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<mods:title id="E04ED62023CF37B32EC34855CFBC66C2">Description of new species of Pseudamaurops Jeannel, 1948 (Coleoptera: Staphylinidae: Pselaphinae) from Greece and North Macedonia</mods:title>
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<treatment id="038268210E62DC72FF2AFB404AEEF93F" ID-DOI="http://doi.org/10.5281/zenodo.5921253" ID-GBIF-Taxon="159812138" ID-Zenodo-Dep="5921253" LSID="urn:lsid:plazi:treatment:038268210E62DC72FF2AFB404AEEF93F" httpUri="http://treatment.plazi.org/id/038268210E62DC72FF2AFB404AEEF93F" lastPageNumber="405" pageId="6" pageNumber="405">
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<paragraph id="8B94D9370E62DC72FF2AFB404D50FB23" blockId="6.[151,546,1160,1223]" box="[151,546,1160,1187]" pageId="6" pageNumber="405">
<heading id="D0DC6E5B0E62DC72FF2AFB404D50FB23" bold="true" box="[151,546,1160,1187]" fontSize="11" level="1" pageId="6" pageNumber="405" reason="1">
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Key to males of
<taxonomicName id="4C2BA2B40E62DC72FEDDFB414D50FB23" ID-CoL="8XP9L" authorityName="Jeannel" authorityYear="1948" box="[352,546,1160,1187]" class="Insecta" family="Staphylinidae" genus="Pseudamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="B95F05250E62DC72FEDDFB414D50FB23" bold="true" box="[352,546,1160,1187]" italics="true" pageId="6" pageNumber="405">Pseudamaurops</emphasis>
</taxonomicName>
</emphasis>
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</paragraph>
<paragraph id="8B94D9370E62DC72FF2AFB654E90FB47" blockId="6.[151,546,1160,1223]" box="[151,482,1196,1223]" pageId="6" pageNumber="405">
modified from
<bibRefCitation id="EFBAA4C60E62DC72FEFCFB654E90FB47" author="Hlavac, P." box="[321,482,1196,1223]" pageId="6" pageNumber="405" pagination="1 - 4" refId="ref4542" refString="Hlavac, P. (2005) Contribution to the knowledge of the genus Pseudamaurops (Coleoptera: Staphylinidae: Pselaphinae), with a description of three new species from Albania and northern Greece. Entomological Problems, 35, 1 - 4." type="journal article" year="2005">Hlaváč (2005)</bibRefCitation>
</paragraph>
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<keyStep id="30DFC1920E62DC72FF2AFB384AEEFAC1" pageId="6" pageNumber="405">
<paragraph id="8B94D9370E62DC72FF2AFB384AEEFAA4" blockId="6.[151,1436,1265,1727]" pageId="6" pageNumber="405">
<keyLead id="30DA7A020E62DC72FF2AFB384AEEFAA4" pageId="6" pageNumber="405">
1. Mesofemora with three spines, two upper spines small, lower spine much longer, antennal segment X longer than wide,
<collectingCountry id="F33C99A70E62DC72FADAFB3B4E58FAA4" name="Serbia" pageId="6" pageNumber="405">Montenegro</collectingCountry>
............................................................
<taxonomicName id="4C2BA2B40E62DC72FBB9FAC64AEEFAA4" authority="Nonveiller &amp; Pavicevic, 2002" authorityName="Nonveiller &amp; Pavicevic" authorityYear="2002" box="[1028,1436,1294,1316]" class="Insecta" family="Staphylinidae" genus="Pseudamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="calcarata">
<emphasis id="B95F05250E62DC72FBB9FAC64B0AFAA4" bold="true" box="[1028,1144,1294,1316]" italics="true" pageId="6" pageNumber="405">P. calcarata</emphasis>
<bibRefCitation id="EFBAA4C60E62DC72FBC3FAC74AEEFAA4" author="Nonveiller, G. &amp; Pavicevic, D." box="[1150,1436,1294,1316]" pageId="6" pageNumber="405" pagination="435 - 442" refId="ref4738" refString="Nonveiller, G. &amp; Pavicevic, D. (2002) Une nouvelle espece de Pseudamaurops Jeannel, 1948 du Montenegro, et remarques sur les genres voisins (Coleoptera: Pselaphinae: Amaruopini). Annales de la Societe Entomologique de France, New Series, 38, 435 - 442. https: // doi. org / 10.1080 / 00379271.2002.10697353" type="journal article" year="2002">Nonveiller &amp; Pavićević, 2002</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="8B94D9370E62DC72FF2AFAE54AEEFAC1" blockId="6.[151,1436,1265,1727]" box="[151,1436,1324,1346]" pageId="6" pageNumber="405">
<keyLead id="30DA7A020E62DC72FF2AFAE54AEEFAC1" box="[151,1436,1324,1346]" pageId="6" pageNumber="405">- Mesofemora with 0-1 spine............................................................................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DFC1920E62DC72FF2AFA804AE7FAFC" pageId="6" pageNumber="405">
<paragraph id="8B94D9370E62DC72FF2AFA804AEEFADE" blockId="6.[151,1436,1265,1727]" box="[151,1436,1353,1375]" pageId="6" pageNumber="405">
<keyLead id="30DA7A020E62DC72FF2AFA804AEEFADE" box="[151,1436,1353,1375]" pageId="6" pageNumber="405">
2 Antennal club slender, antennomeres IX and X elongate, (
<figureCitation id="1310C5B20E62DC72FCB2FA804C26FADF" box="[783,852,1353,1375]" captionStart="FIGURE 1" captionStartId="2.[151,250,1523,1548]" captionTargetBox="[160,1422,451,1495]" captionTargetId="figure@2.[151,1436,445,1503]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Pseudamaurops—a) P. muellerianus (male, holotype); b) P. hristovskii sp. n. (male, holotype). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3516825" httpUri="https://zenodo.org/record/3516825/files/figure.png" pageId="6" pageNumber="405">Fig. 1b</figureCitation>
),
<collectingCountry id="F33C99A70E62DC72FCDAFA804CA0FADF" box="[871,978,1353,1375]" name="Macedonia" pageId="6" pageNumber="405">Macedonia</collectingCountry>
.......................
<emphasis id="B95F05250E62DC72FB5FFA804AEEFADE" bold="true" box="[1250,1436,1353,1375]" pageId="6" pageNumber="405">
<taxonomicName id="4C2BA2B40E62DC72FB5FFA804A12FADF" ID-CoL="8Y8RZ" authority="Bekchiev &amp; Hlaváč, 2019" authorityName="Bekchiev &amp; Hlaváč" authorityYear="2019" box="[1250,1376,1353,1375]" class="Insecta" family="Staphylinidae" genus="Pseudamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="hristovskii" status="sp. nov.">
<emphasis id="B95F05250E62DC72FB5FFA804A12FADF" bold="true" box="[1250,1376,1353,1375]" italics="true" pageId="6" pageNumber="405">P. hristovskii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB85E0E62DC72FADBFA804AEEFADE" box="[1382,1436,1353,1374]" pageId="6" pageNumber="405" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B94D9370E62DC72FF2AFAAE4AE7FAFC" blockId="6.[151,1436,1265,1727]" box="[151,1429,1382,1405]" pageId="6" pageNumber="405">
<keyLead id="30DA7A020E62DC72FF2AFAAE4AE7FAFC" box="[151,1429,1382,1405]" pageId="6" pageNumber="405">
- Antennal club robust, antennomeres IX and X strongly transverse (
<figureCitation id="1310C5B20E62DC72FCD8FAAF4CDAFAFD" box="[869,936,1382,1405]" captionStart="FIGURE 1" captionStartId="2.[151,250,1523,1548]" captionTargetBox="[160,1422,451,1495]" captionTargetId="figure@2.[151,1436,445,1503]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Pseudamaurops—a) P. muellerianus (male, holotype); b) P. hristovskii sp. n. (male, holotype). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3516825" httpUri="https://zenodo.org/record/3516825/files/figure.png" pageId="6" pageNumber="405">Fig. 1a</figureCitation>
)........................................
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DFC1920E62DC72FF2AFA4D4AEEFA54" pageId="6" pageNumber="405">
<paragraph id="8B94D9370E62DC72FF2AFA4D4AEEFA1A" blockId="6.[151,1436,1265,1727]" box="[151,1436,1412,1434]" pageId="6" pageNumber="405">
<keyLead id="30DA7A020E62DC72FF2AFA4D4AEEFA1A" box="[151,1436,1412,1434]" pageId="6" pageNumber="405">3. Pronotum with lateral carinae thin, short or missing.......................................................... 5</keyLead>
</paragraph>
<paragraph id="8B94D9370E62DC72FF2AFA6B4AEEFA54" blockId="6.[151,1436,1265,1727]" pageId="6" pageNumber="405">
<keyLead id="30DA7A020E62DC72FF2AFA6B4AEEFA54" pageId="6" pageNumber="405">- Pronotum with two well defined, anteriorly convergent or parallel lateral carinae, length of carinae exceeding half of pronotal length.............................................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DFC1920E62DC72FF2AFA154AEEF98E" pageId="6" pageNumber="405">
<paragraph id="8B94D9370E62DC72FF2AFA154AEEFA72" blockId="6.[151,1436,1265,1727]" box="[151,1436,1499,1522]" pageId="6" pageNumber="405">
<keyLead id="30DA7A020E62DC72FF2AFA154AEEFA72" box="[151,1436,1499,1522]" pageId="6" pageNumber="405">
4 Pronotal carinae anteriorly convergent.
<collectingCountry id="F33C99A70E62DC72FDE0FA154DDBFA72" box="[605,681,1500,1522]" name="Albania" pageId="6" pageNumber="405">Albania</collectingCountry>
.......................................
<taxonomicName id="4C2BA2B40E62DC72FB3CFA154AEEFA72" ID-CoL="4NJPN" authority="Hlavac, 2005" authorityName="Hlavac" authorityYear="2005" box="[1153,1436,1499,1522]" class="Insecta" family="Staphylinidae" genus="Pseudamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="transversalis">
<emphasis id="B95F05250E62DC72FB3CFA154A64FA71" bold="true" box="[1153,1302,1499,1521]" italics="true" pageId="6" pageNumber="405">P. transversalis</emphasis>
<bibRefCitation id="EFBAA4C60E62DC72FAA1FA154AEEFA72" author="Hlavac, P." box="[1308,1436,1500,1522]" pageId="6" pageNumber="405" pagination="1 - 4" refId="ref4542" refString="Hlavac, P. (2005) Contribution to the knowledge of the genus Pseudamaurops (Coleoptera: Staphylinidae: Pselaphinae), with a description of three new species from Albania and northern Greece. Entomological Problems, 35, 1 - 4." type="journal article" year="2005">Hlaváč, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B94D9370E62DC72FF2AFA334AEEF98E" blockId="6.[151,1436,1265,1727]" box="[151,1436,1529,1551]" pageId="6" pageNumber="405">
<keyLead id="30DA7A020E62DC72FF2AFA334AEEF98E" box="[151,1436,1529,1551]" pageId="6" pageNumber="405">
- Pronotal carinae anteriorly parallel,
<collectingCountry id="F33C99A70E62DC72FD81FA304DF2F98E" box="[572,640,1529,1550]" name="Greece" pageId="6" pageNumber="405">Greece</collectingCountry>
....................................................
<emphasis id="B95F05250E62DC72FB48FA304AEEF98E" bold="true" box="[1269,1436,1529,1551]" pageId="6" pageNumber="405">
<taxonomicName id="4C2BA2B40E62DC72FB48FA304A12F98F" ID-CoL="8Y8S2" authority="Bekchiev &amp; Hlaváč, 2019" authorityName="Bekchiev &amp; Hlaváč" authorityYear="2019" box="[1269,1376,1529,1551]" class="Insecta" family="Staphylinidae" genus="Pseudamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="tymficus" status="sp. nov.">
<emphasis id="B95F05250E62DC72FB48FA304A12F98F" bold="true" box="[1269,1376,1529,1551]" italics="true" pageId="6" pageNumber="405">P. tymficus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB85E0E62DC72FADBFA304AEEF98E" box="[1382,1436,1529,1550]" pageId="6" pageNumber="405" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DFC1920E62DC72FF2AF9DF4AEEF9CA" pageId="6" pageNumber="405">
<paragraph id="8B94D9370E62DC72FF2AF9DF4AEEF9AD" blockId="6.[151,1436,1265,1727]" box="[151,1436,1558,1581]" pageId="6" pageNumber="405">
<keyLead id="30DA7A020E62DC72FF2AF9DF4AEEF9AD" box="[151,1436,1558,1581]" pageId="6" pageNumber="405">
5. Mesofemora with one spine at middle,
<collectingCountry id="F33C99A70E62DC72FDE5F9DF4DD6F9AC" box="[600,676,1558,1580]" name="Albania" pageId="6" pageNumber="405">Albania</collectingCountry>
......................................
<taxonomicName id="4C2BA2B40E62DC72FBD2F9DE4AEEF9AD" ID-CoL="4NJPH" authority="(Apfelbeck, 1907)" baseAuthorityName="Apfelbeck" baseAuthorityYear="1907" box="[1135,1436,1558,1581]" class="Insecta" family="Staphylinidae" genus="Pseudamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="albanicus">
<emphasis id="B95F05250E62DC72FBD2F9DE4B94F9AC" bold="true" box="[1135,1254,1558,1580]" italics="true" pageId="6" pageNumber="405">P. albanicus</emphasis>
(
<bibRefCitation id="EFBAA4C60E62DC72FB4EF9DF4AE7F9AD" author="Apfelbeck, V." box="[1267,1429,1558,1581]" pageId="6" pageNumber="405" pagination="507 - 526" refId="ref4485" refString="Apfelbeck, V. (1907) Neue Koleopteren, gesammelt wahrend einer im Jahre 1905 mit Subvention der kaiserlichen Akademie der Wissenschaften in Wien durchgefuhrten zoologischen Forschungsreise nach Albanien und Montenegro (II. Serie). Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abteilung I, 116, 507 - 526." type="journal article" year="1907">Apfelbeck, 1907</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B94D9370E62DC72FF2AF9FD4AEEF9CA" blockId="6.[151,1436,1265,1727]" box="[151,1436,1588,1610]" pageId="6" pageNumber="405">
<keyLead id="30DA7A020E62DC72FF2AF9FD4AEEF9CA" box="[151,1436,1588,1610]" pageId="6" pageNumber="405">- Mesofemora with one spine at base....................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DFC1920E62DC72FF2AF9984AEEF904" pageId="6" pageNumber="405">
<paragraph id="8B94D9370E62DC72FF2AF9984AEEF9E7" blockId="6.[151,1436,1265,1727]" box="[151,1436,1617,1639]" pageId="6" pageNumber="405">
<keyLead id="30DA7A020E62DC72FF2AF9984AEEF9E7" box="[151,1436,1617,1639]" pageId="6" pageNumber="405">
6. Antennomeres XXI asymmetrical, triangular,
<collectingCountry id="F33C99A70E62DC72FD1BF9984D98F9E6" box="[678,746,1617,1638]" name="Greece" pageId="6" pageNumber="405">Greece</collectingCountry>
......................................
<taxonomicName id="4C2BA2B40E62DC72FB0CF9984AEEF9E7" ID-CoL="4NJPL" authority="Hlavac, 2005" authorityName="Hlavac" authorityYear="2005" box="[1201,1436,1617,1639]" class="Insecta" family="Staphylinidae" genus="Pseudamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="graecus">
<emphasis id="B95F05250E62DC72FB0CF9984A64F9E6" bold="true" box="[1201,1302,1617,1638]" italics="true" pageId="6" pageNumber="405">P. graecus</emphasis>
<bibRefCitation id="EFBAA4C60E62DC72FAA1F9984AEEF9E7" author="Hlavac, P." box="[1308,1436,1617,1639]" pageId="6" pageNumber="405" pagination="1 - 4" refId="ref4542" refString="Hlavac, P. (2005) Contribution to the knowledge of the genus Pseudamaurops (Coleoptera: Staphylinidae: Pselaphinae), with a description of three new species from Albania and northern Greece. Entomological Problems, 35, 1 - 4." type="journal article" year="2005">Hlaváč, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B94D9370E62DC72FF2AF9A64AEEF904" blockId="6.[151,1436,1265,1727]" box="[151,1436,1646,1668]" pageId="6" pageNumber="405">
<keyLead id="30DA7A020E62DC72FF2AF9A64AEEF904" box="[151,1436,1646,1668]" pageId="6" pageNumber="405">
- Antennomeres IXX trapezoidal,
<collectingCountry id="F33C99A70E62DC72FD9BF9A74D00F904" box="[550,626,1646,1668]" name="Albania" pageId="6" pageNumber="405">Albania</collectingCountry>
.................................................................. 7
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<paragraph id="8B94D9370E62DC72FF2AF9454AEEF922" blockId="6.[151,1436,1265,1727]" box="[151,1436,1675,1698]" pageId="6" pageNumber="405">
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7. Mesofemora with one big spine at base, aedeagus as in
<figureCitation id="1310C5B20E62DC72FD4BF9454B18F921" box="[758,1130,1676,1698]" captionStart="FIGURE 3" captionStartId="5.[151,250,1970,1995]" captionTargetBox="[196,1388,173,1936]" captionTargetId="figure@5.[184,1403,166,1950]" captionTargetPageId="5" captionText="FIGURE 3. Aedeagi of Pseudamaurops, ventral and lateral view (Scale: 0.1 mm)—a) P. albanicus; b) P. muellerianus; c) P. hristovski sp. n.; d) P. tyficus sp. n.; e) P. brachati Hlaváč, 2005 (from Hlaváč, 2005, scale—0.2 mm)" figureDoi="http://doi.org/10.5281/zenodo.3516829" httpUri="https://zenodo.org/record/3516829/files/figure.png" pageId="6" pageNumber="405">
Fig.3b........................
<taxonomicName id="4C2BA2B40E62DC72FBE1F9454B18F921" ID-CoL="4NJPM" authority="(Ravasini, 1923)" baseAuthorityName="Ravasini" baseAuthorityYear="1923" box="[1116,1130,1676,1697]" class="Insecta" family="Staphylinidae" genus="Pseudamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="muellerianus">
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. muellerianus (
<bibRefCitation id="EFBAA4C60E62DC72FABEF9454AE7F922" author="Ravasini, C." box="[1283,1429,1676,1698]" pageId="6" pageNumber="405" pagination="123 - 126" refId="ref4894" refString="Ravasini, C. (1923) Risultati scientifici della spedizione Ravasini-Lona in Albania. VI. Pselafidi e Scidmenidi. Bollettino della Societa Entomologica Italiana, 55, 123 - 126." type="journal article" year="1923">Ravasini, 1923</bibRefCitation>
)
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- Mesofemora with one small spine at base, aedeagus as in
<figureCitation id="1310C5B20E62DC72FCB1F9604BC8F93E" box="[780,1210,1705,1727]" captionStart="FIGURE 3" captionStartId="5.[151,250,1970,1995]" captionTargetBox="[196,1388,173,1936]" captionTargetId="figure@5.[184,1403,166,1950]" captionTargetPageId="5" captionText="FIGURE 3. Aedeagi of Pseudamaurops, ventral and lateral view (Scale: 0.1 mm)—a) P. albanicus; b) P. muellerianus; c) P. hristovski sp. n.; d) P. tyficus sp. n.; e) P. brachati Hlaváč, 2005 (from Hlaváč, 2005, scale—0.2 mm)" figureDoi="http://doi.org/10.5281/zenodo.3516829" httpUri="https://zenodo.org/record/3516829/files/figure.png" pageId="6" pageNumber="405">
Fig.3e.............................
<taxonomicName id="4C2BA2B40E62DC72FB11F9604BC8F93E" ID-CoL="4NJPJ" authority="Hlavac, 2005" authorityName="Hlavac" authorityYear="2005" box="[1196,1210,1705,1726]" class="Insecta" family="Staphylinidae" genus="Pseudamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="brachati">
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. brachati
<bibRefCitation id="EFBAA4C60E62DC72FAA1F9604AEEF93F" author="Hlavac, P." box="[1308,1436,1705,1727]" pageId="6" pageNumber="405" pagination="1 - 4" refId="ref4542" refString="Hlavac, P. (2005) Contribution to the knowledge of the genus Pseudamaurops (Coleoptera: Staphylinidae: Pselaphinae), with a description of three new species from Albania and northern Greece. Entomological Problems, 35, 1 - 4." type="journal article" year="2005">Hlaváč, 2005</bibRefCitation>
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