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<document id="A8B749D8804FC6F6B1CD48F4B7A4E6C9" ID-CLB-Dataset="5629" ID-DOI="10.11646/zootaxa.3788.1" ID-GBIF-Dataset="80d33caa-2379-4f5e-ba98-6d1cc29a7906" ID-ISSN="1175-5326" ID-Zenodo-Dep="4913761" ID-ZooBank="7484832A-025B-41FA-9696-DF7531C01AA4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623174507327" checkinUser="felipe" docAuthor="Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W. &amp; León-Cortés, Jorge L." docDate="2014" docId="C27E87FFFF806117FF14DA7E21A3FA0C" docLanguage="en" docName="zt03788p063.pdf" docOrigin="Zootaxa 3788 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tylodinus coapillensis Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés 2014" docType="treatment" docVersion="7" lastPageNumber="28" masterDocId="3E47FF87FF9B610CFF83DB32243FFFC6" masterDocTitle="A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico" masterLastPageNumber="63" masterPageNumber="1" pageNumber="28" updateTime="1699005316853" updateUser="ExternalLinkService">
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<mods:title id="08867080A3783F550927613171B7DE0A">A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico</mods:title>
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<mods:namePart id="3B16B0820979E2E166B1DDD95B4DFA23">Luna-Cozar, Jesús</mods:namePart>
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<mods:namePart id="41821DE883E85F5612DA239DB6275186">Anderson, Robert S.</mods:namePart>
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<mods:namePart id="6FF62D8AE96562143A75A41D8C2CA7FF">Jones, Robert W.</mods:namePart>
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<mods:namePart id="C91D0F75CA844F3855D9E45F31BC75ED">León-Cortés, Jorge L.</mods:namePart>
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<mods:title id="190D06E027837DB2F240F2C774D2AF87">Zootaxa</mods:title>
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<mods:date id="216D6618324CA671B37FD4B37382333F">2014</mods:date>
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<mods:identifier id="F603D280EDA9507C02CCFC534DC66E48" type="DOI">10.11646/zootaxa.3788.1</mods:identifier>
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<treatment id="C27E87FFFF806117FF14DA7E21A3FA0C" ID-DOI="http://doi.org/10.5281/zenodo.5080218" ID-GBIF-Taxon="183401718" ID-Zenodo-Dep="5080218" LSID="urn:lsid:plazi:treatment:C27E87FFFF806117FF14DA7E21A3FA0C" httpUri="http://treatment.plazi.org/id/C27E87FFFF806117FF14DA7E21A3FA0C" lastPageNumber="28" pageId="27" pageNumber="28">
<subSubSection id="02CD6562FF806117FF14DA7E26DEFE35" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="4A6836E9FF806117FF14DA7E266CFEA0" blockId="27.[151,595,332,358]" box="[151,595,332,358]" pageId="27" pageNumber="28">
<heading id="11208185FF806117FF14DA7E266CFEA0" bold="true" box="[151,595,332,358]" fontSize="11" level="2" pageId="27" pageNumber="28" reason="4">
<emphasis id="78A3EAFBFF806117FF14DA7E266CFEA0" bold="true" box="[151,595,332,358]" pageId="27" pageNumber="28">
<taxonomicName id="8DD74D6AFF806117FF14DA7E259DFEA0" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[151,418,332,358]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="coapillensis">
<emphasis id="78A3EAFBFF806117FF14DA7E259DFEA0" bold="true" box="[151,418,332,358]" italics="true" pageId="27" pageNumber="28">Tylodinus coapillensis</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
<paragraph id="4A6836E9FF806117FF14DAA026DEFE35" blockId="27.[151,1437,402,1219]" pageId="27" pageNumber="28">
<emphasis id="78A3EAFBFF806117FF14DAA02512FE6D" bold="true" box="[151,301,402,427]" pageId="27" pageNumber="28">Recognition.</emphasis>
Species of this species group are distinguished by the following combination of characters: body size 3.0 mm or less, black, clothed with white scales widely scattered over body, denser along lateral sides of pronotal disc, and on vertices of tubercles of elytral declivity.
</paragraph>
</subSubSection>
<subSubSection id="02CD6562FF806117FF14DACC2558FB05" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="4A6836E9FF806117FF14DACC2558FB05" blockId="27.[151,1437,402,1219]" pageId="27" pageNumber="28">
<emphasis id="78A3EAFBFF806117FF14DACC252EFDD1" bold="true" box="[151,273,510,535]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
Body black, approximately 2.2x longer than wide. Head with brown or dark brown scales, vertex with white scales or scales similar in coloration and tonality to head scales, dense to contiguous or overlapping, distributed along vertex or in a patch on middle of vertex; frons concave or moderately concave; rostrum moderately robust, surface with large, deep punctures, longer than or similar to punctures on head, carinate or moderately carinate, apical area punctuate, eyes moderately convex or flush with outline of head. Pronotum in lateral view moderately convex, outline with anterior constriction, without posterior constriction (
<figureCitation id="D2EC2A6CFF806117FB78D9802150FD0D" box="[1275,1391,690,715]" captionStart="FIGURES 1–11. 1–5" captionStartId="8.[151,264,1743,1765]" captionTargetBox="[293,1295,205,1711]" captionTargetId="figure-208@8.[280,1300,195,1713]" captionTargetPageId="8" captionText="FIGURES 1–11. 1–5. Dorsal outline: 1. Generalized Tylodinus, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 2. Tylodinus variabilis, note elytra basal margin straight. 3. Tylodinus ixchel, note elytra basal margin slightly sinuate. 4. Tylodinus immundus, note elytral basal margin sinuate. 5. Tylodinus elongatus, note elytra basal margin very sinuate. 6–11. Lateral outline: 6. Tylodinus complicatus, male, note ventrites 3–5 in same plane as ventrites 1–2. 7. Tylodinus variabilis, male, note ventrites 3–5 not in same plane as ventrites 1–2. 8. Tylodinus coapillensis, female, note ventrites 1–2 strongly convex, ventrites 3–5 not in same plane as ventrites 1–2. 9. Tylodinus zilchi (schematic). 10. Tylodinus sepulturaensis (schematic) 11. Tylodinus rugosus (schematic)." figureDoi="http://doi.org/10.5281/zenodo.4913763" httpUri="https://zenodo.org/record/4913763/files/figure.png" pageId="27" pageNumber="28">Figure 10</figureCitation>
); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond the middle (
<figureCitation id="D2EC2A6CFF806117FF7DD9C82546FCD5" box="[254,377,762,787]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="27" pageNumber="28">Figure 14</figureCitation>
), somewhat hollowed longitudinally; punctures contiguous or dense on lateral sides, progressively more widely spaced, smaller and shallower towards median section, less conspicuous towards anterior and posterior sections (
<figureCitation id="D2EC2A6CFF806117FD83D870264CFC9D" box="[512,627,834,859]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="27" pageNumber="28">Figure 15</figureCitation>
); whitish, moderately dense scales on lateral sides and flanks, anterior area forming thick longitudinal fringe along middle and narrow strip that diverges from middle base to flanks, and with dark brown scales on middle and posterior areas of disc; surface without granules. Elytra black, with basal margin slightly sinuate (
<figureCitation id="D2EC2A6CFF806117FE2ED89C262FFC01" box="[429,528,942,967]" captionStart="FIGURES 1–11. 1–5" captionStartId="8.[151,264,1743,1765]" captionTargetBox="[293,1295,205,1711]" captionTargetId="figure-208@8.[280,1300,195,1713]" captionTargetPageId="8" captionText="FIGURES 1–11. 1–5. Dorsal outline: 1. Generalized Tylodinus, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 2. Tylodinus variabilis, note elytra basal margin straight. 3. Tylodinus ixchel, note elytra basal margin slightly sinuate. 4. Tylodinus immundus, note elytral basal margin sinuate. 5. Tylodinus elongatus, note elytra basal margin very sinuate. 6–11. Lateral outline: 6. Tylodinus complicatus, male, note ventrites 3–5 in same plane as ventrites 1–2. 7. Tylodinus variabilis, male, note ventrites 3–5 not in same plane as ventrites 1–2. 8. Tylodinus coapillensis, female, note ventrites 1–2 strongly convex, ventrites 3–5 not in same plane as ventrites 1–2. 9. Tylodinus zilchi (schematic). 10. Tylodinus sepulturaensis (schematic) 11. Tylodinus rugosus (schematic)." figureDoi="http://doi.org/10.5281/zenodo.4913763" httpUri="https://zenodo.org/record/4913763/files/figure.png" pageId="27" pageNumber="28">Figure 3</figureCitation>
), with dense whitish scales subbasally and on tubercles at elytral declivity and widely scattered on middle of elytra, with patch of white scales on tubercles in front of elytra declivity or not; striae 10 absent. Metasternum flat or moderately concave, median section glabrous; metasternal tooth absent. Abdomen with ventrite 2 glabrous or with transverse line of scales, flat or convex along median line. Hind femur length not reaching elytra apex, femora armed, tibiae subparallel or wider towards base, premucro absent. Female with tergite eight trapezoidal, apex slightly curved, sternite eight doliform, apex not constricted, median membranous area 2/3 as long as basal plate length, spermatheca very shallowly curved, with the outer margin not convex, ramus and nodulus indistinct.
</paragraph>
</subSubSection>
<subSubSection id="02CD6562FF806117FF14DE2421A3FA0C" pageId="27" pageNumber="28" type="key">
<paragraph id="4A6836E9FF806117FF14DE242715FAF6" blockId="27.[151,810,1302,1328]" box="[151,810,1302,1328]" pageId="27" pageNumber="28">
<emphasis id="78A3EAFBFF806117FF14DE242715FAF6" bold="true" box="[151,810,1302,1328]" pageId="27" pageNumber="28">
Key to species of
<taxonomicName id="8DD74D6AFF806117FEEEDE242645FAF6" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[365,634,1302,1328]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="coapillensis">
<emphasis id="78A3EAFBFF806117FEEEDE242645FAF6" bold="true" box="[365,634,1302,1328]" italics="true" pageId="27" pageNumber="28">Tylodinus coapillensis</emphasis>
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species group
</emphasis>
</paragraph>
<keyStep id="F1232E4CFF806117FF14DE6F21A3FA0C" pageId="27" pageNumber="28">
<key id="3E4661D8FF806117FF14DE6F21A3FA0C" pageId="27" pageNumber="28">
<paragraph id="4A6836E9FF806117FF14DE6F21A3FA49" blockId="27.[151,1436,1373,1482]" pageId="27" pageNumber="28">
<keyLead id="F12695DCFF806117FF14DE6F21A3FA49" pageId="27" pageNumber="28">
1 Vertex with two small patches of white scales; interval 2 at elytral declivity with a small spine-shaped tubercle (
<figureCitation id="D2EC2A6CFF806117FAAEDE6F21A7FAB4" box="[1325,1432,1373,1394]" captionStart="FIGURES 12–19. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 12–19. 12. Tylodinus pusillus head a) male, b) female. 13–14. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 15–17. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="27" pageNumber="28">Figures 18</figureCitation>
,
<figureCitation id="D2EC2A6CFF806117FF62DE482520FA49" box="[225,287,1402,1423]" captionStart="FIGURES 52–59" captionStartId="28.[151,264,1974,1996]" captionTargetBox="[201,1375,209,1931]" captionTargetId="figure-55@28.[196,1390,195,1949]" captionTargetPageId="28" captionText="FIGURES 52–59. Dorsal and lateral habitus: 52–53. Tylodinus coapillensis, 54–55. Tylodinus leoncortesi, 56–57. Tylodinus triumforium, 58–59. Tylodinus sepulturaensis." figureDoi="http://doi.org/10.5281/zenodo.4913775" httpUri="https://zenodo.org/record/4913775/files/figure.png" pageId="27" pageNumber="28">52–53</figureCitation>
).........................................................................
<taxonomicName id="8DD74D6AFF806117FB1ADE4821A3FA49" authority="Luna-Cozar" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[1177,1436,1402,1423]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="coapillensis">
<emphasis id="78A3EAFBFF806117FB1ADE48211CFA49" box="[1177,1315,1402,1423]" italics="true" pageId="27" pageNumber="28">T. coapillensis</emphasis>
Luna-Cozar
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4A6836E9FF806117FF14DEA521A3FA0C" blockId="27.[151,1436,1373,1482]" pageId="27" pageNumber="28">
<keyLead id="F12695DCFF806117FF14DEA521A3FA0C" pageId="27" pageNumber="28">
- Vertex with dark brown scales; tubercle on interval 2 at elytral declivity low and rounded, not spine shaped (
<figureCitation id="D2EC2A6CFF806117FB6CDEA52143FA6A" box="[1263,1404,1431,1452]" captionStart="FIGURES 52–59" captionStartId="28.[151,264,1974,1996]" captionTargetBox="[201,1375,209,1931]" captionTargetId="figure-55@28.[196,1390,195,1949]" captionTargetPageId="28" captionText="FIGURES 52–59. Dorsal and lateral habitus: 52–53. Tylodinus coapillensis, 54–55. Tylodinus leoncortesi, 56–57. Tylodinus triumforium, 58–59. Tylodinus sepulturaensis." figureDoi="http://doi.org/10.5281/zenodo.4913775" httpUri="https://zenodo.org/record/4913775/files/figure.png" pageId="27" pageNumber="28">Figures 54–55</figureCitation>
)..................................................................................
<taxonomicName id="8DD74D6AFF806117FB22DE8721A3FA0C" authority="Luna-Cozar" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[1185,1436,1461,1482]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="leoncortesi">
<emphasis id="78A3EAFBFF806117FB22DE87211CFA0C" box="[1185,1315,1461,1482]" italics="true" pageId="27" pageNumber="28">T. leoncortesi</emphasis>
Luna-Cozar
</taxonomicName>
</keyLead>
</paragraph>
</key>
</keyStep>
</subSubSection>
</treatment>
</document>
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