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<document id="92E0A7E5CD6CF0484FCB7E6CA551F0DE" ID-DOI="10.11646/phytotaxa.206.1.8" ID-ISSN="1179-3163" ID-Zenodo-Dep="13642276" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725349288233" checkinUser="felipe" docAuthor="Wangwasit, Khanit &amp; Chantaranothai, Pranom" docDate="2015" docId="039F1A09A661FF8EEF8883FAFED9FB8C" docLanguage="en" docName="phytotaxa.206.1.8.pdf" docOrigin="Phytotaxa 206 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.206.1.8" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Hedyotis laotica Wangwasit &amp; Chantar. 2015, comb. nov." docType="treatment" docVersion="3" lastPageNumber="51" masterDocId="FFA66271A665FF8AEF008364FFC8FF87" masterDocTitle="Notes on Hedyotis L. (Rubiaceae) from Thailand" masterLastPageNumber="52" masterPageNumber="47" pageNumber="51" updateTime="1725355194868" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="1ECF872330AD4960EA450F1B450DBE33" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3FC6D1A7873C03274747D59C2E7B27C3">
<mods:title id="7D4FC252A1AE8C8E85843B8AB420805D">Notes on Hedyotis L. (Rubiaceae) from Thailand</mods:title>
</mods:titleInfo>
<mods:name id="C79C0C7FAD82B31B081C04CF17AE6D6C" type="personal">
<mods:role id="4DB5A71DCC8A350A465F3E177B776C62">
<mods:roleTerm id="0A55C291E872FB89B95CB1B1535108FC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E4C96049C0ACC55D8317F1CF5921C25">Wangwasit, Khanit</mods:namePart>
<mods:affiliation id="E55CE9F6EC0E3D048E2AFEC59110DA7F">Queen Sirikit Botanic Garden, The Botanical Garden Organization, Mae Rim, Chiang Mai 50180, Thailand</mods:affiliation>
</mods:name>
<mods:name id="50D8672A318CD5BF1820109FB96390DE" type="personal">
<mods:role id="F986BA50083D9C47C0A8519CEED1AC46">
<mods:roleTerm id="A8808DF84B56B37785117938039E122B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0D2635B82BAB56A547334D9904A7DD01">Chantaranothai, Pranom</mods:namePart>
</mods:name>
<mods:typeOfResource id="144DA69132383B99D28231F8578C4868">text</mods:typeOfResource>
<mods:relatedItem id="5B4301F9925E50EEA50E504E14C7F781" type="host">
<mods:titleInfo id="D7588A1B8768E2281F3EC47A406AC72A">
<mods:title id="60F74F29992365C37774FC6B68850D3A">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="EF9646DE8BAA9D039102B5F35C427CF9">
<mods:date id="F9EEC8876EAB3862A002A0452355E86F">2015</mods:date>
<mods:detail id="9C4F992FB8B8E6C55600AC65194DBD3B" type="pubDate">
<mods:number id="DD08CCBF9B7D90E82D9184EA14CD6B19">2015-05-01</mods:number>
</mods:detail>
<mods:detail id="A52AC23DC5EA85A44D417AE3CEDAD94D" type="volume">
<mods:number id="CA236A7299A08702646C72CE52ACAF80">206</mods:number>
</mods:detail>
<mods:detail id="D2F3D9A17130FB3BD6F601E01AB87DDE" type="issue">
<mods:number id="FA08DE1CF50F001586F1F90DA3E6BF82">1</mods:number>
</mods:detail>
<mods:extent id="4AF1F6D4A53BC24B8DA4109DAC22496D" unit="page">
<mods:start id="A1798E49ADF045D93F8B443905B6F79C">47</mods:start>
<mods:end id="BDE9CF66D5FFBF71E936B4AAE5305107">52</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="9866C324D590BA67CD2191CBD3AE6A94">
<mods:url id="9BC7A71CAC6C89D20CF8A46FFBD18838">http://dx.doi.org/10.11646/phytotaxa.206.1.8</mods:url>
</mods:location>
<mods:classification id="7646964F9F1DEEC052D68D175BCFDAEB">journal article</mods:classification>
<mods:identifier id="BCE8AED25A6D70B3416EB1DA833E4126" type="DOI">10.11646/phytotaxa.206.1.8</mods:identifier>
<mods:identifier id="B490D81D5D2CA0C1EC3A22DCE7A81EF4" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="32042F3B30306310DB15721786FD39EF" type="Zenodo-Dep">13642276</mods:identifier>
</mods:mods>
<treatment id="039F1A09A661FF8EEF8883FAFED9FB8C" ID-DOI="http://doi.org/10.5281/zenodo.13642431" ID-Zenodo-Dep="13642431" LSID="urn:lsid:plazi:treatment:039F1A09A661FF8EEF8883FAFED9FB8C" httpUri="http://treatment.plazi.org/id/039F1A09A661FF8EEF8883FAFED9FB8C" lastPageNumber="51" pageId="4" pageNumber="51">
<subSubSection id="C32CF894A661FF8EEF8883FAFCE1FF3E" box="[136,809,158,185]" pageId="4" pageNumber="51" type="nomenclature">
<paragraph id="8B89AB1FA661FF8EEF8883FAFCE1FF3E" blockId="4.[136,809,158,185]" box="[136,809,158,185]" pageId="4" pageNumber="51">
<heading id="D0C11C73A661FF8EEF8883FAFCE1FF3E" box="[136,809,158,185]" fontSize="11" level="2" pageId="4" pageNumber="51" reason="3">
<taxonomicName id="4C36D09CA661FF8EEF8883FAFD6EFF3E" ID-CoL="3K29K" authority="(Pitard) Wangwasit &amp; Chantar." authorityName="Wangwasit &amp; Chantar." authorityYear="2015" baseAuthorityName="Pitard" baseAuthorityYear="1922" box="[136,678,158,185]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="laotica" status="comb. nov.">
<emphasis id="B942770DA661FF8EEF8883FAFE8AFF3E" bold="true" box="[136,322,158,185]" italics="true" pageId="4" pageNumber="51">Hedyotis laotica</emphasis>
(Pitard) Wangwasit &amp; Chantar.
</taxonomicName>
,
<emphasis id="B942770DA661FF8EEDB183C4FCE1FF3E" box="[689,809,160,185]" italics="true" pageId="4" pageNumber="51">
<taxonomicNameLabel id="A271CA76A661FF8EEDB183C4FCE1FF3E" box="[689,809,160,185]" pageId="4" pageNumber="51" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32CF894A661FF8EEF888380FDE6FE9A" pageId="4" pageNumber="51" type="reference_group">
<paragraph id="8B89AB1FA661FF8EEF888380FDE6FE9A" blockId="4.[136,1451,228,285]" pageId="4" pageNumber="51">
Basionym:
<taxonomicName id="4C36D09CA661FF8EEFF98380FD48FF7E" ID-CoL="4936F" authority="Pitard (1922: 123)" authorityName="Pitard" authorityPageNumber="123" authorityYear="1922" box="[249,640,228,249]" class="Magnoliopsida" family="Rubiaceae" genus="Oldenlandia" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="laotica">
<emphasis id="B942770DA661FF8EEFF98380FE76FF7E" box="[249,446,228,249]" italics="true" pageId="4" pageNumber="51">Oldenlandia laotica</emphasis>
Pitard (1922: 123)
</taxonomicName>
. Type:
<emphasis id="B942770DA661FF8EEDC28380FD12FF7E" bold="true" box="[706,730,228,249]" pageId="4" pageNumber="51">—</emphasis>
<collectingCountry id="F321EB8FA661FF8EEDDA8380FCD4FF7E" box="[730,796,228,249]" name="Laos" pageId="4" pageNumber="51">LAOS</collectingCountry>
. expedition du Mékong, anno 1866–1868,
<emphasis id="B942770DA661FF8EEBCB8380FAFDFF7E" box="[1227,1333,228,249]" italics="true" pageId="4" pageNumber="51">Thorel s.n.</emphasis>
P04009758 (
<typeStatus id="548D15BDA661FF8EEFC4826CFED6FE9A" box="[196,286,264,285]" pageId="4" pageNumber="51" type="lectotype">lectotype</typeStatus>
P, image!, designated here).
</paragraph>
</subSubSection>
<subSubSection id="C32CF894A661FF8EEF888222FC51FDDC" pageId="4" pageNumber="51" type="materials_examined">
<paragraph id="8B89AB1FA661FF8EEF888222FC51FDDC" blockId="4.[136,1452,326,1035]" pageId="4" pageNumber="51">
<materialsCitation id="3B5EA142A661FF8EEF888222FE87FE04" collectingDate="1990-10-19" collectorName="P. Chantaranothai &amp; J. Parnell" country="Thailand" location="Laan Sao Ae" municipality="Phu Phan National Park" pageId="4" pageNumber="51" specimenCount="1" stateProvince="Sakon Nakhon">
<emphasis id="B942770DA661FF8EEF888222FF36FED8" bold="true" box="[136,254,326,351]" pageId="4" pageNumber="51">
<collectingCountry id="F321EB8FA661FF8EEF888222FF31FED8" box="[136,249,326,351]" name="Thailand" pageId="4" pageNumber="51">Thailand</collectingCountry>
:
</emphasis>
—
<collectingRegion id="49F265FDA661FF8EEE1B8222FE0FFED8" box="[283,455,326,351]" country="Thailand" name="Sakon Nakhon" pageId="4" pageNumber="51">Sakon Nakhon</collectingRegion>
:
<collectingMunicipality id="6BED3165A661FF8EEED38222FD21FED8" box="[467,745,326,351]" pageId="4" pageNumber="51">Phu Phan National Park</collectingMunicipality>
,
<location id="8EE9FDC4A661FF8EEDF78222FC45FED8" LSID="urn:lsid:plazi:treatment:039F1A09A661FF8EEF8883FAFED9FB8C:8EE9FDC4A661FF8EEDF78222FC45FED8" box="[759,909,326,351]" country="Thailand" municipality="Phu Phan National Park" name="Laan Sao Ae" pageId="4" pageNumber="51" stateProvince="Sakon Nakhon">Laan Sao Ae</location>
,
<date id="FF888DDFA661FF8EEC9B8222FBFCFED8" box="[923,1076,326,351]" pageId="4" pageNumber="51" value="1990-10-19">
<collectingDate id="EFCC7437A661FF8EEC9B8222FBFCFED8" box="[923,1076,326,351]" pageId="4" pageNumber="51" value="1990-10-19">19 Oct. 1990</collectingDate>
</date>
,
<emphasis id="B942770DA661FF8EEB418222FF1EFE04" italics="true" pageId="4" pageNumber="51">
<collectorName id="26C3CEC9A661FF8EEB418222FADAFED8" box="[1089,1298,326,351]" pageId="4" pageNumber="51">P. Chantaranothai</collectorName>
&amp;
<collectorName id="26C3CEC9A661FF8EEA3B8222FA63FED8" box="[1339,1451,326,351]" pageId="4" pageNumber="51">J. Parnell</collectorName>
90/796
</emphasis>
(K, TCD)
</materialsCitation>
;
<materialsCitation id="3B5EA142A661FF8EEE5B820EFBF0FE04" box="[347,1080,362,387]" collectingDate="2010-11-23" collectorName="K. Wangwasit" country="Thailand" location="Laan Sao Ae" municipality="Phu Phan National Park" pageId="4" pageNumber="51" specimenCount="1" stateProvince="Sakon Nakhon">
the same locality,
<date id="FF888DDFA661FF8EED28820EFD0DFE04" box="[552,709,362,387]" pageId="4" pageNumber="51" value="2010-11-23">
<collectingDate id="EFCC7437A661FF8EED28820EFD0DFE04" box="[552,709,362,387]" pageId="4" pageNumber="51" value="2010-11-23">23 Nov. 2010</collectingDate>
</date>
,
<emphasis id="B942770DA661FF8EEDD1820EFC15FE04" box="[721,989,362,387]" italics="true" pageId="4" pageNumber="51">
<collectorName id="26C3CEC9A661FF8EEDD1820EFCA5FE04" box="[721,877,362,387]" pageId="4" pageNumber="51">K. Wangwasit</collectorName>
101123-7
</emphasis>
(KKU)
</materialsCitation>
;
<materialsCitation id="3B5EA142A661FF8EEB45820EFCCAFE20" collectingDate="1984-09-19" collectorName="N. Fukuoka" country="Thailand" location="Nam Phong" pageId="4" pageNumber="51" specimenCount="1" stateProvince="Khon Kaen">
<collectingRegion id="49F265FDA661FF8EEB45820EFB02FE04" box="[1093,1226,362,387]" country="Thailand" name="Khon Kaen" pageId="4" pageNumber="51">Khon Kaen</collectingRegion>
: along route 2, near
<location id="8EE9FDC4A661FF8EEF8882EAFED8FE20" LSID="urn:lsid:plazi:treatment:039F1A09A661FF8EEF8883FAFED9FB8C:8EE9FDC4A661FF8EEF8882EAFED8FE20" box="[136,272,398,423]" country="Thailand" name="Nam Phong" pageId="4" pageNumber="51" stateProvince="Khon Kaen">Nam Phong</location>
,
<date id="FF888DDFA661FF8EEE1C82EAFE73FE20" box="[284,443,398,423]" pageId="4" pageNumber="51" value="1984-09-19">
<collectingDate id="EFCC7437A661FF8EEE1C82EAFE73FE20" box="[284,443,398,423]" pageId="4" pageNumber="51" value="1984-09-19">19 Sept. 1984</collectingDate>
</date>
,
<emphasis id="B942770DA661FF8EEEC782EAFD66FE20" box="[455,686,398,423]" italics="true" pageId="4" pageNumber="51">
<collectorName id="26C3CEC9A661FF8EEEC782EAFD82FE20" box="[455,586,398,423]" pageId="4" pageNumber="51">N. Fukuoka</collectorName>
T-36206
</emphasis>
(BKF)
</materialsCitation>
;
<materialsCitation id="3B5EA142A661FF8EEC0E82EAFF61FE4C" collectingDate="1931-11-02" collectorName="Put" country="Thailand" location="Bua Yai" pageId="4" pageNumber="51" specimenCount="1" stateProvince="Nakhon Ratchasima">
<collectingRegion id="49F265FDA661FF8EEC0E82EAFC3DFE20" box="[782,1013,398,423]" country="Thailand" name="Nakhon Ratchasima" pageId="4" pageNumber="51">Nakhon Ratchasima</collectingRegion>
:
<location id="8EE9FDC4A661FF8EEB0082EAFB95FE20" LSID="urn:lsid:plazi:treatment:039F1A09A661FF8EEF8883FAFED9FB8C:8EE9FDC4A661FF8EEB0082EAFB95FE20" box="[1024,1117,398,423]" country="Thailand" name="Bua Yai" pageId="4" pageNumber="51" stateProvince="Nakhon Ratchasima">Bua Yai</location>
,
<date id="FF888DDFA661FF8EEB6882EAFB3DFE20" box="[1128,1269,398,423]" pageId="4" pageNumber="51" value="1931-11-02">
<collectingDate id="EFCC7437A661FF8EEB6882EAFB3DFE20" box="[1128,1269,398,423]" pageId="4" pageNumber="51" value="1931-11-02">2 Nov. 1931</collectingDate>
</date>
,
<emphasis id="B942770DA661FF8EEA0182EAFAA0FE20" box="[1281,1384,398,423]" italics="true" pageId="4" pageNumber="51">
<collectorName id="26C3CEC9A661FF8EEA0182EAFAE0FE20" box="[1281,1320,398,423]" pageId="4" pageNumber="51">Put</collectorName>
4278
</emphasis>
(BM, K)
</materialsCitation>
;
<materialsCitation id="3B5EA142A661FF8EEFB482D6FC6BFE4C" box="[180,931,434,459]" collectingDate="1976-11-27" collectorName="C. Phengklai" country="Thailand" elevation="130" location="Burirum" pageId="4" pageNumber="51" specimenCount="1">
<location id="8EE9FDC4A661FF8EEFB482D6FEDFFE4C" LSID="urn:lsid:plazi:treatment:039F1A09A661FF8EEF8883FAFED9FB8C:8EE9FDC4A661FF8EEFB482D6FEDFFE4C" box="[180,279,434,459]" country="Thailand" name="Burirum" pageId="4" pageNumber="51">Burirum</location>
: ca.
<quantity id="4CCE06FAA661FF8EEE4882D6FE59FE4C" box="[328,401,434,459]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="51" unit="m" value="130.0">
<elevation id="001B4C2CA661FF8EEE4882D6FE59FE4C" box="[328,401,434,459]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="51" unit="m" value="130.0">130 m</elevation>
</quantity>
,
<date id="FF888DDFA661FF8EEE9C82D6FDFEFE4C" box="[412,566,434,459]" pageId="4" pageNumber="51" value="1976-11-27">
<collectingDate id="EFCC7437A661FF8EEE9C82D6FDFEFE4C" box="[412,566,434,459]" pageId="4" pageNumber="51" value="1976-11-27">27 Nov. 1976</collectingDate>
</date>
,
<emphasis id="B942770DA661FF8EED4182D6FC98FE4C" box="[577,848,434,459]" italics="true" pageId="4" pageNumber="51">
<collectorName id="26C3CEC9A661FF8EED4182D6FD1BFE4C" box="[577,723,434,459]" pageId="4" pageNumber="51">C. Phengklai</collectorName>
et al. 3482
</emphasis>
(BKF)
</materialsCitation>
;
<materialsCitation id="3B5EA142A661FF8EECAE82D6FA41FE68" collectingDate="1984-10-06" collectorName="G. Murata &amp; C. Phengklai &amp; S. Mitsuta &amp; H. Nagamasu &amp; Nantasan" country="Thailand" elevation="190" location="Cambodian" pageId="4" pageNumber="51" specimenCount="1" stateProvince="Surin">
<collectingRegion id="49F265FDA661FF8EECAE82D6FC27FE4C" box="[942,1007,434,459]" country="Thailand" name="Surin" pageId="4" pageNumber="51">Surin</collectingRegion>
: near
<location id="8EE9FDC4A661FF8EEB3082D6FB79FE4C" LSID="urn:lsid:plazi:treatment:039F1A09A661FF8EEF8883FAFED9FB8C:8EE9FDC4A661FF8EEB3082D6FB79FE4C" box="[1072,1201,434,459]" country="Thailand" name="Cambodian" pageId="4" pageNumber="51" stateProvince="Surin">Cambodian</location>
border, ca.
<locationDeviation id="9A3BB23DA661FF8EEA3582D6FEC0FE68" location="Sangkha" pageId="4" pageNumber="51">
<quantity id="4CCE06FAA661FF8EEA3582D6FAB3FE4C" box="[1333,1403,434,459]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="51" unit="km" value="50.0">50 km</quantity>
SW of Sangkha
</locationDeviation>
,
<quantity id="4CCE06FAA661FF8EEE1282B2FE92FE68" box="[274,346,470,495]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="51" unit="m" value="190.0">
<elevation id="001B4C2CA661FF8EEE1282B2FE92FE68" box="[274,346,470,495]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="51" unit="m" value="190.0">190 m</elevation>
</quantity>
,
<date id="FF888DDFA661FF8EEE6582B2FE21FE68" box="[357,489,470,495]" pageId="4" pageNumber="51" value="1984-10-06">
<collectingDate id="EFCC7437A661FF8EEE6582B2FE21FE68" box="[357,489,470,495]" pageId="4" pageNumber="51" value="1984-10-06">6 Oct. 1984</collectingDate>
</date>
,
<emphasis id="B942770DA661FF8EEEF382B2FAF0FE68" box="[499,1336,470,495]" italics="true" pageId="4" pageNumber="51">
<collectorName id="26C3CEC9A661FF8EEEF382B2FDA3FE68" box="[499,619,470,495]" pageId="4" pageNumber="51">G. Murata</collectorName>
,
<collectorName id="26C3CEC9A661FF8EED7582B2FCC0FE68" box="[629,776,470,495]" pageId="4" pageNumber="51">C. Phengklai</collectorName>
,
<collectorName id="26C3CEC9A661FF8EEC1382B2FC4DFE68" box="[787,901,470,495]" pageId="4" pageNumber="51">S. Mitsuta</collectorName>
,
<collectorName id="26C3CEC9A661FF8EEC9082B2FBE1FE68" box="[912,1065,470,495]" pageId="4" pageNumber="51">H. Nagamasu</collectorName>
&amp; N.
<collectorName id="26C3CEC9A661FF8EEB6982B2FB1CFE68" box="[1129,1236,470,495]" pageId="4" pageNumber="51">Nantasan</collectorName>
T-37669
</emphasis>
(BKF)
</materialsCitation>
;
<materialsCitation id="3B5EA142A661FF8EEA9482B2FD4FFDB0" collectingDate="1984-10-10" collectorName="G. Murata &amp; C. Phengklai &amp; Mitsuta &amp; Nagamasu &amp; Nantasan" country="Thailand" elevation="155" location="Si Sa Ket" pageId="4" pageNumber="51" specimenCount="1" stateProvince="Sisaket">
<collectingRegion id="49F265FDA661FF8EEA9482B2FF10FD94" country="Thailand" name="Sisaket" pageId="4" pageNumber="51">Si Sa Ket</collectingRegion>
: en route from
<collectingRegion id="49F265FDA661FF8EEE80829EFE21FD94" box="[384,489,506,531]" country="Thailand" name="Sisaket" pageId="4" pageNumber="51">Si Sa Ket</collectingRegion>
to
<collectingRegion id="49F265FDA661FF8EED0D829EFD13FD94" box="[525,731,506,531]" country="Thailand" name="Ubon Ratchathani" pageId="4" pageNumber="51">Ubon Ratchathani</collectingRegion>
,
<quantity id="4CCE06FAA661FF8EEDE5829EFCAEFD94" box="[741,870,506,531]" metricMagnitude="2" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="4" pageNumber="51" unit="m" value="155.0" valueMax="180.0" valueMin="130.0">
<elevation id="001B4C2CA661FF8EEDE5829EFCAEFD94" box="[741,870,506,531]" metricMagnitude="2" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="4" pageNumber="51" unit="m" value="155.0" valueMax="180.0" valueMin="130.0">130–180 m</elevation>
</quantity>
,
<date id="FF888DDFA661FF8EEC71829EFBCDFD94" box="[881,1029,506,531]" pageId="4" pageNumber="51" value="1984-10-10">
<collectingDate id="EFCC7437A661FF8EEC71829EFBCDFD94" box="[881,1029,506,531]" pageId="4" pageNumber="51" value="1984-10-10">10 Oct. 1984</collectingDate>
</date>
,
<emphasis id="B942770DA661FF8EEB10829EFDFDFDB0" italics="true" pageId="4" pageNumber="51">
<collectorName id="26C3CEC9A661FF8EEB10829EFB41FD94" box="[1040,1161,506,531]" pageId="4" pageNumber="51">G. Murata</collectorName>
,
<collectorName id="26C3CEC9A661FF8EEB93829EFAE0FD94" box="[1171,1320,506,531]" pageId="4" pageNumber="51">C. Phengklai</collectorName>
, S.
<collectorName id="26C3CEC9A661FF8EEA4F829EFA6FFD94" box="[1359,1447,506,531]" pageId="4" pageNumber="51">Mitsuta</collectorName>
, H.
<collectorName id="26C3CEC9A661FF8EEFAA817AFEEAFDB0" box="[170,290,542,567]" pageId="4" pageNumber="51">Nagamasu</collectorName>
&amp; N.
<collectorName id="26C3CEC9A661FF8EEE66817AFE19FDB0" box="[358,465,542,567]" pageId="4" pageNumber="51">Nantasan</collectorName>
T-38149
</emphasis>
(BKF)
</materialsCitation>
;
<materialsCitation id="3B5EA142A661FF8EED93817AFC5CFDDC" collectingDate="1982-12-09" collectorName="H. Koyama &amp; H. Terao &amp; Wongprasert" country="Thailand" elevation="200" location="Muang" pageId="4" pageNumber="51" specimenCount="1" stateProvince="Ubon Ratchathani">
<collectingRegion id="49F265FDA661FF8EED93817AFCAAFDB0" box="[659,866,542,567]" country="Thailand" name="Ubon Ratchathani" pageId="4" pageNumber="51">Ubon Ratchathani</collectingRegion>
: along route 217,
<locationDeviation id="9A3BB23DA661FF8EEB29817AFB3EFDB0" box="[1065,1270,542,567]" location="Ubon" pageId="4" pageNumber="51">
<quantity id="4CCE06FAA661FF8EEB29817AFBB8FDB0" box="[1065,1136,542,567]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="51" unit="km" value="20.0">20 km</quantity>
W of Ubon
</locationDeviation>
,
<location id="8EE9FDC4A661FF8EEA01817AFA9BFDB0" LSID="urn:lsid:plazi:treatment:039F1A09A661FF8EEF8883FAFED9FB8C:8EE9FDC4A661FF8EEA01817AFA9BFDB0" box="[1281,1363,542,567]" country="Thailand" name="Muang" pageId="4" pageNumber="51" stateProvince="Ubon Ratchathani">Muang</location>
,
<quantity id="4CCE06FAA661FF8EEA5E817AFA6EFDB0" box="[1374,1446,542,567]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="51" unit="m" value="200.0">
<elevation id="001B4C2CA661FF8EEA5E817AFA6EFDB0" box="[1374,1446,542,567]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="51" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<date id="FF888DDFA661FF8EEF888126FEDBFDDC" box="[136,275,578,603]" pageId="4" pageNumber="51" value="1982-12-09">
<collectingDate id="EFCC7437A661FF8EEF888126FEDBFDDC" box="[136,275,578,603]" pageId="4" pageNumber="51" value="1982-12-09">9 Dec. 1982</collectingDate>
</date>
,
<emphasis id="B942770DA661FF8EEE1E8126FC89FDDC" box="[286,833,578,603]" italics="true" pageId="4" pageNumber="51">
<collectorName id="26C3CEC9A661FF8EEE1E8126FE68FDDC" box="[286,416,578,603]" pageId="4" pageNumber="51">H. Koyama</collectorName>
,
<collectorName id="26C3CEC9A661FF8EEEAC8126FDC6FDDC" box="[428,526,578,603]" pageId="4" pageNumber="51">H. Terao</collectorName>
&amp; T.
<collectorName id="26C3CEC9A661FF8EED4E8126FD16FDDC" box="[590,734,578,603]" pageId="4" pageNumber="51">Wongprasert</collectorName>
T-30627
</emphasis>
(BKF)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32CF894A661FF8EEFBD8102FED9FB8C" pageId="4" pageNumber="51" type="distribution">
<paragraph id="8B89AB1FA661FF8EEFBD8102FC09FDF8" blockId="4.[136,1452,326,1035]" box="[189,961,614,639]" pageId="4" pageNumber="51">
<emphasis id="B942770DA661FF8EEFBD8102FE91FDF8" bold="true" box="[189,345,614,639]" pageId="4" pageNumber="51">Distribution:</emphasis>
—
<taxonomicName id="4C36D09CA661FF8EEE758102FDE6FDF8" authorityName="Wangwasit &amp; Chantar." authorityYear="2015" baseAuthorityName="Pitard" baseAuthorityYear="1922" box="[373,558,614,639]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="laotica">
<emphasis id="B942770DA661FF8EEE758102FDE6FDF8" box="[373,558,614,639]" italics="true" pageId="4" pageNumber="51">Hedyotis laotica</emphasis>
</taxonomicName>
is distributed in
<collectingCountry id="F321EB8FA661FF8EEDE98102FC85FDF8" box="[745,845,614,639]" name="Thailand" pageId="4" pageNumber="51">Thailand</collectingCountry>
and
<collectingCountry id="F321EB8FA661FF8EEC838102FC75FDF8" box="[899,957,614,639]" name="Laos" pageId="4" pageNumber="51">Laos</collectingCountry>
.
</paragraph>
<paragraph id="8B89AB1FA661FF8EEFBD81EEFB05FD24" blockId="4.[136,1452,326,1035]" box="[189,1229,650,675]" pageId="4" pageNumber="51">
<emphasis id="B942770DA661FF8EEFBD81EEFEEBFD24" bold="true" box="[189,291,650,675]" pageId="4" pageNumber="51">Habitat:</emphasis>
—It grows in open to semi-shaded grassy fields with sandy soil in deciduous forest.
</paragraph>
<paragraph id="8B89AB1FA661FF8EEFBD81CAFD05FD40" blockId="4.[136,1452,326,1035]" box="[189,717,686,711]" pageId="4" pageNumber="51">
<emphasis id="B942770DA661FF8EEFBD81CAFE8BFD40" bold="true" box="[189,323,686,711]" pageId="4" pageNumber="51">Phenology:</emphasis>
—Flowering in August–December.
</paragraph>
<paragraph id="8B89AB1FA661FF8EEFBD81B6FED9FB8C" blockId="4.[136,1452,326,1035]" pageId="4" pageNumber="51">
<emphasis id="B942770DA661FF8EEFBD81B6FF3DFD6C" bold="true" box="[189,245,722,747]" pageId="4" pageNumber="51">Note</emphasis>
:—Two collections at P,
<emphasis id="B942770DA661FF8EED1481B6FD5DFD6C" box="[532,661,722,747]" italics="true" pageId="4" pageNumber="51">Massie s.n.</emphasis>
(with 4 sheets, P04009756, P04009757, P04009759 and P4009760) and
<emphasis id="B942770DA661FF8EEFBB8192FEFEFC88" box="[187,310,758,783]" italics="true" pageId="4" pageNumber="51">Thorel s.n.</emphasis>
(P04009758), are mentioned in the protologue of
<taxonomicName id="4C36D09CA661FF8EEC858192FC31FC88" authorityName="Wangwasit &amp; Chantar." authorityYear="2015" baseAuthorityName="Pitard" baseAuthorityYear="1922" box="[901,1017,758,783]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="laotica">
<emphasis id="B942770DA661FF8EEC858192FC31FC88" box="[901,1017,758,783]" italics="true" pageId="4" pageNumber="51">O. laotica</emphasis>
</taxonomicName>
. The specimen of Thorel’s collection has flowers and fruits that are well-preserved; and therefore it is selected as the
<typeStatus id="548D15BDA661FF8EEB25807EFB44FCB4" box="[1061,1164,794,819]" pageId="4" pageNumber="51" type="lectotype">lectotype</typeStatus>
here.
<taxonomicName id="4C36D09CA661FF8EEBD5807EFA58FCB4" authorityName="Wangwasit &amp; Chantar." authorityYear="2015" baseAuthorityName="Pitard" baseAuthorityYear="1922" box="[1237,1424,794,819]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="laotica">
<emphasis id="B942770DA661FF8EEBD5807EFA58FCB4" box="[1237,1424,794,819]" italics="true" pageId="4" pageNumber="51">Hedyotis laotica</emphasis>
</taxonomicName>
is placed with
<taxonomicName id="4C36D09CA661FF8EEE10805AFEBCFCD0" authorityName="Linnaeus" authorityYear="1753" box="[272,372,830,855]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="genus">
<emphasis id="B942770DA661FF8EEE10805AFEBCFCD0" box="[272,372,830,855]" italics="true" pageId="4" pageNumber="51">Hedyotis</emphasis>
</taxonomicName>
because it has septicidally and loculicidally dehiscent capsules, dorsiventrally compressed seeds with a longitudinal hilar ridge and punctiform hilum, in accordance with Wikström
<emphasis id="B942770DA661FF8EEB4E8006FB40FCFC" box="[1102,1160,866,891]" italics="true" pageId="4" pageNumber="51">et al.</emphasis>
(2013). Furthermore, the species is similar to
<taxonomicName id="4C36D09CA661FF8EEE6880E2FDD3FC18" authorityName="Walpers" authorityYear="1852" baseAuthorityName="Korthals" baseAuthorityYear="1851" box="[360,539,902,927]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="tetrangularis">
<emphasis id="B942770DA661FF8EEE6880E2FDD3FC18" box="[360,539,902,927]" italics="true" pageId="4" pageNumber="51">H. tetrangularis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C36D09CA661FF8EED5180E2FD0BFC18" baseAuthorityName="Pierre ex Pitard" baseAuthorityYear="1922" box="[593,707,902,927]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="ternata">
<emphasis id="B942770DA661FF8EED5180E2FD0BFC18" box="[593,707,902,927]" italics="true" pageId="4" pageNumber="51">H. ternata</emphasis>
</taxonomicName>
, two species placed in
<taxonomicName id="4C36D09CA661FF8EECC380E2FBEFFC18" authorityName="Linnaeus" authorityYear="1753" box="[963,1063,902,927]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="genus" sensu="stricto">
<emphasis id="B942770DA661FF8EECC380E2FBEFFC18" box="[963,1063,902,927]" italics="true" pageId="4" pageNumber="51">Hedyotis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A271CA76A661FF8EEB2C80E2FBA8FC18" box="[1068,1120,902,927]" pageId="4" pageNumber="51" sensu="stricto">s.str.</taxonomicNameLabel>
by molecular data (Wikström
<emphasis id="B942770DA661FF8EEF8880CEFF0BFC44" box="[136,195,938,963]" italics="true" pageId="4" pageNumber="51">et al.</emphasis>
2013). They are erect herbs with linear leaves, narrowly triangular stipules, corymbiform inflorescences, and heterostylous flowers.
<taxonomicName id="4C36D09CA661FF8EEE8C80AAFD80FC60" authorityName="Wangwasit &amp; Chantar." authorityYear="2015" baseAuthorityName="Pitard" baseAuthorityYear="1922" box="[396,584,974,999]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="laotica">
<emphasis id="B942770DA661FF8EEE8C80AAFD80FC60" box="[396,584,974,999]" italics="true" pageId="4" pageNumber="51">Hedyotis laotica</emphasis>
</taxonomicName>
is distinguishable from both
<taxonomicName id="4C36D09CA661FF8EEC9F80AAFB9EFC60" authorityName="Walpers" authorityYear="1852" baseAuthorityName="Korthals" baseAuthorityYear="1851" box="[927,1110,974,999]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="tetrangularis">
<emphasis id="B942770DA661FF8EEC9F80AAFB9EFC60" box="[927,1110,974,999]" italics="true" pageId="4" pageNumber="51">H. tetrangularis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C36D09CA661FF8EEB9380AAFAC1FC60" baseAuthorityName="Pierre ex Pitard" baseAuthorityYear="1922" box="[1171,1289,974,999]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="ternata">
<emphasis id="B942770DA661FF8EEB9380AAFAC1FC60" box="[1171,1289,974,999]" italics="true" pageId="4" pageNumber="51">H. ternata</emphasis>
</taxonomicName>
by its overall pubescence.
</paragraph>
</subSubSection>
</treatment>
</document>
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