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<document id="0A32FFFCF6C46DC5B7AFEE50755F4202" ID-CLB-Dataset="31902" ID-DOI="10.11646/zootaxa.4329.2.1" ID-GBIF-Dataset="7e33ec7d-211b-4042-8d07-344364ee805d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1001915" ID-ZooBank="8226E27D-E3A9-481A-B51B-558643Bbb03A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507106636447" checkinUser="plazi" docAuthor="Tarasov, Sergei" docDate="2017" docId="03C187E1AE46B62CDD9E4603FE51036C" docLanguage="en" docName="zootaxa.4329.2.1.pdf" docOrigin="Zootaxa 4329 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parachorius javanus Tarasov 2017, new combination" docType="treatment" docVersion="12" lastPageNumber="129" masterDocId="FFF8FF99AE5DB630DD09474FFFDB0270" masterDocTitle="A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data" masterLastPageNumber="149" masterPageNumber="101" pageNumber="128" updateTime="1698469310636" updateUser="ExternalLinkService">
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<mods:titleInfo id="2020E2E9D888945D15B4BF384CB95AC8">
<mods:title id="28D2188A20F6F471D52666ACB04A9B54">A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data</mods:title>
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<mods:namePart id="0E286D4BD36C6694D3098C110B44D82A">Tarasov, Sergei</mods:namePart>
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<mods:title id="8D2BC939862EBAA22E5B14F2E6954534">Zootaxa</mods:title>
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<mods:date id="1339E42BEF7E74536D7B865701D0D0BB">2017</mods:date>
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<mods:number id="043306680E33C630AD2B84536F506F2F">2017-10-04</mods:number>
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<mods:number id="4DA3C8A13A37A6332E8DCFD88A08283F">4329</mods:number>
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<mods:identifier id="39867674BC723E57B7EB85F80950EA17" type="DOI">10.11646/zootaxa.4329.2.1</mods:identifier>
<mods:identifier id="051280C8D6E96E9529A87882C085188E" type="GBIF-Dataset">7e33ec7d-211b-4042-8d07-344364ee805d</mods:identifier>
<mods:identifier id="B6EF009B32EC77FE0B1AEE04975C9F56" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C187E1AE46B62CDD9E4603FE51036C" ID-DOI="http://doi.org/10.5281/zenodo.6010828" ID-GBIF-Taxon="135106135" ID-Zenodo-Dep="6010828" LSID="urn:lsid:plazi:treatment:03C187E1AE46B62CDD9E4603FE51036C" httpUri="http://treatment.plazi.org/id/03C187E1AE46B62CDD9E4603FE51036C" lastPageId="28" lastPageNumber="129" pageId="27" pageNumber="128">
<subSubSection id="C372657CAE46B62BDD9E4603FE9B03F7" pageId="27" pageNumber="128" type="nomenclature">
<paragraph id="8BD736F7AE46B62BDD9E4603FCB80316" blockId="27.[151,867,332,391]" box="[151,867,332,358]" pageId="27" pageNumber="128">
<heading id="D09F819BAE46B62BDD9E4603FCB80316" bold="true" box="[151,867,332,358]" fontSize="11" level="1" pageId="27" pageNumber="128" reason="1">
<emphasis id="B91CEAE5AE46B62BDD9E4603FCB80316" bold="true" box="[151,867,332,358]" pageId="27" pageNumber="128">
<taxonomicName id="4C684D74AE46B62BDD9E4603FD5D0316" ID-CoL="6TNM6" authority="(Boucomont, 1914)" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Boucomont" baseAuthorityYear="1914" box="[151,646,332,358]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="javanus" status="comb. nov.">
<emphasis id="B91CEAE5AE46B62BDD9E4603FE490316" bold="true" box="[151,402,332,358]" italics="true" pageId="27" pageNumber="128">Parachorius javanus</emphasis>
(
<bibRefCitation id="EFF94B06AE46B62BDCA84603FDA60316" author="Boucomont" box="[417,637,332,358]" pageId="27" pageNumber="128" refString="Boucomont, A. (1914) Les Coprophages de l'Archipel Malais. Annales de la Societe Entomologique de France, 83, 238 - 350." type="journal article" year="1914">Boucomont, 1914</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22F579EAE46B62BDF844603FCB80316" box="[653,867,332,358]" pageId="27" pageNumber="128">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD736F7AE46B62BDD9E4621FE9B03F7" blockId="27.[151,867,332,391]" box="[151,320,366,391]" pageId="27" pageNumber="128">
(
<figureCitation id="13532A72AE46B62BDD964621FF2703F7" box="[159,252,366,391]" captionStart="FIGURE 14" captionStartId="27.[151,250,1980,2002]" captionTargetBox="[164,1415,1417,1938]" captionTargetId="figure@27.[164,1423,1417,1941]" captionTargetPageId="27" captionText="FIGURE 14. Morphology of Parachorius javanus. (A, E) aedeagus lateral view (male, holotype). (B) habitus (male, holotype). (C – D) protibia (male, holotype)" httpUri="https://zenodo.org/record/1001945/files/figure.png" pageId="27" pageNumber="128">Figs. 14</figureCitation>
,
<figureCitation id="13532A72AE46B62BDC014621FEE203F7" box="[264,313,366,391]" captionStart="FIGURE 26" captionStartId="43.[151,250,1905,1927]" captionTargetBox="[151,1436,265,1825]" captionTargetId="figure@43.[151,1436,265,1825]" captionTargetPageId="43" captionText="FIGURE 26. Distribution maps of Parachorius species. (A) Parachorius hookeri. (B) Parachorius humeralis. (C) Parachorius javanus. (D) Parachorius longipenis. (E) Parachorius maruyamai. (F) Parachorius newthayerae. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study." httpUri="https://zenodo.org/record/1001969/files/figure.png" pageId="27" pageNumber="128">26C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE46B62BDD9E46F9FDFC039B" pageId="27" pageNumber="128" type="reference_group">
<paragraph id="8BD736F7AE46B62BDD9E46F9FDFC039B" blockId="27.[151,566,438,491]" pageId="27" pageNumber="128">
<treatmentCitationGroup id="AB7811D9AE46B62BDD9E46F9FDFC039B" pageId="27" pageNumber="128">
<taxonomicName id="4C684D74AE46B62BDD9E46F9FDED03BC" ID-CoL="RNHT" authority="Boucomont, 1914: 251" authorityName="Boucomont" authorityPageNumber="251" authorityYear="1914" box="[151,566,438,461]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B91CEAE5AE46B62BDD9E46F9FE9303BD" box="[151,328,438,461]" italics="true" pageId="27" pageNumber="128">Cassolus javanus</emphasis>
<treatmentCitation id="0AC910E6AE46B62BDC4746F8FDED03BC" author="Boucomont" box="[334,566,438,460]" page="251" pageId="27" pageNumber="128" year="1914">
<bibRefCitation id="EFF94B06AE46B62BDC4746F8FDDD03BC" author="Boucomont" box="[334,518,438,460]" pageId="27" pageNumber="128" refString="Boucomont, A. (1914) Les Coprophages de l'Archipel Malais. Annales de la Societe Entomologique de France, 83, 238 - 350." type="journal article" year="1914">Boucomont, 1914</bibRefCitation>
: 251
</treatmentCitation>
</taxonomicName>
<emphasis id="B91CEAE5AE46B62BDD9E469AFE8B039B" box="[151,336,469,491]" italics="true" pageId="27" pageNumber="128">
<taxonomicName id="4C684D74AE46B62BDD9E469AFE91039B" ID-CoL="RNHT" authorityName="Boucomont" authorityYear="1914" box="[151,330,469,491]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="javanus">Cassolus javanus</taxonomicName>
;
</emphasis>
<treatmentCitation id="0AC910E6AE46B62BDC5F469AFDFC039B" author="Balthasar" box="[342,551,469,491]" page="263" pageId="27" pageNumber="128" year="1963">
<bibRefCitation id="EFF94B06AE46B62BDC5F469AFE2C039B" author="Balthasar" box="[342,503,469,491]" pageId="27" pageNumber="128" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Folume 1. Tschechoslowakische Akademie der Wissenschaften, Prague, 391 pp., XXIV pls." type="book" year="1963">Balthasar, 1963</bibRefCitation>
: 263
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE46B62BDD9E4558FDE4005F" box="[151,575,535,560]" pageId="27" pageNumber="128" type="materials_examined">
<paragraph id="8BD736F7AE46B62BDD9E4558FDE4005F" blockId="27.[151,1437,535,1352]" box="[151,575,535,560]" pageId="27" pageNumber="128">
<materialsCitation id="3B003CAAAE46B62BDD9E4558FDE4005F" ID-GBIF-Occurrence="1658411496" box="[151,575,535,560]" location="Sokabumi" pageId="27" pageNumber="128" specimenCount="1" stateProvince="West Java" typeStatus="holotype">
<emphasis id="B91CEAE5AE46B62BDD9E4558FEF60040" bold="true" box="[151,301,535,560]" pageId="27" pageNumber="128">
<typeStatus id="54D38855AE46B62BDD9E4558FF090040" box="[151,210,535,560]" pageId="27" pageNumber="128">Type</typeStatus>
locality
</emphasis>
.
<location id="8EB7602CAE46B62BDC354558FE750040" LSID="urn:lsid:plazi:treatment:03C187E1AE46B62CDD9E4603FE51036C:8EB7602CAE46B62BDC354558FE750040" box="[316,430,535,560]" name="Sokabumi" pageId="27" pageNumber="128" stateProvince="West Java">Sokabumi</location>
[
<collectingRegion id="49ACF815AE46B62BDCB34557FDEF005F" box="[442,564,536,559]" country="Indonesia" name="Jawa Barat" pageId="27" pageNumber="128">West Java</collectingRegion>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE46B62BDDCE4573FC300105" pageId="27" pageNumber="128" type="diagnosis">
<paragraph id="8BD736F7AE46B62BDDCE4573FA830095" blockId="27.[151,1437,535,1352]" pageId="27" pageNumber="128">
<emphasis id="B91CEAE5AE46B62BDDCE4573FE9A0025" bold="true" box="[199,321,572,597]" pageId="27" pageNumber="128">Diagnosis.</emphasis>
Along with
<taxonomicName id="4C684D74AE46B62BDCE84572FDB30025" authorityName="S. Tarasov" authorityYear="2017" box="[481,616,572,597]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="schuelkei">
<emphasis id="B91CEAE5AE46B62BDCE84572FDB30025" box="[481,616,572,597]" italics="true" pageId="27" pageNumber="128">P. schuelkei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D74AE46B62BDFA64572FCAC0025" authorityName="S. Tarasov" authorityYear="2017" box="[687,887,572,597]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="B91CEAE5AE46B62BDFA64572FCAC0025" box="[687,887,572,597]" italics="true" pageId="27" pageNumber="128">P. pseudojavanus</emphasis>
</taxonomicName>
, this species can be separated from all other
<taxonomicName id="4C684D74AE46B62BDD9E4510FEF80008" box="[151,291,607,632]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAE5AE46B62BDD9E4510FEF80008" box="[151,291,607,632]" italics="true" pageId="27" pageNumber="128">Parachorius</emphasis>
</taxonomicName>
the two distinctly produced clypeal teeth whose outer margin is not notched basally.
<taxonomicName id="4C684D74AE46B62BD8194510FF3500EC" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Boucomont" baseAuthorityYear="1914" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B91CEAE5AE46B62BD8194510FF3500EC" italics="true" pageId="27" pageNumber="128">Parachorius javanus</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C684D74AE46B62BDF2745CAFD6F00ED" authorityName="S. Tarasov" authorityYear="2017" box="[558,692,644,669]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="schuelkei">
<emphasis id="B91CEAE5AE46B62BDF2745CAFD6F00ED" box="[558,692,644,669]" italics="true" pageId="27" pageNumber="128">P. schuelkei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D74AE46B62BDFFB45CAFC6D00ED" authorityName="S. Tarasov" authorityYear="2017" box="[754,950,644,669]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="B91CEAE5AE46B62BDFFB45CAFC6D00ED" box="[754,950,644,669]" italics="true" pageId="27" pageNumber="128">P. pseudojavanus</emphasis>
</taxonomicName>
by a combination of characters: (1) body coloration is reddish brown, and (2) parameres are notched basally and claw shaped in lateral view. External morphology and endophallic sclerite characters place
<taxonomicName id="4C684D74AE46B62BDFFB4582FCBF0094" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Boucomont" baseAuthorityYear="1914" box="[754,868,717,740]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B91CEAE5AE46B62BDFFB4582FCBF0094" box="[754,868,717,740]" italics="true" pageId="27" pageNumber="128">P. javanus</emphasis>
</taxonomicName>
as most closely related to
<emphasis id="B91CEAE5AE46B62BD9874582FA830095" box="[1166,1368,716,741]" italics="true" pageId="27" pageNumber="128">
<taxonomicName id="4C684D74AE46B62BD9874582FA880095" authorityName="S. Tarasov" authorityYear="2017" box="[1166,1363,716,741]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="pseudojavanus">P. pseudojavanus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BD736F7AE46B62BDDCE45A0FC300105" blockId="27.[151,1437,535,1352]" pageId="27" pageNumber="128">
<emphasis id="B91CEAE5AE46B62BDDCE45A0FE4C0178" bold="true" box="[199,407,751,776]" pageId="27" pageNumber="128">Taxonomic notes.</emphasis>
The species is known only from the
<typeStatus id="54D38855AE46B62BDE3645A0FC7E0178" box="[831,933,751,776]" pageId="27" pageNumber="128" type="holotype">holotype</typeStatus>
. All other records of
<taxonomicName id="4C684D74AE46B62BD99145BEFAD00178" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Boucomont" baseAuthorityYear="1914" box="[1176,1291,753,776]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B91CEAE5AE46B62BD99145BEFAD00178" box="[1176,1291,753,776]" italics="true" pageId="27" pageNumber="128">P. javanus</emphasis>
</taxonomicName>
are likely to refer to the newly described and closely related
<taxonomicName id="4C684D74AE46B62BDFBF445AFCA3015D" authorityName="S. Tarasov" authorityYear="2017" box="[694,888,788,813]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="B91CEAE5AE46B62BDFBF445AFCA3015D" box="[694,888,788,813]" italics="true" pageId="27" pageNumber="128">P. pseudojavanus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C684D74AE46B62BDE8E445BFBA9015C" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Boucomont" baseAuthorityYear="1914" box="[903,1138,788,813]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B91CEAE5AE46B62BDE8E445BFBA9015C" box="[903,1138,788,813]" italics="true" pageId="27" pageNumber="128">Parachorius javanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D74AE46B62BD9A3445AFAB7015D" authorityName="S. Tarasov" authorityYear="2017" box="[1194,1388,788,813]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="pseudojavanus">
<emphasis id="B91CEAE5AE46B62BD9A3445AFAB7015D" box="[1194,1388,788,813]" italics="true" pageId="27" pageNumber="128">P. pseudojavanus</emphasis>
</taxonomicName>
can only be reliably separated by the shape of aedeagus; the females of the two species are undistinguishable. Due to this, many females from various localities in
<collectingRegion id="49ACF815AE46B62BDF994412FCDA0104" box="[656,769,861,884]" country="Indonesia" name="Jawa Barat" pageId="27" pageNumber="128">West Java</collectingRegion>
remain unidentified.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE46B62BDDCE4430FBAC0738" pageId="27" pageNumber="128" type="description">
<paragraph id="8BD736F7AE46B62BDDCE4430FBBC01E8" blockId="27.[151,1437,535,1352]" box="[199,1127,895,920]" pageId="27" pageNumber="128">
<emphasis id="B91CEAE5AE46B62BDDCE4430FE8201E8" bold="true" box="[199,345,895,920]" pageId="27" pageNumber="128">Description.</emphasis>
<emphasis id="B91CEAE5AE46B62BDC564430FE4401E8" box="[351,415,895,920]" italics="true" pageId="27" pageNumber="128">Body.</emphasis>
Body reddish brown, length
<quantity id="4C909B12AE46B62BDFEC4430FCE501E7" box="[741,830,895,920]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="27" pageNumber="128" unit="mm" value="6.7">6.7 mm</quantity>
. Elytra uniformly colored.
</paragraph>
<paragraph id="8BD736F7AE46B62BDDCE44EBFB6A01CD" blockId="27.[151,1437,535,1352]" box="[199,1201,932,957]" pageId="27" pageNumber="128">
<emphasis id="B91CEAE5AE46B62BDDCE44EBFED001CD" box="[199,267,932,957]" italics="true" pageId="27" pageNumber="128">Head.</emphasis>
Clypeus with two distinctly produced teeth, their outer margin not notched basally.
</paragraph>
<paragraph id="8BD736F7AE46B62BDDCE4486FB580675" blockId="27.[151,1437,535,1352]" pageId="27" pageNumber="128">
<emphasis id="B91CEAE5AE46B62BDDCE4486FEE60190" box="[199,317,969,992]" italics="true" pageId="27" pageNumber="128">Pronotum.</emphasis>
Pronotum rounded laterally. Frontal angles rounded. Disc covered with punctures separated by 1–4 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra.
</paragraph>
<paragraph id="8BD736F7AE46B62BDDCE4340FD1B0657" blockId="27.[151,1437,535,1352]" box="[199,704,1039,1064]" pageId="27" pageNumber="128">
<emphasis id="B91CEAE5AE46B62BDDCE4340FECE0658" box="[199,277,1039,1064]" italics="true" pageId="27" pageNumber="128">Elytra.</emphasis>
Elytra covered with sparse punctures.
</paragraph>
<paragraph id="8BD736F7AE46B62BDDCE437AFD94063D" blockId="27.[151,1437,535,1352]" box="[199,591,1076,1101]" pageId="27" pageNumber="128">
<emphasis id="B91CEAE5AE46B62BDDCE437AFEC9063C" box="[199,274,1077,1100]" italics="true" pageId="27" pageNumber="128">Wings.</emphasis>
Wings normally developed.
</paragraph>
<paragraph id="8BD736F7AE46B62BDDCE4316FC2F0770" blockId="27.[151,1437,535,1352]" pageId="27" pageNumber="128">
<emphasis id="B91CEAE5AE46B62BDDCE4316FED80600" box="[199,259,1113,1136]" italics="true" pageId="27" pageNumber="128">Legs.</emphasis>
Profemora with almost straight or sinuate anterior margin. Protibia with apical spur approximately reaching middle of protarsomere 3. First tooth notably modified in males, approximately twice as wide as second tooth; first and second teeth distinctly separated from each other; first tooth distinctly separated from denticles located between first and second teeth. Metafemoral posterior margin not serrate, simple, not modified. Metatibia slightly sinuate, inner margin not denticulate, straight, not produced apically.
</paragraph>
<paragraph id="8BD736F7AE46B62BDDCE4242FB1F0755" blockId="27.[151,1437,535,1352]" box="[199,1220,1292,1317]" pageId="27" pageNumber="128">
<emphasis id="B91CEAE5AE46B62BDDCE4242FEB90754" box="[199,354,1293,1316]" italics="true" pageId="27" pageNumber="128">Metasternum.</emphasis>
Transverse basal ridge of metasternum long, at least 1/5 of mesocoxa length.
</paragraph>
<paragraph id="8BD736F7AE46B62BDDCE4260FBAC0738" blockId="27.[151,1437,535,1352]" box="[199,1143,1327,1352]" pageId="27" pageNumber="128">
<emphasis id="B91CEAE5AE46B62BDDCE4260FEE00738" box="[199,315,1327,1352]" italics="true" pageId="27" pageNumber="128">Aedeagus.</emphasis>
Parameres symmetrical, notched basally, and claw shaped in lateral view.
</paragraph>
</subSubSection>
<caption id="DF17667FAE46B62BDD9E40F3FE010581" httpUri="https://zenodo.org/record/1001945/files/figure.png" pageId="27" pageNumber="128" targetBox="[164,1415,1417,1938]" targetPageId="27">
<paragraph id="8BD736F7AE46B62BDD9E40F3FE010581" blockId="27.[151,1437,1980,2033]" pageId="27" pageNumber="128">
<emphasis id="B91CEAE5AE46B62BDD9E40F3FEC405A1" bold="true" box="[151,287,1980,2002]" pageId="27" pageNumber="128">FIGURE 14.</emphasis>
Morphology of
<taxonomicName id="4C684D74AE46B62BDCCD40F3FD4D05A2" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Boucomont" baseAuthorityYear="1914" box="[452,662,1980,2002]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="128" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B91CEAE5AE46B62BDCCD40F3FD4D05A2" box="[452,662,1980,2002]" italics="true" pageId="27" pageNumber="128">Parachorius javanus</emphasis>
</taxonomicName>
. (A, E) aedeagus lateral view (male, holotype). (B) habitus (male, holotype). (C–D) protibia (male, holotype)
</paragraph>
</caption>
<subSubSection id="C372657CAE41B62CDDCE47D8FE51036C" pageId="28" pageNumber="129" type="materials_examined">
<paragraph id="8BD736F7AE41B62CDDCE47D8FE51036C" blockId="28.[151,1436,151,285]" pageId="28" pageNumber="129">
<materialsCitation id="3B003CAAAE41B62CDDCE47D8FA4302DF" ID-GBIF-Occurrence="1658411509" box="[199,1432,151,176]" collectionCode="MUSEO" collectorName="H. Fruhstorfer" location="Sukabumi" pageId="28" pageNumber="129" specimenCount="1" specimenCount-male="1" stateProvince="Java" typeStatus="holotype">
<emphasis id="B91CEAE5AE41B62CDDCE47D8FDFE02C0" bold="true" box="[199,549,151,176]" pageId="28" pageNumber="129">
Material examined.
<typeStatus id="54D38855AE41B62CDCB147D8FDFE02C0" box="[440,549,151,176]" pageId="28" pageNumber="129" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="9D6EFD7EAE41B62CDF3D47D8FD5902C0" box="[564,642,151,176]" pageId="28" pageNumber="129" type="male">
<emphasis id="B91CEAE5AE41B62CDF3D47D8FD5902C0" bold="true" box="[564,642,151,176]" pageId="28" pageNumber="129">1 male</emphasis>
</specimenCount>
, &quot;
<collectingRegion id="49ACF815AE41B62CDF9047D7FD1502C0" box="[665,718,152,176]" country="Indonesia" name="Jawa Timur" pageId="28" pageNumber="129">Java</collectingRegion>
occident
<location id="8EB7602CAE41B62CDE3547D8FC7502C0" LSID="urn:lsid:plazi:treatment:03C187E1AE46B62CDD9E4603FE51036C:8EB7602CAE41B62CDE3547D8FC7502C0" box="[828,942,151,176]" name="Sukabumi" pageId="28" pageNumber="129" stateProvince="Java">Sukabumi</location>
2000 1983
<collectorName id="269D5321AE41B62CD93A47D7FB0F02C0" box="[1075,1236,151,176]" pageId="28" pageNumber="129">H. Fruhstorfer</collectorName>
/ EX
<collectionCode id="ED79AE32AE41B62CD81E47D7FAA202C0" box="[1303,1401,152,176]" pageId="28" pageNumber="129">MUSEO</collectionCode>
N
</materialsCitation>
.
<materialsCitation id="3B003CAAAE41B62CDD9E47F3FE5D036C" ID-GBIF-Occurrence="1658411498" collectionCode="VAN, POLL, MUSEUM, PARIS, ZMUA, MNHN" collectorName="Boucomont &amp; J. Krikken &amp; Rufous &amp; Bogor" determinerName="Boucomont" latitude="-6.918333" location="Amsterdam" longLatPrecision="129" longitude="106.92667" pageId="28" pageNumber="129" specimenCount="1" stateProvince="Java" typeStatus="holotype">
<collectionCode id="ED79AE32AE41B62CDD9E47F3FF0B02A4" box="[151,208,188,212]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14075" name="Soci<63>t<EFBFBD> Polymathique du Morbihan" pageId="28" pageNumber="129">VAN</collectionCode>
DE
<collectionCode id="ED79AE32AE41B62CDC0347F3FE8802A4" box="[266,339,188,212]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13857" name="Pfalzmuseum f<>r Naturkunde" pageId="28" pageNumber="129">POLL</collectionCode>
] /
<typeStatus id="54D38855AE41B62CDC7F47F3FE1402A4" box="[374,463,188,212]" pageId="28" pageNumber="129">TYPUS</typeStatus>
/
<determinerName id="7E0BDA33AE41B62CDCE547F2FDB402A4" box="[492,623,189,212]" pageId="28" pageNumber="129">Boucomont</determinerName>
det. 1914
<taxonomicName id="4C684D74AE41B62CDFE647F3FC6802A5" authority="Tarasov, 2017" authorityName="Tarasov" authorityYear="2017" box="[751,947,188,213]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="javanus" status="sp. nov.">Cassolus javanus</taxonomicName>
<taxonomicNameLabel id="A22F579EAE41B62CDEB447F2FC2902A4" box="[957,1010,189,212]" pageId="28" pageNumber="129" rank="species">n.sp.</taxonomicNameLabel>
/
<collectionCode id="ED79AE32AE41B62CD90747F3FB3A02A4" box="[1038,1249,188,212]" pageId="28" pageNumber="129">MUSEUM PARIS</collectionCode>
<collectorName id="269D5321AE41B62CD9E247F2FAB502A4" box="[1259,1390,189,212]" pageId="28" pageNumber="129">Boucomont</collectorName>
/
<collectorName id="269D5321AE41B62CD88347F2FF280288" pageId="28" pageNumber="129">J. Krikken</collectorName>
1969 compared with 5 PM (
<location id="8EB7602CAE41B62CDF494790FD130288" LSID="urn:lsid:plazi:treatment:03C187E1AE46B62CDD9E4603FE51036C:8EB7602CAE41B62CDF494790FD130288" box="[576,712,223,248]" latitude="-6.918333" longLatPrecision="129" longitude="106.92667" name="Amsterdam" pageId="28" pageNumber="129" stateProvince="Java">Amsterdam</location>
:
<collectionCode id="ED79AE32AE41B62CDFC24790FCFA0288" box="[715,801,223,248]" country="Greece" httpUri="http://grbio.org/cool/c09s-jk3y" name="Zoological Museum, University of Athens" pageId="28" pageNumber="129">ZMUA</collectionCode>
) which is comp[letely].
<collectorName id="269D5321AE41B62CD9324790FB570288" box="[1083,1164,223,248]" pageId="28" pageNumber="129">Rufous</collectorName>
from
<collectorName id="269D5321AE41B62CD9DC47AFFAFB0288" box="[1237,1312,224,248]" pageId="28" pageNumber="129">Bogor</collectorName>
&quot;,
<geoCoordinate id="EE5C5030AE41B62CD83F4790FA4C0288" box="[1334,1431,223,248]" direction="south" orientation="latitude" pageId="28" pageNumber="129" precision="92" value="-6.918333">6°55.1'S</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE41B62CDD9E464BFEC8036D" box="[151,275,260,285]" direction="east" orientation="longitude" pageId="28" pageNumber="129" precision="92" value="106.92667">106°55.6'E</geoCoordinate>
(
<collectionCode id="ED79AE32AE41B62CDC2B464BFEA5036C" box="[290,382,260,284]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="129">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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