,215]" pageId="23" pageNumber="24" type="description">
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<paragraph id="8BA336BD4F201569FF5178C3FEE3FF52" blockId="23.[151,689,153,215]" box="[151,296,189,215]" pageId="23" pageNumber="24">
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(
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<figureCitation id="13272A384F201569FF6678C3FF25FF52" box="[160,238,189,215]" captionStart="FIGURE 17" captionStartId="22.[151,250,1941,1966]" captionTargetBox="[309,1278,885,1916]" captionTargetId="figure@22.[309,1278,885,1916]" captionTargetPageId="22" captionText="FIGURE 17. A–E, Anomphalus cf. vanescens Yochelson, 1956, A–C, ESKU-19-LP 30. D–E, ESKU-19-LP 50. F–L, Anomphalus? blancus Kues & Batten, 2001, F–I, ESKU-19-LP 67. J–L, ESKU-19-LP 96. Scale bars represent: 2 mm (A–C); 1 mm (D–L)." figureDoi="http://doi.org/10.5281/zenodo.3763941" httpUri="https://zenodo.org/record/3763941/files/figure.png" pageId="23" pageNumber="24">Fig. 17</figureCitation>
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F–L)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F201569FF51797CFCD4FE9F" box="[151,799,257,282]" pageId="23" pageNumber="24" type="reference_group">
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<paragraph id="8BA336BD4F201569FF51797CFCD4FE9F" blockId="23.[151,799,257,282]" box="[151,799,257,282]" pageId="23" pageNumber="24">
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<treatmentCitationGroup id="AB0C11934F201569FF51797CFCD4FE9F" box="[151,799,257,282]" pageId="23" pageNumber="24">
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<taxonomicName id="4C1C4D3E4F201569FF51797CFEDDFE9F" ID-CoL="7NQ4L" authorityName="Meek & Worthen" authorityYear="1866" box="[151,278,258,282]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F201569FF51797CFEDDFE9F" box="[151,278,258,282]" italics="true" pageId="23" pageNumber="24">Anomphalus</emphasis>
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</taxonomicName>
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?
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<emphasis id="B968EAAF4F201569FEE1797CFEBDFE9F" box="[295,374,258,282]" italics="true" pageId="23" pageNumber="24">blancus</emphasis>
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<treatmentCitation id="0ABD10AC4F201569FEBB797CFDB7FE9F" author="Kues, B. S. & Batten, R. L." box="[381,636,258,282]" page="44" pageId="23" pageNumber="24" year="2001">
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<bibRefCitation id="EF8D4B4C4F201569FEBB797CFDB7FE9F" author="Kues & Batten" box="[381,636,258,282]" firstAuthor="Kues" pageId="23" pageNumber="24" pagination="1 - 95" refId="ref25729" refString="Kues, B. S. & Batten, R. L. (2001) Middle Pennsylvanian gastropods from the Flechado Formation, north-central New Mexico. Journal of Paleontology, 75 (Supplement 1), 1 - 95. https: // doi. org / 10.1666 / 0022 - 3360 (2001) 75 [1: MPGFTF] 2.0. CO; 2" type="journal article" year="2001">Kues & Batten, 2001: 44</bibRefCitation>
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</treatmentCitation>
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, figs 8.23–8.28.
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C30665364F201569FF517939FDFCFE2F" pageId="23" pageNumber="24" type="materials_examined">
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<paragraph id="8BA336BD4F201569FF517939FD7EFEE7" blockId="23.[151,1437,327,1075]" box="[151,693,327,355]" pageId="23" pageNumber="24">
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<emphasis id="B968EAAF4F201569FF517939FECDFEE7" bold="true" box="[151,262,327,354]" pageId="23" pageNumber="24">Material.</emphasis>
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Two specimens: ESKU-19-
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<collectionCode id="ED0DAE784F201569FDF97936FDAAFEE7" box="[575,609,328,354]" country="Russia" name="Laboratory of Palaeontology" pageId="23" pageNumber="24">LP</collectionCode>
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67, 96.
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</paragraph>
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<paragraph id="8BA336BD4F201569FF017912FDFCFE2F" blockId="23.[151,1437,327,1075]" pageId="23" pageNumber="24">
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Dimensions (mm): ESKU-19-
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<collectionCode id="ED0DAE784F201569FDD17912FDF2FE03" box="[535,569,364,390]" country="Russia" name="Laboratory of Palaeontology" pageId="23" pageNumber="24">LP</collectionCode>
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67: height = 2.5; width = 3.0; apical angle = 124º. ESKU-19-
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<collectionCode id="ED0DAE784F201569FB2D7912FAC6FE03" box="[1259,1293,364,390]" country="Russia" name="Laboratory of Palaeontology" pageId="23" pageNumber="24">LP</collectionCode>
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96: height = 2.3; width = 2.6; apical angle = 125º.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F201569FF0179CDFCB6FDBF" pageId="23" pageNumber="24" type="description">
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<paragraph id="8BA336BD4F201569FF0179CDFCB6FDBF" blockId="23.[151,1437,327,1075]" pageId="23" pageNumber="24">
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<emphasis id="B968EAAF4F201569FF0179CDFE92FE4B" bold="true" box="[199,345,435,462]" pageId="23" pageNumber="24">Description.</emphasis>
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Shell small, low-spired but spire clearly elevated, last whorl much higher than spire, turbiniform to naticiform, consisting of two to four whorls; whorls smooth, rounded, convex; whorls embrace at or slightly below periphery at mid-whorl; suture shallow but distinct; aperture somewhat circular, columellar lip straight to gently arched joining outer lip; base convex, minutely phaneromphalous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F201569FF017A3AFDEFFBB7" pageId="23" pageNumber="24" type="discussion">
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<paragraph id="8BA336BD4F201569FF017A3AFDEFFBB7" blockId="23.[151,1437,327,1075]" pageId="23" pageNumber="24">
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<emphasis id="B968EAAF4F201569FF017A3AFEF0FDDB" bold="true" box="[199,315,580,606]" pageId="23" pageNumber="24">Remarks.</emphasis>
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These specimens are very similar to the illustrations of
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<taxonomicName id="4C1C4D3E4F201569FC777A3AFBF6FDDB" authorityName="Meek & Worthen" authorityYear="1866" box="[945,1085,580,606]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F201569FC777A3AFBF6FDDB" box="[945,1085,580,606]" italics="true" pageId="23" pageNumber="24">Anomphalus</emphasis>
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</taxonomicName>
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?
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<emphasis id="B968EAAF4F201569FB967A3AFB6CFDDB" box="[1104,1191,580,606]" italics="true" pageId="23" pageNumber="24">blancus</emphasis>
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<bibRefCitation id="EF8D4B4C4F201569FB697A3BFA57FDDB" author="Kues & Batten" box="[1199,1436,580,607]" firstAuthor="Kues" pageId="23" pageNumber="24" pagination="1 - 95" refId="ref25729" refString="Kues, B. S. & Batten, R. L. (2001) Middle Pennsylvanian gastropods from the Flechado Formation, north-central New Mexico. Journal of Paleontology, 75 (Supplement 1), 1 - 95. https: // doi. org / 10.1666 / 0022 - 3360 (2001) 75 [1: MPGFTF] 2.0. CO; 2" type="journal article" year="2001">Kues & Batten, 2001</bibRefCitation>
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(p. 44, fig. 8.23–8.28) from the Middle Pennsylvanian of New Mexico in shell shape and the whorl expansion rate.
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<bibRefCitation id="EF8D4B4C4F201569FF517AF3FE47FD23" author="Kues & Batten" box="[151,396,652,679]" firstAuthor="Kues" pageId="23" pageNumber="24" pagination="1 - 95" refId="ref25729" refString="Kues, B. S. & Batten, R. L. (2001) Middle Pennsylvanian gastropods from the Flechado Formation, north-central New Mexico. Journal of Paleontology, 75 (Supplement 1), 1 - 95. https: // doi. org / 10.1666 / 0022 - 3360 (2001) 75 [1: MPGFTF] 2.0. CO; 2" type="journal article" year="2001">Kues & Batten (2001)</bibRefCitation>
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assigned
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<emphasis id="B968EAAF4F201569FE3E7AF2FDC2FD23" box="[504,521,652,678]" italics="true" pageId="23" pageNumber="24">A</emphasis>
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||
.?
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<emphasis id="B968EAAF4F201569FDE57AF2FDB1FD23" box="[547,634,652,678]" italics="true" pageId="23" pageNumber="24">blancus</emphasis>
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questionably to the genus
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<taxonomicName id="4C1C4D3E4F201569FC667AF2FBE7FD23" authorityName="Meek & Worthen" authorityYear="1866" box="[928,1068,652,678]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F201569FC667AF2FBE7FD23" box="[928,1068,652,678]" italics="true" pageId="23" pageNumber="24">Anomphalus</emphasis>
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</taxonomicName>
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due to its higher spired and more inflated shell which differs from typical species of
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<taxonomicName id="4C1C4D3E4F201569FD1E7ACEFCAFFD4F" authorityName="Meek & Worthen" authorityYear="1866" box="[728,868,688,714]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F201569FD1E7ACEFCAFFD4F" box="[728,868,688,714]" italics="true" pageId="23" pageNumber="24">Anomphalus</emphasis>
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</taxonomicName>
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in these respects that have low-spired or planispiral shells. The studied specimens also resemble the specimens assigned to
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<taxonomicName id="4C1C4D3E4F201569FC327AAAFB4BFD6B" authorityName="Meek & Worthen" authorityYear="1866" box="[1012,1152,724,750]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F201569FC327AAAFB4BFD6B" box="[1012,1152,724,750]" italics="true" pageId="23" pageNumber="24">Anomphalus</emphasis>
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</taxonomicName>
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sp. from the Permian of
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<collectingCountry id="F30B762D4F201569FF517A86FF35FC97" box="[151,254,760,786]" name="Malaysia" pageId="23" pageNumber="24">Malaysia</collectingCountry>
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by
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<bibRefCitation id="EF8D4B4C4F201569FEEC7A87FE74FC97" author="Batten" box="[298,447,760,787]" firstAuthor="Batten" pageId="23" pageNumber="24" pagination="1 - 26" refId="ref23086" refString="Batten, R. L. (1979) Permian gastropods from Perak, Malaysia. Part 2. The trochids, patellids, and neritids. American Museum Novitates, 2685, 1 - 26." type="journal article" year="1979">Batten (1979</bibRefCitation>
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, p. 8, figs 10–11). However, the present specimens are more high-spired.
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<bibRefCitation id="EF8D4B4C4F201569FAC77A87FA57FC97" author="Batten" box="[1281,1436,760,787]" firstAuthor="Batten" pageId="23" pageNumber="24" pagination="1 - 26" refId="ref23086" refString="Batten, R. L. (1979) Permian gastropods from Perak, Malaysia. Part 2. The trochids, patellids, and neritids. American Museum Novitates, 2685, 1 - 26." type="journal article" year="1979">Batten (1979)</bibRefCitation>
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and
|
||
<bibRefCitation id="EF8D4B4C4F201569FF0E7B63FE0FFCB3" author="Kues & Batten" box="[200,452,796,823]" firstAuthor="Kues" pageId="23" pageNumber="24" pagination="1 - 95" refId="ref25729" refString="Kues, B. S. & Batten, R. L. (2001) Middle Pennsylvanian gastropods from the Flechado Formation, north-central New Mexico. Journal of Paleontology, 75 (Supplement 1), 1 - 95. https: // doi. org / 10.1666 / 0022 - 3360 (2001) 75 [1: MPGFTF] 2.0. CO; 2" type="journal article" year="2001">Kues & Batten (2001)</bibRefCitation>
|
||
discussed that their specimens closely resemble the genus
|
||
<taxonomicName id="4C1C4D3E4F201569FBA37B62FB32FCB3" authorityName="de Koninck" authorityYear="1881" box="[1125,1273,796,822]" class="Gastropoda" family="Anomphalidae" genus="Turbinilopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trochida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F201569FBA37B62FB32FCB3" box="[1125,1273,796,822]" italics="true" pageId="23" pageNumber="24">Turbinilopsis</emphasis>
|
||
</taxonomicName>
|
||
in shell shape and proportions of the shell, but
|
||
<taxonomicName id="4C1C4D3E4F201569FDD67B3EFD6FFCDF" authorityName="de Koninck" authorityYear="1881" box="[528,676,832,858]" class="Gastropoda" family="Anomphalidae" genus="Turbinilopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trochida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F201569FDD67B3EFD6FFCDF" box="[528,676,832,858]" italics="true" pageId="23" pageNumber="24">Turbinilopsis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished in having an obvious callus within the umbilicus. Other species of
|
||
<taxonomicName id="4C1C4D3E4F201569FE937B1AFE22FCFB" authorityName="de Koninck" authorityYear="1881" box="[341,489,868,894]" class="Gastropoda" family="Anomphalidae" genus="Turbinilopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trochida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F201569FE937B1AFE22FCFB" box="[341,489,868,894]" italics="true" pageId="23" pageNumber="24">Turbinilopsis</emphasis>
|
||
</taxonomicName>
|
||
which have been reported from the Permian of Southeast Asia differ distinctly from the present material.
|
||
<taxonomicName id="4C1C4D3E4F201569FE427BF7FD6AFC27" authority="Delpey, 1941" authorityName="Delpey" authorityYear="1941" box="[388,673,904,931]" class="Gastropoda" family="Anomphalidae" genus="Turbinilopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trochida" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="rotundus">
|
||
<emphasis id="B968EAAF4F201569FE427BF7FE5FFC27" box="[388,404,905,930]" italics="true" pageId="23" pageNumber="24">T</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F201569FE647BF6FDCEFC27" box="[418,517,904,930]" italics="true" pageId="23" pageNumber="24">rotundus</emphasis>
|
||
<bibRefCitation id="EF8D4B4C4F201569FDCA7BF6FD6AFC27" author="Delpey" box="[524,673,904,930]" firstAuthor="Delpey" pageId="23" pageNumber="24" pagination="345 - 369" refId="ref23861" refString="Delpey, G. (1941) Les Gasteropodes permiens du Cambodge. Journal de Conchyologie, 84, 255 - 278, 345 - 369." type="journal article" year="1941">Delpey, 1941</bibRefCitation>
|
||
</taxonomicName>
|
||
(p. 276, fig. 18) from
|
||
<collectingCountry id="F30B762D4F201569FC667BF6FBD8FC27" box="[928,1043,904,930]" name="Cambodia" pageId="23" pageNumber="24">Cambodia</collectingCountry>
|
||
(see also
|
||
<bibRefCitation id="EF8D4B4C4F201569FB427BF7FAC6FC27" author="Batten" box="[1156,1293,904,931]" firstAuthor="Batten" pageId="23" pageNumber="24" pagination="1 - 26" refId="ref23086" refString="Batten, R. L. (1979) Permian gastropods from Perak, Malaysia. Part 2. The trochids, patellids, and neritids. American Museum Novitates, 2685, 1 - 26." type="journal article" year="1979">Batten 1979</bibRefCitation>
|
||
) has a much more straight whorl face and a blunt apex. The specimens assigned to
|
||
<taxonomicName id="4C1C4D3E4F201569FC5A7BD2FBFBFC43" authorityName="de Koninck" authorityYear="1881" box="[924,1072,940,966]" class="Gastropoda" family="Anomphalidae" genus="Turbinilopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trochida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F201569FC5A7BD2FBFBFC43" box="[924,1072,940,966]" italics="true" pageId="23" pageNumber="24">Turbinilopsis</emphasis>
|
||
</taxonomicName>
|
||
? sp. by
|
||
<bibRefCitation id="EF8D4B4C4F201569FB4C7BD3FAE5FC43" author="Mansuy" box="[1162,1326,940,967]" firstAuthor="Mansuy" pageId="23" pageNumber="24" pagination="1 - 190" refId="ref26085" refString="Mansuy, H. (1914) Nouvelle contribution a la paleontologie du Yunnan. Memoires du Service Geologique de l'Indochine, 3 (2), 1 - 190." type="journal article" year="1914">Mansuy (1914</bibRefCitation>
|
||
, pl. 4, fig. 11) have a lower spire and hence a greater apical angle. The present specimens are placed tentatively in the genus
|
||
<emphasis id="B968EAAF4F201569FF517B8AFEE0FB8B" box="[151,299,1012,1038]" italics="true" pageId="23" pageNumber="24">
|
||
<taxonomicName id="4C1C4D3E4F201569FF517B8AFEECFB8B" authorityName="Meek & Worthen" authorityYear="1866" box="[151,295,1012,1038]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">Anomphalus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
They are rather high-spired for this genus and hence become similar to the genus
|
||
<taxonomicName id="4C1C4D3E4F201569FB107B8AFA84FB8B" authorityName="Knight" authorityYear="1933" box="[1238,1359,1012,1038]" class="Gastropoda" family="Elasmonematidae" genus="Anematina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Vetigastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F201569FB107B8AFA84FB8B" box="[1238,1359,1012,1038]" italics="true" pageId="23" pageNumber="24">Anematina</emphasis>
|
||
</taxonomicName>
|
||
which generally is even more high-spired.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |