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<document id="1848632AD612F934308FF717C4E01B15" ID-DOI="10.1649/0010-065x-69.2.293" ID-ISSN="1938-4394" ID-Zenodo-Dep="10104861" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630170702411" checkinUser="diego" docAuthor="Krinsky, William L." docDate="2015" docId="123687CBFFFB974FFD0215B6947EFD1C" docLanguage="en" docName="0010-065X-69.2.293.pdf.imf" docOrigin="The Coleopterists Bulletin 69 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-69.2.293" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Rhagodera costaefragmenta Krinsky, new species" docType="treatment" docVersion="3" lastPageNumber="294" masterDocId="EE0FFFB3FFFB974DFFC1104A9636FFE7" masterDocTitle="A New Species ofRhagoderaMannerheim (Coleoptera: Zopheridae: Colydiinae) from San Clemente Island, California" masterLastPageNumber="296" masterPageNumber="293" pageNumber="293" updateTime="1699631155159" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="1DC4CE5BFB3AE65672E9A31292759A66" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9C0BE7ED354014E34A13D8C0AC3354C6">
<mods:title id="358948BFE191F0A93992C75462F480F9">A New Species ofRhagoderaMannerheim (Coleoptera: Zopheridae: Colydiinae) from San Clemente Island, California</mods:title>
</mods:titleInfo>
<mods:name id="7E2169F0678C36522A1BC18DC3CF3154" type="personal">
<mods:role id="024EA022E12016F5330788F5F49A21E7">
<mods:roleTerm id="EDAF4118FC6C229E4CC48FA0F3397AC6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="44B8093DA505D9EA1A22C59D5D69FF36">Krinsky, William L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="9B80300183AFE225109D34982B06B1CC">text</mods:typeOfResource>
<mods:relatedItem id="B209DB11AB901A9A6A12C16AD0ED2CC4" type="host">
<mods:titleInfo id="645EC9D7DC433CFD746AE88816F6CD23">
<mods:title id="EB6D23CF9848460F06382F9507B85663">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="818A4D8EEFB49EF519555389C509E3B5">
<mods:date id="1BE38999472AE2E0CEE7F02CB860D332">2015</mods:date>
<mods:detail id="1063F364937127FA6A7C7ADC8AA4D666" type="pubDate">
<mods:number id="2DD2140F7A38E5B62405EFDE8AFE7B27">2015-06-30</mods:number>
</mods:detail>
<mods:detail id="0223E9C088DBEDFA0AEB7EA6B0B44E9A" type="volume">
<mods:number id="C940BBB53A4071C39692D6E1303633CF">69</mods:number>
</mods:detail>
<mods:detail id="C4CD7E5ED50FB535EAEE7534AFD2A66C" type="issue">
<mods:number id="8BAADC2B43EFEA344A1473E384BF693B">2</mods:number>
</mods:detail>
<mods:extent id="363430D3E97CA406A5D6F0E505728F9D" unit="page">
<mods:start id="2A83E63E1A26798EC6E81018EA70E82E">293</mods:start>
<mods:end id="B22EAB645636B71F0913FD95EB108E31">296</mods:end>
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<mods:location id="7155EEC1969374053E7119553FD6CE71">
<mods:url id="4CE9DF30FF16FB34E5BDA2A978689BC5">http://dx.doi.org/10.1649/0010-065x-69.2.293</mods:url>
</mods:location>
<mods:classification id="4C89F74B118A39C48231CF411F438663">journal article</mods:classification>
<mods:identifier id="8A5FC0AF0C93D1238D7C36ABF9A7889A" type="DOI">10.1649/0010-065x-69.2.293</mods:identifier>
<mods:identifier id="51D00DB77A2A7E2D2EC6E09F4E6C02BE" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="23F661871167C18A9BD6117882AEEF4B" type="Zenodo-Dep">10104861</mods:identifier>
</mods:mods>
<treatment id="123687CBFFFB974FFD0215B6947EFD1C" LSID="urn:lsid:plazi:treatment:123687CBFFFB974FFD0215B6947EFD1C" httpUri="http://treatment.plazi.org/id/123687CBFFFB974FFD0215B6947EFD1C" lastPageId="2" lastPageNumber="294" pageId="0" pageNumber="293">
<subSubSection id="D2856556FFFB974DFD0215B695DDF9CB" pageId="0" pageNumber="293" type="nomenclature">
<paragraph id="9A2036DDFFFB974DFD0215B695DDF9CB" blockId="0.[707,1193,1532,1580]" pageId="0" pageNumber="293">
<emphasis id="A8EBEACFFFFB974DFD0215B6929FF9F5" bold="true" box="[707,1193,1532,1554]" pageId="0" pageNumber="293">
<taxonomicName id="5D9F4D5EFFFB974DFD0215B6921DF9F5" authority="Krinsky" authorityName="Krinsky" box="[707,1067,1532,1554]" class="Insecta" family="Zopheridae" genus="Rhagodera" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="293" phylum="Arthropoda" rank="species" species="costaefragmenta" status="sp. nov.">
<emphasis id="A8EBEACFFFFB974DFD0215B695E4F9F5" bold="true" box="[707,978,1532,1554]" italics="true" pageId="0" pageNumber="293">Rhagodera costaefragmenta</emphasis>
Krinsky
</taxonomicName>
,
<taxonomicNameLabel id="B3D857B4FFFB974DFBF715B7929FF9F5" box="[1078,1193,1533,1554]" pageId="0" pageNumber="293" rank="species">new species</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="02A42A58FFFB974DFC46165D95FCF9CB" box="[903,970,1559,1580]" captionStart="Fig" captionStartId="1.[175,210,945,964]" captionTargetBox="[151,637,193,922]" captionTargetId="figure-645@1.[151,637,193,922]" captionTargetPageId="1" captionText="Fig. 1. Rhagodera costaefragmenta, holotype, dorsal habitus." figureDoi="http://doi.org/10.5281/zenodo.10104865" httpUri="https://zenodo.org/record/10104865/files/figure.png" pageId="0" pageNumber="293">Figs. 1</figureCitation>
,
<figureCitation id="02A42A58FFFB974DFC19165D95D2F9CB" box="[984,996,1559,1580]" captionStart="Figs" captionStartId="2.[197,237,574,593]" captionTargetBox="[173,1193,193,551]" captionTargetId="figure-504@2.[173,1193,193,551]" captionTargetPageId="2" captionText="Figs. 2–4. Comparison of costa 2 in three species of Rhagodera. 2) R. tuberculata; 3) R.interrupta; 4) R. costaefragmenta." figureDoi="http://doi.org/10.5281/zenodo.10104867" httpUri="https://zenodo.org/record/10104867/files/figure.png" pageId="0" pageNumber="293">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D2856556FFFB974CFD1A1608950AFB2E" lastPageId="1" lastPageNumber="294" pageId="0" pageNumber="293" type="materials_examined">
<paragraph id="9A2036DDFFFB974CFD1A1608944CFB04" blockId="0.[707,1193,1602,1730]" lastBlockId="1.[151,639,1044,1729]" lastPageId="1" lastPageNumber="294" pageId="0" pageNumber="293">
<emphasis id="A8EBEACFFFFB974DFD1A16089541F9B0" bold="true" box="[731,887,1602,1623]" pageId="0" pageNumber="293">Type Material.</emphasis>
<materialsCitation id="2AF73C80FFFB974CFC441608944CFB04" collectingDate="1968-08-10" collectingDateMax="1972-11-11" collectingDateMin="1968-08-10" collectionCode="CALIF" collectorName="C. L. Remington &amp; L. S. Matlovsky" county="Three" lastPageId="1" lastPageNumber="294" latitude="32.883335" location="Division of Entomology" longLatPrecision="1208" longitude="-118.51667" municipality="All" pageId="0" pageNumber="293" specimenCount="5" stateProvince="Connecticut" typeStatus="holotype">
H o l ot y pe (se x u n k no w n) labeled “
<collectionCode id="FC8EAE18FFFB974DFCDC1616955EF996" box="[797,872,1628,1649]" pageId="0" pageNumber="293">CALIF</collectionCode>
.: San Clemente Is., indian shell mound, el. 1295′.
<date id="EE21101DFFFB974DFCA4163D95E9F96B" box="[869,991,1655,1676]" pageId="0" pageNumber="293" value="1968-08-10">
<collectingDate id="FE65E9F5FFFB974DFCA4163D95E9F96B" box="[869,991,1655,1676]" pageId="0" pageNumber="293" value="1968-08-10">10 Aug.1968</collectingDate>
</date>
. (
<geoCoordinate id="FFAB501AFFFB974DFC2C163D920BF96B" box="[1005,1085,1655,1677]" degrees="32" direction="north" minutes="53" orientation="latitude" pageId="0" pageNumber="293" precision="925" value="32.883335">32°53′N</geoCoordinate>
,
<geoCoordinate id="FFAB501AFFFB974DFB84163D9294F96B" box="[1093,1186,1655,1677]" degrees="118" direction="west" minutes="31" orientation="longitude" pageId="0" pageNumber="293" precision="925" value="-118.51667">118°31′W</geoCoordinate>
) leg.
<collectorName id="376A530BFFFB974DFD3216D895ADF940" box="[755,923,1682,1703]" pageId="0" pageNumber="293">C.L. Remington</collectorName>
&amp;
<collectorName id="376A530BFFFB974DFC0516D89250F940" box="[964,1126,1682,1703]" pageId="0" pageNumber="293">L.S. Matlovsky</collectorName>
” with author’ s red
<typeStatus id="4524887FFFFB974DFCF516E695B2F926" box="[820,900,1708,1729]" pageId="0" pageNumber="293" type="holotype">holotype</typeStatus>
label.
<typeStatus id="4524887FFFFB974DFC0016E69227F926" box="[961,1041,1708,1729]" pageId="0" pageNumber="293" type="allotype">Allotype</typeStatus>
(female) labeled “
<collectionCode id="FC8EAE18FFFA974CFF60145E96DEFBCE" box="[161,232,1044,1065]" pageId="1" pageNumber="294">CALIF</collectionCode>
.: San Clemente Is., highest terrace,
<quantity id="5D679B38FFFA974CFDF1145E9457FBCE" box="[560,609,1044,1065]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="1" pageNumber="294" unit="mi" value="1.0">1 mi.</quantity>
N. of Pyramid Head.
<date id="EE21101DFFFA974CFE971465942DFBA3" box="[342,539,1071,1092]" pageId="1" pageNumber="294" value="1972-11-11">
<collectingDate id="FE65E9F5FFFA974CFE971465942DFBA3" box="[342,539,1071,1092]" pageId="1" pageNumber="294" value="1972-11-11">11 November 1972</collectingDate>
</date>
leg.
<collectorName id="376A530BFFFA974CFD90146596C8FBB9" pageId="1" pageNumber="294">C.L. Remington</collectorName>
” with author’ s red
<typeStatus id="4524887FFFFA974CFE7A14039430FBB9" box="[443,518,1097,1118]" pageId="1" pageNumber="294" type="allotype">allotype</typeStatus>
label.
<collectingCounty id="73414E51FFFA974CFD8914039448FBB9" box="[584,638,1097,1118]" pageId="1" pageNumber="294">Three</collectingCounty>
<typeStatus id="4524887FFFFA974CFF56142E96DAFB9E" box="[151,236,1124,1145]" pageId="1" pageNumber="294" type="paratype">paratypes</typeStatus>
(female plus two of unknown sex) each with same collection data as allotype plus the author’ s
<typeStatus id="4524887FFFFA974CFF5614D396D1FB49" box="[151,231,1177,1198]" pageId="1" pageNumber="294" type="paratype">paratype</typeStatus>
label.
<collectingMunicipality id="7A44ACA7FFFA974CFEEB14D3977EFB49" box="[298,328,1177,1198]" pageId="1" pageNumber="294">All</collectingMunicipality>
types are deposited in the
<location id="9F406006FFFA974CFD8D14D3976CFB2F" LSID="urn:lsid:plazi:treatment:123687CBFFFB974FFD0215B6947EFD1C:9F406006FFFA974CFD8D14D3976CFB2F" county="Three" latitude="32.883335" longLatPrecision="1208" longitude="-118.51667" municipality="All" name="Division of Entomology" pageId="1" pageNumber="294" stateProvince="Connecticut">Division of Entomology</location>
,
<location id="9F406006FFFA974CFEA414F996EBFB04" LSID="urn:lsid:plazi:treatment:123687CBFFFB974FFD0215B6947EFD1C:9F406006FFFA974CFEA414F996EBFB04" county="Three" latitude="32.883335" longLatPrecision="1208" longitude="-118.51667" municipality="All" name="Peabody Museum of Natural History" pageId="1" pageNumber="294" stateProvince="Connecticut">Peabody Museum of Natural History</location>
,
<location id="9F406006FFFA974CFF291484974AFB04" LSID="urn:lsid:plazi:treatment:123687CBFFFB974FFD0215B6947EFD1C:9F406006FFFA974CFF291484974AFB04" box="[232,380,1230,1251]" county="Three" latitude="32.883335" longLatPrecision="1208" longitude="-118.51667" municipality="All" name="Yale University" pageId="1" pageNumber="294" stateProvince="Connecticut">Yale University</location>
,
<location id="9F406006FFFA974CFE47148497CFFB04" LSID="urn:lsid:plazi:treatment:123687CBFFFB974FFD0215B6947EFD1C:9F406006FFFA974CFE47148497CFFB04" box="[390,505,1230,1251]" county="Three" latitude="32.883335" longLatPrecision="1208" longitude="-118.51667" municipality="All" name="New Haven" pageId="1" pageNumber="294" stateProvince="Connecticut">New Haven</location>
,
<collectingRegion id="585BF83FFFFA974CFDC214849440FB04" box="[515,630,1230,1251]" country="United States of America" name="Connecticut" pageId="1" pageNumber="294">Connecticut</collectingRegion>
.
</materialsCitation>
</paragraph>
<caption id="CEE06655FFFA974CFF6E13FB972DFC3B" ID-DOI="http://doi.org/10.5281/zenodo.10104865" ID-Zenodo-Dep="10104865" httpUri="https://zenodo.org/record/10104865/files/figure.png" pageId="1" pageNumber="294" startId="1.[175,210,945,964]" targetBox="[151,637,193,922]" targetPageId="1">
<paragraph id="9A2036DDFFFA974CFF6E13FB972DFC3B" blockId="1.[151,638,945,988]" pageId="1" pageNumber="294">
<emphasis id="A8EBEACFFFFA974CFF6E13FB96C5FC23" bold="true" box="[175,243,945,964]" pageId="1" pageNumber="294">Fig. 1.</emphasis>
<taxonomicName id="5D9F4D5EFFFA974CFEC813FB9420FC23" box="[265,534,945,964]" class="Insecta" family="Zopheridae" genus="Rhagodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="costaefragmenta">
<emphasis id="A8EBEACFFFFA974CFEC813FB9420FC23" box="[265,534,945,964]" italics="true" pageId="1" pageNumber="294">Rhagodera costaefragmenta</emphasis>
</taxonomicName>
, holotype, dorsal habitus.
</paragraph>
</caption>
<paragraph id="9A2036DDFFFA974CFF6E14A392B2FC3E" blockId="1.[151,639,1044,1729]" lastBlockId="1.[685,1173,194,1730]" pageId="1" pageNumber="294">
<emphasis id="A8EBEACFFFFA974CFF6E14A39725FB19" bold="true" box="[175,275,1257,1278]" pageId="1" pageNumber="294">
<typeStatus id="4524887FFFFA974CFF6E14A39738FB19" box="[175,270,1257,1278]" pageId="1" pageNumber="294" type="holotype">Holotype</typeStatus>
.
</emphasis>
(
<figureCitation id="02A42A58FFFA974CFEE914A3975DFB19" box="[296,363,1257,1278]" captionStart="Fig" captionStartId="1.[175,210,945,964]" captionTargetBox="[151,637,193,922]" captionTargetId="figure-645@1.[151,637,193,922]" captionTargetPageId="1" captionText="Fig. 1. Rhagodera costaefragmenta, holotype, dorsal habitus." figureDoi="http://doi.org/10.5281/zenodo.10104865" httpUri="https://zenodo.org/record/10104865/files/figure.png" pageId="1" pageNumber="294">Fig. 1</figureCitation>
). Length 7.0 mm, width across middle of elytra
<quantity id="5D679B38FFFA974CFEBF154997FFFAFF" box="[382,457,1283,1304]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="294" unit="mm" value="2.7">2.7 mm</quantity>
(2.6X longer than wide).
<collectionCode id="FC8EAE18FFFA974CFF1B155496DEFAD4" box="[218,232,1310,1331]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="294">T</collectionCode>
/W = 0.5. Color reddish brown to piceous; without surface luster.
<emphasis id="A8EBEACFFFFA974CFEAF1572979DFAAA" bold="true" box="[366,427,1336,1357]" pageId="1" pageNumber="294">Head:</emphasis>
Subquadrate (
<quantity id="5D679B38FFFA974CFDF415729448FAAA" box="[565,638,1336,1357]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="294" unit="mm" value="1.5">1.5 mm</quantity>
long); covered with small tubercles, each bearing a short, pale, scale-like seta; anterolateral angles strongly lobed over antennal insertion, elevated smaller lobe above each eye (most of eye visible from above), anterior vertex with triangular array of tubercles (slightly larger than those on rest of head) just anterior to higher distinct protuberance (appearing to be composed of a few fused tubercles). Eye posteriorly removed from antennal insertion. Length of third antennal segment equal to horizontal diameter of eye (
<quantity id="5D679B38FFFA974CFE091617942FF995" box="[456,537,1629,1650]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="294" unit="mm" value="0.4">0.4 mm</quantity>
). Antenna 11-segmented with 3-segmented club, last antennal segment emarginate. Subantennal groove distinct.
<emphasis id="A8EBEACFFFFA974CFF5616E6973FF926" bold="true" box="[151,265,1708,1729]" pageId="1" pageNumber="294">Pronotum:</emphasis>
<quantity id="5D679B38FFFA974CFED516E69754F926" box="[276,354,1708,1729]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="294" unit="mm" value="2.2">2.2 mm</quantity>
wide (1.5X length), lateral margins evenly arcuate, denticulate, apical and basal angles right, anterior margin evenly concave except where it is overlaid by rectangular central discal elevation, lateral margins of elevation interrupted by single notch. Anterior margin of elevation emarginate, posteror margin arcuate. Tubercles and lateral denticles each topped by a short, pale, scalelike seta. Few posterior lateral denticles broader, blunter, and more widely spaced than those forming rest of lateral pronotal margin.
<emphasis id="A8EBEACFFFFA974CFC1111FB9221FE21" bold="true" box="[976,1047,433,454]" pageId="1" pageNumber="294">Elytra:</emphasis>
Surface with sutural costa, 3 discal costae, and marginal costa. Sutural costa and discal costae 1 and 3 prominently elevated at their extreme bases, discal costae interrupted to different degrees: costa 1 well-defined on basal 1/3 of elytron, then with 4 interruptions before distal 1/5 of elytron where a well-defined portion begins and continues to the apex and joins apical margin; costa 2 absent and replaced by irregularly spaced, non-linearly arranged tubercles for 4/5 length of elytron, followed by a well-defined distal portion that ends before reaching apical margin; costa 3 well-defined and continuous from base to 4/5 length of elytron, ending at about same level at which distal segments of costae 1 and 2 begin. Intervals between costae with 2 rows of large, round, well-separated punctures.
<emphasis id="A8EBEACFFFFA974CFD27133F951AFC6D" bold="true" box="[742,812,885,906]" pageId="1" pageNumber="294">Venter:</emphasis>
Procoxal cavities widely open behind. Epipleural fold ends near middle of penultimate ventrite, where an irregular carina composed of small tubercles begins and continues to apex of elytron.
</paragraph>
<paragraph id="9A2036DDFFFA974CFD041395950AFB2E" blockId="1.[685,1173,194,1730]" pageId="1" pageNumber="294">
<emphasis id="A8EBEACFFFFA974CFD0413959515FC13" bold="true" box="[709,803,991,1012]" pageId="1" pageNumber="294">
<typeStatus id="4524887FFFFA974CFD0413959529FC13" box="[709,799,991,1012]" pageId="1" pageNumber="294" type="allotype">Allotype</typeStatus>
.
</emphasis>
Female (ovipositor extended). Length
<quantity id="5D679B38FFFA974CFD6C13B094CCFBE8" box="[685,762,1018,1039]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="294" unit="mm" value="6.5">6.5 mm</quantity>
; width
<quantity id="5D679B38FFFA974CFC8513B095A6FBE8" box="[836,912,1018,1039]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="294" unit="mm" value="2.2">2.2 mm</quantity>
. As
<typeStatus id="4524887FFFFA974CFC7F13B09239FBE8" box="[958,1039,1018,1039]" pageId="1" pageNumber="294" type="holotype">holotype</typeStatus>
except in the following respects: color mostly piceous, with reddish brown on pronotal margins and legs. Antennal segment 3 equals 1.5X horizontal diameter of eye. Sutural costae less prominent at base. Distal portion of costae 1 and 2 less well-defined, shorter, and appearing as clumped tubercles, not reaching apical margin.
</paragraph>
</subSubSection>
<subSubSection id="D2856556FFFA974CFD041484952BFA7A" pageId="1" pageNumber="294" type="description">
<paragraph id="9A2036DDFFFA974CFD041484952BFA7A" blockId="1.[685,1173,194,1730]" pageId="1" pageNumber="294">
<emphasis id="A8EBEACFFFFA974CFD041484951CFB04" bold="true" box="[709,810,1230,1251]" pageId="1" pageNumber="294">Variation.</emphasis>
Female and two of unknown sex (
<typeStatus id="4524887FFFFA974CFD7514A39521FB19" box="[692,791,1257,1278]" pageId="1" pageNumber="294" type="paratype">paratypes</typeStatus>
). Length
<quantity id="5D679B38FFFA974CFCB914A395FEFB19" box="[888,968,1257,1278]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="294" unit="mm" value="6.5">6.5 mm</quantity>
; width
<quantity id="5D679B38FFFA974CFBD414A392A6FB19" box="[1045,1168,1256,1278]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="1" pageNumber="294" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.2–2.4 mm</quantity>
. Color piceous with some parts reddish brown (mostly pronotal lateral margins and legs). Antennal segment 3 equals 1.2X horizontal diameter of eye. Poorly defined distal portions of costae 1 and 2 and costa 1 not reaching apical margin similar to
<typeStatus id="4524887FFFFA974CFD0915C2952FFA7A" box="[712,793,1416,1437]" pageId="1" pageNumber="294" type="allotype">allotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="D2856556FFFA974CFD0415E995DEF9C5" pageId="1" pageNumber="294" type="etymology">
<paragraph id="9A2036DDFFFA974CFD0415E995DEF9C5" blockId="1.[685,1173,194,1730]" pageId="1" pageNumber="294">
<emphasis id="A8EBEACFFFFA974CFD0415E9950EFA5F" bold="true" box="[709,824,1443,1464]" pageId="1" pageNumber="294">Etymology.</emphasis>
The name is derived from the Latin
<emphasis id="A8EBEACFFFFA974CFD6C15F794DFFA34" box="[685,745,1469,1491]" italics="true" pageId="1" pageNumber="294">costae</emphasis>
(lit. “ribs”), referring to the raised ridges on the elytra, and the adjective
<emphasis id="A8EBEACFFFFA974CFC07159D920FFA0A" box="[966,1081,1495,1517]" italics="true" pageId="1" pageNumber="294">fragmentum</emphasis>
meaning “fragment” or “remnant”. The name is descriptive of the fragmented elytral costae.
</paragraph>
</subSubSection>
<subSubSection id="D2856556FFFA974FFD04166D947EFD1C" lastPageId="2" lastPageNumber="295" pageId="1" pageNumber="294" type="diagnosis">
<paragraph id="9A2036DDFFFA974FFD04166D947EFD1C" blockId="1.[685,1173,194,1730]" lastBlockId="2.[173,659,635,763]" lastPageId="2" lastPageNumber="295" pageId="1" pageNumber="294">
<emphasis id="A8EBEACFFFFA974CFD04166D9518F9DB" bold="true" box="[709,814,1575,1596]" pageId="1" pageNumber="294">Diagnosis.</emphasis>
<taxonomicName id="5D9F4D5EFFFA974CFCFD166D926CF9DA" box="[828,1114,1575,1597]" class="Insecta" family="Zopheridae" genus="Rhagodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="costaefragmenta">
<emphasis id="A8EBEACFFFFA974CFCFD166D926CF9DA" box="[828,1114,1575,1597]" italics="true" pageId="1" pageNumber="294">Rhagodera costaefragmenta</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="5D9F4D5EFFFA974CFC9D160895E1F9BF" authorityName="Stephan" authorityYear="1989" box="[860,983,1602,1624]" class="Insecta" family="Zopheridae" genus="Rhagodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="interrupta">
<emphasis id="A8EBEACFFFFA974CFC9D160895E1F9BF" box="[860,983,1602,1624]" italics="true" pageId="1" pageNumber="294">R. interrupta</emphasis>
</taxonomicName>
and
<taxonomicName id="5D9F4D5EFFFA974CFBC6160892B8F9BF" authorityName="Mannerheim" authorityYear="1843" box="[1031,1166,1602,1624]" class="Insecta" family="Zopheridae" genus="Rhagodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="A8EBEACFFFFA974CFBC6160892B8F9BF" box="[1031,1166,1602,1624]" italics="true" pageId="1" pageNumber="294">R. tuberculata</emphasis>
</taxonomicName>
, both of which have the epipleural fold ending near the middle of ventrite IV, followed by a thin carina to the apex.
<taxonomicName id="5D9F4D5EFFFA974CFCB916DB92A5F940" box="[888,1171,1681,1703]" class="Insecta" family="Zopheridae" genus="Rhagodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="costaefragmenta">
<emphasis id="A8EBEACFFFFA974CFCB916DB92A5F940" box="[888,1171,1681,1703]" italics="true" pageId="1" pageNumber="294">Rhagodera costaefragmenta</emphasis>
</taxonomicName>
differs from
<taxonomicName id="5D9F4D5EFFFA974CFCE516E6959EF925" authorityName="Mannerheim" authorityYear="1843" box="[804,936,1708,1730]" class="Insecta" family="Zopheridae" genus="Rhagodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="A8EBEACFFFFA974CFCE516E6959EF925" box="[804,936,1708,1730]" italics="true" pageId="1" pageNumber="294">R. tuberculata</emphasis>
</taxonomicName>
and
<taxonomicName id="5D9F4D5EFFFA974CFC1916E69267F925" authorityName="Stephan" authorityYear="1989" box="[984,1105,1708,1730]" class="Insecta" family="Zopheridae" genus="Rhagodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="interrupta">
<emphasis id="A8EBEACFFFFA974CFC1916E69267F925" box="[984,1105,1708,1730]" italics="true" pageId="1" pageNumber="294">R. interrupta</emphasis>
</taxonomicName>
by the extremely fragmented costa 2 (
<figureCitation id="02A42A58FFF9974FFE1F1231940AFD77" box="[478,572,635,657]" captionStart="Figs" captionStartId="2.[197,237,574,593]" captionTargetBox="[173,1193,193,551]" captionTargetId="figure-504@2.[173,1193,193,551]" captionTargetPageId="2" captionText="Figs. 2–4. Comparison of costa 2 in three species of Rhagodera. 2) R. tuberculata; 3) R.interrupta; 4) R. costaefragmenta." figureDoi="http://doi.org/10.5281/zenodo.10104867" httpUri="https://zenodo.org/record/10104867/files/figure.png" pageId="2" pageNumber="295">Figs. 2–4</figureCitation>
). A subantennal groove is present in
<taxonomicName id="5D9F4D5EFFF9974FFE1412DF94A5FD4C" box="[469,659,661,683]" class="Insecta" family="Zopheridae" genus="Rhagodera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="costaefragmenta">
<emphasis id="A8EBEACFFFF9974FFE1412DF94A5FD4C" box="[469,659,661,683]" italics="true" pageId="2" pageNumber="295">R. costaefragmenta</emphasis>
</taxonomicName>
and
<taxonomicName id="5D9F4D5EFFF9974FFF1612FA976AFD21" authorityName="Mannerheim" authorityYear="1843" box="[215,348,688,710]" class="Insecta" family="Zopheridae" genus="Rhagodera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="A8EBEACFFFF9974FFF1612FA976AFD21" box="[215,348,688,710]" italics="true" pageId="2" pageNumber="295">R. tuberculata</emphasis>
</taxonomicName>
but absent in
<emphasis id="A8EBEACFFFF9974FFE2612FA945EFD21" box="[487,616,688,710]" italics="true" pageId="2" pageNumber="295">
<taxonomicName id="5D9F4D5EFFF9974FFE2612FA9452FD21" authorityName="Stephan" authorityYear="1989" box="[487,612,688,710]" class="Insecta" family="Zopheridae" genus="Rhagodera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="interrupta">R. interrupta</taxonomicName>
.
</emphasis>
The following key to species is adapted from Stephan (1989) to accommodate
<emphasis id="A8EBEACFFFF9974FFE4E12AF947EFD1C" box="[399,584,741,763]" italics="true" pageId="2" pageNumber="295">
<taxonomicName id="5D9F4D5EFFF9974FFE4E12AF9472FD1C" box="[399,580,741,763]" class="Insecta" family="Zopheridae" genus="Rhagodera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="costaefragmenta">R. costaefragmenta</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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