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<document id="7B665C63A70AB4E407E9932463C4348E" ID-CLB-Dataset="255744" ID-DOI="10.25223/brad.n37.2019.a24" ID-GBIF-Dataset="bed395ed-3217-45e6-8e37-9d47822f2ded" ID-Zenodo-Dep="7867784" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1681981871443" checkinUser="mdunkwanyana" docAuthor="Smith, Gideon F." docDate="2019" docId="03B587A7FFF82866C8F4FCECFDCDF938" docLanguage="en" docName="Bradleya.37.26-30.pdf.imf" docOrigin="Bradleya 37" docSource="https://www.mendeley.com/catalogue/e8dc1af9-6adc-32e6-86fb-995a2370202f/" docTitle="Aloe spinosissima" docType="treatment" docVersion="9" lastPageNumber="29" masterDocId="FF8CFFDFFFF92862CA1AFF95FFCAFFFE" masterDocTitle="The taxonomy of aloe xspinosissima hort. ex a. berger (asphodelaceae), a popular hybrid aloe from mediterranean Europe" masterLastPageNumber="30" masterPageNumber="26" pageNumber="26" updateTime="1710359486582" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="4F0254736B3797B07F9C23E0B5013B69">The taxonomy of aloe xspinosissima hort. ex a. berger (asphodelaceae), a popular hybrid aloe from mediterranean Europe</mods:title>
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<mods:namePart id="110FB8F05EAD909DEA1E472FFB540ABF">Smith, Gideon F.</mods:namePart>
<mods:affiliation id="1542A4EAE244417D3D6B1D26159EF904">Department of Botany, P. O. Box 77000, Nelson Mandela University, Port Elizabeth, 6031 South Africa</mods:affiliation>
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<treatment id="03B587A7FFF82866C8F4FCECFDCDF938" ID-DOI="http://doi.org/10.5281/zenodo.7867786" ID-GBIF-Taxon="209570527" ID-Zenodo-Dep="7867786" LSID="urn:lsid:plazi:treatment:03B587A7FFF82866C8F4FCECFDCDF938" httpUri="http://treatment.plazi.org/id/03B587A7FFF82866C8F4FCECFDCDF938" lastPageId="4" lastPageNumber="29" pageId="1" pageNumber="26">
<subSubSection id="C306653AFFF82861C8F4FCECFDB8FAB7" lastPageId="3" lastPageNumber="28" pageId="1" pageNumber="26" type="discussion">
<paragraph id="8BA336B1FFF82863C8F4FCECFC96FC54" blockId="1.[750,1294,889,1736]" pageId="1" pageNumber="26">
<emphasis id="B968EAA3FFF82863C8F4FCECFC96FC54" bold="true" pageId="1" pageNumber="26">
Identity of
<taxonomicName id="4C1C4D32FFF82863C98BFCECFB15FC71" authority="Hort." authorityName="Hort." box="[913,1247,889,911]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFF82863C98BFCECFC0DFC71" bold="true" box="[913,967,889,911]" italics="true" pageId="1" pageNumber="26">Aloe</emphasis>
×
<emphasis id="B968EAA3FFF82863C9F3FCEFFB40FC71" bold="true" box="[1001,1162,890,911]" italics="true" pageId="1" pageNumber="26">spinosissima</emphasis>
Hort.
</taxonomicName>
ex A.Berger
</emphasis>
</paragraph>
<paragraph id="8BA336B1FFF82863C909FC24FBC9F9A7" blockId="1.[750,1294,889,1736]" pageId="1" pageNumber="26">
The earliest reference that could be traced to the name
<taxonomicName id="4C1C4D32FFF82863C976FC59FB80FC1C" box="[876,1098,972,994]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFF82863C976FC59FC56FC1F" box="[876,924,972,993]" italics="true" pageId="1" pageNumber="26">Aloe</emphasis>
×
<emphasis id="B968EAA3FFF82863C9A3FC58FB80FC1C" box="[953,1098,973,994]" italics="true" pageId="1" pageNumber="26">spinosissima</emphasis>
</taxonomicName>
is where Alwin Berger (
<date id="FFA21071FFF82863C95DFC7DFC3BFC03" box="[839,1009,1000,1021]" pageId="1" pageNumber="26" value="1871-08-28">28 August 1871</date>
[Möschlitz, Germany]
<date id="FFA21071FFF82863CEE9FC7DFCA8FBE6" pageId="1" pageNumber="26" value="1931-04-20">20 April 1931</date>
[Stuttgart, Germany]) described a hybrid aloe under this name in Ҥ 5
<emphasis id="B968EAA3FFF82863CE88FB8BFACEFBCD" box="[1170,1284,1054,1075]" italics="true" pageId="1" pageNumber="26">Echinatae</emphasis>
” (
<bibRefCitation id="EF8D4B40FFF82863C8ECFBAFFC76FBB1" author="BERGER, A." box="[758,956,1082,1103]" firstAuthor="Berger" journalOrPublisher="Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft" page="183" pageId="1" pageNumber="26" pagination="159 - 326" part="33" refId="ref2409" refString="BERGER, A. (1908). 8. Aloe L. In: A. ENGLER (ED.), Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft 33: 159 - 326. Verlag von Wilhelm Engelmann, Leipzig." title="Aloe L" type="journal article" volumeTitle="A. ENGLER (ED.)" year="1908">Berger, 1908: 183</bibRefCitation>
) (
<figureCitation id="13272A34FFF82863C9CFFBAFFBF5FBB1" box="[981,1087,1082,1104]" captionStart="Figure 3" captionStartId="3.[750,835,838,860]" captionTargetBox="[750,1294,145,829]" captionTargetId="figure-2578@3.[751,1293,145,828]" captionTargetPageId="3" captionText="Figure 3. Aloe ×spinosissima growing in a pedestalled, Italianate container in the Hanbury Gardens at La Mortola on the Ligurian Riviera, Italy. Photograph: Gideon F. Smith." figureDoi="http://doi.org/10.5281/zenodo.7867792" httpUri="https://zenodo.org/record/7867792/files/figure.png" pageId="1" pageNumber="26">Figures 3</figureCitation>
&amp;
<figureCitation id="13272A34FFF82863CE7EFBAFFBBBFBB1" box="[1124,1137,1082,1103]" captionStart="Figure 4" captionStartId="4.[145,229,838,860]" captionTargetBox="[145,1294,145,829]" captionTargetId="figure-2140@4.[146,1292,146,829]" captionTargetPageId="4" captionText="Figure 4. The Palazzo Orengo at the Hanbury Gardens. Alwin Berger, who described Aloe ×spinosissima, was based at the Garden at the time that he received material of this nothospecies from Ludwig Winter. Photograph: Gideon F. Smith." figureDoi="http://doi.org/10.5281/zenodo.7867794" httpUri="https://zenodo.org/record/7867794/files/figure.png" pageId="1" pageNumber="26">4</figureCitation>
). Three years earlier, where
<bibRefCitation id="EF8D4B40FFF82863C98EFBC3FB93FB95" author="BERGER, A." box="[916,1113,1109,1131]" firstAuthor="Berger" journalOrPublisher="Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie" page="58" pageId="1" pageNumber="26" pagination="42 - 68" part="36" publicationUrl="https://www.biodiversitylibrary.org/item/27521#page/56/mode/1up]" refId="ref2353" refString="BERGER, A. (1905). Uber die Systematische Gliederung der Gattung Aloe. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 36: 42 - 68. [see: https: // www. biodiversitylibrary. org / item / 27521 # page / 56 / mode / 1 up]." title="Uber die Systematische Gliederung der Gattung Aloe" type="journal article" year="1905">Berger (1905: 58)</bibRefCitation>
provided a preliminary outline of his proposed infrageneric classification of
<taxonomicName id="4C1C4D32FFF82863C991FB19FC71FB5F" box="[907,955,1164,1185]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="genus">
<emphasis id="B968EAA3FFF82863C991FB19FC71FB5F" box="[907,955,1164,1185]" italics="true" pageId="1" pageNumber="26">Aloe</emphasis>
</taxonomicName>
, he did not reference this nothospecies in § 2.
<emphasis id="B968EAA3FFF82863C9DDFB3DFBEEFB43" box="[967,1060,1192,1213]" italics="true" pageId="1" pageNumber="26">Humiles</emphasis>
, where it would have been grouped. The name was given to what
<bibRefCitation id="EF8D4B40FFF82863C8F4FB4AFC48FB0A" author="BERGER, A." box="[750,898,1247,1268]" firstAuthor="Berger" journalOrPublisher="Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft" pageId="1" pageNumber="26" pagination="159 - 326" part="33" refId="ref2409" refString="BERGER, A. (1908). 8. Aloe L. In: A. ENGLER (ED.), Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft 33: 159 - 326. Verlag von Wilhelm Engelmann, Leipzig." title="Aloe L" type="journal article" volumeTitle="A. ENGLER (ED.)" year="1908">Berger (1908)</bibRefCitation>
regarded as a hybrid between
<taxonomicName id="4C1C4D32FFF82863CED6FB4AFB48FAEE" authority="(Willd.) Baker" authorityName="Baker" baseAuthorityName="Willd." class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="variety" species="humilis" variety="echinata">
<emphasis id="B968EAA3FFF82863CED6FB4AFCEDFAF1" italics="true" pageId="1" pageNumber="26">A. humilis</emphasis>
var.
<emphasis id="B968EAA3FFF82863C974FB6FFC04FAF1" box="[878,974,1274,1295]" italics="true" pageId="1" pageNumber="26">echinata</emphasis>
(Willd.) Baker
</taxonomicName>
and
<taxonomicName id="4C1C4D32FFF82863CEDDFB6EFCA4FAB9" authority="(J. M. Wood &amp; M. S. Evans)" baseAuthorityName="J. M. Wood &amp; M. S. Evans" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="variety" species="arborescens" variety="natalensis">
<emphasis id="B968EAA3FFF82863CEDDFB6EFC92FAD5" italics="true" pageId="1" pageNumber="26">A. arborescens</emphasis>
var.
<emphasis id="B968EAA3FFF82863C9D3FA83FBF6FAD5" box="[969,1084,1302,1323]" italics="true" pageId="1" pageNumber="26">natalensis</emphasis>
(J.M.Wood &amp; M.S.Evans)
</taxonomicName>
A.Berger (
<bibRefCitation id="EF8D4B40FFF82863C9FCFAA7FB56FAB8" author="BERGER, A." box="[998,1180,1329,1351]" firstAuthor="Berger" journalOrPublisher="Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft" page="19" pageId="1" pageNumber="26" pagination="159 - 326" part="33" refId="ref2409" refString="BERGER, A. (1908). 8. Aloe L. In: A. ENGLER (ED.), Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft 33: 159 - 326. Verlag von Wilhelm Engelmann, Leipzig." title="Aloe L" type="journal article" volumeTitle="A. ENGLER (ED.)" year="1908">Berger, 1908: 19</bibRefCitation>
), while on the page where the protologue of the name appeared (
<bibRefCitation id="EF8D4B40FFF82863C953FAFCFBC4FA83" author="BERGER, A." box="[841,1038,1384,1406]" firstAuthor="Berger" journalOrPublisher="Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft" page="183" pageId="1" pageNumber="26" pagination="159 - 326" part="33" refId="ref2409" refString="BERGER, A. (1908). 8. Aloe L. In: A. ENGLER (ED.), Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft 33: 159 - 326. Verlag von Wilhelm Engelmann, Leipzig." title="Aloe L" type="journal article" volumeTitle="A. ENGLER (ED.)" year="1908">Berger, 1908: 183</bibRefCitation>
), he amended his view of the infraspecific classification of the
<taxonomicName id="4C1C4D32FFF82863CED2FA11FC92FA4A" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="B968EAA3FFF82863CED2FA11FC92FA4A" italics="true" pageId="1" pageNumber="26">A. arborescens</emphasis>
</taxonomicName>
parent to
<taxonomicName id="4C1C4D32FFF82863CE17FA0AFCBDFA2E" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="variety" species="arborescens" variety="pachythyrsa">
<emphasis id="B968EAA3FFF82863CE17FA0AFB0FFA4A" box="[1037,1221,1439,1461]" italics="true" pageId="1" pageNumber="26">A. arborescens</emphasis>
var.
<emphasis id="B968EAA3FFF82863C8F4FA2EFCBDFA2E" box="[750,887,1467,1488]" italics="true" pageId="1" pageNumber="26">pachythyrsa</emphasis>
</taxonomicName>
A.Berger. Note though that both
<taxonomicName id="4C1C4D32FFF82863C8F4FA43FCA1FA15" box="[750,875,1494,1516]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="species" species="humilis">
<emphasis id="B968EAA3FFF82863C8F4FA43FCA1FA15" box="[750,875,1494,1516]" italics="true" pageId="1" pageNumber="26">A. humilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D32FFF82863C9A9FA43FB96FA15" box="[947,1116,1494,1516]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="B968EAA3FFF82863C9A9FA43FB96FA15" box="[947,1116,1494,1516]" italics="true" pageId="1" pageNumber="26">A. arborescens</emphasis>
</taxonomicName>
are nowadays treated as variable species where the recognition of infraspecific taxa is hardly warranted (
<bibRefCitation id="EF8D4B40FFF82863CEFAF998FBE4F9C0" author="VAN WYK, B-E. &amp; SMITH, G. F." firstAuthor="Van Wyk" journalOrPublisher="3 rd edition. Briza Publications, Queenswood, Pretoria." page="290 - 291" pageId="1" pageNumber="26" refId="ref3690" refString="VAN WYK, B-E. [BEN-ERIK] &amp; SMITH, G. F. (2014). Guide to the aloes of South Africa. 3 rd edition. Briza Publications, Queenswood, Pretoria." title="Guide to the aloes of South Africa" type="book" year="2014">Van Wyk &amp; Smith, 2014: 290291</bibRefCitation>
;
<bibRefCitation id="EF8D4B40FFF82863CE21F9BDFACDF9C0" author="CROUCH, N. R. &amp; SMITH, G. F. &amp; KLOPPER, R. R." box="[1083,1287,1576,1598]" etAl="et al." firstAuthor="Crouch" journalOrPublisher="Bothalia" pageId="1" pageNumber="26" pagination="70 - 71" part="38" refId="ref2458" refString="CROUCH, N. R., SMITH, G. F. &amp; KLOPPER, R. R. (2008). Notes on African plants. Asphodelaceae. Notes on the nomenclature and typification of Aloe natalensis (Alooideae). Bothalia 38 (1): 70 - 71." title="Notes on African plants. Asphodelaceae. Notes on the nomenclature and typification of Aloe natalensis (Alooideae)" type="journal article" year="2008">
Crouch
<emphasis id="B968EAA3FFF82863CE8AF9BCFB75F9C3" box="[1168,1215,1576,1598]" italics="true" pageId="1" pageNumber="26">et al</emphasis>
., 2008
</bibRefCitation>
;
<bibRefCitation id="EF8D4B40FFF82863C8F4F9D1FC3EF9A7" author="SMITH, G. F. &amp; KLOPPER, R. R. &amp; FIGUEIREDO, E. &amp; CROUCH, N. R." box="[750,1012,1604,1626]" etAl="et al." firstAuthor="Smith" journalOrPublisher="Bradleya" pageId="1" pageNumber="26" pagination="127 - 137" part="30" refId="ref3639" refString="SMITH, G. F., KLOPPER, R. R., FIGUEIREDO, E. &amp; CROUCH, N. R. (2012). Aspects of the taxonomy of Aloe arborescens Mill. (Asphodelaceae: Alooideae). Bradleya 30: 127 - 137." title="Aspects of the taxonomy of Aloe arborescens Mill. (Asphodelaceae: Alooideae)" type="journal article" year="2012">
Smith
<emphasis id="B968EAA3FFF82863C922F9D0FCA2F9A7" box="[824,872,1604,1626]" italics="true" pageId="1" pageNumber="26">et al</emphasis>
., 2008, 2012
</bibRefCitation>
).
</paragraph>
<paragraph id="8BA336B1FFF82861C909F9CAFDF6FD13" blockId="1.[750,1294,889,1736]" lastBlockId="3.[145,690,700,1354]" lastPageId="3" lastPageNumber="28" pageId="1" pageNumber="26">
Natural hybrids have been recorded between
<taxonomicName id="4C1C4D32FFF82863C8F4F9EEFCB5F96E" box="[750,895,1659,1680]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="species" species="humilis">
<emphasis id="B968EAA3FFF82863C8F4F9EEFCB5F96E" box="[750,895,1659,1680]" italics="true" pageId="1" pageNumber="26">Aloe humilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D32FFF82863C9A4F9EEFB05F96E" authority="(Aiton) Haw." authorityName="Haw." baseAuthorityName="Aiton" box="[958,1231,1659,1681]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="species" species="lineata">
<emphasis id="B968EAA3FFF82863C9A4F9EEFBFAF96E" box="[958,1072,1659,1681]" italics="true" pageId="1" pageNumber="26">A. lineata</emphasis>
(Aiton) Haw.
</taxonomicName>
from near Port Elizabeth (Eastern Cape Province, South Africa), and between
<taxonomicName id="4C1C4D32FFF82863CE3BF927FB5EF939" box="[1057,1172,1714,1736]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="species" species="humilis">
<emphasis id="B968EAA3FFF82863CE3BF927FB5EF939" box="[1057,1172,1714,1736]" italics="true" pageId="1" pageNumber="26">A. humilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D32FFF82861CED0F927FE4FFD2C" authority="Salm-Dyck" authorityName="Salm-Dyck" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" lastPageId="3" lastPageNumber="28" order="Asparagales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="species" species="microstigma">
<emphasis id="B968EAA3FFF82861CED0F927FF35FD2C" italics="true" lastPageId="3" lastPageNumber="28" pageId="1" pageNumber="26">A. microstigma</emphasis>
Salm-Dyck
</taxonomicName>
from near Oudtshoorn in the Klein Karoo (
<bibRefCitation id="EF8D4B40FFFA2861CB4AFD42FDE7FD13" author="REYNOLDS, G. W." box="[336,557,727,749]" firstAuthor="Reynolds" journalOrPublisher="Johannesburg" page="179" pageId="3" pageNumber="28" refId="ref3117" refString="REYNOLDS, G. W. (1950). The aloes of South Africa. The Aloes of South Africa Book Fund, Johannesburg." title="The aloes of South Africa. The Aloes of South Africa Book Fund" type="book" year="1950">Reynolds, 1950: 179</bibRefCitation>
).
</paragraph>
<caption id="DF636639FFFB2860CA8BFC63FAC4FBD6" ID-DOI="http://doi.org/10.5281/zenodo.7867788" ID-Zenodo-Dep="7867788" httpUri="https://zenodo.org/record/7867788/files/figure.png" pageId="2" pageNumber="27" startId="2.[145,229,1014,1036]" targetBox="[145,1294,145,1007]" targetPageId="2" targetType="figure">
<paragraph id="8BA336B1FFFB2860CA8BFC63FAC4FBD6" blockId="2.[145,1294,1014,1064]" pageId="2" pageNumber="27">
<emphasis id="B968EAA3FFFB2860CA8BFC63FECAFBF5" bold="true" box="[145,256,1014,1036]" pageId="2" pageNumber="27">Figure 1.</emphasis>
<taxonomicName id="4C1C4D32FFFB2860CB1CFC63FE12FBF2" box="[262,472,1014,1036]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFFB2860CB1CFC63FEFCFBF5" box="[262,310,1014,1035]" italics="true" pageId="2" pageNumber="27">Aloe</emphasis>
×
<emphasis id="B968EAA3FFFB2860CB50FC62FE12FBF2" box="[330,472,1015,1036]" italics="true" pageId="2" pageNumber="27">spinosissima</emphasis>
</taxonomicName>
in full flower in the Glasshouse at the Royal Horticultural Society Garden Wisley, Woking, Surrey, United Kingdom. Picture taken on 20th February 2008 by Susan Carter.
</paragraph>
</caption>
<caption id="DF636639FFFB2860CA8BFBD2FB5FFBA3" ID-Table-UUID="DF636639FFFB2860CA8BFBD2FB5FFBA3" box="[145,1173,1095,1117]" httpUri="http://table.plazi.org/id/DF636639FFFB2860CA8BFBD2FB5FFBA3" pageId="2" pageNumber="27" startId="2.[145,215,1095,1116]" targetBox="[155,1222,1151,1638]" targetIsTable="true" targetPageId="2" targetType="table">
<paragraph id="8BA336B1FFFB2860CA8BFBD2FB5FFBA3" blockId="2.[145,1173,1095,1117]" box="[145,1173,1095,1117]" pageId="2" pageNumber="27">
<emphasis id="B968EAA3FFFB2860CA8BFBD2FF38FBA2" bold="true" box="[145,242,1095,1116]" pageId="2" pageNumber="27">Table 1.</emphasis>
Chronology of the interpretation of the name
<taxonomicName id="4C1C4D32FFFB2860C8EAFBD2FBCFFBA3" authority="Hort." authorityName="Hort." box="[752,1029,1095,1117]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFFB2860C8EAFBD2FCEAFBA2" box="[752,800,1095,1116]" italics="true" pageId="2" pageNumber="27">Aloe</emphasis>
×
<emphasis id="B968EAA3FFFB2860C92FFBDDFC09FBA3" box="[821,963,1096,1117]" italics="true" pageId="2" pageNumber="27">spinosissima</emphasis>
Hort.
</taxonomicName>
ex A.Berger.
</paragraph>
</caption>
<paragraph id="8BA336B1FFFB2860CAD6FBEAFCCCF99B" pageId="2" pageNumber="27">
<table id="F91CC411FFFBD79DCA81FBEAFB0CF998" box="[155,1222,1151,1638]" gridcols="2" gridrows="7" pageId="2" pageNumber="27">
<tr id="352C34F3FFFBD79DCA81FBEAFB0CFB6B" box="[155,1222,1151,1173]" gridrow="0" pageId="2" pageNumber="27">
<th id="76FD5D8FFFFBD79DCA81FBEAFD99FB6B" box="[155,595,1151,1173]" gridcol="0" gridrow="0" pageId="2" pageNumber="27">
<emphasis id="B968EAA3FFFB2860CAD6FBEAFDECFB6B" bold="true" box="[204,550,1151,1173]" pageId="2" pageNumber="27">Author (date: page number)</emphasis>
</th>
<th id="76FD5D8FFFFBD79DC872FBEAFB0CFB6B" box="[616,1222,1151,1173]" gridcol="1" gridrow="0" pageId="2" pageNumber="27">
<emphasis id="B968EAA3FFFB2860C92FFBEAFBFCFB6B" bold="true" box="[821,1078,1151,1173]" pageId="2" pageNumber="27">Suggested taxonomy</emphasis>
</th>
</tr>
<tr id="352C34F3FFFBD79DCA81FB25FB0CFB01" box="[155,1222,1200,1279]" gridrow="1" pageId="2" pageNumber="27">
<td id="76FD5D8FFFFBD79DCA81FB25FD99FB01" box="[155,595,1200,1279]" gridcol="0" gridrow="1" pageId="2" pageNumber="27">
<bibRefCitation id="EF8D4B40FFFB2860CA81FB58FE99FB1C" author="BERGER, A." box="[155,339,1229,1250]" firstAuthor="Berger" journalOrPublisher="Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft" page="19" pageId="2" pageNumber="27" pagination="159 - 326" part="33" refId="ref2409" refString="BERGER, A. (1908). 8. Aloe L. In: A. ENGLER (ED.), Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft 33: 159 - 326. Verlag von Wilhelm Engelmann, Leipzig." title="Aloe L" type="journal article" volumeTitle="A. ENGLER (ED.)" year="1908">Berger (1908: 19</bibRefCitation>
, 183, 184)*
</td>
<td id="76FD5D8FFFFBD79DC872FB25FB0CFB01" box="[616,1222,1200,1279]" gridcol="1" gridrow="1" pageId="2" pageNumber="27">
A hybrid between
<taxonomicName id="4C1C4D32FFFB2860C928FB25FB9CFB3B" box="[818,1110,1200,1222]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="variety" species="humilis" variety="echinata">
<emphasis id="B968EAA3FFFB2860C928FB25FC77FB3B" box="[818,957,1200,1221]" italics="true" pageId="2" pageNumber="27">Aloe humilis</emphasis>
var.
<emphasis id="B968EAA3FFFB2860C9EDFB25FB9CFB3B" box="[1015,1110,1200,1221]" italics="true" pageId="2" pageNumber="27">echinata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D32FFFB2860C872FB58FC7BFB1C" box="[616,945,1229,1250]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="variety" species="arborescens" variety="natalensis">
<emphasis id="B968EAA3FFFB2860C872FB58FCCCFB1C" box="[616,774,1229,1250]" italics="true" pageId="2" pageNumber="27">A. arborescens</emphasis>
var.
<emphasis id="B968EAA3FFFB2860C95AFB58FC7BFB1C" box="[832,945,1229,1250]" italics="true" pageId="2" pageNumber="27">natalensis</emphasis>
</taxonomicName>
(p. 19)
<taxonomicName id="4C1C4D32FFFB2860CE1EFB58FD0DFB00" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="variety" species="humilis" variety="echinata">
<emphasis id="B968EAA3FFFB2860CE1EFB58FB5AFB1C" box="[1028,1168,1229,1250]" italics="true" pageId="2" pageNumber="27">Aloe humilis</emphasis>
var.
<emphasis id="B968EAA3FFFB2860C872FB7CFD0DFB00" box="[616,711,1257,1278]" italics="true" pageId="2" pageNumber="27">echinata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D32FFFB2860C8E4FB7CFB96FB00" box="[766,1116,1257,1278]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="variety" species="arborescens" variety="pachythyrsa">
<emphasis id="B968EAA3FFFB2860C8E4FB7CFC56FB00" box="[766,924,1257,1278]" italics="true" pageId="2" pageNumber="27">A. arborescens</emphasis>
var.
<emphasis id="B968EAA3FFFB2860C9CFFB7CFB96FB00" box="[981,1116,1257,1278]" italics="true" pageId="2" pageNumber="27">pachythyrsa</emphasis>
</taxonomicName>
(p. 183)
</td>
</tr>
<tr id="352C34F3FFFBD79DCA81FA82FB0CFAB6" box="[155,1222,1303,1352]" gridrow="2" pageId="2" pageNumber="27">
<td id="76FD5D8FFFFBD79DCA81FA82FD99FAB6" box="[155,595,1303,1352]" gridcol="0" gridrow="2" pageId="2" pageNumber="27">
<bibRefCitation id="EF8D4B40FFFB2860CA81FAB0FEBEFAC4" author="Jacobsen" box="[155,372,1317,1339]" firstAuthor="Jacobsen" page="97" pageId="2" pageNumber="27" refId="ref2873" refString="JACOBSEN, H. (1977). Lexicon of succulent plants. Short descriptions, habitats and synonymy of succulent plants other than Cactaceae. 2 nd edn. Blandford Press Ltd, Poole, Dorset." type="book" year="1977">Jacobsen (1977: 97)</bibRefCitation>
**
</td>
<td id="76FD5D8FFFFBD79DC872FA82FB0CFAB6" box="[616,1222,1303,1352]" gridcol="1" gridrow="2" pageId="2" pageNumber="27">
A hybrid between
<taxonomicName id="4C1C4D32FFFB2860C928FA82FC77FAD2" box="[818,957,1303,1324]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="species" species="humilis">
<emphasis id="B968EAA3FFFB2860C928FA82FC77FAD2" box="[818,957,1303,1324]" italics="true" pageId="2" pageNumber="27">Aloe humilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D32FFFB2860C9EFFA82FD25FAB6" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="variety" species="arborescens" variety="pachythyrsa">
<emphasis id="B968EAA3FFFB2860C9EFFA82FB59FAD2" box="[1013,1171,1303,1325]" italics="true" pageId="2" pageNumber="27">A. arborescens</emphasis>
var.
<emphasis id="B968EAA3FFFB2860C872FAA6FD25FAB6" box="[616,751,1331,1352]" italics="true" pageId="2" pageNumber="27">pachythyrsa</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="352C34F3FFFBD79DCA81FAF1FB0CFA6B" box="[155,1222,1380,1429]" gridrow="3" pageId="2" pageNumber="27">
<td id="76FD5D8FFFFBD79DCA81FAF1FD99FA6B" box="[155,595,1380,1429]" gridcol="0" gridrow="3" pageId="2" pageNumber="27">
<bibRefCitation id="EF8D4B40FFFB2860CA81FAE7FE4BFA79" author="JACOBSEN, H." box="[155,385,1394,1416]" firstAuthor="Jacobsen" journalOrPublisher="Blandford Press Ltd, Poole, Dorset" page="194" pageId="2" pageNumber="27" refId="ref2911" refString="JACOBSEN, H. (1986). A handbook of succulent plants. Volume 1. Abromeitiella to Euphorbia. Blandford Press Ltd, Poole, Dorset." title="A handbook of succulent plants. Volume 1. Abromeitiella to Euphorbia" type="book" year="1986">Jacobsen (1986: 194)</bibRefCitation>
***
</td>
<td id="76FD5D8FFFFBD79DC872FAF1FB0CFA6B" box="[616,1222,1380,1429]" gridcol="1" gridrow="3" pageId="2" pageNumber="27">
A hybrid between
<taxonomicName id="4C1C4D32FFFB2860C928FAF1FB9CFA87" box="[818,1110,1380,1401]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="variety" species="humilis" variety="echinata">
<emphasis id="B968EAA3FFFB2860C928FAF1FB9CFA87" box="[818,1110,1380,1401]" italics="true" pageId="2" pageNumber="27">
<emphasis id="B968EAA3FFFB2860C928FAF1FC77FA87" box="[818,957,1380,1401]" italics="true" pageId="2" pageNumber="27">Aloe humilis</emphasis>
var.
<emphasis id="B968EAA3FFFB2860C9EDFAF1FB9CFA87" box="[1015,1110,1380,1401]" italics="true" pageId="2" pageNumber="27">echinata</emphasis>
</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D32FFFB2860C872FA15FC0DFA6B" box="[616,967,1408,1429]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="variety" species="arborescens" variety="pachythyrsa">
<emphasis id="B968EAA3FFFB2860C872FA15FC0DFA6B" box="[616,967,1408,1429]" italics="true" pageId="2" pageNumber="27">
<emphasis id="B968EAA3FFFB2860C872FA15FCCCFA6B" box="[616,774,1408,1429]" italics="true" pageId="2" pageNumber="27">A. arborescens</emphasis>
var.
<emphasis id="B968EAA3FFFB2860C95AFA15FC0DFA6B" box="[832,967,1408,1429]" italics="true" pageId="2" pageNumber="27">pachythyrsa</emphasis>
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="352C34F3FFFBD79DCA81FA26FB0CFA37" box="[155,1222,1459,1481]" gridrow="4" pageId="2" pageNumber="27">
<td id="76FD5D8FFFFBD79DCA81FA26FD99FA37" box="[155,595,1459,1481]" gridcol="0" gridrow="4" pageId="2" pageNumber="27">
<bibRefCitation id="EF8D4B40FFFB2860CA81FA21FEB9FA37" author="NEWTON, L. E." box="[155,371,1460,1481]" firstAuthor="Newton" journalOrPublisher="Springer-Verlag, Berlin" page="104" pageId="2" pageNumber="27" pagination="103 - 186" refId="ref3077" refString="NEWTON, L. E. (2001). Aloe. In: U. EGGLI (ED.), Illustrated handbook of succulent plants: Monocotyledons, pp. 103 - 186. Springer-Verlag, Berlin." title="Aloe" type="book chapter" volumeTitle="U. EGGLI (ED.), Illustrated handbook of succulent plants: Monocotyledons" year="2001">Newton (2001: 104)</bibRefCitation>
</td>
<td id="76FD5D8FFFFBD79DC872FA26FB0CFA37" box="[616,1222,1459,1481]" gridcol="1" gridrow="4" pageId="2" pageNumber="27">An intrageneric hybrid</td>
</tr>
<tr id="352C34F3FFFBD79DCA81FA72FB0CF9E7" box="[155,1222,1511,1561]" gridrow="5" pageId="2" pageNumber="27">
<td id="76FD5D8FFFFBD79DCA81FA72FD99F9E7" box="[155,595,1511,1561]" gridcol="0" gridrow="5" pageId="2" pageNumber="27">
<bibRefCitation id="EF8D4B40FFFB2860CA81FA60FE51F9F4" author="GRACE, O. M. &amp; KLOPPER, R. R. &amp; FIGUEIREDO, E. &amp; SMITH, G. F." box="[155,411,1525,1547]" etAl="et al." firstAuthor="Grace" journalOrPublisher="Strelitzia 29. South African National Biodiversity Institute, Pretoria, and Royal Botanic Gardens, Kew." page="145" pageId="2" pageNumber="27" refId="ref2618" refString="GRACE, O. M., KLOPPER, R. R., FIGUEIREDO, E. &amp; SMITH, G. F. (2011). The aloe names book. Strelitzia 29. South African National Biodiversity Institute, Pretoria, and Royal Botanic Gardens, Kew." title="The aloe names book" type="book" year="2011">
Grace
<emphasis id="B968EAA3FFFB2860CAF9FA63FED0F9F5" box="[227,282,1525,1547]" italics="true" pageId="2" pageNumber="27">et al.</emphasis>
(2011: 145)
</bibRefCitation>
</td>
<td id="76FD5D8FFFFBD79DC872FA72FB0CF9E7" box="[616,1222,1511,1561]" gridcol="1" gridrow="5" pageId="2" pageNumber="27">
A hybrid between
<taxonomicName id="4C1C4D32FFFB2860C928FA72FB9CFA02" box="[818,1110,1511,1532]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="variety" species="humilis" variety="echinata">
<emphasis id="B968EAA3FFFB2860C928FA72FC77FA02" box="[818,957,1511,1532]" italics="true" pageId="2" pageNumber="27">Aloe humilis</emphasis>
var.
<emphasis id="B968EAA3FFFB2860C9EDFA72FB9CFA02" box="[1015,1110,1511,1532]" italics="true" pageId="2" pageNumber="27">echinata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D32FFFB2860C872F996FCCCF9E6" box="[616,774,1539,1561]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="B968EAA3FFFB2860C872F996FCCCF9E6" box="[616,774,1539,1561]" italics="true" pageId="2" pageNumber="27">A. arborescens</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="352C34F3FFFBD79DCA81F9A6FB0CF998" box="[155,1222,1587,1638]" gridrow="6" pageId="2" pageNumber="27">
<td id="76FD5D8FFFFBD79DCA81F9A6FD99F998" box="[155,595,1587,1638]" gridcol="0" gridrow="6" pageId="2" pageNumber="27">
<bibRefCitation id="EF8D4B40FFFB2860CA81F9D7FE02F9A9" author="SMITH, G. F. &amp; FIGUEIREDO, E." box="[155,456,1602,1624]" firstAuthor="Smith" journalOrPublisher="Johannesburg" pageId="2" pageNumber="27" refId="ref3354" refString="SMITH, G. F. &amp; FIGUEIREDO, E. (2015 a). Garden aloes. Growing and breeding cultivars and hybrids. Jacana Media (Pty) Ltd, Auckland Park, Johannesburg." title="Garden aloes. Groaeing and breeding cultioars and hybrids. Jacana Media (Pty) Ltd, Auckland Park" type="book" year="2015" yearSuffix="a">Smith &amp; Figueiredo (2015a</bibRefCitation>
: xii, xiii, 51)
</td>
<td id="76FD5D8FFFFBD79DC872F9A6FB0CF998" box="[616,1222,1587,1638]" gridcol="1" gridrow="6" pageId="2" pageNumber="27">
A hybrid between
<taxonomicName id="4C1C4D32FFFB2860C928F9A1FC77F9B7" box="[818,957,1588,1609]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="species" species="humilis">
<emphasis id="B968EAA3FFFB2860C928F9A1FC77F9B7" box="[818,957,1588,1609]" italics="true" pageId="2" pageNumber="27">Aloe humilis</emphasis>
</taxonomicName>
[
<taxonomicName id="4C1C4D32FFFB2860C9D6F9A1FB94F9B7" box="[972,1118,1588,1609]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="variety" species="humilis" variety="echinata">
var.
<emphasis id="B968EAA3FFFB2860C9E5F9A1FB94F9B7" box="[1023,1118,1588,1609]" italics="true" pageId="2" pageNumber="27">echinata</emphasis>
</taxonomicName>
] and
<taxonomicName id="4C1C4D32FFFB2860C872F9C5FCCCF99B" box="[616,774,1616,1638]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="species" species="arborescens">A. arborescens</taxonomicName>
</td>
</tr>
</table>
</paragraph>
<footnote id="E8072ABFFFFB2860CA8BF9EBFE12F96D" box="[145,472,1662,1684]" pageId="2" pageNumber="27">
<paragraph id="8BA336B1FFFB2860CA8BF9EBFE12F96D" blockId="2.[145,549,1662,1740]" box="[145,472,1662,1684]" pageId="2" pageNumber="27">
*As “
<taxonomicName id="4C1C4D32FFFB2860CAD3F9EBFE05F96D" authority="Hort." authorityName="Hort." box="[201,463,1662,1684]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFFB2860CAD3F9EBFE47F96A" box="[201,397,1662,1684]" italics="true" pageId="2" pageNumber="27">Aloe spinosissima</emphasis>
Hort.
</taxonomicName>
</paragraph>
</footnote>
<footnote id="E8072ABFFFFB2860CA8BF90FFE7FF951" box="[145,437,1690,1712]" pageId="2" pageNumber="27">
<paragraph id="8BA336B1FFFB2860CA8BF90FFE7FF951" blockId="2.[145,549,1662,1740]" box="[145,437,1690,1712]" pageId="2" pageNumber="27">
**As “
<taxonomicName id="4C1C4D32FFFB2860CACFF90FFE66F94E" authority=" " authorityName=" " box="[213,428,1690,1712]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFFB2860CACFF90FFF24F951" bold="true" box="[213,238,1690,1711]" pageId="2" pageNumber="27">A.</emphasis>
cv. Spinosissima
</taxonomicName>
</paragraph>
</footnote>
<footnote id="E8072ABFFFFB2860CA8BF923FDEFF935" box="[145,549,1718,1740]" pageId="2" pageNumber="27">
<paragraph id="8BA336B1FFFB2860CA8BF923FDEFF935" blockId="2.[145,549,1662,1740]" box="[145,549,1718,1740]" pageId="2" pageNumber="27">
***As “
<taxonomicName id="4C1C4D32FFFB2860CAFBF923FDD7F935" authority="HORT." authorityName="HORT." box="[225,541,1718,1740]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="species" species="x spinosissima">
<emphasis id="B968EAA3FFFB2860CAFBF923FED2F935" bold="true" box="[225,280,1718,1739]" pageId="2" pageNumber="27">Aloe</emphasis>
×
<emphasis id="B968EAA3FFFB2860CB2FF923FE1FF932" bold="true" box="[309,469,1718,1740]" pageId="2" pageNumber="27">spinosissima</emphasis>
<smallCapsWord id="8D45A06DFFFB2860CBC1F923FDDCF935" baselines="1734,1735" box="[475,534,1718,1739]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Hort" pageId="2" pageNumber="27">HORT</smallCapsWord>
.
</taxonomicName>
</paragraph>
</footnote>
<caption id="DF636639FFFA2861CA8BFDB7FD7BFD56" ID-DOI="http://doi.org/10.5281/zenodo.7867790" ID-Zenodo-Dep="7867790" httpUri="https://zenodo.org/record/7867790/files/figure.png" pageId="3" pageNumber="28" startId="3.[145,230,546,568]" targetBox="[145,689,145,533]" targetPageId="3" targetType="figure">
<paragraph id="8BA336B1FFFA2861CA8BFDB7FDF4FD72" blockId="3.[145,689,546,680]" pageId="3" pageNumber="28">
<emphasis id="B968EAA3FFFA2861CA8BFDB7FECFFDC9" bold="true" box="[145,261,546,568]" pageId="3" pageNumber="28">Figure 2.</emphasis>
<taxonomicName id="4C1C4D32FFFA2861CB14FDB7FE22FDC6" box="[270,488,546,568]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFFA2861CB14FDB7FEF4FDC9" box="[270,318,546,567]" italics="true" pageId="3" pageNumber="28">Aloe</emphasis>
×
<emphasis id="B968EAA3FFFA2861CB4DFDB6FE22FDC6" box="[343,488,547,568]" italics="true" pageId="3" pageNumber="28">spinosissima</emphasis>
</taxonomicName>
cultivated in the succulent garden of the Botanical Garden of the University of Porto in northwestern coastal Portugal (see
<bibRefCitation id="EF8D4B40FFFA2861CB19FDE3FDE5FD72" author="SMITH, G. F. &amp; FIGUEIREDO, E." box="[259,559,630,652]" firstAuthor="Smith" journalOrPublisher="Cactus &amp; Succulent Journal (U. S.)" pageId="3" pageNumber="28" pagination="99 - 105" part="86" refId="ref3307" refString="SMITH, G. F. &amp; FIGUEIREDO, E. (2014 b). The succulent garden of the Botanical Garden of the University of Porto, Portugal. Cactus &amp; Succulent Journal (U. S.) 86: 99 - 105." title="The succulent garden of the Botanical Garden of the University of Porto, Portugal" type="journal article" year="2014" yearSuffix="b">Smith &amp; Figueiredo, 2014b</bibRefCitation>
).
</paragraph>
<paragraph id="8BA336B1FFFA2861CB7FFD07FD7BFD56" blockId="3.[145,689,546,680]" box="[357,689,658,680]" pageId="3" pageNumber="28">Photograph: Gideon F. Smith.</paragraph>
</caption>
<paragraph id="8BA336B1FFFA2861CAADFD66FD88FB77" blockId="3.[145,690,700,1354]" pageId="3" pageNumber="28">
In the case of
<taxonomicName id="4C1C4D32FFFA2861CB56FD66FDCBFCF6" box="[332,513,755,776]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="B968EAA3FFFA2861CB56FD66FDCBFCF6" box="[332,513,755,776]" italics="true" pageId="3" pageNumber="28">Aloe arborescens</emphasis>
</taxonomicName>
, at least 15 natural hybrids are known (
<bibRefCitation id="EF8D4B40FFFA2861CBA3FC9BFD68FCDA" author="REYNOLDS, G. W." box="[441,674,782,804]" firstAuthor="Reynolds" journalOrPublisher="Johannesburg" page="413" pageId="3" pageNumber="28" refId="ref3117" refString="REYNOLDS, G. W. (1950). The aloes of South Africa. The Aloes of South Africa Book Fund, Johannesburg." title="The aloes of South Africa. The Aloes of South Africa Book Fund" type="book" year="1950">Reynolds, 1950: 413</bibRefCitation>
). However, a natural hybrid between
<taxonomicName id="4C1C4D32FFFA2861C80CFCBFFD7BFCC1" box="[534,689,810,832]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="B968EAA3FFFA2861C80CFCBFFD7BFCC1" box="[534,689,810,832]" italics="true" pageId="3" pageNumber="28">A. arborescens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D32FFFA2861CAD2FCD3FE8EFCA4" box="[200,324,837,859]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="humilis">
<emphasis id="B968EAA3FFFA2861CAD2FCD3FE8EFCA4" box="[200,324,837,859]" italics="true" pageId="3" pageNumber="28">A. humilis</emphasis>
</taxonomicName>
has not yet been found, even though their distribution ranges overlap in the Eastern Cape (see distribution maps in
<bibRefCitation id="EF8D4B40FFFA2861C850FCE8FE92FC50" author="VAN WYK, B-E. &amp; SMITH, G. F." firstAuthor="Van Wyk" journalOrPublisher="3 rd edition. Briza Publications, Queenswood, Pretoria." page="86" pageId="3" pageNumber="28" refId="ref3690" refString="VAN WYK, B-E. [BEN-ERIK] &amp; SMITH, G. F. (2014). Guide to the aloes of South Africa. 3 rd edition. Briza Publications, Queenswood, Pretoria." title="Guide to the aloes of South Africa" type="book" year="2014">Van Wyk &amp; Smith, 2014: 86</bibRefCitation>
and 290, respectively).
<taxonomicName id="4C1C4D32FFFA2861C841FC0DFF31FC36" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="B968EAA3FFFA2861C841FC0DFF31FC36" italics="true" pageId="3" pageNumber="28">Aloe arborescens</emphasis>
</taxonomicName>
is known to easily cross with other species of
<taxonomicName id="4C1C4D32FFFA2861CAE7FC5AFEE6FC1A" box="[253,300,975,996]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="B968EAA3FFFA2861CAE7FC5AFEE6FC1A" box="[253,300,975,996]" italics="true" pageId="3" pageNumber="28">Aloe</emphasis>
</taxonomicName>
(and species of other Alooideae genera, for that matter) that have morphologies quite distinct from its robust, shrubby growth form (
<emphasis id="B968EAA3FFFA2861C882FB93FD61FBE5" box="[664,683,1030,1051]" italics="true" pageId="3" pageNumber="28">cf</emphasis>
.
<emphasis id="B968EAA3FFFA2861CA8BFBB4FF63FBC9" box="[145,169,1057,1079]" italics="true" pageId="3" pageNumber="28">A.</emphasis>
×
<emphasis id="B968EAA3FFFA2861CAD0FBB7FEFDFBC9" box="[202,311,1058,1079]" italics="true" pageId="3" pageNumber="28">inopinata</emphasis>
Gideon F.Sm.,
<taxonomicNameLabel id="A25B57D8FFFA2861C81DFBB7FDF0FBC9" box="[519,570,1058,1079]" pageId="3" pageNumber="28" rank="subSpecies">N.R.</taxonomicNameLabel>
Crouch &amp; Oosth., the natural hybrid between
<taxonomicName id="4C1C4D32FFFA2861C80FFBA8FD7BFBAC" box="[533,689,1085,1106]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="B968EAA3FFFA2861C80FFBA8FD7BFBAC" box="[533,689,1085,1106]" italics="true" pageId="3" pageNumber="28">A. arborescens</emphasis>
</taxonomicName>
and the slender aloe,
<taxonomicName id="4C1C4D32FFFA2861CB9CFBCDFEA8FB77" authority="A. Berger var. chortolirioides" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="variety" species="chortolirioides" variety="chortolirioides">
<emphasis id="B968EAA3FFFA2861CB9CFBCDFD8FFB93" box="[390,581,1112,1134]" italics="true" pageId="3" pageNumber="28">A. chortolirioides</emphasis>
A.Berger var.
<emphasis id="B968EAA3FFFA2861CAD9FBE1FEA8FB77" box="[195,354,1140,1161]" italics="true" pageId="3" pageNumber="28">chortolirioides</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF8D4B40FFFA2861CB6AFBE1FDF9FB74" author="SMITH, G. F. &amp; FIGUEIREDO, E. &amp; CROUCH, N. R. &amp; OOSTHUIZEN, D. &amp; KLOPPER, R. R." box="[368,563,1140,1162]" etAl="et al." firstAuthor="Smith" journalOrPublisher="Haseltonia" pageId="3" pageNumber="28" pagination="55 - 63" part="22" refId="ref3427" refString="SMITH, G. F., FIGUEIREDO, E., CROUCH, N. R., OOSTHUIZEN, D. &amp; KLOPPER, R. R. (2016). Aloe × inopinata Gideon F. SM., N. R. Crouch &amp; Oosth., (Asphodelaceae) [Aloe arborescens Mill. × Aloe chortolirioides A. Berger var. chortolirioides]: a nothospecies from the Barberton Centre of Endemism, Eastern South Africa. Haseltonia 22: 55 - 63." title="Aloe × inopinata Gideon F. SM., N. R. Crouch &amp; Oosth., (Asphodelaceae) [Aloe arborescens Mill. × Aloe chortolirioides A. Berger var. chortolirioides]: a nothospecies from the Barberton Centre of Endemism, Eastern South Africa" type="journal article" year="2016">
Smith
<emphasis id="B968EAA3FFFA2861CBA0FBE1FE20FB77" box="[442,490,1140,1161]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
., 2016
</bibRefCitation>
).
</paragraph>
<caption id="DF636639FFFA2861C8F4FCD3FAC4FC51" ID-DOI="http://doi.org/10.5281/zenodo.7867792" ID-Zenodo-Dep="7867792" httpUri="https://zenodo.org/record/7867792/files/figure.png" pageId="3" pageNumber="28" startId="3.[750,835,838,860]" targetBox="[750,1294,145,829]" targetPageId="3" targetType="figure">
<paragraph id="8BA336B1FFFA2861C8F4FCD3FAC4FC51" blockId="3.[750,1294,838,944]" pageId="3" pageNumber="28">
<emphasis id="B968EAA3FFFA2861C8F4FCD3FCA3FCA5" bold="true" box="[750,873,838,860]" pageId="3" pageNumber="28">Figure 3.</emphasis>
<taxonomicName id="4C1C4D32FFFA2861C961FCD3FB96FCA2" box="[891,1116,838,860]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFFA2861C961FCD3FC61FCA5" box="[891,939,838,859]" italics="true" pageId="3" pageNumber="28">Aloe</emphasis>
×
<emphasis id="B968EAA3FFFA2861C9D1FCD2FB96FCA2" box="[971,1116,839,860]" italics="true" pageId="3" pageNumber="28">spinosissima</emphasis>
</taxonomicName>
growing in a pedestalled, Italianate container in the Hanbury Gardens at La Mortola on the Ligurian Riviera, Italy. Photograph: Gideon F. Smith.
</paragraph>
</caption>
<paragraph id="8BA336B1FFFA2861CAADFB1AFDB8FAB7" blockId="3.[145,690,700,1354]" pageId="3" pageNumber="28">
Among aloes it is quite common to find species with widely diverging growth forms to be interfertile, with, for example, the tree-like fan aloe [
<taxonomicName id="4C1C4D32FFFA2861CA83FB77FE92FB08" box="[153,344,1249,1271]" class="Liliopsida" family="Asphodelaceae" genus="Kumara" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="plicatilis">
<emphasis id="B968EAA3FFFA2861CA83FB77FE92FB08" box="[153,344,1249,1271]" italics="true" pageId="3" pageNumber="28">Kumara plicatilis</emphasis>
</taxonomicName>
(L.) G.D.Rowley] and low-growing, stemless partridge aloe [
<taxonomicName id="4C1C4D32FFFA2861CBC6FB68FD78FAED" box="[476,690,1277,1299]" class="Liliopsida" family="Asphodelaceae" genus="Gonialoe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="variegata">
<emphasis id="B968EAA3FFFA2861CBC6FB68FD78FAED" box="[476,690,1277,1299]" italics="true" pageId="3" pageNumber="28">Gonialoe variegata</emphasis>
</taxonomicName>
(L.) Boatwr. &amp; J.C.Manning] also having been successfully hybridised (
<bibRefCitation id="EF8D4B40FFFA2861CBBAFAA1FDA9FAB7" author="SMITH, G. F. &amp; FIGUEIREDO, E. &amp; MOLTENO, S." box="[416,611,1332,1354]" etAl="et al." firstAuthor="Smith" journalOrPublisher="Bradleya" pageId="3" pageNumber="28" pagination="53 - 60" part="36" refId="ref3518" refString="SMITH, G. F., FIGUEIREDO, E. &amp; MOLTENO, S. (2018). × Gonimara Gideon F. Sm. &amp; Molteno (Asphodelaceae): a new nothogenus name for the artificial hybrid, × Gonimara corderoyi (A. Berger) Gideon F. Sm. &amp; Molteno, between the alooid species Gonialoe variegata and Kumara plicatilis, two southern African endemics. Bradleya 36: 53 - 60." title="× Gonimara Gideon F. Sm. &amp; Molteno (Asphodelaceae): a new nothogenus name for the artificial hybrid, × Gonimara corderoyi (A. Berger) Gideon F. Sm. &amp; Molteno, between the alooid species Gonialoe oariegata and Kumara plicatilis, two southern African endemics." type="journal article" year="2018">
Smith
<emphasis id="B968EAA3FFFA2861CBF0FAA0FDD0FAB7" box="[490,538,1332,1354]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
., 2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C306653AFFFA2861CA8BFAFEFD99FA7E" box="[145,595,1386,1409]" pageId="3" pageNumber="28" type="nomenclature">
<paragraph id="8BA336B1FFFA2861CA8BFAFEFD99FA7E" blockId="3.[145,690,1386,1656]" box="[145,595,1386,1409]" pageId="3" pageNumber="28">
<emphasis id="B968EAA3FFFA2861CA8BFAFEFD99FA7E" bold="true" box="[145,595,1386,1409]" pageId="3" pageNumber="28">
Nomenclature of
<taxonomicName id="4C1C4D32FFFA2861CB72FAFFFD99FA7E" ID-CoL="8WQJ5" ID-ENA="1429993" authority="Hort." box="[360,595,1386,1408]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFFA2861CB72FAFFFE57FA7E" bold="true" box="[360,413,1386,1408]" italics="true" pageId="3" pageNumber="28">Aloe</emphasis>
×
<emphasis id="B968EAA3FFFA2861CBAEFAFEFD99FA7E" bold="true" box="[436,595,1387,1408]" italics="true" pageId="3" pageNumber="28">spinosissima</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C306653AFFFA2861CA8BFA13FE2BF9DB" pageId="3" pageNumber="28" type="reference_group">
<paragraph id="8BA336B1FFFA2861CA8BFA13FE2BF9DB" blockId="3.[145,690,1386,1656]" pageId="3" pageNumber="28">
<treatmentCitationGroup id="AB0C119FFFFA2861CA8BFA13FE2BF9DB" pageId="3" pageNumber="28">
<treatmentCitation id="0ABD10A0FFFA2861CA8BFA13FD40FA65" box="[145,650,1414,1436]" pageId="3" pageNumber="28">
<emphasis id="B968EAA3FFFA2861CA8BFA13FD40FA65" bold="true" box="[145,650,1414,1436]" pageId="3" pageNumber="28">
<taxonomicName id="4C1C4D32FFFA2861CA8BFA13FE16FA65" ID-CoL="8WQJ5" ID-ENA="1429993" authority="Hort." authorityName="Hort." box="[145,476,1414,1436]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFFA2861CA8BFA13FF0DFA62" bold="true" box="[145,199,1414,1436]" italics="true" pageId="3" pageNumber="28">Aloe</emphasis>
×
<emphasis id="B968EAA3FFFA2861CAF2FA13FE43FA65" bold="true" box="[232,393,1414,1435]" italics="true" pageId="3" pageNumber="28">spinosissima</emphasis>
Hort.
</taxonomicName>
ex A.Berger
</emphasis>
</treatmentCitation>
in
<emphasis id="B968EAA3FFFA2861CA8BFA37FDB7FA49" box="[145,637,1442,1463]" italics="true" pageId="3" pageNumber="28">Pflanzenr. (Engler) Liliac.-Asphodel.-Aloin</emphasis>
.
<emphasis id="B968EAA3FFFA2861C894FA37FD60FA49" bold="true" box="[654,682,1442,1463]" pageId="3" pageNumber="28">33</emphasis>
: 183 (1908).
<treatmentCitation id="0ABD10A0FFFA2861CB0EFA28FDC3FA2D" author="Hort." box="[276,521,1469,1491]" page="570" pageId="3" pageNumber="28" score="1.97023359760351E-6" volumeTitle="Jahandiez" year="1933">
<bibCitation id="5E363EDDFFFA2861CB0EFA28FDC3FA2D" box="[276,521,1469,1491]" page="570" pageId="3" pageNumber="28" score="1.97023359760351E-6" volumeTitle="Jahandiez" year="1933">
<bibRefCitation id="EF8D4B40FFFA2861CB0EFA28FDCAFA2C" author="JAHANDIEZ, E." box="[276,512,1469,1491]" firstAuthor="Jahandiez" journalOrPublisher="Revue Horticole; resume de tout ce qui parait d'interessant en jardinage [etc.]. (Paris)" pageId="3" pageNumber="28" pagination="570 - 571" part="23" publicationUrl="http://bibliotheque-numerique.hortalia.org/files/original/cc8a632c3d5cc42fea5cc7875f21e1a7.pdf]" refId="ref2940" refString="JAHANDIEZ, E. ([1932 -] 1933). Les plantes grasses autres que les cactees. Les aloes et genres voisins (suite). Revue Horticole; resume de tout ce qui parait d'interessant en jardinage [etc.]. (Paris). New series. 23 (105 of whole series): 570 - 571. [Aloe × spinosissima on p. 570 - 571. See: http: // bibliotheque-numerique. hortalia. org / files / original / cc 8 a 632 c 3 d 5 cc 42 fea 5 cc 7875 f 21 e 1 a 7. pdf]" title="Les plantes grasses autres que les cactees. Les aloes et genres voisins (suite)" type="url" year="1932">Jahandiez: 570 (1933</bibRefCitation>
)
</bibCitation>
</treatmentCitation>
;
<treatmentCitation id="0ABD10A0FFFA2861C80DFA28FF12FA10" author="Hort." page="194" pageId="3" pageNumber="28" score="1.97023359760351E-6" volumeTitle="Jacobsen" year="1977">
<bibCitation id="5E363EDDFFFA2861C80DFA28FF12FA10" page="194" pageId="3" pageNumber="28" score="1.97023359760351E-6" volumeTitle="Jacobsen" year="1977">Jacobsen: 194 (1977)</bibCitation>
</treatmentCitation>
;
<treatmentCitation id="0ABD10A0FFFA2861CAF3FA4CFE3CFA10" author="Hort." box="[233,502,1497,1519]" page="" pageId="3" pageNumber="28">
<bibRefCitation id="EF8D4B40FFFA2861CAF3FA4CFE3CFA10" author="GRAF, A. B." box="[233,502,1497,1519]" firstAuthor="Graf" journalOrPublisher="Roehrs Company Inc" pageId="3" pageNumber="28" part="10 th edition" refId="ref2670" refString="GRAF, A. B. (1980 [January]). Exotica series 3. Pictorial cyclopedia of exotic plants from tropical regions. [12, 000 illustrations, 204 plants in colour, guide to care of plants indoors, horticultural colour guide, plant geography]. 10 th edition. Roehrs Company Inc., E. Rutherford, N. J. 07073, USA." title="Pictorial cyclopedia of exotic plants from tropical regions" type="book" volumeTitle="Exotica series 3" year="1980">
<bibCitation id="5E363EDDFFFA2861CAF3FA4CFE3CFA10" box="[233,502,1497,1519]" pageId="3" pageNumber="28">Graf: 1048, 1530 (1980</bibCitation>
</bibRefCitation>
</treatmentCitation>
);
<treatmentCitation id="0ABD10A0FFFA2861C80BFA4CFF12F9F4" author="Hort." page="197" pageId="3" pageNumber="28" score="1.97023359760351E-6" volumeTitle="Jacobsen" year="1986">
<bibCitation id="5E363EDDFFFA2861C80BFA4CFF12F9F4" page="197" pageId="3" pageNumber="28" score="1.97023359760351E-6" volumeTitle="Jacobsen" year="1986">Jacobsen: 197 (1986)</bibCitation>
</treatmentCitation>
;
<treatmentCitation id="0ABD10A0FFFA2861CAE9FA61FDE2F9F4" box="[243,552,1524,1546]" pageId="3" pageNumber="28">
<bibRefCitation id="EF8D4B40FFFA2861CAE9FA61FDEAF9F7" author="GRACE, O. M. &amp; KLOPPER, R. R. &amp; FIGUEIREDO, E. &amp; SMITH, G. F." box="[243,544,1524,1546]" firstAuthor="Grace" journalOrPublisher="South African National Biodiversity Institute, Pretoria, and Royal Botanic Gardens, Kew." pageId="3" pageNumber="28" part="29" refId="ref2618" refString="GRACE, O. M., KLOPPER, R. R., FIGUEIREDO, E. &amp; SMITH, G. F. (2011). The aloe names book. Strelitzia 29. South African National Biodiversity Institute, Pretoria, and Royal Botanic Gardens, Kew." title="The aloe names book" type="book" volumeTitle="Strelitzia" year="2011">
Grace
<emphasis id="B968EAA3FFFA2861CB50FA60FE43F9F7" box="[330,393,1524,1546]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
. 145 (2011
</bibRefCitation>
)
</treatmentCitation>
;
<treatmentCitation id="0ABD10A0FFFA2861C85EFA61FE11F9DB" pageId="3" pageNumber="28">
<bibRefCitation id="EF8D4B40FFFA2861C85EFA61FE18F9DB" author="SMITH, G. F. &amp; FIGUEIREDO, E." firstAuthor="Smith" journalOrPublisher="Johannesburg" pageId="3" pageNumber="28" refId="ref3354" refString="SMITH, G. F. &amp; FIGUEIREDO, E. (2015 a). Garden aloes. Growing and breeding cultivars and hybrids. Jacana Media (Pty) Ltd, Auckland Park, Johannesburg." title="Garden aloes. Growing and breeding cultivars and hybrids. Jacana Media (Pty) Ltd, Auckland Park" type="book" year="2015">Smith &amp; Figueiredo: xii, xiii, 51 (2015a</bibRefCitation>
)
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C306653AFFFA2861CBFDF985FED4F986" pageId="3" pageNumber="28" type="materials_examined">
<paragraph id="8BA336B1FFFA2861CBFDF985FED4F986" blockId="3.[145,690,1386,1656]" pageId="3" pageNumber="28">
<materialsCitation id="3B743CECFFFA2861CBFDF985FD9EF9A2" ID-GBIF-Occurrence="4092166301" collectingDate="2018-08-24" collectionCode="PRU" collectorName="G. F. Smith &amp; E. Figueiredo" country="South Africa" location="Pretoria" pageId="3" pageNumber="28" specimenCount="66" typeStatus="neotype">
<emphasis id="B968EAA3FFFA2861CBFDF985FD87F9DB" bold="true" box="[487,589,1552,1573]" pageId="3" pageNumber="28">
<typeStatus id="54A78813FFFA2861CBFDF985FD87F9DB" box="[487,589,1552,1573]" pageId="3" pageNumber="28" type="neotype">Neotype</typeStatus>
</emphasis>
:
<collectingCountry id="F30B7621FFFA2861C843F985FF3BF9BE" name="South Africa" pageId="3" pageNumber="28" value="South Africa">SOUTH AFRICA</collectingCountry>
.
<location id="8EC3606AFFFA2861CB1DF9BEFEAEF9BF" LSID="urn:lsid:plazi:treatment:03B587A7FFF82866C8F4FCECFDCDF938:8EC3606AFFFA2861CB1DF9BEFEAEF9BF" box="[263,356,1579,1601]" country="South Africa" name="Pretoria" pageId="3" pageNumber="28">Pretoria</location>
. Ex hort.
<date id="FFA21071FFFA2861CBF1F9BEFD60F9BE" box="[491,682,1579,1600]" pageId="3" pageNumber="28" value="2018-08-24">
<collectingDate id="EFE6E999FFFA2861CBF1F9BEFD60F9BE" box="[491,682,1579,1600]" pageId="3" pageNumber="28" value="2018-08-24">24 August 2018</collectingDate>
</date>
.
<emphasis id="B968EAA3FFFA2861CA8BF9D2FDC8F9A5" box="[145,514,1606,1628]" italics="true" pageId="3" pageNumber="28">
<collectorName id="26E95367FFFA2861CA8BF9D2FE11F9A2" box="[145,475,1607,1628]" pageId="3" pageNumber="28">G.F. Smith &amp; E. Figueiredo</collectorName>
<specimenCount id="9D1AFD38FFFA2861CBFDF9D3FDC8F9A5" box="[487,514,1606,1627]" pageId="3" pageNumber="28" type="generic">66</specimenCount>
</emphasis>
(
<collectionCode id="ED0DAE74FFFA2861C80CF9D2FD81F9A2" box="[534,587,1607,1628]" collectionName="PRU" pageId="3" pageNumber="28">PRU</collectionCode>
)
</materialsCitation>
,
<emphasis id="B968EAA3FFFA2861C87CF9D3FEDDF989" bold="true" pageId="3" pageNumber="28">designated here</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C306653AFFFA2861CA8BF90CFD5FF934" pageId="3" pageNumber="28" type="reference_group">
<paragraph id="8BA336B1FFFA2861CA8BF90CFF25F950" blockId="3.[145,661,1689,1738]" box="[145,239,1689,1710]" pageId="3" pageNumber="28">Parents:</paragraph>
<paragraph id="8BA336B1FFFA2861CA8BF921FD5FF934" blockId="3.[145,661,1689,1738]" box="[145,661,1716,1738]" pageId="3" pageNumber="28">
<taxonomicName id="4C1C4D32FFFA2861CA8BF921FE48F934" ID-CoL="C3DN" authority="Mill." authorityName="Mill." box="[145,386,1716,1738]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="B968EAA3FFFA2861CA8BF921FE8DF937" box="[145,327,1716,1737]" italics="true" pageId="3" pageNumber="28">Aloe arborescens</emphasis>
Mill.
</taxonomicName>
and
<taxonomicName id="4C1C4D32FFFA2861CBA3F920FD5FF934" ID-CoL="C3PY" authority="(L.) Mill." authorityName="Mill." baseAuthorityName="L." box="[441,661,1716,1738]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="humilis">
<emphasis id="B968EAA3FFFA2861CBA3F920FDE7F937" box="[441,557,1716,1738]" italics="true" pageId="3" pageNumber="28">A. humilis</emphasis>
(L.) Mill.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C306653AFFFA2861C8F4FC5DFAC0FA87" pageId="3" pageNumber="28" type="discussion">
<paragraph id="8BA336B1FFFA2861C8F4FC5DFC93FB0E" blockId="3.[750,1294,968,1402]" pageId="3" pageNumber="28">
Nomenclatural notes. Herbarium Mortolense (HMGBH) has specimens of
<taxonomicName id="4C1C4D32FFFA2861CE2BFC76FACDFC07" box="[1073,1287,995,1017]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFFA2861CE2BFC76FBABFC06" box="[1073,1121,995,1016]" italics="true" pageId="3" pageNumber="28">Aloe</emphasis>
×
<emphasis id="B968EAA3FFFA2861CE62FC71FACDFC07" box="[1144,1287,996,1017]" italics="true" pageId="3" pageNumber="28">spinosissima</emphasis>
</taxonomicName>
, but I was informed that these were not prepared during the time of Alwin Bergers employment at the Hanbury Gardens (Alessandro Guiggi, personal communication). The specimens of
<taxonomicName id="4C1C4D32FFFA2861C8F4FBF8FC62FB7C" box="[750,936,1133,1154]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFFA2861C8F4FBF8FD35FB7C" box="[750,767,1133,1154]" italics="true" pageId="3" pageNumber="28">A</emphasis>
. ×
<emphasis id="B968EAA3FFFA2861C900FBF8FC62FB7C" box="[794,936,1133,1154]" italics="true" pageId="3" pageNumber="28">spinosissima</emphasis>
</taxonomicName>
held at (HMGBH) are not available for examination online. A
<typeStatus id="54A78813FFFA2861CEB5FB1DFACDFB63" box="[1199,1287,1160,1181]" pageId="3" pageNumber="28" type="neotype">neotype</typeStatus>
,
<emphasis id="B968EAA3FFFA2861C8F4FB31FB8EFB46" box="[750,1092,1187,1209]" italics="true" pageId="3" pageNumber="28">G.F. Smith &amp; E. Figueiredo 66</emphasis>
, consisting of material cultivated in South Africa was therefore prepared.
</paragraph>
<paragraph id="8BA336B1FFFA2861C909FB63FAC0FA87" blockId="3.[750,1294,968,1402]" pageId="3" pageNumber="28">
Although the name
<taxonomicName id="4C1C4D32FFFA2861C9F0FB63FB73FAF2" box="[1002,1209,1270,1292]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFFA2861C9F0FB63FBD3FAF5" box="[1002,1049,1270,1291]" italics="true" pageId="3" pageNumber="28">Aloe</emphasis>
×
<emphasis id="B968EAA3FFFA2861CE37FB62FB73FAF2" box="[1069,1209,1271,1292]" italics="true" pageId="3" pageNumber="28">spinosissima</emphasis>
</taxonomicName>
is often noted as having been published by
<bibRefCitation id="EF8D4B40FFFA2861CE6FFA84FB2DFAD9" author="JAHANDIEZ, E." box="[1141,1255,1297,1319]" firstAuthor="Jahandiez" journalOrPublisher="Revue Horticole; resume de tout ce qui parait d'interessant en jardinage [etc.]. (Paris)" pageId="3" pageNumber="28" pagination="570 - 571" part="23" publicationUrl="http://bibliotheque-numerique.hortalia.org/files/original/cc8a632c3d5cc42fea5cc7875f21e1a7.pdf]" refId="ref2940" refString="JAHANDIEZ, E. ([1932 -] 1933). Les plantes grasses autres que les cactees. Les aloes et genres voisins (suite). Revue Horticole; resume de tout ce qui parait d'interessant en jardinage [etc.]. (Paris). New series. 23 (105 of whole series): 570 - 571. [Aloe × spinosissima on p. 570 - 571. See: http: // bibliotheque-numerique. hortalia. org / files / original / cc 8 a 632 c 3 d 5 cc 42 fea 5 cc 7875 f 21 e 1 a 7. pdf]" title="Les plantes grasses autres que les cactees. Les aloes et genres voisins (suite)" type="url" year="1932">Jahandiez</bibRefCitation>
, as “Jahand. in
<emphasis id="B968EAA3FFFA2861C968FAB8FBF8FABC" box="[882,1074,1325,1346]" italics="true" pageId="3" pageNumber="28">Rev. Hort. [Paris]</emphasis>
23 (105 of whole series): 570 (1933)”, the name was clearly validly published 25 years earlier by
<bibRefCitation id="EF8D4B40FFFA2861CE2CFAF1FAC9FA87" author="BERGER, A." box="[1078,1283,1380,1401]" firstAuthor="Berger" journalOrPublisher="Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft" page="183" pageId="3" pageNumber="28" pagination="159 - 326" part="33" refId="ref2409" refString="BERGER, A. (1908). 8. Aloe L. In: A. ENGLER (ED.), Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft 33: 159 - 326. Verlag von Wilhelm Engelmann, Leipzig." title="Aloe L" type="journal article" volumeTitle="A. ENGLER (ED.)" year="1908">Berger (1908: 183)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C306653AFFFA2866C8F4FA0FFEE5FC1F" lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28" type="description">
<paragraph id="8BA336B1FFFA2861C8F4FA0FFC4BFA4E" blockId="3.[750,1294,1434,1731]" box="[750,897,1434,1456]" pageId="3" pageNumber="28">
<heading id="D0EB81DDFFFA2861C8F4FA0FFC4BFA4E" bold="true" box="[750,897,1434,1456]" fontSize="9" level="5" pageId="3" pageNumber="28" reason="6">
<emphasis id="B968EAA3FFFA2861C8F4FA0FFC4BFA4E" bold="true" box="[750,897,1434,1456]" pageId="3" pageNumber="28">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA336B1FFFA2866C909FA23FEE5FC1F" blockId="3.[750,1294,1434,1731]" lastBlockId="4.[145,689,944,993]" lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28">
Perennial, small to medium-sized, shrubby, rosulate, leaf succulent, up to
<quantity id="4CE49B54FFFA2861CE54FA47FBBBFA19" box="[1102,1137,1490,1511]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="28" unit="m" value="1.0">1m</quantity>
tall, usually shorter, excluding inflorescence.
<emphasis id="B968EAA3FFFA2861CE6CFA78FB70F9FC" box="[1142,1210,1517,1538]" italics="true" pageId="3" pageNumber="28">Stems</emphasis>
many, sprouting from the base.
<emphasis id="B968EAA3FFFA2861CE2CF99CFB4BF9E0" box="[1078,1153,1545,1566]" italics="true" pageId="3" pageNumber="28">Leaves</emphasis>
lanceolate, spreading with upper ½ to 3 usually gracefully recurved,
<quantity id="4CE49B54FFFA2861C944F9D5FC0AF9AB" box="[862,960,1600,1621]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="28" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">2030cm</quantity>
long, glaucous to light green;
<emphasis id="B968EAA3FFFA2861C8F4F9CEFC68F98E" box="[750,930,1627,1648]" italics="true" pageId="3" pageNumber="28">adaxial surface</emphasis>
sparingly soft-spiny, concave;
<emphasis id="B968EAA3FFFA2861C8F4F9E3FC50F972" box="[750,922,1654,1676]" italics="true" pageId="3" pageNumber="28">abaxial surface</emphasis>
densely soft-spiny, convex;
<emphasis id="B968EAA3FFFA2861CEC2F9E2FCD7F956" italics="true" pageId="3" pageNumber="28">margins</emphasis>
adorned with white, somewhat bony but harmless teeth.
<emphasis id="B968EAA3FFFA2861C9B1F938FBF6F93C" box="[939,1084,1709,1730]" italics="true" pageId="3" pageNumber="28">Inflorescence</emphasis>
a conical raceme, very rarely paniculate, to
<quantity id="4CE49B54FFFD2866CBCEFC25FDC5FC3B" box="[468,527,944,966]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="29" unit="cm" value="50.0">50cm</quantity>
tall.
<emphasis id="B968EAA3FFFD2866C843FC25FD7BFC3B" box="[601,689,944,965]" italics="true" pageId="4" pageNumber="29">Flowers</emphasis>
bright orange.
</paragraph>
</subSubSection>
<caption id="DF636639FFFD2866CA8BFCD3FAC4FC6A" ID-DOI="http://doi.org/10.5281/zenodo.7867794" ID-Zenodo-Dep="7867794" httpUri="https://zenodo.org/record/7867794/files/figure.png" pageId="4" pageNumber="29" startId="4.[145,229,838,860]" targetBox="[145,1294,145,829]" targetPageId="4">
<paragraph id="8BA336B1FFFD2866CA8BFCD3FAC4FC6A" blockId="4.[145,1294,838,916]" pageId="4" pageNumber="29">
<emphasis id="B968EAA3FFFD2866CA8BFCD3FEC8FCA5" bold="true" box="[145,258,838,860]" pageId="4" pageNumber="29">Figure 4.</emphasis>
The Palazzo Orengo at the Hanbury Gardens. Alwin Berger, who described
<taxonomicName id="4C1C4D32FFFD2866CE7DFCD3FF0DFC86" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFFD2866CE7DFCD3FF0DFC86" italics="true" pageId="4" pageNumber="29">Aloe ×spinosissima</emphasis>
</taxonomicName>
, was based at the Garden at the time that he received material of this nothospecies from Ludwig Winter. Photograph: Gideon F. Smith.
</paragraph>
</caption>
<subSubSection id="C306653AFFFD2866CA8BFB97FE8DFB94" pageId="4" pageNumber="29" type="etymology">
<paragraph id="8BA336B1FFFD2866CA8BFB97FECDFBE9" blockId="4.[145,689,1026,1131]" box="[145,263,1026,1047]" pageId="4" pageNumber="29">
<heading id="D0EB81DDFFFD2866CA8BFB97FECDFBE9" bold="true" box="[145,263,1026,1047]" fontSize="9" level="5" pageId="4" pageNumber="29" reason="6">
<emphasis id="B968EAA3FFFD2866CA8BFB97FECDFBE9" bold="true" box="[145,263,1026,1047]" pageId="4" pageNumber="29">Eponymy</emphasis>
</heading>
</paragraph>
<paragraph id="8BA336B1FFFD2866CAADFB8BFE8DFB94" blockId="4.[145,689,1026,1131]" pageId="4" pageNumber="29">The epithet spinosissima means very spiny and is derived from the Latin word spinosus [English: spiny].</paragraph>
</subSubSection>
<subSubSection id="C306653AFFFD2866CA8BFB1EFDCDF938" pageId="4" pageNumber="29" type="description">
<paragraph id="8BA336B1FFFD2866CA8BFB1EFF28FB5F" blockId="4.[145,690,1163,1735]" box="[145,226,1163,1185]" pageId="4" pageNumber="29">
<heading id="D0EB81DDFFFD2866CA8BFB1EFF28FB5F" bold="true" box="[145,226,1163,1185]" fontSize="9" level="5" pageId="4" pageNumber="29" reason="6">
<emphasis id="B968EAA3FFFD2866CA8BFB1EFF28FB5F" bold="true" box="[145,226,1163,1185]" pageId="4" pageNumber="29">Origin</emphasis>
</heading>
</paragraph>
<paragraph id="8BA336B1FFFD2866CAADFB32FDCDF938" blockId="4.[145,690,1163,1735]" pageId="4" pageNumber="29">
According to
<bibRefCitation id="EF8D4B40FFFD2866CB57FB32FDEFFB42" author="Jacobsen" box="[333,549,1191,1213]" firstAuthor="Jacobsen" page="97" pageId="4" pageNumber="29" refId="ref2873" refString="JACOBSEN, H. (1977). Lexicon of succulent plants. Short descriptions, habitats and synonymy of succulent plants other than Cactaceae. 2 nd edn. Blandford Press Ltd, Poole, Dorset." type="book" year="1977">Jacobsen (1977: 97</bibRefCitation>
,
<bibRefCitation id="EF8D4B40FFFD2866C82EFB32FD68FB42" author="JACOBSEN, H." box="[564,674,1191,1213]" firstAuthor="Jacobsen" journalOrPublisher="Blandford Press Ltd, Poole, Dorset" page="197" pageId="4" pageNumber="29" refId="ref2911" refString="JACOBSEN, H. (1986). A handbook of succulent plants. Volume 1. Abromeitiella to Euphorbia. Blandford Press Ltd, Poole, Dorset." title="A handbook of succulent plants. Volume 1. Abromeitiella to Euphorbia" type="book" year="1986">1986: 197</bibRefCitation>
),
<taxonomicName id="4C1C4D32FFFD2866CA8BFB57FEAEFB26" box="[145,356,1218,1240]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="species" species="spinosissima">
<emphasis id="B968EAA3FFFD2866CA8BFB57FF0BFB29" box="[145,193,1218,1239]" italics="true" pageId="4" pageNumber="29">Aloe</emphasis>
×
<emphasis id="B968EAA3FFFD2866CACCFB56FEAEFB26" box="[214,356,1219,1240]" italics="true" pageId="4" pageNumber="29">spinosissima</emphasis>
</taxonomicName>
is a hybrid that was raised at La Mortola, the Hanbury Gardens (Giardini Botanici Hanbury) on the Italian Riviera. However,
<bibRefCitation id="EF8D4B40FFFD2866CACAFA80FE69FAD4" author="BERGER, A." box="[208,419,1301,1322]" firstAuthor="Berger" journalOrPublisher="Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft" page="183" pageId="4" pageNumber="29" pagination="159 - 326" part="33" refId="ref2409" refString="BERGER, A. (1908). 8. Aloe L. In: A. ENGLER (ED.), Liliaceae-Asphodeloideae-Aloineae. Das Pflanzenreich IV. 38. III. II. Heft 33: 159 - 326. Verlag von Wilhelm Engelmann, Leipzig." title="Aloe L" type="journal article" volumeTitle="A. ENGLER (ED.)" year="1908">Berger (1908: 183)</bibRefCitation>
stated that “Ich erhielt die Pflanze von L. Winter in Bordighera, über ihren Ursprung ist mir nichts bekannt geworden.” [English: “I received the plant from L. Winter in Bordighera, I know nothing of its origin.”]. Following employment for the development of the Hanbury Garden, La Mortola, in northwestern Italy, from 1868 to 1875 the German horticulturalist and botanist Ludwig Jonathon Winter (
<date id="FFA21071FFFD2866C86CFA64FECBF9DC" pageId="4" pageNumber="29" value="1846-08-09">9 August 1846</date>
[Heidelberg, Germany]
<date id="FFA21071FFFD2866C807F999FD7BF9DF" box="[541,689,1548,1570]" pageId="4" pageNumber="29" value="1912-07-12">12 July 1912</date>
[Mannheim, Germany]) was based close by in Bordighera, Italy. Alwin Berger took up the curatorship position at La Mortola in 1897, and while based there did considerable work on several succulent plant families (
<bibRefCitation id="EF8D4B40FFFD2866CBB8F903FDFEF955" author="MOORE, A." box="[418,564,1686,1707]" firstAuthor="Moore" journalOrPublisher="Cadogan Guides, London" pageId="4" pageNumber="29" refId="ref3054" refString="MOORE, A. (2004). La Mortola. In the footsteps of Thomas Hanbury. Cadogan Guides, London." title="La Mortola. In the footsteps of Thomas Hanbury" type="book" volumeTitle="La Mortola. In the footsteps of Thomas Hanbury. Cadogan Guides" year="2004">Moore, 2004</bibRefCitation>
;
<bibRefCitation id="EF8D4B40FFFD2866C850F900FE62F939" author="SMITH, G. F. &amp; FIGUEIREDO, E." firstAuthor="Smith" journalOrPublisher="Struik Lifestyle (an imprint of Random House Struik (Pty) Ltd), Cape Town." page="94 - 105" pageId="4" pageNumber="29" refId="ref3217" refString="SMITH, G. F. &amp; FIGUEIREDO, E. (2013). Succulent paradise. Twelve great gardens of the world. Struik Lifestyle (an imprint of Random House Struik (Pty) Ltd), Cape Town." title="Succulent paradise. Twelve great gardens of the world" type="book" year="2013">Smith &amp; Figueiredo, 2013: 94105</bibRefCitation>
,
<bibRefCitation id="EF8D4B40FFFD2866CBACF924FE32F938" author="SMITH, G. F. &amp; FIGUEIREDO, E." box="[438,504,1713,1734]" firstAuthor="Smith" journalOrPublisher="Cactus &amp; Succulent Journal (U. S.)" pageId="4" pageNumber="29" pagination="12 - 19" part="86" refId="ref3260" refString="SMITH, G. F. &amp; FIGUEIREDO, E. (2014 a). The Hanbury Gardens at La Mortola in Italy: a perfect setting for succulents. Cactus &amp; Succulent Journal (U. S.) 86: 12 - 19." title="The Hanbury Gardens at La Mortola in Italy: a perfect setting for succulents" type="journal article" year="2014" yearSuffix="a">2014a</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>