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<mods:title id="A42B6B5FDBD59BD86324181D5413BE9F">Taxonomic revision of Paraphasma Redtenbacher, 1906 (Phasmatodea, Pseudophasmatidae) based on phallic and external morphology</mods:title>
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<emphasis id="B968EAAEFF93FF8EFF2AFB55FE40F330" bold="true" box="[151,437,1263,1290]" italics="true" pageId="70" pageNumber="71">Pseudophasma amabile</emphasis>
(
<bibRefCitation id="EF8D4B4DFF93FF8EFE79FB4AFD4BF330" author="Redtenbacher, J." box="[452,702,1264,1290]" pageId="70" pageNumber="71" pagination="1 - 18" refId="ref47604" refString="Redtenbacher, J. (1906) I. Tribus Bacillini, II. Tribus Obrimini, III. Tribus Pygirhynchini, IV. Tribus Ascepasmini, V. Tribus Anisomorphini, VI. Tribus Phasmini, VII. Tribus Heteropterygini, VIII. Tribus Phyllini. In: Brunner von Wattenwyl, K. &amp; Redtenbacher, J. (Eds.), 1906 - 1908, Die Insektenfamilie der Phasmiden. Wilhelm Engelmann, Leipzig, pp. 1 - 18, tafel I - VI." type="book chapter" year="1906">Redtenbacher, 1906</bibRefCitation>
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.
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<bibRefCitation id="EF8D4B4DFF93FF8EFECEFAE2FD9AF34A" author="Redtenbacher, J." box="[371,623,1368,1392]" pageId="70" pageNumber="71" pagination="1 - 18" refId="ref47604" refString="Redtenbacher, J. (1906) I. Tribus Bacillini, II. Tribus Obrimini, III. Tribus Pygirhynchini, IV. Tribus Ascepasmini, V. Tribus Anisomorphini, VI. Tribus Phasmini, VII. Tribus Heteropterygini, VIII. Tribus Phyllini. In: Brunner von Wattenwyl, K. &amp; Redtenbacher, J. (Eds.), 1906 - 1908, Die Insektenfamilie der Phasmiden. Wilhelm Engelmann, Leipzig, pp. 1 - 18, tafel I - VI." type="book chapter" year="1906">Redtenbacher, 1906: 115</bibRefCitation>
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Delfosse
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, 2019: 189
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, 2324, 26, 36, figs 13C,D, 18D, 20B, 21E, 24E, 25, 26.
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(here designated):
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,
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, Marcapata (NHMW) (
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).
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<typeStatus id="54A7881EFF93FF8EFB40FA2CFA69F394" box="[1277,1436,1430,1454]" pageId="70" pageNumber="71" type="paralectotype">Paralectotypes</typeStatus>
(here designated):
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,
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, Marcapata (NHMW) (
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);
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,
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, Marcapata (ZMUH);
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,
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,
<collectingCountry id="F30B762CFF93FF8EFAF6FA0EFA6DF3F6" box="[1355,1432,1460,1484]" name="Bolivia" pageId="70" pageNumber="71">Bolivia</collectingCountry>
, Yungas de
<collectingRegion id="49D8F85EFF93FF8EFE8BFA6EFE89F3D1" box="[310,380,1492,1515]" country="Bolivia" name="La Paz" pageId="70" pageNumber="71">La Paz</collectingRegion>
(ZMUH);
<specimenCount id="9D1AFD35FF93FF8EFE56FA69FDF9F3D1" box="[491,524,1491,1515]" pageId="70" pageNumber="71" type="male">1♂</specimenCount>
,
<collectingCountry id="F30B762CFF93FF8EFDAAFA69FD91F3D1" box="[535,612,1491,1515]" name="Bolivia" pageId="70" pageNumber="71">Bolivia</collectingCountry>
, Yungas de
<collectingRegion id="49D8F85EFF93FF8EFD60FA6EFCD6F3D1" box="[733,803,1492,1515]" country="Bolivia" name="La Paz" pageId="70" pageNumber="71">La Paz</collectingRegion>
(MNHN).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA336BCFF93FF8EFF2AFA48FC91F013" blockId="70.[151,1437,1368,1607]" pageId="70" pageNumber="71">
<treatmentCitationGroup id="AB0C1192FF93FF8EFF2AFA48FC91F013" pageId="70" pageNumber="71">
<treatmentCitation id="0ABD10ADFF93FF8EFF2AFA48FD45F030" author="Gunther, K." box="[151,688,1522,1546]" page="563" pageId="70" pageNumber="71" year="1930">
<taxonomicName id="4C1C4D3FFF93FF8EFF2AFA48FD45F030" ID-CoL="4P6WH" authority="Gunther, 1930: 563" authorityName="Gunther" authorityPageNumber="563" authorityYear="1930" box="[151,688,1522,1546]" class="Insecta" family="Pseudophasmatidae" genus="Pseudophasma" kingdom="Animalia" order="Phasmida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="xanthotaenidium">
<emphasis id="B968EAAEFF93FF8EFF2AFA48FE14F030" box="[151,481,1522,1546]" italics="true" pageId="70" pageNumber="71">Pseudophasma xanthotaenidium</emphasis>
<bibRefCitation id="EF8D4B4DFF93FF8EFE55FA48FD45F030" author="Gunther, K." box="[488,688,1522,1546]" pageId="70" pageNumber="71" pagination="559 - 570" refId="ref46012" refString="Gunther, K. (1930) Neue und wenig bekannte Phasmoiden von Sudamerika. Mitteilungen aus dem Zoologischen Museum in Berlin, 15, 559 - 570." type="journal article" year="1930">Günther, 1930: 563</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 6;
<treatmentCitation id="0ABD10ADFF93FF8EFD42FA48FC5CF030" author="Klante, H." box="[767,937,1522,1546]" page="90" pageId="70" pageNumber="71" year="1960">
<bibRefCitation id="EF8D4B4DFF93FF8EFD42FA48FC5CF030" author="Klante, H." box="[767,937,1522,1546]" pageId="70" pageNumber="71" pagination="89 - 101" refId="ref46942" refString="Klante, H. (1960) Stabheuschrecken (Insecta, Phasmatodea) aus dem Naturkundemuseum Gorlitz. Abhandlungen und Berichte des Naturkundemuseums Gorlitz, 36, 89 - 101." type="journal article" year="1960">Klante, 1960: 90</bibRefCitation>
</treatmentCitation>
[=
<taxonomicName id="4C1C4D3FFF93FF8EFC70FA48FA66F030" ID-CoL="4D7XT" authority="Redtenbacher, 1906" authorityName=", Redtenbacher" authorityYear="1906" box="[973,1427,1522,1546]" class="Insecta" family="Pseudophasmatidae" genus="Paraphasma" kingdom="Animalia" order="Phasmida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="marginale">
<emphasis id="B968EAAEFF93FF8EFC70FA48FB48F030" box="[973,1213,1522,1546]" italics="true" pageId="70" pageNumber="71">Paraphasma marginale</emphasis>
<bibRefCitation id="EF8D4B4DFF93FF8EFB7EFA48FA66F030" author="Redtenbacher, J." box="[1219,1427,1522,1546]" pageId="70" pageNumber="71" pagination="1 - 18" refId="ref47604" refString="Redtenbacher, J. (1906) I. Tribus Bacillini, II. Tribus Obrimini, III. Tribus Pygirhynchini, IV. Tribus Ascepasmini, V. Tribus Anisomorphini, VI. Tribus Phasmini, VII. Tribus Heteropterygini, VIII. Tribus Phyllini. In: Brunner von Wattenwyl, K. &amp; Redtenbacher, J. (Eds.), 1906 - 1908, Die Insektenfamilie der Phasmiden. Wilhelm Engelmann, Leipzig, pp. 1 - 18, tafel I - VI." type="book chapter" year="1906">Redtenbacher, 1906</bibRefCitation>
</taxonomicName>
].
<emphasis id="B968EAAEFF93FF8EFF7AF9AAFE6EF013" bold="true" box="[199,411,1552,1577]" pageId="70" pageNumber="71">
Syn. nov.
<typeStatus id="54A7881EFF93FF8EFE92F9ABFE60F013" box="[303,405,1553,1577]" pageId="70" pageNumber="71" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D1AFD35FF93FF8EFE1CF9ABFE43F012" box="[417,438,1553,1576]" pageId="70" pageNumber="71" type="male"></specimenCount>
,
<collectingCountry id="F30B762CFF93FF8EFE7DF9ABFE1BF012" box="[448,494,1553,1576]" name="Peru" pageId="70" pageNumber="71">Peru</collectingCountry>
(MNB).
<emphasis id="B968EAAEFF93FF8EFDF1F9ABFD4DF013" bold="true" box="[588,696,1553,1577]" pageId="70" pageNumber="71">
<typeStatus id="54A7881EFF93FF8EFDF1F9ABFD47F013" box="[588,690,1553,1577]" pageId="70" pageNumber="71" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D1AFD35FF93FF8EFD03F9ABFD24F012" box="[702,721,1553,1576]" pageId="70" pageNumber="71" type="female"></specimenCount>
,
<collectingCountry id="F30B762CFF93FF8EFD60F9ABFCFEF012" box="[733,779,1553,1576]" name="Peru" pageId="70" pageNumber="71">Peru</collectingCountry>
(MNB).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA336BCFF93FF8EFF2AF995FC5EF07D" blockId="70.[151,1437,1368,1607]" box="[151,939,1583,1607]" pageId="70" pageNumber="71">
<treatmentCitationGroup id="AB0C1192FF93FF8EFF2AF995FC5EF07D" box="[151,939,1583,1607]" pageId="70" pageNumber="71">
<treatmentCitation id="0ABD10ADFF93FF8EFF2AF995FD68F07D" author="Zompro, O." box="[151,669,1583,1607]" page="159" pageId="70" pageNumber="71" year="2004">
<taxonomicName id="4C1C4D3FFF93FF8EFF2AF995FD68F07D" ID-CoL="4P6WH" authority=", Zompro, 2004: 159" authorityName="Zompro" authorityPageNumber="159" authorityYear="2004" box="[151,669,1583,1607]" class="Insecta" family="Pseudophasmatidae" genus="Paraphasma" kingdom="Animalia" order="Phasmida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="xanthotaenidium">
<emphasis id="B968EAAEFF93FF8EFF2AF995FE3CF07D" box="[151,457,1583,1607]" italics="true" pageId="70" pageNumber="71">Paraphasma xanthotaenidium</emphasis>
,
<bibRefCitation id="EF8D4B4DFF93FF8EFE6BF98AFD68F07D" author="Zompro, O." box="[470,669,1583,1607]" pageId="70" pageNumber="71" pagination="1 - 327" refId="ref48943" refString="Zompro, O. (2004) Revision of the genera of the Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg, Neue Folge, 37, 1 - 327." type="journal article" year="2004">Zompro, 2004: 159</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0ABD10ADFF93FF8EFD15F995FC52F07D" author="Otte, D. &amp; Brock, P. D." box="[680,935,1583,1607]" page="252" pageId="70" pageNumber="71" year="2005">
<bibRefCitation id="EF8D4B4DFF93FF8EFD15F995FC52F07D" author="Otte, D. &amp; Brock, P. D." box="[680,935,1583,1607]" pageId="70" pageNumber="71" refId="ref47359" refString="Otte, D. &amp; Brock, P. D. (2005) Phasmida Species File. Catalog of stick and leaf insects of the world. Insect Diversity Association, Philadelphia, 414 pp." type="book" year="2005">Otte &amp; Brock, 2005: 252</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3066537FF93FF8FFF2AF9CFFCA5F6C3" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71" type="discussion">
<paragraph id="8BA336BCFF93FF8EFF2AF9CFFB6CF15D" blockId="70.[151,1437,1653,2039]" pageId="70" pageNumber="71">
<emphasis id="B968EAAEFF93FF8EFF2AF9CFFEFEF0B5" bold="true" box="[151,267,1653,1679]" pageId="70" pageNumber="71">Remarks.</emphasis>
The phylogenetic analysis of
<taxonomicName id="4C1C4D3FFF93FF8EFDEAF9CFFD13F0B5" baseAuthorityName="Chiquetto-Machado &amp; Cancello" baseAuthorityYear="2021" box="[599,742,1653,1679]" class="Insecta" family="Pseudophasmatidae" genus="Paraphasma" kingdom="Animalia" order="Phasmida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAAEFF93FF8EFDEAF9CFFD13F0B5" box="[599,742,1653,1679]" italics="true" pageId="70" pageNumber="71">Paraphasma</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8D4B4DFF93FF8EFD49F9CFFB6BF0B5" author="Chiquetto-Machado, P. I. &amp; Cancello, E. M." box="[756,1182,1653,1679]" pageId="70" pageNumber="71" pagination="158 - 198" refId="ref44573" refString="Chiquetto-Machado, P. I. &amp; Cancello, E. M. (2021) Cladistic analysis of Paraphasma (Phasmatodea: Pseudophasmatidae) highlights the importance of the phallic organ for phasmid systematics. Zoological Journal of the Linnean Society, 193, 158 - 198. https: // doi. org / 10.1093 / zoolinnean / zlab 004" type="journal article" year="2021">Chiquetto-Machado &amp; Cancello 2021</bibRefCitation>
: fig. 25) resulted in the placement of
<taxonomicName id="4C1C4D3FFF93FF8EFE92F923FDEBF089" authorityName="Redtenbacher" authorityYear="1906" box="[303,542,1689,1715]" class="Insecta" family="Pseudophasmatidae" genus="Paraphasma" kingdom="Animalia" order="Phasmida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="amabile">
<emphasis id="B968EAAEFF93FF8EFE92F923FDEBF089" box="[303,542,1689,1715]" italics="true" pageId="70" pageNumber="71">Paraphasma amabile</emphasis>
</taxonomicName>
as the sister of
<taxonomicName id="4C1C4D3FFF93FF8EFD76F923FB61F089" authority="Kirby, 1904 b" authorityName="Kirby" authorityYear="1904" box="[715,1172,1689,1715]" class="Insecta" family="Pseudophasmatidae" genus="Pseudophasma" kingdom="Animalia" order="Phasmida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="cambridgei">
<emphasis id="B968EAAEFF93FF8EFD76F923FC0EF089" box="[715,1019,1689,1715]" italics="true" pageId="70" pageNumber="71">Pseudophasma cambridgei</emphasis>
<bibRefCitation id="EF8D4B4DFF93FF8EFBBCF923FB61F089" author="Kirby, W. F." box="[1025,1172,1689,1715]" pageId="70" pageNumber="71" pagination="372 - 377" refId="ref46877" refString="Kirby, W. F. (1904 b) Notes on Phasmidae in the collection of the British Museum (Natural History), South Kensington, with descriptions of new species - No. I. The Annals and Magazine of Natural History, Series 7, 13, 372 - 377. https: // doi. org / 10.1080 / 00222930408562458" type="journal article" year="1904">Kirby, 1904b</bibRefCitation>
</taxonomicName>
in a clade supported by seven synapomorphies. Two of them were exclusive of this clade, i.e. the male subgenital plate with a single lateral projection, on the right side, and the presence of a basal apodeme in the ventral lobe of the phallic organ. The other five synapomorphies, convergently shared with other clades in the tree, were the following: rudimentary ocelli; meso- and metafemur with distinct carinae; distinctly asymmetric vomer; anterior region of the male subgenital plate strongly convex; lateral projection of the male subgenital plate large and conspicuous.
</paragraph>
<paragraph id="8BA336BCFF93FF8FFF7AF8CBFCA5F6C3" blockId="70.[151,1437,1653,2039]" lastBlockId="71.[151,1436,151,465]" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71">
This result strongly indicates that
<taxonomicName id="4C1C4D3FFF93FF8EFDF1F8CBFCCBF1B1" authorityName="Redtenbacher" authorityYear="1906" box="[588,830,1905,1931]" class="Insecta" family="Pseudophasmatidae" genus="Paraphasma" kingdom="Animalia" order="Phasmida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="amabile">
<emphasis id="B968EAAEFF93FF8EFDF1F8CBFCCBF1B1" box="[588,830,1905,1931]" italics="true" pageId="70" pageNumber="71">Paraphasma amabile</emphasis>
</taxonomicName>
is more closely related to
<taxonomicName id="4C1C4D3FFF93FF8EFBCDF8CBFAEFF1B1" authorityName="Kirby" authorityYear="1896" box="[1136,1306,1905,1931]" class="Insecta" family="Pseudophasmatidae" genus="Pseudophasma" kingdom="Animalia" order="Phasmida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAAEFF93FF8EFBCDF8CBFAEFF1B1" box="[1136,1306,1905,1931]" italics="true" pageId="70" pageNumber="71">Pseudophasma</emphasis>
</taxonomicName>
than to the true representatives of
<taxonomicName id="4C1C4D3FFF93FF8EFE18F82FFDC1F195" baseAuthorityName="Chiquetto-Machado &amp; Cancello" baseAuthorityYear="2021" box="[421,564,1941,1967]" class="Insecta" family="Pseudophasmatidae" genus="Paraphasma" kingdom="Animalia" order="Phasmida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAAEFF93FF8EFE18F82FFDC1F195" box="[421,564,1941,1967]" italics="true" pageId="70" pageNumber="71">Paraphasma</emphasis>
</taxonomicName>
, leading us to propose here the new combination
<taxonomicName id="4C1C4D3FFF93FF8EFB30F82FFE7CF1EE" authority="(Redtenbacher, 1906)" authorityName="Chiquetto-Machado &amp; Morales &amp; Cancello" authorityYear="2022" baseAuthorityName="Redtenbacher" baseAuthorityYear="1906" class="Insecta" family="Pseudophasmatidae" genus="Pseudophasma" kingdom="Animalia" order="Phasmida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="amabile" status="comb. nov.">
<emphasis id="B968EAAEFF93FF8EFB30F82FFA69F195" box="[1165,1436,1941,1967]" italics="true" pageId="70" pageNumber="71">Pseudophasma amabile</emphasis>
(
<bibRefCitation id="EF8D4B4DFF93FF8EFF22F803FE74F1EE" author="Redtenbacher, J." box="[159,385,1977,2004]" pageId="70" pageNumber="71" pagination="1 - 18" refId="ref47604" refString="Redtenbacher, J. (1906) I. Tribus Bacillini, II. Tribus Obrimini, III. Tribus Pygirhynchini, IV. Tribus Ascepasmini, V. Tribus Anisomorphini, VI. Tribus Phasmini, VII. Tribus Heteropterygini, VIII. Tribus Phyllini. In: Brunner von Wattenwyl, K. &amp; Redtenbacher, J. (Eds.), 1906 - 1908, Die Insektenfamilie der Phasmiden. Wilhelm Engelmann, Leipzig, pp. 1 - 18, tafel I - VI." type="book chapter" year="1906">Redtenbacher, 1906</bibRefCitation>
)
</taxonomicName>
<emphasis id="B968EAAEFF93FF8EFE2DF803FDE4F1E9" bold="true" box="[400,529,1977,2003]" pageId="70" pageNumber="71">
<taxonomicNameLabel id="A25B57D5FF93FF8EFE2DF803FDE4F1E9" box="[400,529,1977,2003]" pageId="70" pageNumber="71" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
This transfer is also corroborated by the similarity between the phallic organs of
<taxonomicName id="4C1C4D3FFF93FF8EFF2AF864FEF9F1CD" authorityName="Chiquetto-Machado &amp; Morales &amp; Cancello" authorityYear="2022" baseAuthorityName="Redtenbacher" baseAuthorityYear="1906" box="[151,268,2013,2039]" class="Insecta" family="Pseudophasmatidae" genus="Pseudophasma" kingdom="Animalia" order="Phasmida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="amabile">
<emphasis id="B968EAAEFF93FF8EFF2AF864FEF9F1CD" box="[151,268,2013,2039]" italics="true" pageId="70" pageNumber="71">P. amabile</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D3FFF93FF8EFEFFF864FE28F1CD" authorityName="Kirby" authorityYear="1904" box="[322,477,2013,2039]" class="Insecta" family="Pseudophasmatidae" genus="Pseudophasma" kingdom="Animalia" order="Phasmida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="cambridgei">
<emphasis id="B968EAAEFF93FF8EFEFFF864FE28F1CD" box="[322,477,2013,2039]" italics="true" pageId="70" pageNumber="71">P. cambridgei</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF8D4B4DFF93FF8EFDABF867FC4AF1CD" author="Chiquetto-Machado, P. I. &amp; Cancello, E. M." box="[534,959,2013,2039]" pageId="70" pageNumber="71" pagination="158 - 198" refId="ref44573" refString="Chiquetto-Machado, P. I. &amp; Cancello, E. M. (2021) Cladistic analysis of Paraphasma (Phasmatodea: Pseudophasmatidae) highlights the importance of the phallic organ for phasmid systematics. Zoological Journal of the Linnean Society, 193, 158 - 198. https: // doi. org / 10.1093 / zoolinnean / zlab 004" type="journal article" year="2021">Chiquetto-Machado &amp; Cancello 2021</bibRefCitation>
: fig. 13AD) and by the general terminalia morphology of both sexes of
<taxonomicName id="4C1C4D3FFF92FF8FFE56FF2DFD96F68B" authorityName="Chiquetto-Machado &amp; Morales &amp; Cancello" authorityYear="2022" baseAuthorityName="Redtenbacher" baseAuthorityYear="1906" box="[491,611,151,177]" class="Insecta" family="Pseudophasmatidae" genus="Pseudophasma" kingdom="Animalia" order="Phasmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="amabile">
<emphasis id="B968EAAEFF92FF8FFE56FF2DFD96F68B" box="[491,611,151,177]" italics="true" pageId="71" pageNumber="72">P. amabile</emphasis>
</taxonomicName>
, which is typical of many representatives of
<taxonomicName id="4C1C4D3FFF92FF8FFBD6FF2DFAE0F68B" authorityName="Kirby" authorityYear="1896" box="[1131,1301,151,177]" class="Insecta" family="Pseudophasmatidae" genus="Pseudophasma" kingdom="Animalia" order="Phasmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAAEFF92FF8FFBD6FF2DFAE0F68B" box="[1131,1301,151,177]" italics="true" pageId="71" pageNumber="72">Pseudophasma</emphasis>
</taxonomicName>
. It is worth noting, however, that the species-rich and heterogeneous
<taxonomicName id="4C1C4D3FFF92FF8FFCAAFF01FC34F6EF" authorityName="Kirby" authorityYear="1896" box="[791,961,187,213]" class="Insecta" family="Pseudophasmatidae" genus="Pseudophasma" kingdom="Animalia" order="Phasmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAAEFF92FF8FFCAAFF01FC34F6EF" box="[791,961,187,213]" italics="true" pageId="71" pageNumber="72">Pseudophasma</emphasis>
</taxonomicName>
is probably polyphyletic and in great need of revision (
<bibRefCitation id="EF8D4B4DFF92FF8FFEA3FF65FDB7F6C3" author="Chiquetto-Machado, P. I." box="[286,578,223,249]" pageId="71" pageNumber="72" pagination="392 - 402" refId="ref44455" refString="Chiquetto-Machado, P. I. (2018) Redescription of the Brazilian stick insect Pseudophasma cambridgei Kirby (Phasmatodea: Pseudophasmatidae), with first description of the female and egg. Austral Entomology, 57, 392 - 402. https: // doi. org / 10.1111 / aen. 12287" type="journal article" year="2018">Chiquetto-Machado 2018</bibRefCitation>
;
<bibRefCitation id="EF8D4B4DFF92FF8FFDF3FF65FCB1F6C3" author="Robertson, J. A. &amp; Bradler, S. &amp; Whiting, M. F." box="[590,836,223,249]" pageId="71" pageNumber="72" pagination="216" refId="ref47816" refString="Robertson, J. A., Bradler, S. &amp; Whiting, M. F. (2018) Evolution of oviposition techniques in stick and leaf insects (Phasmatodea). Frontiers in Ecology and Evolution, 6, 216. https: // doi. org / 10.3389 / fevo. 2018.00216" type="journal article" year="2018">
Robertson
<emphasis id="B968EAAEFF92FF8FFD75FF65FCF5F6C3" box="[712,768,223,249]" italics="true" pageId="71" pageNumber="72">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3066537FF92FF8FFF7AFEB9FF3DF797" pageId="71" pageNumber="72" type="reference_group">
<paragraph id="8BA336BCFF92FF8FFF7AFEB9FF3DF797" blockId="71.[151,1436,151,465]" pageId="71" pageNumber="72">
<taxonomicName id="4C1C4D3FFF92FF8FFF7AFEB9FD1AF727" authority="Gunther, 1930" authorityName="Gunther" authorityYear="1930" box="[199,751,259,285]" class="Insecta" family="Pseudophasmatidae" genus="Pseudophasma" kingdom="Animalia" order="Phasmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="xanthotaenidium">
<emphasis id="B968EAAEFF92FF8FFF7AFEB9FDCFF727" box="[199,570,259,285]" italics="true" pageId="71" pageNumber="72">Pseudophasma xanthotaenidium</emphasis>
<bibRefCitation id="EF8D4B4DFF92FF8FFDF5FEB9FD1AF727" author="Gunther, K." box="[584,751,259,285]" pageId="71" pageNumber="72" pagination="559 - 570" refId="ref46012" refString="Gunther, K. (1930) Neue und wenig bekannte Phasmoiden von Sudamerika. Mitteilungen aus dem Zoologischen Museum in Berlin, 15, 559 - 570." type="journal article" year="1930">Günther, 1930</bibRefCitation>
</taxonomicName>
was erroneously regarded as a synonym of
<taxonomicName id="4C1C4D3FFF92FF8FFAB0FEB9FEFCF77B" authorityName=", Redtenbacher" authorityYear="1906" class="Insecta" family="Pseudophasmatidae" genus="Paraphasma" kingdom="Animalia" order="Phasmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marginale">
<emphasis id="B968EAAEFF92FF8FFAB0FEB9FEFCF77B" italics="true" pageId="71" pageNumber="72">Paraphasma marginale</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF8D4B4DFF92FF8FFE93FE9DFE3DF778" author="Klante, H." box="[302,456,295,322]" pageId="71" pageNumber="72" pagination="89 - 101" refId="ref46942" refString="Klante, H. (1960) Stabheuschrecken (Insecta, Phasmatodea) aus dem Naturkundemuseum Gorlitz. Abhandlungen und Berichte des Naturkundemuseums Gorlitz, 36, 89 - 101." type="journal article" year="1960">Klante (1960)</bibRefCitation>
and was later listed as a valid species of
<taxonomicName id="4C1C4D3FFF92FF8FFCC7FE9DFBFCF77B" baseAuthorityName="Chiquetto-Machado &amp; Cancello" baseAuthorityYear="2021" box="[890,1033,295,321]" class="Insecta" family="Pseudophasmatidae" genus="Paraphasma" kingdom="Animalia" order="Phasmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAAEFF92FF8FFCC7FE9DFBFCF77B" box="[890,1033,295,321]" italics="true" pageId="71" pageNumber="72">Paraphasma</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF8D4B4DFF92FF8FFB93FE9DFB2CF77B" author="Zompro, O." box="[1070,1241,295,321]" pageId="71" pageNumber="72" pagination="1 - 327" refId="ref48943" refString="Zompro, O. (2004) Revision of the genera of the Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg, Neue Folge, 37, 1 - 327." type="journal article" year="2004">Zompro (2004)</bibRefCitation>
, who also pointed out that it was probably a synonym of
<emphasis id="B968EAAEFF92FF8FFDFFFEF1FCCCF75F" box="[578,825,331,357]" italics="true" pageId="71" pageNumber="72">
<taxonomicName id="4C1C4D3FFF92FF8FFDFFFEF1FCC0F75F" authorityName="Redtenbacher" authorityYear="1906" box="[578,821,331,357]" class="Insecta" family="Pseudophasmatidae" genus="Paraphasma" kingdom="Animalia" order="Phasmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="amabile">Paraphasma amabile</taxonomicName>
.
</emphasis>
Comparisons between the
<typeStatus id="54A7881EFF92FF8FFBDBFEF1FB54F75F" box="[1126,1185,331,357]" pageId="71" pageNumber="72">types</typeStatus>
of
<taxonomicName id="4C1C4D3FFF92FF8FFB78FEF1FA69F75F" authorityName="Gunther" authorityYear="1930" box="[1221,1436,331,357]" class="Insecta" family="Pseudophasmatidae" genus="Pseudophasma" kingdom="Animalia" order="Phasmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="xanthotaenidium">
<emphasis id="B968EAAEFF92FF8FFB78FEF1FA69F75F" box="[1221,1436,331,357]" italics="true" pageId="71" pageNumber="72">P. xanthotaenidium</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D3FFF92FF8FFF7BFED5FECEF7B3" authorityName="Redtenbacher" authorityYear="1906" box="[198,315,367,393]" class="Insecta" family="Pseudophasmatidae" genus="Paraphasma" kingdom="Animalia" order="Phasmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="amabile">
<emphasis id="B968EAAEFF92FF8FFF7BFED5FECEF7B3" box="[198,315,367,393]" italics="true" pageId="71" pageNumber="72">P. amabile</emphasis>
</taxonomicName>
confirmed this hypothesis, so this species is here synonymized under
<taxonomicName id="4C1C4D3FFF92FF8FFBF9FED5FAB8F7B3" authorityName="Chiquetto-Machado &amp; Morales &amp; Cancello" authorityYear="2022" baseAuthorityName="Redtenbacher" baseAuthorityYear="1906" box="[1092,1357,367,393]" class="Insecta" family="Pseudophasmatidae" genus="Pseudophasma" kingdom="Animalia" order="Phasmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="amabile" status="comb. nov.">
<emphasis id="B968EAAEFF92FF8FFBF9FED5FAB8F7B3" box="[1092,1357,367,393]" italics="true" pageId="71" pageNumber="72">Pseudophasma amabile</emphasis>
</taxonomicName>
<emphasis id="B968EAAEFF92FF8FFAE9FED5FF3DF797" bold="true" pageId="71" pageNumber="72">
<taxonomicNameLabel id="A25B57D5FF92FF8FFAE9FED5FF3DF797" pageId="71" pageNumber="72" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3066537FF92FF8FFF7AFE0CFBB1F7EB" box="[199,1092,438,465]" pageId="71" pageNumber="72" type="materials_examined">
<paragraph id="8BA336BCFF92FF8FFF7AFE0CFBB1F7EB" blockId="71.[151,1436,151,465]" box="[199,1092,438,465]" pageId="71" pageNumber="72">
<emphasis id="B968EAAEFF92FF8FFF7AFE0CFDC1F7EA" bold="true" box="[199,564,438,464]" pageId="71" pageNumber="72">Additional material examined.</emphasis>
<materialsCitation id="3B743CE1FF92FF8FFD86FE0DFBB5F7EB" box="[571,1088,438,465]" collectionCode="NHMW" country="Peru" location="Cusco" pageId="71" pageNumber="72" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cusco">
<emphasis id="B968EAAEFF92FF8FFD86FE0DFD12F7EA" bold="true" box="[571,743,438,465]" pageId="71" pageNumber="72">
<collectingCountry id="F30B762CFF92FF8FFD86FE0DFD7CF7EB" box="[571,649,439,465]" name="Peru" pageId="71" pageNumber="72">PERU</collectingCountry>
.
<collectingRegion id="49D8F85EFF92FF8FFD28FE0CFD17F7EA" box="[661,738,438,464]" country="Peru" name="Cusco" pageId="71" pageNumber="72">Cusco</collectingRegion>
:
</emphasis>
<specimenCount id="9D1AFD35FF92FF8FFD52FE0DFCE6F7EB" box="[751,787,439,465]" pageId="71" pageNumber="72" type="male">1♂</specimenCount>
,
<specimenCount id="9D1AFD35FF92FF8FFC9DFE0DFCB6F7EB" box="[800,835,439,465]" pageId="71" pageNumber="72" type="female">1♀</specimenCount>
, Marcapata (
<collectionCode id="ED0DAE79FF92FF8FFC6BFE0DFBCDF7EB" box="[982,1080,439,465]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="71" pageNumber="72" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>