141 lines
18 KiB
XML
141 lines
18 KiB
XML
<document id="44F1FFD35B8B88F24ECAAAD923B39C13" ID-CLB-Dataset="27341" ID-DOI="10.11646/zootaxa.4585.1.2" ID-GBIF-Dataset="8547d24d-5ff0-4946-9593-946f24086548" ID-ISSN="1175-5326" ID-Zenodo-Dep="2636187" ID-ZooBank="1D7B13B8-C6D0-44F4-AFD2-47733E01E08E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554964596229" checkinUser="plazi" docAuthor="Ulitzka, Manfred R." docDate="2019" docId="03C187ED4041FFA207A45FCFFADD5EA0" docLanguage="en" docName="zootaxa.4585.1.2.pdf" docOrigin="Zootaxa 4585 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rohrthrips jiewenae Ulitzka 2019, sp. n." docType="treatment" docVersion="9" lastPageNumber="30" masterDocId="FFF8FF954042FFA107335F58FF98587F" masterDocTitle="Five new species of Rohrthrips (Thysanoptera: Rohrthripidae) from Burmese amber, and the evolution of Tubulifera wings" masterLastPageNumber="40" masterPageNumber="27" pageNumber="30" updateTime="1698719504840" updateUser="ExternalLinkService">
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<mods:title id="A0454C777305AAD4CE6F80E609BA3191">Five new species of Rohrthrips (Thysanoptera: Rohrthripidae) from Burmese amber, and the evolution of Tubulifera wings</mods:title>
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<mods:namePart id="C6F01EE1B29E857506F351374066A85C">Ulitzka, Manfred R.</mods:namePart>
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<mods:title id="CB9F18A593877E3BBCB44CC5F907734C">Zootaxa</mods:title>
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<mods:date id="E6A5B59D0FC4EEC36A2BD7A6325A9CDE">2019</mods:date>
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<treatment id="03C187ED4041FFA207A45FCFFADD5EA0" ID-DOI="http://doi.org/10.5281/zenodo.5933850" ID-GBIF-Taxon="156147061" ID-Zenodo-Dep="5933850" LSID="urn:lsid:plazi:treatment:03C187ED4041FFA207A45FCFFADD5EA0" httpUri="http://treatment.plazi.org/id/03C187ED4041FFA207A45FCFFADD5EA0" lastPageNumber="30" pageId="3" pageNumber="30">
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<subSubSection id="C37265704041FFA207A45FCFFE4D58CD" box="[151,469,151,178]" pageId="3" pageNumber="30" type="nomenclature">
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<paragraph id="8BD736FB4041FFA207A45FCFFE4D58CD" blockId="3.[151,469,151,211]" box="[151,469,151,178]" pageId="3" pageNumber="30">
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<heading id="D09F81974041FFA207A45FCFFE4D58CD" bold="true" box="[151,469,151,178]" fontSize="11" level="1" pageId="3" pageNumber="30" reason="1">
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<emphasis id="B91CEAE94041FFA207A45FCFFE4D58CD" bold="true" box="[151,469,151,178]" pageId="3" pageNumber="30">
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<taxonomicName id="4C684D784041FFA207A45FCFFE1458CE" authority="Ulitzka, 2019" authorityName="Ulitzka" authorityYear="2019" box="[151,396,151,177]" class="Insecta" family="Phlaeothripidae" genus="Rohrthrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="jiewenae" status="sp. nov.">
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<emphasis id="B91CEAE94041FFA207A45FCFFE1458CE" bold="true" box="[151,396,151,177]" italics="true" pageId="3" pageNumber="30">Rohrthrips jiewenae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22F57924041FFA206A05FC0FE4D58CD" box="[403,469,152,178]" pageId="3" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37265704041FFA207A45FE2FC035E78" pageId="3" pageNumber="30" type="description">
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<paragraph id="8BD736FB4041FFA207A45FE2FF7358AC" blockId="3.[151,469,151,211]" box="[151,235,186,211]" pageId="3" pageNumber="30">
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(
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<figureCitation id="13532A7E4041FFA207AC5FE2FF7A58AC" box="[159,226,186,211]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1937,1960]" captionTargetBox="[151,1436,235,1893]" captionTargetId="figure@4.[151,1436,235,1893]" captionTargetPageId="4" captionText="FIGURES 1–4. Rohrthrips spp. R. breviceps sp. n. holotype male 1–2: (1) dorsal view; (2) pterothoracic sternal plates (green: mesopresternum, yellow: mesosternum, red: trochantin, violet: mesokatepisternum, blue: metasternum). (3) R. jiewenae sp. n. holotype female, dorsal view, indicated: fungal spores. (4) R. maryae sp. n. holotype male, pterothoracic sternal plates." httpUri="https://zenodo.org/record/2636189/files/figure.png" pageId="3" pageNumber="30">Fig. 3</figureCitation>
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)
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</paragraph>
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<paragraph id="8BD736FB4041FFA207A45E5BFEF95940" blockId="3.[151,1437,258,1759]" pageId="3" pageNumber="30">
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<emphasis id="B91CEAE94041FFA207A45E5BFF735964" box="[151,235,259,283]" italics="true" pageId="3" pageNumber="30">Female</emphasis>
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. Body slightly contracted; legs extended; right wings and left hind wing overlapping body, right fore wing spread to the side.
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</paragraph>
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<paragraph id="8BD736FB4041FFA207F45E12FC1959F8" blockId="3.[151,1437,258,1759]" pageId="3" pageNumber="30">
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Colour (
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<figureCitation id="13532A7E4041FFA206105E12FEFD591C" box="[291,357,330,355]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1937,1960]" captionTargetBox="[151,1436,235,1893]" captionTargetId="figure@4.[151,1436,235,1893]" captionTargetPageId="4" captionText="FIGURES 1–4. Rohrthrips spp. R. breviceps sp. n. holotype male 1–2: (1) dorsal view; (2) pterothoracic sternal plates (green: mesopresternum, yellow: mesosternum, red: trochantin, violet: mesokatepisternum, blue: metasternum). (3) R. jiewenae sp. n. holotype female, dorsal view, indicated: fungal spores. (4) R. maryae sp. n. holotype male, pterothoracic sternal plates." httpUri="https://zenodo.org/record/2636189/files/figure.png" pageId="3" pageNumber="30">Fig. 3</figureCitation>
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) dark brown including antennae and legs; all major setae and wing fringes dark; wing veins light brown but much darker at base; wing membrane shaded brownish.
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</paragraph>
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<paragraph id="8BD736FB4041FFA207F45ECAFDD55C99" blockId="3.[151,1437,258,1759]" pageId="3" pageNumber="30">
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Head mainly smooth, about as long as wide; slightly produced in front of anterior margin of eyes, with cheeks rounded behind eyes; cheeks lacking setae; one pair of long and strong ocellar setae close to base of antennae. Eyes not prolonged ventrally. Ocelli large; hind ocelli very close to compound eyes; far apart from each other. Vertex with two postocular setae far behind eyes. Antennae 9-segmented; segment I barrel-shaped; segment II basally bended outwards with a very short pedicle, then similar to III–VII asymmetrical inverse cone-shaped, distal margin inclined to the exterior side; VIII spindle-shaped; IX style-shaped. Segment II with a small distal sensorium; number of sense cones of the following segments difficult to assess (distorted) but at least one inner and one outer on III–V. Mouth cone short and rounded. Maxillary palps clearly 3-segmented. Pronotum about as long as wide, trapezoidal; mediolateral and posteroangular setae long and pointed, anteroangular setae as well as anteromarginal and posteromarginal setae minute. Anterior half of mesonotum sculptured coarsely reticulate; mesonotal setae not visible. Prosternal plates not assessable; mesosternellum and metasternum fused; sternacosta meeting each of the mesothoracic coxae with a conspicuous process. Fore wings (
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<figureCitation id="13532A7E4041FFA204615C46FC0D5B48" box="[850,917,798,823]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1937,1960]" captionTargetBox="[151,1436,235,1893]" captionTargetId="figure@4.[151,1436,235,1893]" captionTargetPageId="4" captionText="FIGURES 1–4. Rohrthrips spp. R. breviceps sp. n. holotype male 1–2: (1) dorsal view; (2) pterothoracic sternal plates (green: mesopresternum, yellow: mesosternum, red: trochantin, violet: mesokatepisternum, blue: metasternum). (3) R. jiewenae sp. n. holotype female, dorsal view, indicated: fungal spores. (4) R. maryae sp. n. holotype male, pterothoracic sternal plates." httpUri="https://zenodo.org/record/2636189/files/figure.png" pageId="3" pageNumber="30">Fig. 3</figureCitation>
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) becoming progressively larger in distal half; anterior vein complete, reaching costa near apex, bearing one median and two distal setae as well as one short basal seta located anteriorly to the vein; second vein not developed; membrane without microtrichia. Fringe cilia straight; arising from sockets; duplicated cilia present around apical margin of wing; clavus not clearly visible. Hind wings without microtrichia; without any veins; sockets of wing fringes smaller than on fore wing. Fore legs with femora stout; fore tarsi with a strong tooth and a very strong hamus. Mid and hind legs slender. All femora with one long ventral and all tibiae with one long dorsal seta; mid and hind tibiae terminally each with three spines; mid and hind tarsi two-segmented, with hamus. Abdominal tergites smooth, without wing-retaining setae; lateral setae long but delicate, pointed; setae s3 on V–VII longer than other lateral setae, not conspicuously bent inwards as in other species. Abdominal segment I (pelta) not assessable; IX conical, with setae s3 very long, reaching tip of tube. Sternite IX with structures interpretable as gynosternal plates and fustis (cf.
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<bibRefCitation id="EFF94B0A4041FFA204C05BDEFB115CE0" author="Bhatti, J. S." box="[1011,1161,1158,1183]" pageId="3" pageNumber="30" pagination="285 - 352" refId="ref10160" refString="Bhatti, J. S. (1998 b) New structural features in the Order Tubulifera (Insecta). 4. The ovispan and other abdominal structures. Zoology (Journal of Pure and Applied Zoology), 5, 285 - 352." type="journal article" year="1998">Bhatti 1998b</bibRefCitation>
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). Abdominal segment X (tube) long and slightly extended at base; its sculpture not assessable; apparently without campaniform sensilla; with terminal crown of long anal setae.
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</paragraph>
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<paragraph id="8BD736FB4041FFA207F45BAAFC035E78" blockId="3.[151,1437,258,1759]" pageId="3" pageNumber="30">
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<emphasis id="B91CEAE94041FFA207F45BAAFEEE5D74" bold="true" box="[199,374,1266,1291]" pageId="3" pageNumber="30">Measurements</emphasis>
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. Female (in microns): Body, length 1548. Head, length 166; width 176. Eyes, length 63; width 38. Ocelli, diameter 19; distance between the hind ocelli 47. Ocellar setae 37. Prothorax, length 239; width 365; posteroangular setae 76; mediolateral setae about 65 (not clearly visible); epimeral setae 120. Mesonotum, length 120; width not measurable. Pterothorax, largest width 347. Abdomen, length 911; largest width 490 (segment IV); segment X (tube), length 176; basal width 50. Setae s3 on tergite IX about 255. Antennae, length 334 (segments partially retracted); length (largest width) of segment
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<date id="FFD6103B4041FFA205C35AFFFC845DC0" box="[752,796,1446,1471]" pageId="3" pageNumber="30" value="1950-01">I 50</date>
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(31),
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<date id="FFD6103B4041FFA204535AFFFC0E5DC0" box="[864,918,1447,1471]" pageId="3" pageNumber="30" value="1943-02">II 43</date>
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(31),
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<date id="FFD6103B4041FFA204EA5AFFFB805DC0" box="[985,1048,1446,1471]" pageId="3" pageNumber="30" value="1950-03">III 50</date>
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(34),
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<date id="FFD6103B4041FFA2036F5AFFFB055DC0" box="[1116,1181,1447,1471]" pageId="3" pageNumber="30" value="1940-04">IV 40</date>
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(36),
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<date id="FFD6103B4041FFA203D35AFFFA805DC0" box="[1248,1304,1447,1471]" pageId="3" pageNumber="30" value="1937-05">V 37</date>
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(28),
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<date id="FFD6103B4041FFA202685AFFFA045DC0" box="[1371,1436,1447,1471]" pageId="3" pageNumber="30" value="1943-06">VI 43</date>
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(19),
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<date id="FFD6103B4041FFA207E05A93FE855D9C" box="[211,285,1483,1507]" pageId="3" pageNumber="30" value="1931-07">VII 31</date>
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(16),
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<date id="FFD6103B4041FFA206535A93FE2B5D9C" box="[352,435,1483,1507]" pageId="3" pageNumber="30" value="1931-08">VIII 31</date>
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(12),
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<date id="FFD6103B4041FFA206C55A93FDB15D9C" box="[502,553,1483,1507]" pageId="3" pageNumber="30">IX 9</date>
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(3). Sense cones, length of inner (outer) on segment
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<date id="FFD6103B4041FFA2034A5A93FB325D9C" box="[1145,1194,1482,1507]" pageId="3" pageNumber="30">III 6</date>
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(6),
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<date id="FFD6103B4041FFA203EC5A93FA8A5D9C" box="[1247,1298,1483,1507]" pageId="3" pageNumber="30">IV 9</date>
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(6),
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<date id="FFD6103B4041FFA202745A93FAE65D9C" box="[1351,1406,1483,1507]" pageId="3" pageNumber="30" value="2010-05">V 10</date>
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(–); basal width about 3–4. Fore wings, length 1196; largest width 225.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37265704041FFA207F4594AFEA05EE8" pageId="3" pageNumber="30" type="materials_examined">
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<paragraph id="8BD736FB4041FFA207F4594AFED85E30" blockId="3.[151,1437,258,1759]" pageId="3" pageNumber="30">
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<emphasis id="B91CEAE94041FFA207F4594AFE0A5E54" bold="true" box="[199,402,1554,1579]" pageId="3" pageNumber="30">Material studied</emphasis>
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.
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<materialsCitation id="3B003CA64041FFA20697594AFED85E30" ID-GBIF-Occurrence="2235579716" location="Inclusion in Burmese" pageId="3" pageNumber="30" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54D388594041FFA20697594AFD945E54" box="[420,524,1554,1579]" pageId="3" pageNumber="30" type="holotype">Holotype</typeStatus>
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female MU-Fos-89/1.
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<location id="8EB760204041FFA2042E594AFB8E5E55" LSID="urn:lsid:plazi:treatment:03C187ED4041FFA207A45FCFFADD5EA0:8EB760204041FFA2042E594AFB8E5E55" box="[797,1046,1554,1579]" name="Inclusion in Burmese" pageId="3" pageNumber="30">Inclusion in Burmese</location>
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amber, donated to the author by Yanzhen Zhao.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BD736FB4041FFA207F45902FEA05EE8" blockId="3.[151,1437,258,1759]" pageId="3" pageNumber="30">
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<emphasis id="B91CEAE94041FFA207F45902FEF35E0C" bold="true" box="[199,363,1626,1651]" pageId="3" pageNumber="30">Syninclusions</emphasis>
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. Two midges and parts of a fly, one unknown insect larva and some fungal spores close to the thrips (
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<figureCitation id="13532A7E4041FFA207D55926FEB25EE8" box="[230,298,1662,1687]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1937,1960]" captionTargetBox="[151,1436,235,1893]" captionTargetId="figure@4.[151,1436,235,1893]" captionTargetPageId="4" captionText="FIGURES 1–4. Rohrthrips spp. R. breviceps sp. n. holotype male 1–2: (1) dorsal view; (2) pterothoracic sternal plates (green: mesopresternum, yellow: mesosternum, red: trochantin, violet: mesokatepisternum, blue: metasternum). (3) R. jiewenae sp. n. holotype female, dorsal view, indicated: fungal spores. (4) R. maryae sp. n. holotype male, pterothoracic sternal plates." httpUri="https://zenodo.org/record/2636189/files/figure.png" pageId="3" pageNumber="30">Fig. 3</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37265704041FFA207F459FAFADD5EA0" pageId="3" pageNumber="30" type="etymology">
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<paragraph id="8BD736FB4041FFA207F459FAFADD5EA0" blockId="3.[151,1437,258,1759]" pageId="3" pageNumber="30">
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<emphasis id="B91CEAE94041FFA207F459FAFEDE5EC4" bold="true" box="[199,326,1698,1723]" pageId="3" pageNumber="30">Etymology</emphasis>
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. The species
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<taxonomicName id="4C684D784041FFA206D359FBFDFB5EC5" authorityName="Ulitzka" authorityYear="2019" box="[480,611,1699,1722]" class="Insecta" family="Phlaeothripidae" genus="Rohrthrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="jiewenae">
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<emphasis id="B91CEAE94041FFA206D359FBFDFB5EC5" box="[480,611,1699,1722]" italics="true" pageId="3" pageNumber="30">R. jiewenae</emphasis>
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</taxonomicName>
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is dedicated to Jiewen Zhao, granddaughter of Yanzhen Zhao who found the
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<typeStatus id="54D388594041FFA207F3599EFEBA5EA0" box="[192,290,1734,1759]" pageId="3" pageNumber="30" type="holotype">holotype</typeStatus>
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female in a mine of the Hukawng Valley,
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<collectingRegion id="49ACF8194041FFA205C8599EFCD45EA0" box="[763,844,1734,1759]" country="Myanmar" name="Kachin" pageId="3" pageNumber="30">Kachin</collectingRegion>
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State,
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<collectingCountry id="F37F766B4041FFA204AB599FFB9D5EA0" box="[920,1029,1735,1759]" name="Myanmar" pageId="3" pageNumber="30">Myanmar</collectingCountry>
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and donated it to the author.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |