494 lines
65 KiB
XML
494 lines
65 KiB
XML
<document id="801985579D69595FA8C0F3DB58D1ED78" ID-CLB-Dataset="32385" ID-DOI="10.11646/zootaxa.4306.3.10" ID-GBIF-Dataset="9e74e9ee-be1b-425e-90f4-f233191c55fe" ID-ISSN="1175-5326" ID-Zenodo-Dep="844474" ID-ZooBank="F44D9AF1-767E-474C-9E68-1044ECCF1B7D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1502961447104" checkinUser="plazi" docAuthor="Makbun, Noppadon" docDate="2017" docId="0394383FFFDBB95DFF6EFF4EFF7BFCCB" docLanguage="en" docName="zootaxa.4306.3.10.pdf" docOrigin="Zootaxa 4306 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anisogomphus yingsaki Makbun 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="442" masterDocId="FFAD4047FFDAB958FFF9FFD9FF94FFF3" masterDocTitle="Anisogomphus yingsaki (Odonata: Gomphidae) sp. nov., a new gomphid species from Thailand" masterLastPageNumber="443" masterPageNumber="437" pageNumber="438" updateTime="1698466723027" updateUser="ExternalLinkService">
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<mods:titleInfo id="FD94BCFE864D0AC414AFB1441A17C864">
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<mods:title id="63F62F74D2CA79458F28196793E38434">Anisogomphus yingsaki (Odonata: Gomphidae) sp. nov., a new gomphid species from Thailand</mods:title>
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<mods:name id="9E98F704D30481C991ACA1BBAA8BDA5D" type="personal">
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<mods:roleTerm id="2B08AA699BC351322168B60ABC912393">Author</mods:roleTerm>
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<mods:namePart id="6D59F48812312666640D5ED5AD377C96">Makbun, Noppadon</mods:namePart>
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<mods:title id="8B55516D9A6393678E6B3B9DED18B06A">Zootaxa</mods:title>
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<mods:part id="A67D5F3ED3BCF05C66076D2EBB5E5727">
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<mods:date id="175F2E8D5A6AB763B227C35F66EA4717">2017</mods:date>
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<mods:number id="B815759FE62EA47D9AFC647F0FAB264F">4306</mods:number>
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<mods:identifier id="2127E9897D0DE5F11328D439C6B651DB" type="DOI">10.11646/zootaxa.4306.3.10</mods:identifier>
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<treatment id="0394383FFFDBB95DFF6EFF4EFF7BFCCB" ID-DOI="http://doi.org/10.5281/zenodo.6029923" ID-GBIF-Taxon="132785277" ID-Zenodo-Dep="6029923" LSID="urn:lsid:plazi:treatment:0394383FFFDBB95DFF6EFF4EFF7BFCCB" httpUri="http://treatment.plazi.org/id/0394383FFFDBB95DFF6EFF4EFF7BFCCB" lastPageId="5" lastPageNumber="442" pageId="1" pageNumber="438">
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<subSubSection id="C327DAA2FFDBB959FF6EFF4EFEB4FF20" pageId="1" pageNumber="438" type="nomenclature">
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<paragraph id="8B828929FFDBB959FF6EFF4EFD89FF41" blockId="1.[151,541,151,211]" box="[151,541,151,178]" pageId="1" pageNumber="438">
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<heading id="D0CA3E45FFDBB959FF6EFF4EFD89FF41" bold="true" box="[151,541,151,178]" fontSize="11" level="1" pageId="1" pageNumber="438" reason="1">
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<emphasis id="B949553BFFDBB959FF6EFF4EFD89FF41" bold="true" box="[151,541,151,178]" pageId="1" pageNumber="438">
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<taxonomicName id="4C3DF2AAFFDBB959FF6EFF4EFE2CFF42" ID-CoL="9WZZL" authority="Makbun, 2017" authorityName="Makbun" authorityYear="2017" box="[151,440,151,177]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="yingsaki" status="sp. nov.">
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<emphasis id="B949553BFFDBB959FF6EFF4EFE2CFF42" bold="true" box="[151,440,151,177]" italics="true" pageId="1" pageNumber="438">Anisogomphus yingsaki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27AE840FFDBB959FE46FF41FD89FF41" box="[447,541,152,178]" pageId="1" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8B828929FFDBB959FF6EFF63FEB4FF20" blockId="1.[151,541,151,211]" box="[151,288,186,211]" pageId="1" pageNumber="438">
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(
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<figureCitation id="130695ACFFDBB959FF66FF63FE83FF20" box="[159,279,186,211]" captionStart-0="FIGURE 1 – 2" captionStart-1="FIGURE 3 – 12" captionStart-2="FIGURE 13 – 16" captionStartId-0="2.[151,250,1962,1984]" captionStartId-1="3.[151,250,1921,1943]" captionStartId-2="4.[151,250,985,1007]" captionTargetBox-0="[153,1433,233,1915]" captionTargetBox-1="[202,1391,193,1900]" captionTargetBox-2="[153,1433,183,968]" captionTargetId-0="figure@2.[153,1433,233,1915]" captionTargetId-1="figure@3.[202,1391,193,1906]" captionTargetId-2="figure@4.[153,1433,183,968]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1 – 2. Anisogomphus yingsaki sp. nov., photographed in nature by Noppadon Makbun, 22 - vi- 2016, Ban Na Kha, Ban Muang, Sakon Nakhon province: (1) male; (2) female." captionText-1="FIGURE 3 – 12. Anisogomphus yingsaki sp. nov., holotype male: (3) head in frontal view; (4) synthorax in dorsal view; (5) synthorax in lateral view; (6) left fore- and hindwing; (7) accessory genitalia in lateral view; (8) accessory genitalia in ventral view; (9) vesica spermalis in lateral view; (10) marking on S 7; (11) anal appendages in lateral view; (12) anal appendages in dorsal view." captionText-2="FIGURE 13 – 16. Anisogomphus yingsaki sp. nov., paratype female: (13) head in frontal view; (14) anal appendages in lateral view; (15) anal appendages in ventral view; (16) valvula vulvae in ventral view." httpUri-0="https://zenodo.org/record/844476/files/figure.png" httpUri-1="https://zenodo.org/record/844478/files/figure.png" httpUri-2="https://zenodo.org/record/844480/files/figure.png" pageId="1" pageNumber="438">Figs. 1–16</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C327DAA2FFDBB959FF6EFEDBFBD0FE58" pageId="1" pageNumber="438" type="materials_examined">
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||
<paragraph id="8B828929FFDBB959FF6EFEDBFEB1FECC" blockId="1.[151,1437,258,2011]" pageId="1" pageNumber="438">
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||
<materialsCitation id="3B558374FFDBB959FF6EFEDBFEB1FECC" ID-GBIF-Occurrence="1586201887" collectingDate="2016-06-22" collectorName="N. Makbun" country="Thailand" elevation="173" location="Ban Na Kha" municipality="Ban Muang" pageId="1" pageNumber="438" specimenCount="1" specimenCount-male="1" stateProvince="Sakon Nakhon province" typeStatus="holotype">
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||
<typeStatus id="5486378BFFDBB959FF6EFEDBFE90FEE8" box="[151,260,258,283]" pageId="1" pageNumber="438" type="holotype">
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<emphasis id="B949553BFFDBB959FF6EFEDBFE90FEE8" bold="true" box="[151,260,258,283]" pageId="1" pageNumber="438">Holotype</emphasis>
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||
</typeStatus>
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||
:
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<specimenCount id="9D3B42A0FFDBB959FEECFEDAFEB8FEE9" box="[277,300,259,282]" pageId="1" pageNumber="438" type="male">♂</specimenCount>
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,
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<collectingCountry id="F32AC9B9FFDBB959FEC3FEDBFE34FEE8" box="[314,416,258,283]" name="Thailand" pageId="1" pageNumber="438">Thailand</collectingCountry>
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,
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<collectingRegion id="49F947CBFFDBB959FE54FEDBFD50FEE8" box="[429,708,258,283]" country="Thailand" name="Sakon Nakhon" pageId="1" pageNumber="438">Sakon Nakhon province</collectingRegion>
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,
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<collectingMunicipality id="6BE61353FFDBB959FD28FEDAFCCDFEE8" box="[721,857,259,283]" pageId="1" pageNumber="438">Ban Muang</collectingMunicipality>
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||
,
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<location id="8EE2DFF2FFDBB959FC9EFEDAFC6DFEE8" LSID="urn:lsid:plazi:treatment:0394383FFFDBB95DFF6EFF4EFF7BFCCB:8EE2DFF2FFDBB959FC9EFEDAFC6DFEE8" box="[871,1017,258,283]" country="Thailand" municipality="Ban Muang" name="Ban Na Kha" pageId="1" pageNumber="438" stateProvince="Sakon Nakhon province">Ban Na Kha</location>
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, altitude
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<elevation id="00106E1AFFDBB959FB9AFEDBFB72FEE9" box="[1123,1254,258,283]" metricMagnitude="2" metricUnit="m" metricValue="1.725" metricValueMax="1.75" metricValueMin="1.7" pageId="1" pageNumber="438" unit="m" value="172.5" valueMax="175.0" valueMin="170.0">
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<quantity id="4CC524CCFFDBB959FB9AFEDBFB72FEE9" box="[1123,1254,258,283]" metricMagnitude="2" metricUnit="m" metricValue="1.725" metricValueMax="1.75" metricValueMin="1.7" pageId="1" pageNumber="438" unit="m" value="172.5" valueMax="175.0" valueMin="170.0">170–175 m</quantity>
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</elevation>
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,
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<collectingDate id="EFC75601FFDBB959FB0CFEDBFAE7FEE8" box="[1269,1395,258,283]" pageId="1" pageNumber="438" value="2016-06-22">
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||
<date id="FF83AFE9FFDBB959FB0CFEDBFAE7FEE8" box="[1269,1395,258,283]" pageId="1" pageNumber="438" value="2016-06-22">22-vi-2016</date>
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</collectingDate>
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||
,
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<collectorName id="26C8ECFFFFDBB959FA78FEDAFF60FECC" pageId="1" pageNumber="438">N. Makbun</collectorName>
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||
leg.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8B828929FFDBB959FF3EFE93FBD0FE58" blockId="1.[151,1437,258,2011]" pageId="1" pageNumber="438">
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||
<materialsCitation id="3B558374FFDBB959FF3EFE93FBD0FE58" ID-GBIF-Occurrence="1586201886" collectingDate="2016-06-22" collectingDateMax="2016-06-23" collectingDateMin="2016-06-22" collectionCode="THNHM" collectorName="N. Makbun & The" country="Thailand" pageId="1" pageNumber="438" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Pathum Thani" typeStatus="paratype">
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||
<typeStatus id="5486378BFFDBB959FF3EFE93FEABFE90" box="[199,319,330,355]" pageId="1" pageNumber="438" type="paratype">
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||
<emphasis id="B949553BFFDBB959FF3EFE93FEABFE90" bold="true" box="[199,319,330,355]" pageId="1" pageNumber="438">Paratypes</emphasis>
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||
</typeStatus>
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||
(from the same location as in
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||
<typeStatus id="5486378BFFDBB959FD5BFE93FC9EFE90" box="[674,778,330,355]" pageId="1" pageNumber="438" type="holotype">holotype</typeStatus>
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||
):
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||
<specimenCount id="9D3B42A0FFDBB959FCE7FE93FCD5FE90" box="[798,833,330,355]" pageId="1" pageNumber="438" type="female">1♀</specimenCount>
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||
,
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||
<collectingDate id="EFC75601FFDBB959FCA8FE93FC45FE90" box="[849,977,330,355]" pageId="1" pageNumber="438" value="2016-06-22">
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||
<date id="FF83AFE9FFDBB959FCA8FE93FC45FE90" box="[849,977,330,355]" pageId="1" pageNumber="438" value="2016-06-22">22-vi-2016</date>
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||
</collectingDate>
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||
;
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||
<specimenCount id="9D3B42A0FFDBB959FC26FE93FB8DFE90" box="[991,1049,330,355]" pageId="1" pageNumber="438" type="male">2♂♂</specimenCount>
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||
,
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||
<collectingDate id="EFC75601FFDBB959FBDEFE93FB32FE90" box="[1063,1190,330,355]" pageId="1" pageNumber="438" value="2016-06-23">
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<date id="FF83AFE9FFDBB959FBDEFE93FB32FE90" box="[1063,1190,330,355]" pageId="1" pageNumber="438" value="2016-06-23">23-vi-2016</date>
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</collectingDate>
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,
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<collectorName id="26C8ECFFFFDBB959FB4AFE92FAA1FE90" box="[1203,1333,330,355]" pageId="1" pageNumber="438">N. Makbun</collectorName>
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leg.
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<collectorName id="26C8ECFFFFDBB959FA88FE93FA09FE90" box="[1393,1437,330,355]" pageId="1" pageNumber="438">The</collectorName>
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<typeStatus id="5486378BFFDBB959FF6EFEB7FF6DFE74" box="[151,249,366,391]" pageId="1" pageNumber="438" type="holotype">holotype</typeStatus>
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and one female
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<typeStatus id="5486378BFFDBB959FE3AFEB6FDB7FE74" box="[451,547,367,391]" pageId="1" pageNumber="438" type="paratype">paratype</typeStatus>
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||
are deposited in the
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<collectingCountry id="F32AC9B9FFDBB959FCD8FEB7FC11FE74" box="[801,901,366,391]" name="Thailand" pageId="1" pageNumber="438">Thailand</collectingCountry>
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Natural History Museum (
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<collectionCode id="ED2C11ECFFDBB959FB3AFEB6FAA4FE74" box="[1219,1328,367,391]" pageId="1" pageNumber="438">THNHM</collectionCode>
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),
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<collectingRegion id="49F947CBFFDBB959FABEFEB7FF4FFE58" country="Thailand" name="Pathum Thani" pageId="1" pageNumber="438">Pathum Thani</collectingRegion>
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||
,
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||
<collectingCountry id="F32AC9B9FFDBB959FF1FFE4BFED9FE58" box="[230,333,402,427]" name="Thailand" pageId="1" pageNumber="438">Thailand</collectingCountry>
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. The other specimens are retained in the author’s private collection.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C327DAA2FFDBB959FF3EFE6FFBD2FDC8" pageId="1" pageNumber="438" type="etymology">
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<paragraph id="8B828929FFDBB959FF3EFE6FFBD2FDC8" blockId="1.[151,1437,258,2011]" pageId="1" pageNumber="438">
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||
<emphasis id="B949553BFFDBB959FF3EFE6FFEDAFE3C" bold="true" box="[199,334,438,463]" pageId="1" pageNumber="438">Etymology.</emphasis>
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The species name, a noun in the genitive case, honours Mr. Yingsak Paweenpermsuk, a dragonfly enthusiast who happened to find this species, took their photographs and sent them to the author for the identification. After scrutinizing his photos closely, the author realised that it was new species. Apart from that, he kindly took the author and his colleague to the
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<typeStatus id="5486378BFFDBB959FD5CFDFAFD41FDC8" box="[677,725,547,571]" pageId="1" pageNumber="438">type</typeStatus>
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locality for specimen collection.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C327DAA2FFDBB95CFF3EFD9FFEF6F9D1" lastPageId="4" lastPageNumber="441" pageId="1" pageNumber="438" type="description">
|
||
<paragraph id="8B828929FFDBB959FF3EFD9FFD1DFDAC" blockId="1.[151,1437,258,2011]" box="[199,649,582,607]" pageId="1" pageNumber="438">
|
||
<emphasis id="B949553BFFDBB959FF3EFD9FFE49FDAC" bold="true" box="[199,477,582,607]" pageId="1" pageNumber="438">
|
||
Description of
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||
<typeStatus id="5486378BFFDBB959FE8FFD9FFE49FDAC" box="[374,477,582,607]" pageId="1" pageNumber="438" type="holotype">holotype</typeStatus>
|
||
</emphasis>
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||
(
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||
<figureCitation id="130695ACFFDBB959FE15FD9FFDAEFDAC" box="[492,570,582,607]" captionStart="FIGURE 1 – 2" captionStartId="2.[151,250,1962,1984]" captionTargetBox="[153,1433,233,1915]" captionTargetId="figure@2.[153,1433,233,1915]" captionTargetPageId="2" captionText="FIGURE 1 – 2. Anisogomphus yingsaki sp. nov., photographed in nature by Noppadon Makbun, 22 - vi- 2016, Ban Na Kha, Ban Muang, Sakon Nakhon province: (1) male; (2) female." httpUri="https://zenodo.org/record/844476/files/figure.png" pageId="1" pageNumber="438">Figs. 1</figureCitation>
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||
,
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||
<figureCitation id="130695ACFFDBB959FDB1FD9EFD15FDAC" box="[584,641,583,607]" captionStart="FIGURE 3 – 12" captionStartId="3.[151,250,1921,1943]" captionTargetBox="[202,1391,193,1900]" captionTargetId="figure@3.[202,1391,193,1906]" captionTargetPageId="3" captionText="FIGURE 3 – 12. Anisogomphus yingsaki sp. nov., holotype male: (3) head in frontal view; (4) synthorax in dorsal view; (5) synthorax in lateral view; (6) left fore- and hindwing; (7) accessory genitalia in lateral view; (8) accessory genitalia in ventral view; (9) vesica spermalis in lateral view; (10) marking on S 7; (11) anal appendages in lateral view; (12) anal appendages in dorsal view." httpUri="https://zenodo.org/record/844478/files/figure.png" pageId="1" pageNumber="438">3–12</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B828929FFDBB959FF3EFDB3FADFFCA8" blockId="1.[151,1437,258,2011]" pageId="1" pageNumber="438">
|
||
<emphasis id="B949553BFFDBB959FF3EFDB3FE90FD70" box="[199,260,618,643]" italics="true" pageId="1" pageNumber="438">Head</emphasis>
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||
. Eyes green in life and becoming brown after preservation; face black with whitish yellow pattern (
|
||
<figureCitation id="130695ACFFDBB959FA96FDB3FF32FD54" captionStart="FIGURE 3 – 12" captionStartId="3.[151,250,1921,1943]" captionTargetBox="[202,1391,193,1900]" captionTargetId="figure@3.[202,1391,193,1906]" captionTargetPageId="3" captionText="FIGURE 3 – 12. Anisogomphus yingsaki sp. nov., holotype male: (3) head in frontal view; (4) synthorax in dorsal view; (5) synthorax in lateral view; (6) left fore- and hindwing; (7) accessory genitalia in lateral view; (8) accessory genitalia in ventral view; (9) vesica spermalis in lateral view; (10) marking on S 7; (11) anal appendages in lateral view; (12) anal appendages in dorsal view." httpUri="https://zenodo.org/record/844478/files/figure.png" pageId="1" pageNumber="438">Fig. 3</figureCitation>
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); occiput whitish yellow with black anterior margin, posterior margin slightly raised with a row of setae across the subapical area; vertex whitish yellow with black margin, slightly depressed in the middle; a pair of blunt tubercles behind lateral ocelli; antennae black; dorsal surface of frons entirely whitish yellow; anterior part of frons black with whitish yellow margin; postclypeus whitish yellow with two black markings, its margin black and concave centrally; anteclypeus and genae black; labrum whitish yellow with black margin, deep concavity in middle; mandible black with basal whitish yellow marking; labium, lateral and median lobe of labium pale.
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</paragraph>
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||
<paragraph id="8B828929FFDBB959FF3EFCBFFB3BFB89" blockId="1.[151,1437,258,2011]" pageId="1" pageNumber="438">
|
||
<emphasis id="B949553BFFDBB959FF3EFCBFFE8CFC8C" box="[199,280,870,895]" italics="true" pageId="1" pageNumber="438">Thorax</emphasis>
|
||
. Prothorax black with whitish yellow anterior margin, slightly expanded in middle; median lobe black with a pair of elongate yellowish lateral spots, the anterior one larger than the posterior one; posterior lobe whitish yellow with black margin, depressed centrally; synthorax black with whitish yellow markings (
|
||
<figureCitation id="130695ACFFDBB959FAE3FC77FA1BFC34" box="[1306,1423,942,967]" captionStart="FIGURE 3 – 12" captionStartId="3.[151,250,1921,1943]" captionTargetBox="[202,1391,193,1900]" captionTargetId="figure@3.[202,1391,193,1906]" captionTargetPageId="3" captionText="FIGURE 3 – 12. Anisogomphus yingsaki sp. nov., holotype male: (3) head in frontal view; (4) synthorax in dorsal view; (5) synthorax in lateral view; (6) left fore- and hindwing; (7) accessory genitalia in lateral view; (8) accessory genitalia in ventral view; (9) vesica spermalis in lateral view; (10) marking on S 7; (11) anal appendages in lateral view; (12) anal appendages in dorsal view." httpUri="https://zenodo.org/record/844478/files/figure.png" pageId="1" pageNumber="438">Figs. 4–5</figureCitation>
|
||
); antehumeral stripes almost straight, broader in the middle, slightly tapering at both blunt ends; humeral stripe reduced to small triangular spot under antealar sinus; mesothoracic collar whitish yellow; mesepimeron with yellow lateral stripe, expanded anteriorly at its apex, near its anterior margin, covered about a half of its width; metepisternum with yellow band, connected with metinfraepisternum stripe; metepimeron yellow with black margin, except some basal and apical areas; metinfraepisternum black with large yellow spot.
|
||
</paragraph>
|
||
<paragraph id="8B828929FFDBB959FF3EFB5EFBDDFB14" blockId="1.[151,1437,258,2011]" pageId="1" pageNumber="438">
|
||
Legs black except coxae which are wholly pale, inner area of profemur pale, posterior of profemur with pale spot and metafemur with basal 2/3 pale; metafemur with two rows of small spines at base, and large widely spaced ones at distal half, the distal end of which reaches as far as the middle of S1 (
|
||
<figureCitation id="130695ACFFDBB959FC0EFB17FBAFFB14" box="[1015,1083,1230,1255]" captionStart="FIGURE 3 – 12" captionStartId="3.[151,250,1921,1943]" captionTargetBox="[202,1391,193,1900]" captionTargetId="figure@3.[202,1391,193,1906]" captionTargetPageId="3" captionText="FIGURE 3 – 12. Anisogomphus yingsaki sp. nov., holotype male: (3) head in frontal view; (4) synthorax in dorsal view; (5) synthorax in lateral view; (6) left fore- and hindwing; (7) accessory genitalia in lateral view; (8) accessory genitalia in ventral view; (9) vesica spermalis in lateral view; (10) marking on S 7; (11) anal appendages in lateral view; (12) anal appendages in dorsal view." httpUri="https://zenodo.org/record/844478/files/figure.png" pageId="1" pageNumber="438">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B828929FFDBB959FF3EFB2DFDC7FA84" blockId="1.[151,1437,258,2011]" pageId="1" pageNumber="438">
|
||
<emphasis id="B949553BFFDBB959FF3EFB2DFE9EFAF8" box="[199,266,1268,1291]" italics="true" pageId="1" pageNumber="438">Wings</emphasis>
|
||
. Hyaline, basally tinted with brownish (
|
||
<figureCitation id="130695ACFFDBB959FD33FB2BFC9AFAF8" box="[714,782,1266,1291]" captionStart="FIGURE 3 – 12" captionStartId="3.[151,250,1921,1943]" captionTargetBox="[202,1391,193,1900]" captionTargetId="figure@3.[202,1391,193,1906]" captionTargetPageId="3" captionText="FIGURE 3 – 12. Anisogomphus yingsaki sp. nov., holotype male: (3) head in frontal view; (4) synthorax in dorsal view; (5) synthorax in lateral view; (6) left fore- and hindwing; (7) accessory genitalia in lateral view; (8) accessory genitalia in ventral view; (9) vesica spermalis in lateral view; (10) marking on S 7; (11) anal appendages in lateral view; (12) anal appendages in dorsal view." httpUri="https://zenodo.org/record/844478/files/figure.png" pageId="1" pageNumber="438">Fig. 6</figureCitation>
|
||
); primary antenodals first and fifth. Ax 12 (left), 13 (right) on Fw, 9 both on Hw, Px 10 both on Fw, 9 (left), 10 (right) on Hw; anal triangle 3-celled; triangles not crossed; one crossvein in cubital space in both wings; two crossveins between Arc and R1–R4 junction in FW; pterostigma brown, braced, covered 3.5 cells below.
|
||
</paragraph>
|
||
<paragraph id="8B828929FFDBB959FF3EFA5BFB2EF9D8" blockId="1.[151,1437,258,2011]" pageId="1" pageNumber="438">
|
||
<emphasis id="B949553BFFDBB959FF3EFA5BFEA5FA68" box="[199,305,1410,1435]" italics="true" pageId="1" pageNumber="438">Abdomen</emphasis>
|
||
. Black with pale markings (
|
||
<figureCitation id="130695ACFFDBB959FD84FA5BFD54FA68" box="[637,704,1410,1435]" captionStart="FIGURE 1 – 2" captionStartId="2.[151,250,1962,1984]" captionTargetBox="[153,1433,233,1915]" captionTargetId="figure@2.[153,1433,233,1915]" captionTargetPageId="2" captionText="FIGURE 1 – 2. Anisogomphus yingsaki sp. nov., photographed in nature by Noppadon Makbun, 22 - vi- 2016, Ban Na Kha, Ban Muang, Sakon Nakhon province: (1) male; (2) female." httpUri="https://zenodo.org/record/844476/files/figure.png" pageId="1" pageNumber="438">Fig. 1</figureCitation>
|
||
) as follows: S1 with dorsal and lateral spots; S2 dorsally with triangular spot and laterally with square spot covering auricle; S3 with basal pale yellow ring and small lateral spot; S4–6 with basal ring and middorsal stripe; S7 gradually swollen apically from the middle with basal ring connected to
|
||
<emphasis id="B949553BFFDBB959FF4CFA37FE89F9F4" box="[181,285,1518,1543]" italics="true" pageId="1" pageNumber="438">dumbbell</emphasis>
|
||
-shaped lateral spot (
|
||
<figureCitation id="130695ACFFDBB959FDF4FA37FDF6F9F4" box="[525,610,1518,1543]" captionStart="FIGURE 3 – 12" captionStartId="3.[151,250,1921,1943]" captionTargetBox="[202,1391,193,1900]" captionTargetId="figure@3.[202,1391,193,1906]" captionTargetPageId="3" captionText="FIGURE 3 – 12. Anisogomphus yingsaki sp. nov., holotype male: (3) head in frontal view; (4) synthorax in dorsal view; (5) synthorax in lateral view; (6) left fore- and hindwing; (7) accessory genitalia in lateral view; (8) accessory genitalia in ventral view; (9) vesica spermalis in lateral view; (10) marking on S 7; (11) anal appendages in lateral view; (12) anal appendages in dorsal view." httpUri="https://zenodo.org/record/844478/files/figure.png" pageId="1" pageNumber="438">Fig. 10</figureCitation>
|
||
); S8 with two pale small lateral spots; S9 apically with small middorsal spot; postero-dorsal edge of S8–9 slightly projected when viewed laterally; S10 entirely black.
|
||
</paragraph>
|
||
<paragraph id="8B828929FFDBB959FF3EF9EFFF57F92C" blockId="1.[151,1437,258,2011]" pageId="1" pageNumber="438">
|
||
<emphasis id="B949553BFFDBB959FF3EF9EFFE19F9BC" box="[199,397,1590,1615]" italics="true" pageId="1" pageNumber="438">Anal appendages</emphasis>
|
||
. Length of cerci the same as S10 but longer than epiproct; in lateral view cerci whitish yellow, acutely cone-shaped, slightly curved, tip paler, sharp, and slightly bent upward, its ventral margin at slightly posterior of the middle with a small tooth pointing downward. Cerci slightly convergent in dorsal view; epiproct internally whitish yellow with darker outer margin and apex, diverging and bent inward apically (
|
||
<figureCitation id="130695ACFFDBB959FACDF97BFF22F92C" captionStart="FIGURE 3 – 12" captionStartId="3.[151,250,1921,1943]" captionTargetBox="[202,1391,193,1900]" captionTargetId="figure@3.[202,1391,193,1906]" captionTargetPageId="3" captionText="FIGURE 3 – 12. Anisogomphus yingsaki sp. nov., holotype male: (3) head in frontal view; (4) synthorax in dorsal view; (5) synthorax in lateral view; (6) left fore- and hindwing; (7) accessory genitalia in lateral view; (8) accessory genitalia in ventral view; (9) vesica spermalis in lateral view; (10) marking on S 7; (11) anal appendages in lateral view; (12) anal appendages in dorsal view." httpUri="https://zenodo.org/record/844478/files/figure.png" pageId="1" pageNumber="438">Figs. 11– 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B828929FFDBB959FF3EF932FA89F8D4" blockId="1.[151,1437,258,2011]" pageId="1" pageNumber="438">
|
||
<emphasis id="B949553BFFDBB959FF3EF932FE37F8F0" box="[199,419,1770,1795]" italics="true" pageId="1" pageNumber="438">Accessory genitalia</emphasis>
|
||
. Anterior lamina pale semicircular shape covered with long hairs; posterior hamulus pale with slightly narrower obtuse apex, anterior side apically concave with small subapically tooth (
|
||
<figureCitation id="130695ACFFDBB959FB33F8D7FA9BF8D4" box="[1226,1295,1806,1831]" captionStart="FIGURE 3 – 12" captionStartId="3.[151,250,1921,1943]" captionTargetBox="[202,1391,193,1900]" captionTargetId="figure@3.[202,1391,193,1906]" captionTargetPageId="3" captionText="FIGURE 3 – 12. Anisogomphus yingsaki sp. nov., holotype male: (3) head in frontal view; (4) synthorax in dorsal view; (5) synthorax in lateral view; (6) left fore- and hindwing; (7) accessory genitalia in lateral view; (8) accessory genitalia in ventral view; (9) vesica spermalis in lateral view; (10) marking on S 7; (11) anal appendages in lateral view; (12) anal appendages in dorsal view." httpUri="https://zenodo.org/record/844478/files/figure.png" pageId="1" pageNumber="438">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B828929FFDBB959FF3EF8EDFE1FF860" blockId="1.[151,1437,258,2011]" pageId="1" pageNumber="438">
|
||
<emphasis id="B949553BFFDBB959FF3EF8EDFE11F8B8" box="[199,389,1842,1867]" italics="true" pageId="1" pageNumber="438">Vesica spermalis</emphasis>
|
||
. Vesicle triangular in shape; second segment elongate tubular and inflated distally; median segment expanded forming an elliptic shape; distal segment triangular shaped with apical part modified to a short flagellum (
|
||
<figureCitation id="130695ACFFDBB959FEE8F8A3FEEBF860" box="[273,383,1914,1939]" captionStart="FIGURE 3 – 12" captionStartId="3.[151,250,1921,1943]" captionTargetBox="[202,1391,193,1900]" captionTargetId="figure@3.[202,1391,193,1906]" captionTargetPageId="3" captionText="FIGURE 3 – 12. Anisogomphus yingsaki sp. nov., holotype male: (3) head in frontal view; (4) synthorax in dorsal view; (5) synthorax in lateral view; (6) left fore- and hindwing; (7) accessory genitalia in lateral view; (8) accessory genitalia in ventral view; (9) vesica spermalis in lateral view; (10) marking on S 7; (11) anal appendages in lateral view; (12) anal appendages in dorsal view." httpUri="https://zenodo.org/record/844478/files/figure.png" pageId="1" pageNumber="438">Figs. 8–9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B828929FFDBB959FF3EF847FED9F828" blockId="1.[151,1437,258,2011]" pageId="1" pageNumber="438">
|
||
<emphasis id="B949553BFFDBB959FF3EF847FE5BF844" bold="true" box="[199,463,1950,1975]" pageId="1" pageNumber="438">Measurements (mm):</emphasis>
|
||
Hw 27, abdomen (including anal appendages) 38, total length (with head and appendages) 53.
|
||
</paragraph>
|
||
<caption id="DF42D9A1FFD8B95AFF6EF873FD54F82D" httpUri="https://zenodo.org/record/844476/files/figure.png" pageId="2" pageNumber="439" targetBox="[153,1433,233,1915]" targetPageId="2">
|
||
<paragraph id="8B828929FFD8B95AFF6EF873FD54F82D" blockId="2.[151,1436,1962,2014]" pageId="2" pageNumber="439">
|
||
<emphasis id="B949553BFFD8B95AFF6EF873FEB8F84C" bold="true" box="[151,300,1962,1984]" pageId="2" pageNumber="439">FIGURE 1–2.</emphasis>
|
||
<taxonomicName id="4C3DF2AAFFD8B95AFECAF873FDB6F833" authority="Makbun, 2017" authorityName="Makbun" authorityYear="2017" box="[307,546,1962,1984]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="439" phylum="Arthropoda" rank="species" species="yingsaki" status="sp. nov.">
|
||
<emphasis id="B949553BFFD8B95AFECAF873FDB6F833" box="[307,546,1962,1984]" italics="true" pageId="2" pageNumber="439">Anisogomphus yingsaki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27AE840FFD8B95AFDD0F873FDEEF84C" box="[553,634,1962,1983]" pageId="2" pageNumber="439" rank="species">
|
||
<emphasis id="B949553BFFD8B95AFDD0F873FDEEF84C" bold="true" box="[553,634,1962,1983]" pageId="2" pageNumber="439">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, photographed in nature by Noppadon Makbun, 22-vi-2016, Ban Na Kha, Ban Muang, Sakon Nakhon province: (1) male; (2) female.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF42D9A1FFD9B95BFF6EF858FE86F800" httpUri="https://zenodo.org/record/844478/files/figure.png" pageId="3" pageNumber="440" targetBox="[202,1391,193,1900]" targetPageId="3">
|
||
<paragraph id="8B828929FFD9B95BFF6EF858FE86F800" blockId="3.[151,1436,1921,2035]" pageId="3" pageNumber="440">
|
||
<emphasis id="B949553BFFD9B95BFF6EF858FEAFF865" bold="true" box="[151,315,1921,1943]" pageId="3" pageNumber="440">FIGURE 3–12.</emphasis>
|
||
<taxonomicName id="4C3DF2AAFFD9B95BFEBAF858FDA2F864" authority="Makbun, 2017" authorityName="Makbun" authorityYear="2017" box="[323,566,1921,1943]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="440" phylum="Arthropoda" rank="species" species="yingsaki" status="sp. nov.">
|
||
<emphasis id="B949553BFFD9B95BFEBAF858FDA2F864" box="[323,566,1921,1943]" italics="true" pageId="3" pageNumber="440">Anisogomphus yingsaki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27AE840FFD9B95BFDC4F858FD04F865" box="[573,656,1921,1942]" pageId="3" pageNumber="440" rank="species">
|
||
<emphasis id="B949553BFFD9B95BFDC4F858FD04F865" bold="true" box="[573,656,1921,1942]" pageId="3" pageNumber="440">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male: (3) head in frontal view; (4) synthorax in dorsal view; (5) synthorax in lateral view; (6) left fore- and hindwing; (7) accessory genitalia in lateral view; (8) accessory genitalia in ventral view; (9) vesica spermalis in lateral view; (10) marking on S7; (11) anal appendages in lateral view; (12) anal appendages in dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF42D9A1FFDEB95CFF6EFC00FC54FBFD" httpUri="https://zenodo.org/record/844480/files/figure.png" pageId="4" pageNumber="441" targetBox="[153,1433,183,968]" targetPageId="4">
|
||
<paragraph id="8B828929FFDEB95CFF6EFC00FC54FBFD" blockId="4.[151,1435,985,1038]" pageId="4" pageNumber="441">
|
||
<emphasis id="B949553BFFDEB95CFF6EFC00FED2FC1D" bold="true" box="[151,326,985,1007]" pageId="4" pageNumber="441">FIGURE 13–16.</emphasis>
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFEB4FC00FDA9FC1C" authority="Makbun, 2017" authorityName="Makbun" authorityYear="2017" box="[333,573,985,1007]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="yingsaki" status="sp. nov.">
|
||
<emphasis id="B949553BFFDEB95CFEB4FC00FDA9FC1C" box="[333,573,985,1007]" italics="true" pageId="4" pageNumber="441">Anisogomphus yingsaki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27AE840FFDEB95CFDBCFC00FD02FC1D" box="[581,662,985,1006]" pageId="4" pageNumber="441" rank="species">
|
||
<emphasis id="B949553BFFDEB95CFDBCFC00FD02FC1D" bold="true" box="[581,662,985,1006]" pageId="4" pageNumber="441">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype female: (13) head in frontal view; (14) anal appendages in lateral view; (15) anal appendages in ventral view; (16) valvula vulvae in ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B828929FFDEB95CFF3EFBECFC6AFB2D" blockId="4.[151,1437,1077,2038]" pageId="4" pageNumber="441">
|
||
<emphasis id="B949553BFFDEB95CFF3EFBECFDBEFBBD" bold="true" box="[199,554,1077,1102]" pageId="4" pageNumber="441">
|
||
Variation in
|
||
<typeStatus id="5486378BFFDEB95CFE9EFBECFE46FBBD" box="[359,466,1077,1102]" pageId="4" pageNumber="441" type="paratype">paratype</typeStatus>
|
||
males.
|
||
</emphasis>
|
||
One of the two
|
||
<typeStatus id="5486378BFFDEB95CFD00FBEFFCCEFBBD" box="[761,858,1078,1102]" pageId="4" pageNumber="441" type="paratype">paratype</typeStatus>
|
||
males is smaller and the other same size as the
|
||
<typeStatus id="5486378BFFDEB95CFF6EFB80FF68FB81" box="[151,252,1113,1138]" pageId="4" pageNumber="441" type="holotype">holotype</typeStatus>
|
||
, Hw 27–28, abdomen (including anal appendages) 35.5–38, total length (with head and appendages) 49.5–53. Ax 12–13 on Fw, 9–10 on Hw, Px 8–10 on Fw, 8–9 on Hw. S7 marking of one of the
|
||
<typeStatus id="5486378BFFDEB95CFB31FBA7FAA7FB65" box="[1224,1331,1150,1174]" pageId="4" pageNumber="441" type="paratype">paratypes</typeStatus>
|
||
variable, the posterior spot of dumbbell-shaped lateral spot not connected to the rest. Two male
|
||
<typeStatus id="5486378BFFDEB95CFB85FB7BFB73FB49" box="[1148,1255,1186,1210]" pageId="4" pageNumber="441" type="paratype">paratypes</typeStatus>
|
||
possess a blunt flagellum. This is likely to be a result from penile organ extraction procedure.
|
||
</paragraph>
|
||
<paragraph id="8B828929FFDEB95CFF3EFB30FA85FA29" blockId="4.[151,1437,1077,2038]" pageId="4" pageNumber="441">
|
||
<emphasis id="B949553BFFDEB95CFF3EFB30FE1EFAF1" bold="true" box="[199,394,1257,1282]" pageId="4" pageNumber="441">
|
||
<typeStatus id="5486378BFFDEB95CFF3EFB30FEA0FAF1" box="[199,308,1257,1282]" pageId="4" pageNumber="441" type="paratype">Paratype</typeStatus>
|
||
female
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="130695ACFFDEB95CFE63FB30FE7EFAF2" box="[410,490,1257,1282]" captionStart="FIGURE 1 – 2" captionStartId="2.[151,250,1962,1984]" captionTargetBox="[153,1433,233,1915]" captionTargetId="figure@2.[153,1433,233,1915]" captionTargetPageId="2" captionText="FIGURE 1 – 2. Anisogomphus yingsaki sp. nov., photographed in nature by Noppadon Makbun, 22 - vi- 2016, Ban Na Kha, Ban Muang, Sakon Nakhon province: (1) male; (2) female." httpUri="https://zenodo.org/record/844476/files/figure.png" pageId="4" pageNumber="441">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="130695ACFFDEB95CFE0FFB30FDD4FAF1" box="[502,576,1257,1282]" captionStart="FIGURE 13 – 16" captionStartId="4.[151,250,985,1007]" captionTargetBox="[153,1433,183,968]" captionTargetId="figure@4.[153,1433,183,968]" captionTargetPageId="4" captionText="FIGURE 13 – 16. Anisogomphus yingsaki sp. nov., paratype female: (13) head in frontal view; (14) anal appendages in lateral view; (15) anal appendages in ventral view; (16) valvula vulvae in ventral view." httpUri="https://zenodo.org/record/844480/files/figure.png" pageId="4" pageNumber="441">13–16</figureCitation>
|
||
). Similar to male, slightly stouter and shorter, black with yellow pattern, but differs as follows; occiput black; postclypeus pale bordered with black and broken in the middle; wing slightly tinted with yellow; antenodals 13 on Fw, 8 (right), 9 (left) on Hw; postnodals 9 (right), 10 (left) on Fw, 9 (right), 10 (left) on Hw; abdomen almost evenly cylindrical, without obvious expansion in distal segments; S3 with lateral spot and small middorsal spot; S4–5 with smaller lateral spots and without middorsal spot; S6 with basal ring and the smallest lateral spot; S8 with lateral spot near posterior margin; cerci pale, about the same length as S10, with gradually tapering, pointed tip (
|
||
<figureCitation id="130695ACFFDEB95CFE00FA18FD16FA29" box="[505,642,1473,1498]" captionStart="FIGURE 13 – 16" captionStartId="4.[151,250,985,1007]" captionTargetBox="[153,1433,183,968]" captionTargetId="figure@4.[153,1433,183,968]" captionTargetPageId="4" captionText="FIGURE 13 – 16. Anisogomphus yingsaki sp. nov., paratype female: (13) head in frontal view; (14) anal appendages in lateral view; (15) anal appendages in ventral view; (16) valvula vulvae in ventral view." httpUri="https://zenodo.org/record/844480/files/figure.png" pageId="4" pageNumber="441">Figs. 14–15</figureCitation>
|
||
); valvula vulvae simple and reaching beyond S9 (
|
||
<figureCitation id="130695ACFFDEB95CFB48FA18FA90FA29" box="[1201,1284,1473,1498]" captionStart="FIGURE 13 – 16" captionStartId="4.[151,250,985,1007]" captionTargetBox="[153,1433,183,968]" captionTargetId="figure@4.[153,1433,183,968]" captionTargetPageId="4" captionText="FIGURE 13 – 16. Anisogomphus yingsaki sp. nov., paratype female: (13) head in frontal view; (14) anal appendages in lateral view; (15) anal appendages in ventral view; (16) valvula vulvae in ventral view." httpUri="https://zenodo.org/record/844480/files/figure.png" pageId="4" pageNumber="441">Fig. 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B828929FFDEB95CFF3EFA3CFEF6F9D1" blockId="4.[151,1437,1077,2038]" pageId="4" pageNumber="441">
|
||
<emphasis id="B949553BFFDEB95CFF3EFA3CFE58FA0D" bold="true" box="[199,460,1509,1534]" pageId="4" pageNumber="441">Measurements (mm):</emphasis>
|
||
Hw 29, abdomen (including caudal appendages) 35.5, total length (with head and appendages) 50.5.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327DAA2FFDEB95DFF3EF9F4FA1BFE07" lastPageId="5" lastPageNumber="442" pageId="4" pageNumber="441" type="diagnosis">
|
||
<paragraph id="8B828929FFDEB95CFF3EF9F4FC3DF8B1" blockId="4.[151,1437,1077,2038]" pageId="4" pageNumber="441">
|
||
<emphasis id="B949553BFFDEB95CFF3EF9F4FE5AF9B5" bold="true" box="[199,462,1581,1606]" pageId="4" pageNumber="441">Differential diagnosis.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFDABF9F4FD63F9B5" baseAuthorityName="Schorr & Paulson" baseAuthorityYear="2017" box="[594,759,1581,1606]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949553BFFDEB95CFDABF9F4FD63F9B5" box="[594,759,1581,1606]" italics="true" pageId="4" pageNumber="441">Anisogomphus</emphasis>
|
||
</taxonomicName>
|
||
presently comprises of 16 species, of which two species,
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFA7DF9F7FE15F999" authority="Laidlaw, 1922" authorityName="Laidlaw" authorityYear="1922" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="orites">
|
||
<emphasis id="B949553BFFDEB95CFA7DF9F7FF43F99A" italics="true" pageId="4" pageNumber="441">A. orites</emphasis>
|
||
<bibRefCitation id="EFACF4D8FFDEB95CFF26F988FE15F999" author="Laidlaw" box="[223,385,1617,1642]" pageId="4" pageNumber="441" refString="Laidlaw, F. F. (1922) A list of the dragonflies recorded from the Indian Empire with special reference to the collection of the Indian Museum. Part V. The subfamily Gomphinae. Records of the Indian Museum, 24 (3), 367 - 414." type="journal article" year="1922">Laidlaw, 1922</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFE45F98BFCD2F999" authority="Yousuf & Yunus, 1977" authorityName="Yousuf & Yunus" authorityYear="1977" box="[444,838,1617,1642]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="vulvalis">
|
||
<emphasis id="B949553BFFDEB95CFE45F98BFDA0F999" box="[444,564,1617,1642]" italics="true" pageId="4" pageNumber="441">A. vulvalis</emphasis>
|
||
Yousuf & Yunus, 1977
|
||
</taxonomicName>
|
||
, are known only from female specimens and a third one,
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFF35F9AFFDFEF97D" authority="(Hagen in Selys, 1878)" authorityName="Hagen in Selys" authorityYear="1878" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1878" box="[204,618,1653,1678]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="ceylonicus">
|
||
<emphasis id="B949553BFFDEB95CFF35F9AFFEF5F97D" box="[204,353,1653,1678]" italics="true" pageId="4" pageNumber="441">A. ceylonicus</emphasis>
|
||
(Hagen in Selys, 1878)
|
||
</taxonomicName>
|
||
, is known from a crushed immature male, a female, and exuvia (Bedjanič & van der
|
||
<bibRefCitation id="EFACF4D8FFDEB95CFEE5F943FE2FF941" author="Poorten" box="[284,443,1690,1714]" pageId="4" pageNumber="441" refString="Bedjanic, M. & van der Poorten, N. (2013) On the synonymy of two enigmatic endemic clubtails from Sri Lanka (Anisoptera: Gomphidae). Agrion, 17 (2), 44 - 47." type="journal article" year="2013">Poorten 2013</bibRefCitation>
|
||
). Members of this genus can be divided into two groups by the shape of male anal appendages (
|
||
<bibRefCitation id="EFACF4D8FFDEB95CFED0F964FDD8F925" author="Karube" box="[297,588,1725,1750]" pageId="4" pageNumber="441" refString="Karube, H. & Kompier, T. (2016) Occurrence of a new gomphid dragonfly Anisogomphus neptunus sp. nov., from northern Vietnam. Tombo - Acta Odonatologica Japonica, 58, 35 - 39." type="journal article" year="2016">Karube & Kompier 2016</bibRefCitation>
|
||
). The first group has simple rod-like cerci and the other has more complex cerci, which are joined or almost joined to each other.
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFD04F938FB93F909" authority="Makbun, 2017" authorityName="Makbun" authorityYear="2017" box="[765,1031,1761,1786]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="yingsaki" status="sp. nov.">
|
||
<emphasis id="B949553BFFDEB95CFD04F938FB93F909" box="[765,1031,1761,1786]" italics="true" pageId="4" pageNumber="441">Anisogomphus yingsaki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27AE840FFDEB95CFBF6F938FBFDF909" box="[1039,1129,1761,1786]" pageId="4" pageNumber="441" rank="species">
|
||
<emphasis id="B949553BFFDEB95CFBF6F938FBFDF909" bold="true" box="[1039,1129,1761,1786]" pageId="4" pageNumber="441">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is in the first group, which includes species such as
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFE48F8DFFD4FF8ED" authority="(Selys, 1854)" authorityName="Selys" authorityYear="1854" baseAuthorityName="Selys" baseAuthorityYear="1854" box="[433,731,1797,1822]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="bivittatus">
|
||
<emphasis id="B949553BFFDEB95CFE48F8DFFDAEF8ED" box="[433,570,1797,1822]" italics="true" pageId="4" pageNumber="441">A. bivittatus</emphasis>
|
||
(Selys, 1854)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFD11F8DFFCEBF8ED" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1878" box="[744,895,1797,1822]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="ceylonicus">
|
||
<emphasis id="B949553BFFDEB95CFD11F8DFFCEBF8ED" box="[744,895,1797,1822]" italics="true" pageId="4" pageNumber="441">A. ceylonicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFC77F8DFFB25F8ED" authority="Klots, 1947" authorityName="Klots" authorityYear="1947" box="[910,1201,1797,1822]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="flavifacies">
|
||
<emphasis id="B949553BFFDEB95CFC77F8DFFBB5F8ED" box="[910,1057,1797,1822]" italics="true" pageId="4" pageNumber="441">A. flavifacies</emphasis>
|
||
<bibRefCitation id="EFACF4D8FFDEB95CFBD3F8DCFB25F8ED" author="Klots" box="[1066,1201,1797,1822]" pageId="4" pageNumber="441" refString="Klots, E. B. (1947) Chinese dragonflies (Odonata) in the American Museum of Natural History. American Museum Novitates, 1341, 1 - 15." type="journal article" year="1947">Klots, 1947</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFB44F8DFFF4EF8B1" authority="(Morton, 1928)" authorityName="Morton" authorityYear="1928" baseAuthorityName="Morton" baseAuthorityYear="1928" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="forresti">
|
||
<emphasis id="B949553BFFDEB95CFB44F8DFFABBF8ED" box="[1213,1327,1798,1822]" italics="true" pageId="4" pageNumber="441">A. forresti</emphasis>
|
||
(
|
||
<bibRefCitation id="EFACF4D8FFDEB95CFAC6F8DFFF46F8B1" author="Morton" pageId="4" pageNumber="441" refString="Morton, K. J. (1928) Notes on the Odonata of Yunnan, with descriptions of new species. Transactions of the Entomological Society of London, 1928, 109 - 118. https: // doi. org / 10.1111 / j. 1365 - 2311.1928. tb 01194. x" type="journal article" year="1928">Morton, 1928</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFF1FF8F3FE61F8B1" authority="(Selys, 1872)" authorityName="Selys" authorityYear="1872" baseAuthorityName="Selys" baseAuthorityYear="1872" box="[230,501,1833,1858]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="maacki">
|
||
<emphasis id="B949553BFFDEB95CFF1FF8F3FEC2F8B1" box="[230,342,1833,1858]" italics="true" pageId="4" pageNumber="441">A. maacki</emphasis>
|
||
(Selys, 1872)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFDC9F8F3FC31F8B1" authority="Yang & Davies, 1993" authorityName="Yang & Davies" authorityYear="1993" box="[560,933,1833,1858]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="resortus">
|
||
<emphasis id="B949553BFFDEB95CFDC9F8F3FD3EF8B2" box="[560,682,1834,1857]" italics="true" pageId="4" pageNumber="441">A. resortus</emphasis>
|
||
<bibRefCitation id="EFACF4D8FFDEB95CFD48F8F3FC31F8B1" author="Yang" box="[689,933,1833,1858]" pageId="4" pageNumber="441" refString="Yang, B. & Davies, D. A. L. (1993) Gomphid dragonflies of Yunnan, China, with descriptions of new species and some views on the origin of the group (Anisoptera: Gomphidae). Odonatologica, 22 (1), 45 - 62." type="journal article" year="1993">Yang & Davies, 1993</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B828929FFDEB95CFF3EF894FB26F805" blockId="4.[151,1437,1077,2038]" pageId="4" pageNumber="441">
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFF3EF894FE46F895" authority="Makbun, 2017" authorityName="Makbun" authorityYear="2017" box="[199,466,1869,1894]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="yingsaki" status="sp. nov.">
|
||
<emphasis id="B949553BFFDEB95CFF3EF894FE46F895" box="[199,466,1869,1894]" italics="true" pageId="4" pageNumber="441">Anisogomphus yingsaki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27AE840FFDEB95CFE25F894FDADF895" box="[476,569,1869,1894]" pageId="4" pageNumber="441" rank="species">
|
||
<emphasis id="B949553BFFDEB95CFE25F894FDADF895" bold="true" box="[476,569,1869,1894]" pageId="4" pageNumber="441">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is most similar to
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFCEFF897FC34F895" baseAuthorityName="Selys" baseAuthorityYear="1854" box="[790,928,1869,1894]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="bivittatus">
|
||
<emphasis id="B949553BFFDEB95CFCEFF897FC34F895" box="[790,928,1869,1894]" italics="true" pageId="4" pageNumber="441">A. bivittatus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F32AC9B9FFDEB95CFC13F894FBB0F895" box="[1002,1060,1869,1894]" name="India" pageId="4" pageNumber="441">India</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32AC9B9FFDEB95CFBA6F894FB31F895" box="[1119,1189,1869,1894]" name="Nepal" pageId="4" pageNumber="441">Nepal</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFB4AF897FADDF895" authorityName="Klots" authorityYear="1947" box="[1203,1353,1869,1894]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="flavifacies">
|
||
<emphasis id="B949553BFFDEB95CFB4AF897FADDF895" box="[1203,1353,1869,1894]" italics="true" pageId="4" pageNumber="441">A. flavifacies</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3DF2AAFFDEB95CFA7DF897FF66F87A" authorityName="Yang & Davies" authorityYear="1993" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="resortus">
|
||
<emphasis id="B949553BFFDEB95CFA7DF897FF66F87A" italics="true" pageId="4" pageNumber="441">A. resortus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F32AC9B9FFDEB95CFECFF8A8FEE8F879" box="[310,380,1905,1930]" name="China" pageId="4" pageNumber="441">China</collectingCountry>
|
||
, with which it shares similar shape of anal appendages. However, it can be distinguished from them by a combination of the following characters: antehumeral stripes on mesepisternum almost straight and broader in the middle, S7 with basal ring connected to
|
||
<emphasis id="B949553BFFDEB95CFCF0F860FCE5F821" box="[777,881,1977,2002]" italics="true" pageId="4" pageNumber="441">dumbbell</emphasis>
|
||
-shaped lateral spot, vesica spermalis with apical part modified to a short flagellum, and female valvula vulvae simple and reaching beyond S9.
|
||
</paragraph>
|
||
<paragraph id="8B828929FFDFB95DFF3EFF41FAA0FEB3" blockId="5.[151,1436,151,824]" pageId="5" pageNumber="442">
|
||
It should be noted that a short flagellum of vesica spermalis of
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFC54FF41FBB9FF43" authority="Makbun, 2017" authorityName="Makbun" authorityYear="2017" box="[941,1069,151,176]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="yingsaki" status="sp. nov.">
|
||
<emphasis id="B949553BFFDFB95DFC54FF41FBB9FF43" box="[941,1069,151,176]" italics="true" pageId="5" pageNumber="442">A. yingsaki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27AE840FFDFB95DFBCEFF4EFB00FF43" box="[1079,1172,151,176]" pageId="5" pageNumber="442" rank="species">
|
||
<emphasis id="B949553BFFDFB95DFBCEFF4EFB00FF43" bold="true" box="[1079,1172,151,176]" pageId="5" pageNumber="442">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is unique amongst the members whose vesica spermalis were illustrated (
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFD29FF65FCF7FF26" authorityName="Klots" authorityYear="1947" box="[720,867,188,213]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="flavifacies">
|
||
<emphasis id="B949553BFFDFB95DFD29FF65FCF7FF26" box="[720,867,188,213]" italics="true" pageId="5" pageNumber="442">A. flavifacies</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFC88FF65FC7FFF27" authorityName="Yang & Davies" authorityYear="1993" box="[881,1003,188,212]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="resortus">
|
||
<emphasis id="B949553BFFDFB95DFC88FF65FC7FFF27" box="[881,1003,188,212]" italics="true" pageId="5" pageNumber="442">A. resortus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFC00FF65FBFDFF26" baseAuthorityName="Selys" baseAuthorityYear="1872" box="[1017,1129,188,213]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="maacki">
|
||
<emphasis id="B949553BFFDFB95DFC00FF65FBFDFF26" box="[1017,1129,188,213]" italics="true" pageId="5" pageNumber="442">A. maacki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFB8EFF65FF46FF0B" authority="Yang & Davies, 1993" authorityName="Yang & Davies" authorityYear="1993" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="nitidus">
|
||
<emphasis id="B949553BFFDFB95DFB8EFF65FB76FF26" box="[1143,1250,188,213]" italics="true" pageId="5" pageNumber="442">A. nitidus</emphasis>
|
||
<bibRefCitation id="EFACF4D8FFDFB95DFB10FF64FF46FF0B" author="Yang" pageId="5" pageNumber="442" refString="Yang, B. & Davies, D. A. L. (1993) Gomphid dragonflies of Yunnan, China, with descriptions of new species and some views on the origin of the group (Anisoptera: Gomphidae). Odonatologica, 22 (1), 45 - 62." type="journal article" year="1993">Yang & Davies, 1993</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFF1BFF39FDC4FF0B" authority="(Karube, 2001)" authorityName="Karube" authorityYear="2001" baseAuthorityName="Karube" baseAuthorityYear="2001" box="[226,592,223,248]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="tamdaoensis">
|
||
<emphasis id="B949553BFFDFB95DFF1BFF39FE06FF0B" box="[226,402,223,248]" italics="true" pageId="5" pageNumber="442">A. tamdaoensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFACF4D8FFDFB95DFE5CFF06FDDCFF0B" author="Karube" box="[421,584,223,248]" pageId="5" pageNumber="442" refString="Karube, H. (2001) Three new species of Gomphidae from Vietnam (Anisoptera). Odonatologica, 30 (3), 271 - 279." type="journal article" year="2001">Karube, 2001</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFD98FF39FBB2FF0B" authority="Karube & Kompier, 2016" authorityName="Karube & Kompier" authorityYear="2016" box="[609,1062,223,248]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="neptunus">
|
||
<emphasis id="B949553BFFDFB95DFD98FF39FD7DFF0B" box="[609,745,224,248]" italics="true" pageId="5" pageNumber="442">A. neptunus</emphasis>
|
||
<bibRefCitation id="EFACF4D8FFDFB95DFD0CFF06FBB2FF0B" author="Karube" box="[757,1062,223,248]" pageId="5" pageNumber="442" refString="Karube, H. & Kompier, T. (2016) Occurrence of a new gomphid dragonfly Anisogomphus neptunus sp. nov., from northern Vietnam. Tombo - Acta Odonatologica Japonica, 58, 35 - 39." type="journal article" year="2016">Karube & Kompier, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFBCFFF39FB73FF0B" box="[1078,1255,224,248]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="yanagisawai">
|
||
<emphasis id="B949553BFFDFB95DFBCFFF39FB73FF0B" box="[1078,1255,224,248]" italics="true" pageId="5" pageNumber="442">A. yanagisawai</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFAD6FF39FEDDFEEF" authority="Lieftinck, 1948" authorityName="Lieftinck" authorityYear="1948" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="anderi">
|
||
<emphasis id="B949553BFFDFB95DFAD6FF39FA08FF0B" box="[1327,1436,223,248]" italics="true" pageId="5" pageNumber="442">A. anderi</emphasis>
|
||
Lieftinck, 1948
|
||
</taxonomicName>
|
||
). Distal part of vesica spermalis of those members possess a blunt end but
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFB59FEDDFA89FEEE" authority="Makbun, 2017" authorityName="Makbun" authorityYear="2017" box="[1184,1309,260,285]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="yingsaki" status="sp. nov.">
|
||
<emphasis id="B949553BFFDFB95DFB59FEDDFA89FEEE" box="[1184,1309,260,285]" italics="true" pageId="5" pageNumber="442">A. yingsaki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27AE840FFDFB95DFADFFEDDFA15FEEE" box="[1318,1409,260,285]" pageId="5" pageNumber="442" rank="species">
|
||
<emphasis id="B949553BFFDFB95DFADFFEDDFA15FEEE" bold="true" box="[1318,1409,260,285]" pageId="5" pageNumber="442">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is the first member in the genus that has a short flagellum, a condition not previously reported in this genus.
|
||
</paragraph>
|
||
<paragraph id="8B828929FFDFB95DFF3EFE95FA1BFE07" blockId="5.[151,1436,151,824]" pageId="5" pageNumber="442">
|
||
<emphasis id="B949553BFFDFB95DFF3EFE95FDD9FE96" bold="true" box="[199,589,332,357]" pageId="5" pageNumber="442">Notes on habitat and behaviour.</emphasis>
|
||
While many
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFD12FE95FC04FE96" baseAuthorityName="Schorr & Paulson" baseAuthorityYear="2017" box="[747,912,332,357]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949553BFFDFB95DFD12FE95FC04FE96" box="[747,912,332,357]" italics="true" pageId="5" pageNumber="442">Anisogomphus</emphasis>
|
||
</taxonomicName>
|
||
species are found near montane streams (e.g.
|
||
<bibRefCitation id="EFACF4D8FFDFB95DFF6EFEA9FEB1FE7B" author="Fraser" box="[151,293,368,392]" pageId="5" pageNumber="442" refString="Fraser, F. C. (1934) The Fauna of British India, including Ceylon and Burma. Odonata. Uol. II. Taylor and Francis, London, 398 pp., 4 col. pls." type="book" year="1934">Fraser 1934</bibRefCitation>
|
||
),
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFEC4FEA9FE2AFE7B" authority="Makbun, 2017" authorityName="Makbun" authorityYear="2017" box="[317,446,367,392]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="yingsaki" status="sp. nov.">
|
||
<emphasis id="B949553BFFDFB95DFEC4FEA9FE2AFE7B" box="[317,446,367,392]" italics="true" pageId="5" pageNumber="442">A. yingsaki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27AE840FFDFB95DFE33FEB6FDBCFE7B" box="[458,552,367,392]" pageId="5" pageNumber="442" rank="species">
|
||
<emphasis id="B949553BFFDFB95DFE33FEB6FDBCFE7B" bold="true" box="[458,552,367,392]" pageId="5" pageNumber="442">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was found perching on grass near large river with a sandy bottom. These dragonflies often perched on leaf surfaces, 20–30 centimetres above the ground, on sunny days and disappeared when it was clouded. They would perch nearby or higher when threatened. Other interesting species found in this area include
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFEDFFE05FD72FE06" authority="Asahina, 1986" authorityName="Asahina" authorityYear="1986" box="[294,742,476,501]" class="Insecta" family="Gomphidae" genus="Macrogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="matsukii">
|
||
<emphasis id="B949553BFFDFB95DFEDFFE05FDAFFE06" box="[294,571,476,501]" italics="true" pageId="5" pageNumber="442">Macrogomphus matsukii</emphasis>
|
||
Asahina, 1986
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFD08FE05FB14FE07" authority="Selys, 1871" authorityName="Selys" authorityYear="1871" box="[753,1152,476,501]" class="Insecta" family="Corduliidae" genus="Epophthalmia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="frontalis">
|
||
<emphasis id="B949553BFFDFB95DFD08FE05FC61FE06" box="[753,1013,476,501]" italics="true" pageId="5" pageNumber="442">Epophthalmia frontalis</emphasis>
|
||
Selys, 1871
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFB45FE05FAFDFE06" authorityName="Carle" authorityYear="1986" box="[1212,1385,476,501]" class="Insecta" family="Gomphidae" genus="Nychogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949553BFFDFB95DFB45FE05FAFDFE06" box="[1212,1385,476,501]" italics="true" pageId="5" pageNumber="442">Nychogomphus</emphasis>
|
||
</taxonomicName>
|
||
sp.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327DAA2FFDFB95DFF3EFE26FB0CFD54" pageId="5" pageNumber="442" type="description">
|
||
<paragraph id="8B828929FFDFB95DFF3EFE26FB0CFD54" blockId="5.[151,1436,151,824]" pageId="5" pageNumber="442">
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFF3EFE26FE45FDEB" authority="Makbun, 2017" authorityName="Makbun" authorityYear="2017" box="[199,465,511,536]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="yingsaki" status="sp. nov.">
|
||
<emphasis id="B949553BFFDFB95DFF3EFE26FE45FDEB" box="[199,465,511,536]" italics="true" pageId="5" pageNumber="442">Anisogomphus yingsaki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27AE840FFDFB95DFE23FE26FDA1FDEB" box="[474,565,511,536]" pageId="5" pageNumber="442" rank="species">
|
||
<emphasis id="B949553BFFDFB95DFE23FE26FDA1FDEB" bold="true" box="[474,565,511,536]" pageId="5" pageNumber="442">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
adults were also found patrolling along the river as dusk fell (from 17.00 to 19.00) on
|
||
<date id="FF83AFE9FFDFB95DFEF5FDFCFE4EFDCE" box="[268,474,548,573]" pageId="5" pageNumber="442" value="2016-06-21" valueMax="2016-06-23" valueMin="2016-06-21">June 21–23, 2016</date>
|
||
. My colleague and I observed both males and females (but mostly males) patrolling over the water surface and hovering for few minutes before patrolling again. After doing this for some time, some of them perched on nearby bushes. We also once witnessed a hovering male grasping a female and flying into the forest near the river. This crepuscular activity was observed also in
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFC72FD48FB9BFD5B" authorityName="Asahina" authorityYear="1986" box="[907,1039,655,680]" class="Insecta" family="Gomphidae" genus="Macrogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="matsukii">
|
||
<emphasis id="B949553BFFDFB95DFC72FD48FB9BFD5B" box="[907,1039,655,680]" italics="true" pageId="5" pageNumber="442">M. matsukii</emphasis>
|
||
</taxonomicName>
|
||
in this area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327DAA2FFDFB95DFF3EFD6DFF7BFCCB" pageId="5" pageNumber="442" type="distribution">
|
||
<paragraph id="8B828929FFDFB95DFF3EFD6DFF7BFCCB" blockId="5.[151,1436,151,824]" pageId="5" pageNumber="442">
|
||
<emphasis id="B949553BFFDFB95DFF3EFD6DFEF4FD3E" bold="true" box="[199,352,692,717]" pageId="5" pageNumber="442">Distribution.</emphasis>
|
||
Besides its
|
||
<typeStatus id="5486378BFFDFB95DFE16FD6CFD8BFD3E" box="[495,543,693,717]" pageId="5" pageNumber="442">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="49F947CBFFDFB95DFD5FFD6DFCDBFD3E" box="[678,847,692,717]" country="Thailand" name="Sakon Nakhon" pageId="5" pageNumber="442">Sakon Nakhon</collectingRegion>
|
||
in the northeast
|
||
<collectingCountry id="F32AC9B9FFDFB95DFBEDFD6DFBEEFD3E" box="[1044,1146,692,717]" name="Thailand" pageId="5" pageNumber="442">Thailand</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C3DF2AAFFDFB95DFB71FD6DFA93FD3E" authority="Makbun, 2017" authorityName="Makbun" authorityYear="2017" box="[1160,1287,692,717]" class="Insecta" family="Gomphidae" genus="Anisogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="yingsaki" status="sp. nov.">
|
||
<emphasis id="B949553BFFDFB95DFB71FD6DFA93FD3E" box="[1160,1287,692,717]" italics="true" pageId="5" pageNumber="442">A. yingsaki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27AE840FFDFB95DFAE8FD6DFAF9FD3E" box="[1297,1389,692,717]" pageId="5" pageNumber="442" rank="species">
|
||
<emphasis id="B949553BFFDFB95DFAE8FD6DFAF9FD3E" bold="true" box="[1297,1389,692,717]" pageId="5" pageNumber="442">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
has been recently photographed in
|
||
<collectingRegion id="49F947CBFFDFB95DFE0DFD0EFDE0FD03" box="[500,628,727,752]" country="Thailand" name="Chiang Rai" pageId="5" pageNumber="442">Chiang Rai</collectingRegion>
|
||
(A.J. Pierce, pers. comm.) and in
|
||
<collectingRegion id="49F947CBFFDFB95DFC0FFD0EFB11FD03" box="[1014,1157,727,752]" country="Thailand" name="Prachin Buri" pageId="5" pageNumber="442">Prachin Buri</collectingRegion>
|
||
(J. Sim, pers. comm.) in the North and South-east, respectively. According to Mr. Yingsak Paweenpermsuk, the flight period is from May to August.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |