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<document id="79704AB10E142D56DD3C694F6C9970F4" ID-CLB-Dataset="281838" ID-DOI="10.11646/phytotaxa.630.2.7" ID-GBIF-Dataset="950f23bf-a521-464b-81bb-68da9d09e8bc" ID-ISSN="1179-3163" ID-Zenodo-Dep="10376850" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1702550719214" checkinUser="plazi" docAuthor="Tong, Yi-Hua, Ni, Jing-Bo &amp; Wang, Bing-Mou" docDate="2023" docId="03A98E3BFFBEF35AFF129187FCE5FDBE" docLanguage="en" docName="phytotaxa.630.2.7.pdf" docOrigin="Phytotaxa 630 (2)" docSource="https://phytotaxa.mapress.com/pt/article/download/phytotaxa.630.2.7/51371" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Primulina liangwaniae B. M. Wang &amp; Y. H. Tong 2023, sp. nov." docType="treatment" docVersion="4" lastPageNumber="157" masterDocId="FF90F643FFBFF359FF9A9013FF81FFD6" masterDocTitle="Primulina liangwaniae (Gesneriaceae), a new species from Guangdong, China" masterLastPageNumber="158" masterPageNumber="154" pageNumber="155" updateTime="1702666079599" updateUser="karina" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="EC20931FA7934DD70D5882BE1ED71CF4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="56951F997C71DF5ACA10573080D5B7DC">
<mods:title id="2F05D17AAD7AF1C6C96043D0DA1C25D0">Primulina liangwaniae (Gesneriaceae), a new species from Guangdong, China</mods:title>
</mods:titleInfo>
<mods:name id="B35A38278C60EDEBE61C0F19725E8AFD" type="personal">
<mods:role id="F759A7FEF234DBDDFFCB5BF02C593648">
<mods:roleTerm id="EB4814B4A774B89DEDB41027BB9AD8B4">Author</mods:roleTerm>
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<mods:namePart id="F1F7F93D946390C0680E221EA5051202">Tong, Yi-Hua</mods:namePart>
<mods:affiliation id="D757E4E2D49A454468AE22C8A21E3E19">State Key Laboratory of Plant Diversity and Specialty Crops &amp; Key Laboratory of Digital Botanical Garden of Guangdong Province, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China. &amp; South China National Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China. &amp; Panyu Central Hospital, Guangzhou 511402, China.</mods:affiliation>
</mods:name>
<mods:name id="3AF5FA1C7505B6B2AAA0A2462F757A38" type="personal">
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</mods:role>
<mods:namePart id="7DB10577386F5D82F2AE574888D0D820">Ni, Jing-Bo</mods:namePart>
<mods:nameIdentifier id="1A4302F47901C2536460CD5899528D89" type="ORCID">0000-0002-1695-6141</mods:nameIdentifier>
<mods:affiliation id="2CA2702D3880F5481BB2295255CC7EA0">State Key Laboratory of Plant Diversity and Specialty Crops &amp; Key Laboratory of Digital Botanical Garden of Guangdong Province, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China. &amp; South China National Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China.</mods:affiliation>
<mods:nameIdentifier id="92950F8E92CC840220AAC79AF176B990" type="email">zzxx8424b@126.com</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="06A7AD24195F3AD4BAE35DA32A30FC61">Wang, Bing-Mou</mods:namePart>
<mods:nameIdentifier id="281EC15271B56D24F161C9CE3A6D4665" type="ORCID">0000-0002-1103-2599</mods:nameIdentifier>
<mods:affiliation id="300333E79DB93DEC3F3DB8548C9BDD3B">Panyu Central Hospital, Guangzhou 511402, China.</mods:affiliation>
<mods:nameIdentifier id="AB07C8123D0A17A61F7D1265571B9353" type="email">451535625@qq.com</mods:nameIdentifier>
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<mods:title id="C1102A9D2E2E1C6BB20C268677233CEA">Phytotaxa</mods:title>
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<mods:part id="52FF898889EC38A17F298C4976F6E0AC">
<mods:date id="2D554CB99597B89FF2435B7B4C801C9C">2023</mods:date>
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<mods:number id="EE4FFC2EF8E62E09F825DE628748A28C">2023-12-14</mods:number>
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<mods:number id="361234381B85A9E9753B8266AF09D5F5">630</mods:number>
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<mods:url id="680EE516462CB6ED9C0AB5A4887EBD2E">https://phytotaxa.mapress.com/pt/article/download/phytotaxa.630.2.7/51371</mods:url>
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<mods:classification id="D01B66572CBFE30D88A35CE8C07C0BB6">journal article</mods:classification>
<mods:identifier id="3CF2463BF91718DAA9E709D05B688A45" type="CLB-Dataset">281838</mods:identifier>
<mods:identifier id="0011E58E2251403C8FA3D12EF4844CD2" type="DOI">10.11646/phytotaxa.630.2.7</mods:identifier>
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<treatment id="03A98E3BFFBEF35AFF129187FCE5FDBE" ID-DOI="http://doi.org/10.5281/zenodo.10376662" ID-GBIF-Taxon="219861888" ID-Zenodo-Dep="10376662" LSID="urn:lsid:plazi:treatment:03A98E3BFFBEF35AFF129187FCE5FDBE" httpUri="http://treatment.plazi.org/id/03A98E3BFFBEF35AFF129187FCE5FDBE" lastPageId="3" lastPageNumber="157" pageId="1" pageNumber="155">
<subSubSection id="C31A6CA6FFBEF358FF129187FCE6FE78" box="[136,871,404,431]" pageId="1" pageNumber="155" type="nomenclature">
<paragraph id="8BBF3F2DFFBEF358FF129187FCE6FE78" blockId="1.[136,871,404,431]" box="[136,871,404,431]" pageId="1" pageNumber="155">
<heading id="D0F78841FFBEF358FF129187FCE6FE78" box="[136,871,404,431]" fontSize="11" level="2" pageId="1" pageNumber="155" reason="3">
<taxonomicName id="4C0044AEFFBEF358FF129187FD24FE79" authority="B.M.Wang &amp; Y.H.Tong" authorityName="B. M. Wang &amp; Y. H. Tong" authorityYear="2023" box="[136,677,404,431]" class="Magnoliopsida" family="Gesneriaceae" genus="Primulina" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="155" phylum="Tracheophyta" rank="species" species="liangwaniae" status="sp. nov.">
<emphasis id="B974E33FFFBEF358FF129187FE12FE79" bold="true" box="[136,403,404,431]" italics="true" pageId="1" pageNumber="155">Primulina liangwaniae</emphasis>
B.M.Wang &amp; Y.H.Tong
</taxonomicName>
,
<emphasis id="B974E33FFFBEF358FD2B9186FC87FE78" box="[689,774,405,430]" italics="true" pageId="1" pageNumber="155">
<taxonomicNameLabel id="A2475E44FFBEF358FD2B9186FC87FE78" box="[689,774,405,430]" pageId="1" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="133B23A8FFBEF358FC8E9187FCD7FE78" box="[788,854,404,430]" captionStart="FIGURE 1" captionStartId="2.[136,229,1848,1870]" captionTargetBox="[164,1423,190,1824]" captionTargetId="figure-19@2.[164,1423,190,1824]" captionTargetPageId="2" captionText="FIGURE 1. Primulina liangwaniae. A. Habitat; B. Flowering plant, top view; C. Flowering plant, side view; D. Inflorescence, front view; E. Inflorescence, side view; F. Opened corolla; G. Stamens; H. Disc, part of ovary and three calyx lobes; I. Stigma and part of style; J. Capsule with persistent calyx. Photographed by Yi-Hua Tong." figureDoi="http://doi.org/10.5281/zenodo.10376852" httpUri="https://zenodo.org/record/10376852/files/figure.png" pageId="1" pageNumber="155">Fig. 1</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31A6CA6FFBEF358FF1291F3FE65FDEB" pageId="1" pageNumber="155" type="materials_examined">
<paragraph id="8BBF3F2DFFBEF358FF1291F3FE65FDEB" blockId="1.[136,1452,480,718]" pageId="1" pageNumber="155">
<materialsCitation id="3B683570FFBEF358FF1291F3FED6FDEB" collectedFrom="Y. H. Tong, J. B. Ni, B. M. Wang &amp; J. Cai" collectingDate="2022-03-21" collectionCode="IBSC, IBK" country="China" elevation="528" location="Guangdong Province" pageId="1" pageNumber="155" specimenCode="TYH-2568" specimenCount="2" stateProvince="Guangdong Province" typeStatus="holotype">
<emphasis id="B974E33FFFBEF358FF1291F3FF43FE23" bold="true" box="[136,194,480,501]" pageId="1" pageNumber="155">Type:</emphasis>
—
<collectingCountry id="F3177FBDFFBEF358FF4191F3FEA8FE23" box="[219,297,480,501]" name="China" pageId="1" pageNumber="155">CHINA</collectingCountry>
.
<collectingRegion id="49C4F1CFFFBEF358FEAC91F3FD89FE20" box="[310,520,480,502]" country="China" name="Guangdong" pageId="1" pageNumber="155">Guangdong Province</collectingRegion>
: cultivated in South China Botanical Garden, introduced from Yingde City, Huanghua Town, limestone hills, on rocks in shaded places, elev. ca.
<quantity id="4CF892C8FFBEF358FD359217FD6FFDCF" box="[687,750,516,538]" metricMagnitude="2" metricUnit="m" metricValue="5.279999999999999" pageId="1" pageNumber="155" unit="m" value="528.0">
<elevation id="002DD81EFFBEF358FD359217FD6FFDCF" box="[687,750,516,538]" metricMagnitude="2" metricUnit="m" metricValue="5.279999999999999" pageId="1" pageNumber="155" unit="m" value="528.0">528 m</elevation>
</quantity>
,
<date id="FFBE19EDFFBEF358FD629217FC0DFDCF" box="[760,908,516,538]" pageId="1" pageNumber="155" value="2022-03-21">
<collectingDate id="EFFAE005FFBEF358FD629217FC0DFDCF" box="[760,908,516,538]" pageId="1" pageNumber="155" value="2022-03-21">21 March 2022</collectingDate>
</date>
(fl.),
<emphasis id="B974E33FFFBEF358FC599217FA2DFDCF" box="[963,1452,516,537]" italics="true" pageId="1" pageNumber="155">
<collectedFrom id="381C7CD0FFBEF358FC599217FAC0FDCF" box="[963,1345,516,537]" pageId="1" pageNumber="155">Y.H. Tong, J.B. Ni, B.M. Wang &amp; J. Cai</collectedFrom>
<specimenCode id="DBA69756FFBEF358FADD9217FA2DFDCF" box="[1351,1452,516,537]" pageId="1" pageNumber="155">TYH-2568</specimenCode>
</emphasis>
(
<typeStatus id="54BB818FFFBEF358FF5E923BFE9BFDE8" box="[196,282,552,574]" pageId="1" pageNumber="155" type="holotype">holotype</typeStatus>
<collectionCode id="ED11A7E8FFBEF358FE85923BFED6FDEB" box="[287,343,552,573]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15606" lsid="urn:lsid:biocol.org:col:15606" name="South China Botanical Garden" pageId="1" pageNumber="155" type="Herbarium">IBSC</collectionCode>
</materialsCitation>
<materialsCitation id="3B683570FFBEF358FECD923BFE61FDEB" box="[343,480,552,574]" collectedFrom="Y. H. Tong, J. B. Ni, B. M. Wang &amp; J. Cai" collectingDate="2022-03-21" collectionCode="IBSC, IBK" country="China" elevation="528" location="Guangdong Province" pageId="1" pageNumber="155" specimenCode="TYH-2568" specimenCount="2" stateProvince="Guangdong Province" typeStatus="holotype">
,
<typeStatus id="54BB818FFFBEF358FEFB923BFE27FDE8" box="[353,422,552,574]" pageId="1" pageNumber="155" type="isotype">isotype</typeStatus>
<collectionCode id="ED11A7E8FFBEF358FE36923BFE58FDEB" box="[428,473,552,573]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13150" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" pageId="1" pageNumber="155" type="Herbarium">IBK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A6CA6FFBEF358FF129258FB14FD18" pageId="1" pageNumber="155" type="diagnosis">
<paragraph id="8BBF3F2DFFBEF358FF129258FB14FD18" blockId="1.[136,1452,480,718]" pageId="1" pageNumber="155">
<emphasis id="B974E33FFFBEF358FF129258FF72FDB7" bold="true" box="[136,243,587,609]" pageId="1" pageNumber="155">Diagnosis:</emphasis>
—Being close to
<taxonomicName id="4C0044AEFFBEF358FE02925FFD8EFDB7" box="[408,527,588,609]" class="Magnoliopsida" family="Gesneriaceae" genus="Primulina" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="155" phylum="Tracheophyta" rank="species" species="versicolor">
<emphasis id="B974E33FFFBEF358FE02925FFD8EFDB7" box="[408,527,588,609]" italics="true" pageId="1" pageNumber="155">P. versicolor</emphasis>
</taxonomicName>
in having similar shape and color of corollas, but it mainly differs in the smaller (5.4–9.5 ×
<quantity id="4CF892C8FFBEF358FAEF925FFF5BFD53" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="7.0" metricValueMin="4.5" pageId="1" pageNumber="155" unit="cm" value="5.75" valueMax="7.0" valueMin="4.5">4.5–7 cm</quantity>
<emphasis id="B974E33FFFBEF358FF789263FF77FD53" box="[226,246,624,645]" italics="true" pageId="1" pageNumber="155">vs</emphasis>
. 8–18 ×
<quantity id="4CF892C8FFBEF358FED49263FE48FD53" box="[334,457,624,646]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.65" metricValueMin="0.65" pageId="1" pageNumber="155" unit="cm" value="11.5" valueMax="16.5" valueMin="6.5">6.5–16.5 cm</quantity>
) leaf blades with a crenate (vs. entire) margin, much shorter peduncles (
<quantity id="4CF892C8FFBEF358FB109263FB52FD53" box="[1162,1235,624,645]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="1" pageNumber="155" unit="cm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 cm</quantity>
<emphasis id="B974E33FFFBEF358FB409263FB6FFD53" box="[1242,1262,624,645]" italics="true" pageId="1" pageNumber="155">vs</emphasis>
.
<quantity id="4CF892C8FFBEF358FB619263FAD2FD53" box="[1275,1363,624,645]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="1.8" metricValueMin="0.8" pageId="1" pageNumber="155" unit="cm" value="13.0" valueMax="18.0" valueMin="8.0">8–18 cm</quantity>
), smaller bracts (2.2–2.8 ×
<quantity id="4CF892C8FFBEF358FEF69287FE58FD7F" box="[364,473,660,682]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="1" pageNumber="155" unit="cm" value="1.85" valueMax="2.2" valueMin="1.5">1.5–2.2 cm</quantity>
<emphasis id="B974E33FFFBEF358FE7B9287FE74FD7F" box="[481,501,660,681]" italics="true" pageId="1" pageNumber="155">vs</emphasis>
. 5–5.5 ×
<quantity id="4CF892C8FFBEF358FDCC9287FD32FD7F" box="[598,691,660,682]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" metricValueMax="5.0" metricValueMin="4.4" pageId="1" pageNumber="155" unit="cm" value="4.7" valueMax="5.0" valueMin="4.4">4.4–5 cm</quantity>
) with margin being crenate on upper part and entire on lower part (vs. entire thoroughly), much shorter pedicels (
<quantity id="4CF892C8FFBEF358FD8092ABFDEBFD1B" box="[538,618,696,717]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="1" pageNumber="155" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 mm</quantity>
<emphasis id="B974E33FFFBEF358FDEB92ABFD04FD1B" box="[625,645,696,717]" italics="true" pageId="1" pageNumber="155">vs</emphasis>
.
<quantity id="4CF892C8FFBEF358FD0B92ABFD7BFD1B" box="[657,762,696,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.2" metricValueMin="1.0" pageId="1" pageNumber="155" unit="mm" value="16.0" valueMax="22.0" valueMin="10.0">10–22 mm</quantity>
) and narrowly cordate (vs. forked) stigma.
</paragraph>
</subSubSection>
<subSubSection id="C31A6CA6FFBEF358FF1292EFFB24F9A1" pageId="1" pageNumber="155" type="description">
<paragraph id="8BBF3F2DFFBEF358FF1292EFFB24F9A1" blockId="1.[136,1452,764,2087]" pageId="1" pageNumber="155">
<emphasis id="B974E33FFFBEF358FF1292EFFE9DFCC0" bold="true" box="[136,284,764,790]" pageId="1" pageNumber="155">Description:</emphasis>
—Perennial herbs, rhizomes short, ca.
<quantity id="4CF892C8FFBEF358FD5792EFFC98FCC1" box="[717,793,764,791]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="155" unit="cm" value="1.5">1.5 cm</quantity>
long, ca.
<quantity id="4CF892C8FFBEF358FC1D92EFFC49FCC1" box="[903,968,764,791]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="155" unit="mm" value="8.0">8 mm</quantity>
in diam. Leaves all basal, opposite; petiole cross section sub-semicircular,
<quantity id="4CF892C8FFBEF358FE709333FDBFFCED" box="[490,574,800,827]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="1" pageNumber="155" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 cm</quantity>
long,
<quantity id="4CF892C8FFBEF358FD1D9333FD64FCED" box="[647,741,800,827]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="155" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
thick, together with leaf blade and peduncle densely white- or purplish-villous and pubescent, base broadened; blade thickly carnose, dark green or tinged purplish adaxially, green tinged purplish, purplish or purple abaxially, ovate or broadly ovate, 5.4–9.5 ×
<quantity id="4CF892C8FFBEF358FB8D937BFB05FC55" box="[1047,1156,872,899]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="7.0" metricValueMin="4.5" pageId="1" pageNumber="155" unit="cm" value="5.75" valueMax="7.0" valueMin="4.5">4.5–7 cm</quantity>
, apex obtuse or subacute, base cuneate or truncate, sometimes slightly cordate, margin crenate, lateral veins 4–6 pairs, impressed adaxially, raised abaxially. Cymes axillary, unbranched or with 1–2 branches, (2–)4–9-flowered; peduncles light green,
<quantity id="4CF892C8FFBEF358FAC293A3FA2AFC1D" box="[1368,1451,944,971]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="1" pageNumber="155" unit="cm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 cm</quantity>
long,
<quantity id="4CF892C8FFBEF358FF5393C7FEBAFC39" box="[201,315,980,1007]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="155" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
thick; bracts 2, opposite, greenish, tinged purple at apex, concave, ovate or broadly ovate, 2.2–2.8 ×
<quantity id="4CF892C8FFBEF358FF1293EBFE89FBC5" box="[136,264,1016,1043]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="1" pageNumber="155" unit="cm" value="1.85" valueMax="2.2" valueMin="1.5">1.5–2.2 cm</quantity>
, margin crenate on upper part, entire on lower part, apex acute, outside densely white- or purplish-villous and pubescent, inside glabrous. Pedicels
<quantity id="4CF892C8FFBEF358FDC3940FFD39FBE1" box="[601,696,1052,1079]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="1" pageNumber="155" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 mm</quantity>
long, densely puberulent. Calyx 5-parted nearly to the base, lobes linear, 9–10 × ca.
<quantity id="4CF892C8FFBEF358FECC9453FE1AFB8D" box="[342,411,1088,1115]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="155" unit="mm" value="2.0">2 mm</quantity>
, margin entire or with 1–3 inconspicuous teeth each side, puberulent on both sides. Corolla yellowish or yellowish white, with three patches at the sinuses of the two upper lip lobes, patches purple outside and dark purple inside, sometimes absent outside and yellow inside, and with 2 purple bands on the two sutures of the middle lobe of the lower lip to the position where fertile stamens inserted and 9 dark purple stripes,
<quantity id="4CF892C8FFBEF358FADB94BFFA2AFB11" box="[1345,1451,1196,1223]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="4.0" metricValueMin="3.2" pageId="1" pageNumber="155" unit="cm" value="3.6" valueMax="4.0" valueMin="3.2">3.2–4 cm</quantity>
long, outside densely glandular-pubescent and puberulent, inside glabrous on tube and densely glandular-pubescent on lobes; tube infundibuliform,
<quantity id="4CF892C8FFBEF358FE5394E7FDC4FAD9" box="[457,581,1268,1295]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="1" pageNumber="155" unit="cm" value="2.7" valueMax="2.8" valueMin="2.6">2.6–2.8 cm</quantity>
long, ca.
<quantity id="4CF892C8FFBEF358FD2894E7FD7FFAD9" box="[690,766,1268,1295]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="155" unit="cm" value="1.2">1.2 cm</quantity>
in diameter at the mouth; limb distinctly 2-lipped, adaxial lip 2-lobed to over half the length, lobes ovate, round at apex, 5–6 × ca.
<quantity id="4CF892C8FFBEF358FC11950BFC4EFAE5" box="[907,975,1304,1331]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="155" unit="mm" value="9.0">9 mm</quantity>
; abaxial lip 3-lobed, lobes oblong with the central one smaller, round at apex, 8–12 ×
<quantity id="4CF892C8FFBEF358FDFB952FFD40FA81" box="[609,705,1340,1367]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="1" pageNumber="155" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
. Stamens 2, adnate to ca.
<quantity id="4CF892C8FFBEF358FC47952FFBA8FA81" box="[989,1065,1340,1367]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="155" unit="cm" value="1.3">1.3 cm</quantity>
above the base of the corolla tube, filaments white, base inflated, ca.
<quantity id="4CF892C8FFBEF358FD949573FDC7FAAD" box="[526,582,1376,1403]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="155" unit="cm" value="1.0">1 cm</quantity>
long, geniculate below the middle, densely glandular-pubescent throughout; anthers elliptic, ca.
<quantity id="4CF892C8FFBEF358FEFF9597FE26FA49" box="[357,423,1412,1439]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="155" unit="mm" value="4.0">4 mm</quantity>
long; staminodes 3, lateral ones tinged purple at apex, adnate to ca.
<quantity id="4CF892C8FFBEF358FB2A9597FB7CFA49" box="[1200,1277,1412,1439]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="155" unit="cm" value="1.2">1.2 cm</quantity>
above the base of corolla tube,
<quantity id="4CF892C8FFBEF358FEA395BBFE16FA15" box="[313,407,1448,1475]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="155" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
long, densely glandular-pubescent, apex bended, the central one very small, adnate to ca.
<quantity id="4CF892C8FFBEF358FA1295BBFF35FA31" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="155" unit="mm" value="2.5">2.5 mm</quantity>
above the base of corolla tube, ca.
<quantity id="4CF892C8FFBEF358FDA195DFFD10FA31" box="[571,657,1484,1511]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="155" unit="mm" value="0.5">0.5 mm</quantity>
long, glabrous. Disc annular, ca.
<quantity id="4CF892C8FFBEF358FB9F95DFFBC4FA31" box="[1029,1093,1484,1511]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="155" unit="mm" value="1.0">1 mm</quantity>
high, glabrous. Pistil ca.
<quantity id="4CF892C8FFBEF358FAC595DFFA2AFA31" box="[1375,1451,1484,1511]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="155" unit="cm" value="3.1">3.1 cm</quantity>
long; ovary ca.
<quantity id="4CF892C8FFBEF358FEA295E3FE05F9DD" box="[312,388,1520,1547]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="155" unit="cm" value="1.3">1.3 cm</quantity>
long, densely white-villous and pubescent; style ca.
<quantity id="4CF892C8FFBEF358FC4D95E3FBA5F9DD" box="[983,1060,1520,1547]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="155" unit="cm" value="1.5">1.5 cm</quantity>
long, densely glandular-pubescent or puberulous; stigma narrowly cordate,
<quantity id="4CF892C8FFBEF358FDCA9607FD43F9F9" box="[592,706,1556,1583]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="1" pageNumber="155" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
long, shallowly lobed, lobes ca.
<quantity id="4CF892C8FFBEF358FBA89607FB08F9F9" box="[1074,1161,1556,1583]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="155" unit="mm" value="0.5">0.5 mm</quantity>
long. Capsule pale green, linear,
<quantity id="4CF892C8FFBEF358FF49962BFEBBF985" box="[211,314,1592,1619]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="6.5" metricValueMin="4.0" pageId="1" pageNumber="155" unit="cm" value="5.25" valueMax="6.5" valueMin="4.0">4–6.5 cm</quantity>
×
<quantity id="4CF892C8FFBEF358FECF962BFE49F985" box="[341,456,1592,1619]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="1" pageNumber="155" unit="mm" value="2.75" valueMax="3.5" valueMin="2.0">2–3.5 mm</quantity>
, with persistent calyx lobes at base and style at apex, densely white-villous and pubescent, persistent calyx lobes 11–13 ×
<quantity id="4CF892C8FFBEF358FE79964FFDD9F9A1" box="[483,600,1628,1655]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="155" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
. Seeds small, sub-turbinate, 0.3–0.4 ×
<quantity id="4CF892C8FFBEF358FB93964FFB20F9A1" box="[1033,1185,1628,1655]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="1" pageNumber="155" unit="mm" value="0.125" valueMax="0.15" valueMin="0.1">0.1–0.15 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A6CA6FFBEF358FF279693FC4AF8FD" pageId="1" pageNumber="155" type="distribution">
<paragraph id="8BBF3F2DFFBEF358FF279693FE50F8D1" blockId="1.[136,1452,764,2087]" pageId="1" pageNumber="155">
<emphasis id="B974E33FFFBEF358FF279693FE6FF94C" bold="true" box="[189,494,1664,1690]" pageId="1" pageNumber="155">Distribution and ecology:</emphasis>
—
<taxonomicName id="4C0044AEFFBEF358FD909693FC8CF94C" authorityName="B. M. Wang &amp; Y. H. Tong" authorityYear="2023" box="[522,781,1664,1690]" class="Magnoliopsida" family="Gesneriaceae" genus="Primulina" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="155" phylum="Tracheophyta" rank="species" species="liangwaniae">
<emphasis id="B974E33FFFBEF358FD909693FC8CF94C" box="[522,781,1664,1690]" italics="true" pageId="1" pageNumber="155">Primulina liangwaniae</emphasis>
</taxonomicName>
is only known from the
<typeStatus id="54BB818FFFBEF358FBB99692FBD2F94D" box="[1059,1107,1665,1691]" pageId="1" pageNumber="155">type</typeStatus>
locality in Huanghua Town of Yingde City,
<collectingCountry id="F3177FBDFFBEF358FEBA96B7FEE6F968" box="[288,359,1700,1726]" name="China" pageId="1" pageNumber="155">China</collectingCountry>
. It grows on limestone rocks at an elevation of ca.
<specimenCount id="9D06F4A4FFBEF358FC2996B7FB82F969" box="[947,1027,1700,1727]" count="530" pageId="1" pageNumber="155" type="generic">
<quantity id="4CF892C8FFBEF358FC2996B7FC7EF969" box="[947,1023,1700,1727]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="155" unit="m" value="530.0">530 m</quantity>
.
</specimenCount>
The associated species include some plants also adapted to limestone, such as
<taxonomicName id="4C0044AEFFBEF358FDFD96DAFCF4F935" authority="Hance" authorityName="Hance" box="[615,885,1736,1763]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="155" phylum="Tracheophyta" rank="species" species="leprosa">
<emphasis id="B974E33FFFBEF358FDFD96DAFCA0F934" box="[615,801,1736,1762]" italics="true" pageId="1" pageNumber="155">Begonia leprosa</emphasis>
Hance
</taxonomicName>
,
<taxonomicName id="4C0044AEFFBEF358FC1996DBFBEBF935" authority="Hance" authorityName="Hance" box="[899,1130,1736,1763]" class="Magnoliopsida" family="Urticaceae" genus="Pilea" kingdom="Plantae" order="Rosales" pageId="1" pageNumber="155" phylum="Tracheophyta" rank="species" species="peltata">
<emphasis id="B974E33FFFBEF358FC1996DBFB97F934" box="[899,1046,1736,1762]" italics="true" pageId="1" pageNumber="155">Pilea peltata</emphasis>
Hance
</taxonomicName>
,
<taxonomicName id="4C0044AEFFBEF358FBE296DBFAF7F934" box="[1144,1398,1736,1762]" class="Polypodiopsida" family="Nephrolepidaceae" genus="Nephrolepis" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="155" phylum="Tracheophyta" rank="species" species="cordifolia">
<emphasis id="B974E33FFFBEF358FBE296DBFAF7F934" box="[1144,1398,1736,1762]" italics="true" pageId="1" pageNumber="155">Nephrolepis cordifolia</emphasis>
</taxonomicName>
(L.) C.Presl,
<taxonomicName id="4C0044AEFFBEF358FF7296FFFE1CF8D1" box="[232,413,1772,1799]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="155" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B974E33FFFBEF358FF7296FFFEE7F8D0" box="[232,358,1772,1798]" italics="true" pageId="1" pageNumber="155">Selaginella</emphasis>
spp.
</taxonomicName>
, etc.
</paragraph>
<paragraph id="8BBF3F2DFFBEF358FF279703FC4AF8FD" blockId="1.[136,1452,764,2087]" box="[189,971,1808,1835]" pageId="1" pageNumber="155">
<emphasis id="B974E33FFFBEF358FF279703FEC2F8FC" bold="true" box="[189,323,1808,1834]" pageId="1" pageNumber="155">Phenology:</emphasis>
—Flowering in late March to early April; fruiting in May.
</paragraph>
</subSubSection>
<subSubSection id="C31A6CA6FFBEF35BFF279727FBC7F7F0" lastPageId="2" lastPageNumber="156" pageId="1" pageNumber="155" type="etymology">
<paragraph id="8BBF3F2DFFBEF358FF279727FD90F840" blockId="1.[136,1452,764,2087]" pageId="1" pageNumber="155">
<emphasis id="B974E33FFFBEF358FF279727FEC9F898" bold="true" box="[189,328,1844,1870]" pageId="1" pageNumber="155">Etymology:</emphasis>
—The specific epithet is derived from the name of the third author’s mother, Ms. Liang-Wan Zhang, who encouraged her son to maintain continuous enthusiasm with plants, but sadly passed away in 2020. The Chinese name is given as “
<emphasis id="B974E33FFFBEF358FECF976EFE7FF841" bold="true" box="[341,510,1917,1943]" pageId="1" pageNumber="155">̊șẫǝüë</emphasis>
”.
</paragraph>
<paragraph id="8BBF3F2DFFBEF358FF2797B3FCEDF7F0" blockId="1.[136,1452,764,2087]" pageId="1" pageNumber="155">
<emphasis id="B974E33FFFBEF358FF2797B3FDC0F86C" bold="true" box="[189,577,1952,1978]" pageId="1" pageNumber="155">Provisional conservation status:</emphasis>
—We only found one small population with ca. 50 individuals growing on limestone rocks of the entrance of a cave at the
<typeStatus id="54BB818FFFBEF358FD3A97D6FD51F809" box="[672,720,1989,2015]" pageId="1" pageNumber="155">type</typeStatus>
locality. Due to lack of time and inconvenient traffic, a thorough population assessment of this species in that limestone area has not been made, so it is best classified as ‘Data Deficient’ (DD) at present (
<bibRefCitation id="EF9142DCFFBEF358FEDF981FFCE1F7F0" author="IUCN Standards &amp; Petitions Committee" box="[325,864,2060,2086]" pageId="1" pageNumber="155" refId="ref2786" refString="IUCN Standards and Petitions Committee. (2022) Guidelines for Using the IUCN Red List Categories and Criteria. Version 15.1. Prepared by the Standards and Petitions Committee. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed:" type="url" year="2022">IUCN Standards and Petitions Committee 2022</bibRefCitation>
).
</paragraph>
<caption id="DF7F6FA5FFBDF35BFF12972BFD5AF843" ID-DOI="http://doi.org/10.5281/zenodo.10376852" ID-Zenodo-Dep="10376852" httpUri="https://zenodo.org/record/10376852/files/figure.png" pageId="2" pageNumber="156" startId="2.[136,229,1848,1870]" targetBox="[164,1423,190,1824]" targetPageId="2" targetType="figure">
<paragraph id="8BBF3F2DFFBDF35BFF12972BFD5AF843" blockId="2.[136,1452,1848,1942]" pageId="2" pageNumber="156">
<emphasis id="B974E33FFFBDF35BFF12972BFF7CF89B" bold="true" box="[136,253,1848,1870]" pageId="2" pageNumber="156">FIGURE 1.</emphasis>
<taxonomicName id="4C0044AEFFBDF35BFE98972BFE5EF89B" authorityName="B. M. Wang &amp; Y. H. Tong" authorityYear="2023" box="[258,479,1848,1869]" class="Magnoliopsida" family="Gesneriaceae" genus="Primulina" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="156" phylum="Tracheophyta" rank="species" species="liangwaniae">
<emphasis id="B974E33FFFBDF35BFE98972BFE5EF89B" box="[258,479,1848,1869]" italics="true" pageId="2" pageNumber="156">Primulina liangwaniae</emphasis>
</taxonomicName>
. A. Habitat; B. Flowering plant, top view; C. Flowering plant, side view; D. Inflorescence, front view; E. Inflorescence, side view; F. Opened corolla; G. Stamens; H. Disc, part of ovary and three calyx lobes; I. Stigma and part of style; J. Capsule with persistent calyx. Photographed by Yi-Hua Tong.
</paragraph>
</caption>
<paragraph id="8BBF3F2DFFBDF35BFF2797D7FBC7F7F0" blockId="2.[136,1452,1988,2087]" pageId="2" pageNumber="156">
<emphasis id="B974E33FFFBDF35BFF2797D7FD5FF808" bold="true" box="[189,734,1988,2014]" pageId="2" pageNumber="156">
Additional specimens examined (
<typeStatus id="54BB818FFFBDF35BFDC597D7FD50F808" box="[607,721,1988,2014]" pageId="2" pageNumber="156" type="paratype">paratype</typeStatus>
):
</emphasis>
—
<collectingCountry id="F3177FBDFFBDF35BFD6697D7FCD6F808" box="[764,855,1988,2014]" name="China" pageId="2" pageNumber="156">CHINA</collectingCountry>
.
<collectingRegion id="49C4F1CFFFBDF35BFCF397D7FBE2F808" box="[873,1123,1988,2014]" country="China" name="Guangdong" pageId="2" pageNumber="156">Guangdong Province</collectingRegion>
: cultivated in South China Botanical Garden, introduced from Yingde City, Huanghua Town, limestone hills, on rocks in shaded places, elev. ca.
<quantity id="4CF892C8FFBDF35BFF12981FFF50F7F1" box="[136,209,2060,2087]" metricMagnitude="2" metricUnit="m" metricValue="5.279999999999999" pageId="2" pageNumber="156" unit="m" value="528.0">528 m</quantity>
,
<date id="FFBE19EDFFBDF35BFF47981FFEF3F7F0" box="[221,370,2060,2087]" pageId="2" pageNumber="156" value="2022-05-22">22 May 2022</date>
(fr.),
<emphasis id="B974E33FFFBDF35BFE28981EFC66F7F0" box="[434,999,2060,2087]" italics="true" pageId="2" pageNumber="156">Y.H. Tong, J.B. Ni, B.M. Wang &amp; J. Cai TYH-2569</emphasis>
(IBSC).
</paragraph>
</subSubSection>
<subSubSection id="C31A6CA6FFBCF35AFF27908DFCE5FDBE" pageId="3" pageNumber="157" type="discussion">
<paragraph id="8BBF3F2DFFBCF35AFF27908DFCE5FDBE" blockId="3.[136,1452,158,617]" pageId="3" pageNumber="157">
<emphasis id="B974E33FFFBCF35AFF27908DFEC4FF6E" bold="true" box="[189,325,158,184]" pageId="3" pageNumber="157">Discussion:</emphasis>
—
<taxonomicName id="4C0044AEFFBCF35AFEFB908DFDE4FF6E" authorityName="B. M. Wang &amp; Y. H. Tong" authorityYear="2023" box="[353,613,158,184]" class="Magnoliopsida" family="Gesneriaceae" genus="Primulina" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="157" phylum="Tracheophyta" rank="species" species="liangwaniae">
<emphasis id="B974E33FFFBCF35AFEFB908DFDE4FF6E" box="[353,613,158,184]" italics="true" pageId="3" pageNumber="157">Primulina liangwaniae</emphasis>
</taxonomicName>
has high ornamental values due to its adorable and relatively small habit and beautiful flowers. This species and its close relative,
<taxonomicName id="4C0044AEFFBCF35AFC8D90D0FC26FF0A" box="[791,935,194,220]" class="Magnoliopsida" family="Gesneriaceae" genus="Primulina" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="157" phylum="Tracheophyta" rank="species" species="versicolor">
<emphasis id="B974E33FFFBCF35AFC8D90D0FC26FF0A" box="[791,935,194,220]" italics="true" pageId="3" pageNumber="157">P. versicolor</emphasis>
</taxonomicName>
, are both distributed in the same county, i.e., Yingde, but in different towns, viz. Huanghua Town and Xiniu Town, and their
<typeStatus id="54BB818FFFBCF35AFB9090F4FBBBFED7" box="[1034,1082,231,257]" pageId="3" pageNumber="157">type</typeStatus>
localities are not far (ca.
<quantity id="4CF892C8FFBCF35AFAC190F5FA25FED6" box="[1371,1444,230,256]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="157" unit="km" value="30.0">30 km</quantity>
) from each other. Besides of the differences mentioned in the diagnosis section, a more detailed comparison between the two species is provided in
<tableCitation id="C6820A96FFBCF35AFE34913DFD83FE9E" box="[430,514,302,328]" captionStart="TABLE 1" captionStartId="3.[136,224,664,688]" captionTargetPageId="3" captionText="TABLE 1 A comparison between Primulina liangwaniae and P. versicolor. The data of P. versicolor is from Pan et al. (2016), and the line drawing and colour plate of P. versicolor in that article are also consulted." pageId="3" pageNumber="157">Table 1</tableCitation>
. It is worth mentioning that their flowering times are also different: this new species flowers from late March to early April, while
<taxonomicName id="4C0044AEFFBCF35AFDE49140FC8BFEBA" box="[638,778,338,364]" class="Magnoliopsida" family="Gesneriaceae" genus="Primulina" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="157" phylum="Tracheophyta" rank="species" species="versicolor">
<emphasis id="B974E33FFFBCF35AFDE49140FC8BFEBA" box="[638,778,338,364]" italics="true" pageId="3" pageNumber="157">P. versicolor</emphasis>
</taxonomicName>
flowers from late June to early July (
<bibRefCitation id="EF9142DCFFBCF35AFB389141FACCFEBA" author="Pan, B. &amp; Wang, B. M. &amp; He, J. Y. &amp; Wen, F." box="[1186,1357,338,364]" pageId="3" pageNumber="157" pagination="25 - 37" refId="ref2990" refString="Pan, B., Wang, B. M., He, J. Y. &amp; Wen, F. (2016) Primulina versicolor and P. alutacea spp. nov. (Gesneriaceae), two new species with yellow flowers from Northern Guangdong, China. Edinburgh Journal of Botany 73 (1): 25 - 37. https: // doi. org / 10.1017 / S 0960428615000268" type="journal article" year="2016">
Pan
<emphasis id="B974E33FFFBCF35AFB499140FA8BFEBA" box="[1235,1290,338,364]" italics="true" pageId="3" pageNumber="157">et al.</emphasis>
2016
</bibRefCitation>
). Except these two species, there are also other four sympatrically distributed congeneric species, viz.
<taxonomicName id="4C0044AEFFBCF35AFB0D9164FE15FE62" authority="Z. L. Ning, M. Kang &amp; X. Y. Zhuang" authorityName="Z. L. Ning, M. Kang &amp; X. Y. Zhuang" class="Magnoliopsida" family="Gesneriaceae" genus="Primulina" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="157" phylum="Tracheophyta" rank="species" species="yingdeensis">
<emphasis id="B974E33FFFBCF35AFB0D9164FAB7FE46" box="[1175,1334,374,400]" italics="true" pageId="3" pageNumber="157">P. yingdeensis</emphasis>
Z.L.Ning, M.Kang &amp; X.Y.Zhuang
</taxonomicName>
in
<bibRefCitation id="EF9142DCFFBCF35AFE219189FD44FE62" author="Ning, Z. L. &amp; Pan, B. &amp; Wen, F. &amp; Kang, M. &amp; Zhuang, X. Y." box="[443,709,410,436]" pageId="3" pageNumber="157" pagination="399 - 409" refId="ref2842" refString="Ning, Z. L., Pan, B., Wen, F., Kang, M. &amp; Zhuang, X. Y. (2016) Primulina yingdeensis, a new species from Guangdong, China, and P. rosulata, a new combination (Gesneriaceae), based on morphological and molecular evidence. Willdenowia 46 (3): 399 - 409. https: // doi. org / 10.3372 / wi. 46.46308" type="journal article" year="2016">
Ning
<emphasis id="B974E33FFFBCF35AFE619188FDB4FE62" box="[507,565,410,436]" italics="true" pageId="3" pageNumber="157">et al.</emphasis>
(2016: 401)
</bibRefCitation>
,
<taxonomicName id="4C0044AEFFBCF35AFD489188FB02FE63" authority="Z. L. Ning &amp; M. Kang" authorityName="Z. L. Ning &amp; M. Kang" box="[722,1155,410,437]" class="Magnoliopsida" family="Gesneriaceae" genus="Primulina" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="157" phylum="Tracheophyta" rank="species" species="qingyuanensis">
<emphasis id="B974E33FFFBCF35AFD489188FC0EFE62" box="[722,911,411,436]" italics="true" pageId="3" pageNumber="157">P. qingyuanensis</emphasis>
Z.L.Ning &amp; M.Kang
</taxonomicName>
in
<bibRefCitation id="EF9142DCFFBCF35AFB319189FA26FE62" author="Ning, Z. L. &amp; Wang, J. &amp; Smith, J. F. &amp; Kang, M." box="[1195,1447,410,436]" pageId="3" pageNumber="157" pagination="48 - 52" refId="ref2925" refString="Ning, Z. L., Wang, J., Smith, J. F. &amp; Kang, M. (2013) Primulina qingyuanensis (Gesneriaceae), a new species from limestone areas in Guangdong, China. Phytotaxa 137 (1): 48 - 52. https: // doi. org / 10.11646 / phytotaxa. 137.1.5" type="journal article" year="2013">
Ning
<emphasis id="B974E33FFFBCF35AFB719188FAA4FE62" box="[1259,1317,410,436]" italics="true" pageId="3" pageNumber="157">et al.</emphasis>
(2013: 48)
</bibRefCitation>
,
<taxonomicName id="4C0044AEFFBCF35AFF1291ACFDC2FE0F" authority="F. Wen, B. Pan &amp; B. M. Wang" authorityName="F. Wen, B. Pan &amp; B. M. Wang" box="[136,579,446,473]" class="Magnoliopsida" family="Gesneriaceae" genus="Primulina" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="157" phylum="Tracheophyta" rank="species" species="alutacea">
<emphasis id="B974E33FFFBCF35AFF1291ACFE85FE0E" box="[136,260,446,472]" italics="true" pageId="3" pageNumber="157">P. alutacea</emphasis>
F.Wen, B.Pan &amp; B.M.Wang
</taxonomicName>
in
<bibRefCitation id="EF9142DCFFBCF35AFDF291ACFCCFFE0E" author="Pan, B. &amp; Wang, B. M. &amp; He, J. Y. &amp; Wen, F." box="[616,846,446,473]" pageId="3" pageNumber="157" pagination="25 - 37" refId="ref2990" refString="Pan, B., Wang, B. M., He, J. Y. &amp; Wen, F. (2016) Primulina versicolor and P. alutacea spp. nov. (Gesneriaceae), two new species with yellow flowers from Northern Guangdong, China. Edinburgh Journal of Botany 73 (1): 25 - 37. https: // doi. org / 10.1017 / S 0960428615000268" type="journal article" year="2016">
Pan
<emphasis id="B974E33FFFBCF35AFD0391ACFD53FE0E" box="[665,722,446,472]" italics="true" pageId="3" pageNumber="157">et al.</emphasis>
(2016: 30)
</bibRefCitation>
and
<taxonomicName id="4C0044AEFFBCF35AFC1F91ACFF6DFE2A" authority="(W. T. Wang 1982: 55) Mich. Moller &amp; A. Weber" authorityName="Mich. Moller &amp; A. Weber" baseAuthorityName="W. T. Wang" baseAuthorityPageNumber="55" baseAuthorityYear="1982" class="Magnoliopsida" family="Gesneriaceae" genus="Primulina" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="157" phylum="Tracheophyta" rank="species" species="lobulata">
<emphasis id="B974E33FFFBCF35AFC1F91ACFC7FFE0E" box="[901,1022,446,472]" italics="true" pageId="3" pageNumber="157">P. lobulata</emphasis>
(W.T.
<bibRefCitation id="EF9142DCFFBCF35AFBDA91ADFB71FE0F" author="Wang, W. T." box="[1088,1264,446,473]" pageId="3" pageNumber="157" pagination="37 - 64" refId="ref3325" refString="Wang, W. T. (1982) Notulae de Gesneriaceis Sinensibus. Bulletin of Botanical Research, Harbin 2 (4): 37 - 64." type="journal article" year="1982">Wang 1982: 55</bibRefCitation>
) Mich.Möller &amp; A.Weber
</taxonomicName>
in
<bibRefCitation id="EF9142DCFFBCF35AFE8891F1FDABFE2A" author="Weber, A. &amp; Middleton, D. J. &amp; Forrest, A. &amp; Kiew, R. &amp; Lim, C. L. &amp; Rafidah, A. R. &amp; Sontag, S. &amp; Triboun, P. &amp; Wei, Y. G. &amp; Yao, T. L. &amp; Moller, M." box="[274,554,482,508]" pageId="3" pageNumber="157" pagination="767 - 790" refId="ref3440" refString="Weber, A., Middleton, D. J., Forrest, A., Kiew, R., Lim, C. L., Rafidah, A. R., Sontag, S., Triboun, P., Wei, Y. G., Yao, T. L. &amp; Moller, M. (2011) Molecular systematics and remodelling of Chirita and associated genera (Gesneriaceae). Taxon 60 (3): 767 - 790. https: // doi. org / 10.1002 / tax. 603012" type="journal article" year="2011">
Weber
<emphasis id="B974E33FFFBCF35AFEF891F0FE1DFE2A" box="[354,412,482,508]" italics="true" pageId="3" pageNumber="157">et al.</emphasis>
(2011: 783)
</bibRefCitation>
, as well as some gesneriads of other genera, e.g.,
<taxonomicName id="4C0044AEFFBCF35AFBFF91F1FE8EFDF6" authority="Hance (1883: 167)" authorityName="Hance" authorityPageNumber="167" authorityYear="1883" class="Magnoliopsida" family="Gesneriaceae" genus="Petrocodon" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="157" phylum="Tracheophyta" rank="species" species="dealbatus">
<emphasis id="B974E33FFFBCF35AFBFF91F1FADDFE2A" box="[1125,1372,482,508]" italics="true" pageId="3" pageNumber="157">Petrocodon dealbatus</emphasis>
<bibRefCitation id="EF9142DCFFBCF35AFAFE91F0FE8EFDF6" author="Hance, H. F." pageId="3" pageNumber="157" pagination="165 - 170" refId="ref2651" refString="Hance, H. F. (1883) New Chinese Cyrtandreae. Journal of Botany 21: 165 - 170." type="journal article" year="1883">Hance (1883: 167)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C0044AEFFBCF35AFED79215FD4EFDF6" authority="C. B. Clarke" authorityName="C. B. Clarke" box="[333,719,518,544]" class="Magnoliopsida" family="Gesneriaceae" genus="Hemiboea" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="157" phylum="Tracheophyta" rank="species" species="follicularis">
<emphasis id="B974E33FFFBCF35AFED79215FDC7FDF6" box="[333,582,518,544]" italics="true" pageId="3" pageNumber="157">Hemiboea follicularis</emphasis>
C.B.Clarke
</taxonomicName>
in
<bibRefCitation id="EF9142DCFFBCF35AFD609215FC1FFDF6" author="Hooker, J. D." box="[762,926,518,544]" pageId="3" pageNumber="157" refId="ref2673" refString="Hooker, J. D. (1888) Hooker's Icones Plantarum, vol. 18. Williams &amp; Norgate, London &amp; Edinburgh, L. Reeve &amp; Co., London." type="book" year="1888">Hooker (1888</bibRefCitation>
: t. 1798), which indicates that this limestone area harbours a relatively high biodiversity of gesneriads. It is believed that more new species will be discovered if botanical investigations are continuously conducted in the future.
</paragraph>
</subSubSection>
</treatment>
</document>
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