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<mods:title id="3445D9304A435B2638840968B5AEFB05">Review of the Oriental genus Indocloeon Müller-Liebenau 1982 (Ephemeroptera: Baetidae) with descriptions of two new species</mods:title>
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<mods:namePart id="087C7058E2D467EB1F3327277AAAC0B9">Kluge, Nikita J.</mods:namePart>
<mods:affiliation id="12CAC422BBC5FC3B8011254D7B1B36CE">Department of Entomology, Saint Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia Website: http: // www. insecta. bio. spbu. ru. n. kluge @ spbu. ru; https: // orcid. org / 0000 - 0001 - 9741 - 7790</mods:affiliation>
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<mods:namePart id="5DFD90D617786BA372C83434D91AFD35">Suttinun, Chanaporn</mods:namePart>
<mods:affiliation id="1E9F25D853DDAA29D5C314943C648B88">Animal Systematics and Ecology Speciality Research Unit (ASESRU), Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand. chana. suttinun @ gmail. com; https: // orcid. org / 0000 - 0002 - 0380 - 1554</mods:affiliation>
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<treatment id="03B587A2F21CFF8EECABF9D20F42F841" ID-DOI="http://doi.org/10.5281/zenodo.3853360" ID-GBIF-Taxon="164248612" ID-Zenodo-Dep="3853360" LSID="urn:lsid:plazi:treatment:03B587A2F21CFF8EECABF9D20F42F841" httpUri="http://treatment.plazi.org/id/03B587A2F21CFF8EECABF9D20F42F841" lastPageNumber="478" pageId="27" pageNumber="478">
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<heading id="D0EB81D8F21CFF8EECABF9D20E99F98C" bold="true" box="[151,758,1563,1590]" fontSize="11" level="1" pageId="27" pageNumber="478" reason="1">
<taxonomicName id="4C1C4D37F21CFF8EECABF9D20E99F98C" ID-CoL="3PN47" authority="Kluge 2012" authorityName="Kluge" authorityYear="2012" box="[151,758,1563,1590]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="indonesiae" subGenus="Hindocloeon">
<emphasis id="B968EAA6F21CFF8EECABF9D20E99F98C" bold="true" box="[151,758,1563,1590]" pageId="27" pageNumber="478">
<emphasis id="B968EAA6F21CFF8EECABF9D20D4FF98F" bold="true" box="[151,288,1563,1590]" italics="true" pageId="27" pageNumber="478">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F21CFF8EED0DF9D20DA2F98F" bold="true" box="[305,461,1563,1590]" italics="true" pageId="27" pageNumber="478">Hindocloeon</emphasis>
)
<emphasis id="B968EAA6F21CFF8EEDE3F9D20E0EF98F" bold="true" box="[479,609,1563,1590]" italics="true" pageId="27" pageNumber="478">indonesiae</emphasis>
<bibRefCitation id="EF8D4B45F21CFF8EEE54F9D50E99F98C" author="Kluge, N. J." box="[616,758,1563,1590]" pageId="27" pageNumber="478" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012</bibRefCitation>
</emphasis>
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<paragraph id="8BA336B4F21CFF8EECABF9F60DD8F9E0" blockId="27.[151,758,1563,1625]" box="[151,439,1599,1625]" pageId="27" pageNumber="478">
(
<figureCitation id="13272A31F21CFF8EECA3F9F60D54F9E0" box="[159,315,1599,1625]" captionStart="FIGURES 106111" captionStartId="30.[151,264,1633,1658]" captionTargetBox="[151,1436,237,1608]" captionTargetId="figure@30.[151,1436,237,1608]" captionTargetPageId="30" captionText="FIGURES 106111. Eggs of Hindocloeon. 106108, Indocloeon (Hindocloeon) continentale sp. n.: 106107, specimen from Erawan (Thailand); 108, specimen from Erumeli (India). 109110, Indocloeon (Hindocloeon) indonesiae; 111, Indocloeon (Hindocloeon) sp. «Java»." figureDoi="http://doi.org/10.5281/zenodo.3839375" httpUri="https://zenodo.org/record/3839375/files/figure.png" pageId="27" pageNumber="478">Figs 109110</figureCitation>
,
<figureCitation id="13272A31F21CFF8EED7BF9F60DC3F9E0" box="[327,428,1599,1625]" captionStart="FIGURES 112114" captionStartId="31.[151,264,1857,1882]" captionTargetBox="[166,1436,885,1832]" captionTargetId="figure@31.[151,1436,885,1832]" captionTargetPageId="31" captionText="FIGURES 112114. Indocloeon (Hindocloeon) indonesiae. 112113, female imago; 114, male imago (holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839377" httpUri="https://zenodo.org/record/3839377/files/figure.png" pageId="27" pageNumber="478">112114</figureCitation>
).
</paragraph>
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<paragraph id="8BA336B4F21CFF8EECABF9B70F8AF92F" blockId="27.[151,997,1662,1687]" box="[151,997,1662,1687]" pageId="27" pageNumber="478">
<treatmentCitationGroup id="AB0C119AF21CFF8EECABF9B70F8AF92F" box="[151,997,1662,1687]" pageId="27" pageNumber="478">
<treatmentCitation id="0ABD10A5F21CFF8EECABF9B70E41F92E" author="Kluge, N. J." box="[151,558,1662,1687]" page="363" pageId="27" pageNumber="478" year="2012">
<taxonomicName id="4C1C4D37F21CFF8EECABF9B70E41F92E" ID-CoL="3PN47" authority="Kluge 2012: 363" authorityName="Kluge" authorityPageNumber="363" authorityYear="2012" box="[151,558,1662,1687]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="indonesiae">
<emphasis id="B968EAA6F21CFF8EECABF9B70D12F92F" box="[151,381,1662,1686]" italics="true" pageId="27" pageNumber="478">Indocloeon indonesiae</emphasis>
<bibRefCitation id="EF8D4B45F21CFF8EEDB8F9B70E41F92E" author="Kluge, N. J." box="[388,558,1662,1687]" pageId="27" pageNumber="478" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012: 363</bibRefCitation>
</taxonomicName>
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(larva, subimago,
<specimenCount id="9D1AFD3DF21CFF8EEED7F9B60E91F92F" box="[747,766,1663,1686]" pageId="27" pageNumber="478" type="male"></specimenCount>
and
<specimenCount id="9D1AFD3DF21CFF8EEF13F9B60F2DF92F" box="[815,834,1663,1686]" pageId="27" pageNumber="478" type="female"></specimenCount>
imagines, egg);
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C306653FF21CFF8EECABF977086EF809" pageId="27" pageNumber="478" type="materials_examined">
<paragraph id="8BA336B4F21CFF8EECABF9770F6FF944" blockId="27.[151,1437,1726,2040]" pageId="27" pageNumber="478">
<emphasis id="B968EAA6F21CFF8EECABF9770D14F960" bold="true" box="[151,379,1726,1753]" pageId="27" pageNumber="478">Material examined</emphasis>
(
<collectionCode id="ED0DAE71F21CFF8EEDB0F9760DAFF960" box="[396,448,1727,1753]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="27" pageNumber="478" type="Museum">ZIN</collectionCode>
).
<materialsCitation id="3B743CE9F21CFF8EEDE8F9760E93F944" ID-GBIF-Occurrence="2622593314" collectingDate="2009-09-19" collectingDateMax="2009-09-26" collectingDateMin="2009-09-19" collectorName="N. Kluge &amp; L. Sheyko" country="Indonesia" location="Lombok Island" municipality="Senaru" pageId="27" pageNumber="478" specimenCount="4" specimenCount-female="1" specimenCount-larva="2" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54A78816F21CFF8EEDE8F9760E53F960" box="[468,572,1727,1753]" pageId="27" pageNumber="478" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54A78816F21CFF8EEE4AF9090E89F963" box="[630,742,1728,1754]" pageId="27" pageNumber="478" type="paratype">paratypes</typeStatus>
:
<collectingCountry id="F30B7624F21CFF8EEECEF9760FE5F960" box="[754,906,1727,1753]" name="Indonesia" pageId="27" pageNumber="478">INDONESIA</collectingCountry>
,
<location id="8EC3606FF21CFF8EEFA6F9760827F960" LSID="urn:lsid:plazi:treatment:03B587A2F21CFF8EECABF9D20F42F841:8EC3606FF21CFF8EEFA6F9760827F960" box="[922,1096,1727,1753]" country="Indonesia" municipality="Senaru" name="Lombok Island" pageId="27" pageNumber="478">Lombok Island</location>
,
<collectingMunicipality id="6BC7ACCEF21CFF8EE869F97608C9F960" box="[1109,1190,1727,1753]" pageId="27" pageNumber="478">Senaru</collectingMunicipality>
,
<date id="FFA21074F21CFF8EE88EF9760931F960" box="[1202,1374,1727,1753]" pageId="27" pageNumber="478" value="2009-09-19" valueMax="2009-09-26" valueMin="2009-09-19">
<collectingDate id="EFE6E99CF21CFF8EE88EF9760931F960" box="[1202,1374,1727,1753]" pageId="27" pageNumber="478" value="2009-09-19" valueMax="2009-09-26" valueMin="2009-09-19">1926.IX.2009</collectingDate>
</date>
, coll.
<collectorName id="26E95362F21CFF8EECABF92D0C92F944" box="[151,253,1763,1790]" pageId="27" pageNumber="478">N. Kluge</collectorName>
and
<collectorName id="26E95362F21CFF8EED08F92D0DC4F944" box="[308,427,1763,1790]" pageId="27" pageNumber="478">L. Sheyko</collectorName>
: 2 L-S-I
<specimenCount id="9D1AFD3DF21CFF8EEE30F92A0E70F944" box="[524,543,1763,1789]" pageId="27" pageNumber="478" type="male"></specimenCount>
, 1 L-S-I
<specimenCount id="9D1AFD3DF21CFF8EEEBAF92A0EFBF944" box="[646,660,1763,1789]" pageId="27" pageNumber="478" type="female"></specimenCount>
,
<specimenCount id="9D1AFD3DF21CFF8EEEA3F92A0E93F944" box="[671,764,1763,1789]" pageId="27" pageNumber="478" type="larva">2 larvae</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA336B4F21CFF8EECFBF8CE086EF809" blockId="27.[151,1437,1726,2040]" pageId="27" pageNumber="478">
<emphasis id="B968EAA6F21CFF8EECFBF8CE0EF7F898" bold="true" box="[199,664,1798,1825]" pageId="27" pageNumber="478">Notes appended to original description.</emphasis>
The statement that the imaginal antenna has «scapus with dark brown spot near apex, pedicellus with distal half brown» (
<bibRefCitation id="EF8D4B45F21CFF8EEEE9F8E30F33F8FC" author="Kluge, N. J." box="[725,860,1834,1861]" pageId="27" pageNumber="478" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012</bibRefCitation>
) is not quite correct: actually scapus has two spots, one on the lateral side and another on the median side; the pedicellus does not have the whole distal half brown, but only its median and lateral sides (as in
<figureCitation id="13272A31F21CFF8EEE70F8BB0EF2F835" box="[588,669,1906,1932]" captionStart="FIGURES 9195" captionStartId="25.[151,264,1887,1912]" captionTargetBox="[151,1436,181,1861]" captionTargetId="figure@25.[151,1436,181,1861]" captionTargetPageId="25" captionText="FIGURES 9195. Indocloeon (Hindocloeon) continentale sp. n. 91, fragment of subimaginal exuviae with postsubalar sclerite; 92, antennae of male imago; 93, male imago; 94, its metathorax and base of abdomen; 95, male subimago (arrow shows brown hypodermal spot just ventrad of metathoracic postnotum)." figureDoi="http://doi.org/10.5281/zenodo.3839367" httpUri="https://zenodo.org/record/3839367/files/figure.png" pageId="27" pageNumber="478">Fig. 92</figureCitation>
) [see
<taxonomicName id="4C1C4D37F21CFF8EEEDDF8BB0F1DF835" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[737,882,1906,1932]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="455" phylum="Arthropoda" rank="subGenus" subGenus="Hindocloeon">
<emphasis id="B968EAA6F21CFF8EEEDDF8BB0F1DF835" box="[737,882,1906,1932]" italics="true" pageId="27" pageNumber="478">Hindocloeon</emphasis>
</taxonomicName>
(9)]. Colouration of male and female imagines is as shown in
<figureCitation id="13272A31F21CFF8EED1EF85F0DD2F809" box="[290,445,1942,1968]" captionStart="FIGURES 112114" captionStartId="31.[151,264,1857,1882]" captionTargetBox="[166,1436,885,1832]" captionTargetId="figure@31.[151,1436,885,1832]" captionTargetPageId="31" captionText="FIGURES 112114. Indocloeon (Hindocloeon) indonesiae. 112113, female imago; 114, male imago (holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839377" httpUri="https://zenodo.org/record/3839377/files/figure.png" pageId="27" pageNumber="478">Figs 112114</figureCitation>
. Egg chorion has very fine rugosity (
<figureCitation id="13272A31F21CFF8EEF67F85F0F9AF809" box="[859,1013,1942,1968]" captionStart="FIGURES 106111" captionStartId="30.[151,264,1633,1658]" captionTargetBox="[151,1436,237,1608]" captionTargetId="figure@30.[151,1436,237,1608]" captionTargetPageId="30" captionText="FIGURES 106111. Eggs of Hindocloeon. 106108, Indocloeon (Hindocloeon) continentale sp. n.: 106107, specimen from Erawan (Thailand); 108, specimen from Erumeli (India). 109110, Indocloeon (Hindocloeon) indonesiae; 111, Indocloeon (Hindocloeon) sp. «Java»." figureDoi="http://doi.org/10.5281/zenodo.3839375" httpUri="https://zenodo.org/record/3839375/files/figure.png" pageId="27" pageNumber="478">Fig. 109110</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C306653FF21CFF8EECFBF8700F42F841" pageId="27" pageNumber="478" type="distribution">
<paragraph id="8BA336B4F21CFF8EECFBF8700F42F841" blockId="27.[151,1437,1726,2040]" pageId="27" pageNumber="478">
<emphasis id="B968EAA6F21CFF8EECFBF8700D0CF86D" bold="true" box="[199,355,1977,2004]" pageId="27" pageNumber="478">Distribution:</emphasis>
<collectingRegion id="49D8F856F21CFF8EED51F8730E09F86A" box="[365,614,1977,2004]" country="Indonesia" name="Nusa Tenggara Timur" pageId="27" pageNumber="478">Lesser Sunda Islands</collectingRegion>
: initially described from Lombok (
<bibRefCitation id="EF8D4B45F21CFF8EEFC2F87008E4F86D" author="Kluge, N. J." box="[1022,1163,1977,2004]" pageId="27" pageNumber="478" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012</bibRefCitation>
), subsequently reported from Sumbawa and
<collectingCountry id="F30B7624F21CFF8EED45F8140DD0F84E" box="[377,447,2013,2039]" name="Indonesia" pageId="27" pageNumber="478">Flores</collectingCountry>
(
<bibRefCitation id="EF8D4B45F21CFF8EEDF2F8140F4EF841" author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[462,801,2013,2040]" pageId="27" pageNumber="478" pagination="43 - 60" refId="ref14547" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2017) New species of Indocloeon Muller-Liebenau from South-East Asia (Ephemeroptera, Baetidae). ZooKeys, 723, 43 - 60. https: // doi. org / 10.3897 / zookeys. 723.20578" type="journal article" year="2017">Kaltenbach &amp; Gattolliat 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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