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<document id="D41D6DCD96F47B28D80CE339D56D7361" ID-CLB-Dataset="22188" ID-DOI="10.5852/ejt.2018.471" ID-GBIF-Dataset="28e681c4-4f53-4780-ad09-6a7b3b1b60f1" ID-Zenodo-Dep="3825049" ID-ZooBank="58777EEB-7C5D-4887-8BE2-CEE4A4115CF5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589388819463" checkinUser="valdenar" docAuthor="Agnèse, Jean-François, Chirio, Laurent, Legros, Olivier, Oslisly, Richard &amp; Bhé, Hervé Mvé" docDate="2018" docId="038C87C6FFE7FFBAFE57FEA221F9F9D2" docLanguage="en" docName="ejt-471_agnese_chirico_legros_oslisly_bhe.pdf.imf" docOrigin="European Journal of Taxonomy 471" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aphyosemion aurantiacum Chirio, Legros &amp; Agnese 2018, sp. nov." docType="treatment" docUuid="93840539-567F-4817-B774-3568D388E094" docUuidSource="ZooBank" docVersion="8" lastPageNumber="21" masterDocId="FFB5FFBEFFF4FFAEFFF9FFA9233DFFF1" masterDocTitle="Unexpected discovery of six new species of Aphyosemion (Cyprinodontiformes, Aplocheilidae) in the Wonga-Wongué Presidential Reserve in Gabon" masterLastPageNumber="28" masterPageNumber="1" pageNumber="20" updateTime="1698839408374" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="8B4CE823143EAC289A44E2AEE4C479DC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D08E03AA90131923D8459374416EAC9F">
<mods:title id="E2043CC2758AD91DD7D0E689C070DAD7">Unexpected discovery of six new species of Aphyosemion (Cyprinodontiformes, Aplocheilidae) in the Wonga-Wongué Presidential Reserve in Gabon</mods:title>
</mods:titleInfo>
<mods:name id="D4B69C5D5900DC3528B6CC6106C23D7E" type="personal">
<mods:role id="C5EB1BF820320FB1F61DDC7F43E2D978">
<mods:roleTerm id="202ADFF099768DCD005BF6F95F778745">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="66EEA658BBAA22988A06CD18843ADDD8">Agnèse, Jean-François</mods:namePart>
<mods:nameIdentifier id="857CB8A393E979946048F27E31CAAC56" type="LSID">8AEF0368-6444-4E50-8C72-23CCC65E69A5</mods:nameIdentifier>
<mods:affiliation id="0E61669FD90459F26BAA149BE619A4B4">Univ. Montpellier, IRD, EPHE, Montpellier, France. 14, rue des Roses, 06130 Grasse, France. Floralaan 51, 1501 Buizingen, Belgium. Patrimoines Locaux et Gouvernance UMR 208, IRD, MNHN, Paris, France. Agence Nationale des Parcs Nationaux (ANPN), BP 20379, Libreville, Gabon. Institut de Recherches Agronomiques et Forestières (IRAF), Centre National de la Recherche Scientifique et Technologique (CENAREST), BP 2246, Libreville, Gabon.</mods:affiliation>
<mods:nameIdentifier id="328491AE3851718DE3CA3A46FE98578C" type="email">jean-francois.agnese@ird.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="77D41739B25C510655C179C3A850A787" type="personal">
<mods:role id="6100F9BBC97165A52A2867989589694A">
<mods:roleTerm id="48BCFFFAD731EFDD3D019CF4B6BFFAC2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AC87222999882F9B66F2188463EB3B28">Chirio, Laurent</mods:namePart>
<mods:nameIdentifier id="03EF8527EABD41BD8C98E8FF3A717333" type="LSID">33A241AD-4C70-4A0A-8E30-B137E4FC771F</mods:nameIdentifier>
<mods:nameIdentifier id="0ACEF411A3847453E5E384C673406B44" type="email">lchirio@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="15F7C0C1D9352991B4B96E5F9BE12383" type="personal">
<mods:role id="A347B416059CBB8C8F5B49A76C886039">
<mods:roleTerm id="D7313F0A2903352DB8405201EC7982E0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="030A6DEDA2651EF6BA6A35E43534F6B8">Legros, Olivier</mods:namePart>
<mods:nameIdentifier id="A0CE375B4CF5107AA665B88AEC4ACDCC" type="LSID">F13EB21F-A391-451B-8139-9AA0F90AEF52</mods:nameIdentifier>
<mods:nameIdentifier id="6A1DA0B56D5A31B2D96F1E9DFAD509C8" type="email">olivier.legros@belgacom.net</mods:nameIdentifier>
</mods:name>
<mods:name id="519BDA3A1C5D59852D86572B20E87206" type="personal">
<mods:role id="2F07E50927D70131CE4C0641C27AC6A6">
<mods:roleTerm id="DFA08DABD628D671B40812ECB29096A6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75BC85DF9387507DF3301E84D0C8FD4D">Oslisly, Richard</mods:namePart>
<mods:nameIdentifier id="D39B30D753E54E51A4455784046D1151" type="LSID">1DC16C89-E5DB-43F2-874D-AAB2DC8BAF50</mods:nameIdentifier>
<mods:nameIdentifier id="B6E045225F486EDB24EE6E537FBFDE9D" type="email">richard.oslisly@ird.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="CB42BFE251C485E1D8950EB53D1F5994" type="personal">
<mods:role id="D451C51753834ADF5BF424D75739D92E">
<mods:roleTerm id="5F65F118F085212354646126346F4B66">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B8174F8782F8A2D9A0C8E26A7D1685BC">Bhé, Hervé Mvé</mods:namePart>
<mods:nameIdentifier id="0ADBA452D6636AE6B766A9C77B3DC9CD" type="LSID">2F0F18BD-C1AF-401B-94FE-0CB0AD1A8DE5</mods:nameIdentifier>
<mods:nameIdentifier id="BA98F3A084FB73AD692280A5D226D443" type="email">woleuntem@hotmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="EF4B3DA193E4432C17D144B3729A92A4">text</mods:typeOfResource>
<mods:relatedItem id="4D2B33AFFE280709AA9B226050BDEAB5" type="host">
<mods:titleInfo id="94C1444131B0B57330B8229EA71CD487">
<mods:title id="A5F8BB8AFC8D42022FE794C182026659">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="011CDECE0C4DF7E7117449C1E56557E5">
<mods:date id="6A33D8CBEC40151AFDAF4A4B1EECF0CE">2018</mods:date>
<mods:detail id="C44E05BA62D3D470BB22AD4BBCDD4C33" type="pubDate">
<mods:number id="93A0ADC829742E9FD3F478F131FB59AF">2018-10-30</mods:number>
</mods:detail>
<mods:detail id="D675850C3D9BBCB259340455659F0A42" type="volume">
<mods:number id="0E78585474F95078190C24304E9AA5E2">471</mods:number>
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<mods:start id="CC9A8540D033EA4FBB341C26B344C57F">1</mods:start>
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<mods:classification id="4B1029A28589D925D5B23AB76B4BE167">journal article</mods:classification>
<mods:identifier id="310907380415B7F9494AF8E7C58348E2" type="CLB-Dataset">22188</mods:identifier>
<mods:identifier id="C6EB5ADE623F6D46268B67AAC5637179" type="DOI">10.5852/ejt.2018.471</mods:identifier>
<mods:identifier id="1A852F99E6A3CA51264B4FDA998AA775" type="GBIF-Dataset">28e681c4-4f53-4780-ad09-6a7b3b1b60f1</mods:identifier>
<mods:identifier id="9E757E7966CE17A2A1D32CD7B3D88C6C" type="Zenodo-Dep">3825049</mods:identifier>
<mods:identifier id="48B7C98D8DA1E758154B984DDFC0F04A" type="ZooBank">58777EEB-7C5D-4887-8BE2-CEE4A4115CF5</mods:identifier>
</mods:mods>
<treatment id="038C87C6FFE7FFBAFE57FEA221F9F9D2" ID-DOI="http://doi.org/10.5281/zenodo.3846003" ID-GBIF-Taxon="164130212" ID-Zenodo-Dep="3846003" LSID="urn:lsid:zoobank.org:act:93840539-567F-4817-B774-3568D388E094" httpUri="http://treatment.plazi.org/id/038C87C6FFE7FFBAFE57FEA221F9F9D2" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="C33F655BFFE7FFBDFE57FEA2278EFEBB" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8B9A36D0FFE7FFBDFE57FEA227BBFED7" blockId="19.[385,1203,267,366]" box="[430,1158,267,294]" pageId="19" pageNumber="20">
<heading id="D0D281BCFFE7FFBDFE57FEA227BBFED7" box="[430,1158,267,294]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
<taxonomicName id="4C254D53FFE7FFBDFE57FEA22719FED4" authority="Chirio, Legros &amp; Agnese" authorityName="Chirio, Legros &amp; Agnese" authorityYear="2018" box="[430,1060,267,294]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="aurantiacum" status="sp. nov.">
<emphasis id="B951EAC2FFE7FFBDFE57FEA221CBFED4" bold="true" box="[430,758,267,294]" italics="true" pageId="19" pageNumber="20">Aphyosemion aurantiacum</emphasis>
Chirio, Legros &amp; Agnèse
</taxonomicName>
<taxonomicNameLabel id="A26257B9FFE7FFBDFBD5FEA527BBFED7" box="[1068,1158,268,294]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B9A36D0FFE7FFBDFE78FE86278EFEBB" blockId="19.[385,1203,267,366]" box="[385,1203,303,330]" pageId="19" pageNumber="20">
<uri id="FFB43AD2FFE7FFBDFE78FE86278EFEBB" box="[385,1203,303,330]" pageId="19" pageNumber="20">
urn:lsid:zoobank.org:act:
<uuid id="FF830C05FFE7FFBDFD65FE86278EFEBB" box="[668,1203,303,330]" pageId="19" pageNumber="20">93840539-567F-4817-B774-3568D388E094</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C33F655BFFE7FFBDFD50FEFA20B6FE9F" box="[681,907,339,366]" pageId="19" pageNumber="20" type="description">
<paragraph id="8B9A36D0FFE7FFBDFD50FEFA20B6FE9F" blockId="19.[385,1203,267,366]" box="[681,907,339,366]" pageId="19" pageNumber="20">
<figureCitation id="131E2A55FFE7FFBDFD50FEFA21D1FE9C" box="[681,748,339,365]" captionStart="Fig" captionStartId="21.[189,232,1788,1814]" captionTargetBox="[228,1364,265,1702]" captionTargetId="figure@21.[228,1364,265,1743]" captionTargetPageId="21" captionText="Fig. 6. A–D. Aphyosemion aurantiacum Chirio, Legros &amp; Agnèse sp. nov. A. Adult, ♁, from locality 12, not preserved. Photo O. Buisson. B. Adult, ♀, from locality 12, not preserved. Photo O. Buisson. C. Holotype, adult, ♁, from Wézé Spring (MRAC 2016-019-P-64). D. Paratype, adult, ♀, from Wézé Spring (MRAC 2016-019-P-65-73). E–H. A. rubrogaster Chirio, Legros &amp; Agnèse sp. nov. E. Adult, ♁, from locality 16, not preserved. Photo O. Buisson. F. Adult, ♀, from locality 16, not preserved. Photo O. Buisson. G. Holotype, adult, ♁, from the Niengé River (MRAC 2016-019-P-93). H. Paratype, adult, ♀, from the Niengé River (MRAC 2016-019-P-94-108)." figureDoi="http://doi.org/10.5281/zenodo.3825065" httpUri="https://zenodo.org/record/3825065/files/figure.png" pageId="19" pageNumber="20">Fig. 6</figureCitation>
A–D,
<tableCitation id="C6A7036BFFE7FFBDFCCDFEFA20B6FE9F" box="[820,907,339,366]" captionStart="Table 7" captionStartId="22.[189,258,268,294]" captionText="Table 7. Meristic and morphometric data of Aphyosemion aurantiacum Chirio, Legros &amp; Agnèse sp. nov. See Table 3 for the abbreviations used." pageId="19" pageNumber="20">Table 7</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33F655BFFE7FFBDFF44FE3F271BFE2F" box="[189,1062,406,478]" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8B9A36D0FFE7FFBDFF44FE3F2279FE41" blockId="19.[189,324,406,432]" box="[189,324,406,432]" pageId="19" pageNumber="20">
<heading id="D0D281BCFFE7FFBDFF44FE3F2279FE41" bold="true" box="[189,324,406,432]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B951EAC2FFE7FFBDFF44FE3F2279FE41" bold="true" box="[189,324,406,432]" pageId="19" pageNumber="20">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A36D0FFE7FFBDFF44FE6D271BFE2F" blockId="19.[189,1062,452,478]" box="[189,1062,452,478]" pageId="19" pageNumber="20">The specific epithet refers to the main colour of this species (dark orange).</paragraph>
</subSubSection>
<subSubSection id="C33F655BFFE7FFBDFF44FDAF277FFC7F" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8B9A36D0FFE7FFBDFF44FDAF2297FDD1" blockId="19.[189,426,518,544]" box="[189,426,518,544]" pageId="19" pageNumber="20">
<heading id="D0D281BCFFE7FFBDFF44FDAF2297FDD1" bold="true" box="[189,426,518,544]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B951EAC2FFE7FFBDFF44FDAF2297FDD1" bold="true" box="[189,426,518,544]" pageId="19" pageNumber="20">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B4D3C8DFFE7FFBDFF13FD9C2605FD66" ID-GBIF-Occurrence="2615267302" collectingDate="2014-07-03" collectionCode="CHRSP, MRAC" collectorName="Weze &amp; Laurent Chirio" country="Gabon" elevation="89" latitude="-0.58186" location="Weze" longLatPrecision="1" longitude="9.46718" pageId="19" pageNumber="20" specimenCode="CHRSP5, MRAC 2016-019" specimenCount="1" typeStatus="holotype">
<paragraph id="8B9A36D0FFE7FFBDFF13FD9C2261FDBE" blockId="19.[189,1398,565,663]" box="[234,348,565,591]" pageId="19" pageNumber="20">
<emphasis id="B951EAC2FFE7FFBDFF13FD9C2261FDBE" bold="true" box="[234,348,565,591]" pageId="19" pageNumber="20">
<typeStatus id="549E8872FFE7FFBDFF13FD9C2261FDBE" box="[234,348,565,591]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9A36D0FFE7FFBDFF44FDF12605FD66" blockId="19.[189,1398,565,663]" pageId="19" pageNumber="20">
<collectingCountry id="F3327640FFE7FFBDFF44FDF12217FD83" box="[189,298,600,626]" name="Gabon" pageId="19" pageNumber="20">GABON</collectingCountry>
: adult ♁,
<quantity id="4CDD9B35FFE7FFBDFE5CFDF122C6FD82" box="[421,507,600,627]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="19" pageNumber="20" unit="mm" value="33.0">33 mm</quantity>
SL (
<quantity id="4CDD9B35FFE7FFBDFDC0FDF1219BFD82" box="[569,678,600,627]" metricMagnitude="-2" metricUnit="m" metricValue="3.97" pageId="19" pageNumber="20" unit="mm" value="39.7">39.7 mm</quantity>
TL),
<location id="8EFA600BFFE7FFBDFD16FDF1200FFD83" LSID="urn:lsid:plazi:treatment:038C87C6FFE7FFBAFE57FEA221F9F9D2:8EFA600BFFE7FFBDFD16FDF1200FFD83" box="[751,818,600,626]" country="Gabon" latitude="-0.58186" longLatPrecision="1" longitude="9.46718" name="Weze" pageId="19" pageNumber="20">Wézé</location>
spring,
<geoCoordinate id="EE115017FFE7FFBDFC6EFDF1272EFD83" box="[919,1043,600,627]" degrees="0.58186" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="1" value="-0.58186">0.58186 S</geoCoordinate>
,
<geoCoordinate id="EE115017FFE7FFBDFBDAFDF1279CFD82" box="[1059,1185,600,627]" degrees="9.46718" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="1" value="9.46718">9.46718 E</geoCoordinate>
,
<elevation id="0008D1E3FFE7FFBDFB48FDF1260FFD83" box="[1201,1330,600,627]" metricMagnitude="1" metricUnit="m" metricValue="8.9" pageId="19" pageNumber="20" unit="m" value="89.0">
<quantity id="4CDD9B35FFE7FFBDFB48FDF127CDFD82" box="[1201,1264,600,627]" metricMagnitude="1" metricUnit="m" metricValue="8.9" pageId="19" pageNumber="20" unit="m" value="89.0">89 m</quantity>
a.s.l.
</elevation>
, field reference code
<specimenCode id="DB839EABFFE7FFBDFE89FDD522E0FD66" box="[368,477,636,663]" collectionCode="CHRSP" pageId="19" pageNumber="20">CHRSP5</specimenCode>
-sources
<collectorName id="26D05306FFE7FFBDFDBDFDD521B7FD67" box="[580,650,636,662]" pageId="19" pageNumber="20">Wézé</collectorName>
,
<date id="FF9B1010FFE7FFBDFD6FFDD52026FD67" box="[662,795,636,662]" pageId="19" pageNumber="20" value="2014-07-03">
<collectingDate id="EFDFE9F8FFE7FFBDFD6FFDD52026FD67" box="[662,795,636,662]" pageId="19" pageNumber="20" value="2014-07-03">3 Jul. 2014</collectingDate>
</date>
,
<collectorName id="26D05306FFE7FFBDFCDFFDD420EEFD67" box="[806,979,636,663]" pageId="19" pageNumber="20">Laurent Chirio</collectorName>
leg. (
<specimenCode id="DB839EABFFE7FFBDFBECFDD527D4FD66" box="[1045,1257,636,663]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="19" pageNumber="20" type="Museum">MRAC 2016-019</specimenCode>
-P-64).
</paragraph>
</materialsCitation>
<paragraph id="8B9A36D0FFE7FFBDFF13FD16225AFD28" blockId="19.[189,1399,703,910]" box="[234,359,703,729]" pageId="19" pageNumber="20">
<emphasis id="B951EAC2FFE7FFBDFF13FD16225AFD28" bold="true" box="[234,359,703,729]" pageId="19" pageNumber="20">
<typeStatus id="549E8872FFE7FFBDFF13FD16225AFD28" box="[234,359,703,729]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9A36D0FFE7FFBDFF44FD4A277FFC7F" blockId="19.[189,1399,703,910]" pageId="19" pageNumber="20">
<collectingCountry id="F3327640FFE7FFBDFF44FD4A2217FD0C" box="[189,298,739,765]" name="Gabon" pageId="19" pageNumber="20">GABON</collectingCountry>
: 4 ♁♁,
<specimenCount id="9D23FD59FFE7FFBDFE70FD4B22F0FD0C" box="[393,461,738,765]" pageId="19" pageNumber="20" type="female">5 ♀♀</specimenCount>
,
<quantity id="4CDD9B35FFE7FFBDFE24FD4A21B4FD0C" box="[477,649,739,765]" metricMagnitude="-2" metricUnit="m" metricValue="2.665" metricValueMax="2.9899999999999998" metricValueMin="2.34" pageId="19" pageNumber="20" unit="mm" value="26.65" valueMax="29.9" valueMin="23.4">23.4–29.9 mm</quantity>
SL, same collection data as for
<typeStatus id="549E8872FFE7FFBDFBF3FD4A274CFD0C" box="[1034,1137,739,765]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED34AE15FFE7FFBDFB78FD4A27E4FD0C" box="[1153,1241,739,765]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="19" pageNumber="20" type="Museum">MRAC</collectionCode>
2016-019-P- 65-73); 18 ♁♁,
<quantity id="4CDD9B35FFE7FFBDFE81FCAE2114FCD0" box="[376,553,775,801]" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="3.21" metricValueMin="2.2199999999999998" pageId="19" pageNumber="20" unit="mm" value="27.15" valueMax="32.1" valueMin="22.2">22.2 –32.1 mm</quantity>
SL, northern tributary of Wézé River,
<geoCoordinate id="EE115017FFE7FFBDFC15FCAF2759FCD0" box="[1004,1124,774,801]" degrees="0.56336" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="1" value="-0.56336">0.56336 S</geoCoordinate>
,
<geoCoordinate id="EE115017FFE7FFBDFB88FCAE27D6FCD0" box="[1137,1259,775,801]" degrees="9.33732" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="1" value="9.33732">9.33732 E</geoCoordinate>
,
<quantity id="4CDD9B35FFE7FFBDFB0EFCAF260FFCD0" box="[1271,1330,774,801]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="19" pageNumber="20" unit="m" value="45.0">45 m</quantity>
a.s.l., field reference code
<collectionCode id="ED34AE15FFE7FFBDFE4DFC832128FCB4" box="[436,533,810,837]" pageId="19" pageNumber="20">CHRSP</collectionCode>
5-Wézé2,
<date id="FF9B1010FFE7FFBDFD75FC822019FCB4" box="[652,804,811,837]" pageId="19" pageNumber="20" value="2016-03-01">1 Mar. 2016</date>
, Laurent Chirio leg. (
<collectionCode id="ED34AE15FFE7FFBDFBD0FC8227BCFCB4" box="[1065,1153,811,837]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="19" pageNumber="20" type="Museum">MRAC</collectionCode>
2016-019-P-74-91);
<specimenCount id="9D23FD59FFE7FFBDFF44FCE623D5FC98" box="[189,232,847,873]" pageId="19" pageNumber="20" type="female">1 ♀</specimenCount>
,
<quantity id="4CDD9B35FFE7FFBDFF0CFCE6227BFC98" box="[245,326,847,873]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="19" pageNumber="20" unit="mm" value="23.0">23 mm</quantity>
SL, southern tributary of Wézé River,
<geoCoordinate id="EE115017FFE7FFBDFCFFFCE62040FC98" box="[774,893,847,873]" degrees="0.66401" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="1" value="-0.66401">0.66401 S</geoCoordinate>
,
<geoCoordinate id="EE115017FFE7FFBDFC70FCE7273FFC98" box="[905,1026,846,873]" degrees="9.42652" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="1" value="9.42652">9.42652 E</geoCoordinate>
,
<quantity id="4CDD9B35FFE7FFBDFBF7FCE62775FC98" box="[1038,1096,847,873]" metricMagnitude="1" metricUnit="m" metricValue="8.7" pageId="19" pageNumber="20" unit="m" value="87.0">87 m</quantity>
a.s.l., field reference code
<collectionCode id="ED34AE15FFE7FFBDFF44FCDA2222FC7F" box="[189,287,883,910]" pageId="19" pageNumber="20">CHRSP</collectionCode>
5-Wézé3,
<date id="FF9B1010FFE7FFBDFE6BFCDA2119FC7C" box="[402,548,883,910]" pageId="19" pageNumber="20" value="2016-03-05">5 Mar. 2016</date>
, Laurent Chirio leg. (
<collectionCode id="ED34AE15FFE7FFBDFCE6FCDA204AFC7C" box="[799,887,883,909]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="19" pageNumber="20" type="Museum">MRAC</collectionCode>
2016-019-P-92).
</paragraph>
</subSubSection>
<subSubSection id="C33F655BFFE7FFBAFF44FC1C267AFB0E" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8B9A36D0FFE7FFBDFF44FC1C22F4FC3E" blockId="19.[189,457,949,975]" box="[189,457,949,975]" pageId="19" pageNumber="20">
<heading id="D0D281BCFFE7FFBDFF44FC1C22F4FC3E" bold="true" box="[189,457,949,975]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B951EAC2FFE7FFBDFF44FC1C22F4FC3E" bold="true" box="[189,457,949,975]" pageId="19" pageNumber="20">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A36D0FFE7FFBDFF44FC4A21D0FB7F" blockId="19.[189,1398,995,1166]" pageId="19" pageNumber="20">
Differs from all other species in
<taxonomicName id="4C254D53FFE7FFBDFDA2FC4D21F9FC0C" authorityName="Huber" authorityYear="1998" box="[603,708,995,1022]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="alpha">
<emphasis id="B951EAC2FFE7FFBDFDA2FC4D21F9FC0C" box="[603,708,995,1022]" italics="true" pageId="19" pageNumber="20">A. alpha</emphasis>
</taxonomicName>
group through combination of following features: head orange, flanks beige, ventral region orange (vs never the same colour pattern), orange anal fin with no punctuation (vs blue-orange, purplish-blue or orange with punctuation). Distinguished from all other species of
<taxonomicName id="4C254D53FFE7FFBDFEC6FBE62117FB98" authorityName="Radda" authorityYear="1971" box="[319,554,1103,1129]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="B951EAC2FFE7FFBDFEC6FBE62117FB98" box="[319,554,1103,1129]" italics="true" pageId="19" pageNumber="20">Chromaphyosemion</emphasis>
</taxonomicName>
by an alpha-shaped drawing in pre- and post-opercular region (vs no alpha-shaped drawing, only some red macules).
</paragraph>
<paragraph id="8B9A36D0FFE7FFBDFF44FB1C2156FB21" blockId="19.[189,619,1205,1232]" box="[189,619,1205,1232]" pageId="19" pageNumber="20">
<emphasis id="B951EAC2FFE7FFBDFF44FB1C22CAFB21" bold="true" box="[189,503,1205,1232]" pageId="19" pageNumber="20">Colouration of live males</emphasis>
(
<figureCitation id="131E2A55FFE7FFBDFDFEFB1C215EFB21" box="[519,611,1205,1232]" captionStart="Fig" captionStartId="21.[189,232,1788,1814]" captionTargetBox="[228,1364,265,1702]" captionTargetId="figure@21.[228,1364,265,1743]" captionTargetPageId="21" captionText="Fig. 6. A–D. Aphyosemion aurantiacum Chirio, Legros &amp; Agnèse sp. nov. A. Adult, ♁, from locality 12, not preserved. Photo O. Buisson. B. Adult, ♀, from locality 12, not preserved. Photo O. Buisson. C. Holotype, adult, ♁, from Wézé Spring (MRAC 2016-019-P-64). D. Paratype, adult, ♀, from Wézé Spring (MRAC 2016-019-P-65-73). E–H. A. rubrogaster Chirio, Legros &amp; Agnèse sp. nov. E. Adult, ♁, from locality 16, not preserved. Photo O. Buisson. F. Adult, ♀, from locality 16, not preserved. Photo O. Buisson. G. Holotype, adult, ♁, from the Niengé River (MRAC 2016-019-P-93). H. Paratype, adult, ♀, from the Niengé River (MRAC 2016-019-P-94-108)." figureDoi="http://doi.org/10.5281/zenodo.3825065" httpUri="https://zenodo.org/record/3825065/files/figure.png" pageId="19" pageNumber="20">Fig. 6A</figureCitation>
)
</paragraph>
<paragraph id="8B9A36D0FFE7FFBDFF44FB4D2168FA7C" blockId="19.[189,1399,1251,1422]" pageId="19" pageNumber="20">FLANKS AND VENTER. Flanks beige with two grey, rarely visible lateral stripes, four lines of red dots from opercle to caudal peduncle. Ventral region orange and region between end of anal fin and caudal peduncle intense orange, two rows of blue iridescent scales between end of anal fin and caudal peduncle. Dorsal region brown, with two rows of paradorsal copper scales from operculum to caudal peduncle with two lines of carmine red dots.</paragraph>
<paragraph id="8B9A36D0FFE7FFBDFF44FA1F2001F9CD" blockId="19.[189,1399,1462,1596]" pageId="19" pageNumber="20">HEAD. Orange with premaxilla brown, mandible orange with black lower lips, brown supraorbital region, orange infraorbital region with black macule, postorbital region orange with two black macules, orange pre-opercle with one black macule, orange opercle with two black macules; black macules on pre- and post-opercular region reveal an alpha-shaped drawing.</paragraph>
<paragraph id="8B9A36D0FFE7FFBDFF44F9CC201FF8A7" blockId="19.[189,1399,1636,1879]" pageId="19" pageNumber="20">FINS. Dorsal fin blue, large orange zone on rays, rows of carmine red dots between rays, and carmine red streaks on distal and posterobasal portions of fin. Orange acumen on apex. Anal fin orange, blue zone near submarginal band, regular, red submarginal band, blue marginal band, orange acumen. No punctuation. Caudal fin blue with orange caudal peduncle, 5–6 red dots in median region, very long red streaks in median region and in upper and lower lobes, regular, red submarginal band, blue marginal band, and orange acumens. Pelvic fins orange with no punctuation, submarginal red band and blue marginal band. Pectoral fins with orange reflections.</paragraph>
<paragraph id="8B9A36D0FFE7FFBDFF44F8D721BDF868" blockId="19.[189,640,1918,1945]" box="[189,640,1918,1945]" pageId="19" pageNumber="20">
<emphasis id="B951EAC2FFE7FFBDFF44F8D72130F869" bold="true" box="[189,525,1918,1944]" pageId="19" pageNumber="20">Colouration of live females</emphasis>
(
<figureCitation id="131E2A55FFE7FFBDFDE4F8D7214AF868" box="[541,631,1918,1945]" captionStart="Fig" captionStartId="21.[189,232,1788,1814]" captionTargetBox="[228,1364,265,1702]" captionTargetId="figure@21.[228,1364,265,1743]" captionTargetPageId="21" captionText="Fig. 6. A–D. Aphyosemion aurantiacum Chirio, Legros &amp; Agnèse sp. nov. A. Adult, ♁, from locality 12, not preserved. Photo O. Buisson. B. Adult, ♀, from locality 12, not preserved. Photo O. Buisson. C. Holotype, adult, ♁, from Wézé Spring (MRAC 2016-019-P-64). D. Paratype, adult, ♀, from Wézé Spring (MRAC 2016-019-P-65-73). E–H. A. rubrogaster Chirio, Legros &amp; Agnèse sp. nov. E. Adult, ♁, from locality 16, not preserved. Photo O. Buisson. F. Adult, ♀, from locality 16, not preserved. Photo O. Buisson. G. Holotype, adult, ♁, from the Niengé River (MRAC 2016-019-P-93). H. Paratype, adult, ♀, from the Niengé River (MRAC 2016-019-P-94-108)." figureDoi="http://doi.org/10.5281/zenodo.3825065" httpUri="https://zenodo.org/record/3825065/files/figure.png" pageId="19" pageNumber="20">Fig. 6B</figureCitation>
)
</paragraph>
<paragraph id="8B9A36D0FFE7FFBDFF44F804264BF81A" blockId="19.[189,1398,1965,2027]" pageId="19" pageNumber="20">FLANKS AND VENTER. Flanks pale yellow, with four lines of red dots from opercle to caudal peduncle, two greyish lateral stripes, yellow venter, white on basal portion. Two rows of paradorsal golden scales</paragraph>
<paragraph id="8B9A36D0FFE0FFBAFF44FEA2211EFEB8" blockId="20.[189,1399,267,329]" pageId="20" pageNumber="21">in dorsal region from operculum to start of anal fin with red dots, copper scales from start of anal fin to caudal peduncle with red dots.</paragraph>
<paragraph id="8B9A36D0FFE0FFBAFF44FEDB21DFFE22" blockId="20.[189,1399,369,467]" pageId="20" pageNumber="21">HEAD. Premaxilla brown, mandible pale yellow with black lower lips, brown supraorbital region, white infraorbital region with black macule, white postorbital region with two red macules, yellow pre-opercle with red dot, yellow opercle with two red dots.</paragraph>
<paragraph id="8B9A36D0FFE0FFBAFF44FE55205DFD55" blockId="20.[189,1399,507,677]" pageId="20" pageNumber="21">FINS. Dorsal fin yellow-green, with rows of carmine red dots between rays, red streaks on distal portion. Anal fin blue on basal portion with row of red dots between rays, some red streaks between rays on median portion, pale orange on distal portion. Caudal fin yellow-green near caudal peduncle, with some red dots on basal portion, red streaks between rays from median to distal portion. Pelvic fins translucent with blue reflection on the edge. Pectoral fins translucent.</paragraph>
<paragraph id="8B9A36D0FFE0FFBAFF44FD6520DDFD16" blockId="20.[189,992,716,743]" box="[189,992,716,743]" pageId="20" pageNumber="21">
<emphasis id="B951EAC2FFE0FFBAFF44FD652074FD17" bold="true" box="[189,841,716,743]" pageId="20" pageNumber="21">Colouration of ethanol-preserved males and females</emphasis>
(
<figureCitation id="131E2A55FFE0FFBAFCA0FD6520A3FD16" box="[857,926,716,743]" captionStart="Fig" captionStartId="21.[189,232,1788,1814]" captionTargetBox="[228,1364,265,1702]" captionTargetId="figure@21.[228,1364,265,1743]" captionTargetPageId="21" captionText="Fig. 6. A–D. Aphyosemion aurantiacum Chirio, Legros &amp; Agnèse sp. nov. A. Adult, ♁, from locality 12, not preserved. Photo O. Buisson. B. Adult, ♀, from locality 12, not preserved. Photo O. Buisson. C. Holotype, adult, ♁, from Wézé Spring (MRAC 2016-019-P-64). D. Paratype, adult, ♀, from Wézé Spring (MRAC 2016-019-P-65-73). E–H. A. rubrogaster Chirio, Legros &amp; Agnèse sp. nov. E. Adult, ♁, from locality 16, not preserved. Photo O. Buisson. F. Adult, ♀, from locality 16, not preserved. Photo O. Buisson. G. Holotype, adult, ♁, from the Niengé River (MRAC 2016-019-P-93). H. Paratype, adult, ♀, from the Niengé River (MRAC 2016-019-P-94-108)." figureDoi="http://doi.org/10.5281/zenodo.3825065" httpUri="https://zenodo.org/record/3825065/files/figure.png" pageId="20" pageNumber="21">Fig. 6</figureCitation>
C–D)
</paragraph>
<paragraph id="8B9A36D0FFE0FFBAFF44FD52207EFC55" blockId="20.[189,1399,762,932]" pageId="20" pageNumber="21">FLANKS AND VENTER. Flanks of males grey-yellow with no red dots, venter grey-yellow. Two rows of paradorsal black scales in grey-yellow dorsal region from operculum to caudal peduncle, visible greyish upper lateral stripe. Flanks of females beige with two grey lateral stripes and two lines of two to three red dots behind opercle. Venter beige, with two grey lateral stripes. Two rows of paradorsal red scales in grey dorsal region from operculum to caudal peduncle.</paragraph>
<paragraph id="8B9A36D0FFE0FFBAFF44FC65226AFBA0" blockId="20.[189,1398,972,1105]" pageId="20" pageNumber="21">HEAD. Head of male grey-yellow mandible with black lower lips, grey-yellow infraorbital region with small black macule, grey-yellow pre-opercular region, opercle with two black macules; dark macules in post-opercular region. Head of female with black lower lips, infraorbital region with black macule, dark opercle.</paragraph>
<paragraph id="8B9A36D0FFE0FFBAFF44FBD3267AFB0E" blockId="20.[189,1398,1145,1280]" pageId="20" pageNumber="21">FINS. Dorsal fin of male orange with two to three black dots on basal portion, streaks between rays on distal portion. Anal fin orange with dark rays. Caudal fin orange with many red streaks on distal portion, red submarginal band. Dorsal fin of female light orange, rows of red dots between rays. Anal fin greyish with no red dots. Caudal fin light orange, with some red dots on upper portion, red streaks on edge.</paragraph>
</subSubSection>
<subSubSection id="C33F655BFFE0FFBAFF44FA8E21F9F9D2" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8B9A36D0FFE0FFBAFF44FA8E22D3FAB0" blockId="20.[189,494,1319,1345]" box="[189,494,1319,1345]" pageId="20" pageNumber="21">
<heading id="D0D281BCFFE0FFBAFF44FA8E22D3FAB0" bold="true" box="[189,494,1319,1345]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B951EAC2FFE0FFBAFF44FA8E22D3FAB0" bold="true" box="[189,494,1319,1345]" pageId="20" pageNumber="21">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A36D0FFE0FFBAFF44FAFC21F9F9D2" blockId="20.[189,1399,1365,1571]" pageId="20" pageNumber="21">
This species has been found at three localities in the Wézé Basin (
<tableCitation id="C6A7036BFFE0FFBAFC3BFAFC2721FA9E" box="[962,1052,1365,1391]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetPageId="3" captionText="Table 1. Species identification; population number (N°) on the map (Fig. 1) and on the phylogenetic network (Fig. 2); voucher specimen references registered at the Royal Museum for Central Africa (MRAC), Tervuren (Belgium); coordinates." httpUri="http://table.plazi.org/id/DF5A6658FFF7FFADFF44FEA52180FE9D" pageId="20" pageNumber="21" tableUuid="DF5A6658FFF7FFADFF44FEA52180FE9D">Table 1</tableCitation>
,
<figureCitation id="131E2A55FFE0FFBAFBD3FAFC2750FA9E" box="[1066,1133,1365,1391]" captionStart="Fig" captionStartId="2.[189,232,1858,1884]" captionTargetBox="[189,1398,321,1815]" captionTargetId="figure@2.[189,1398,321,1815]" captionTargetPageId="2" captionText="Fig. 1. Collecting sites of the different populations. Black circles: Aphyosemion alpha Huber, 1998; Black triangles: A. flavocyaneum Chirio, Legros &amp; Agnèse sp. nov.; red triangle: A. flammulatum Chirio, Legros &amp; Agnèse sp. nov.; yellow triangles: A. barakoniense Chirio, Legros &amp; Agnèse sp. nov.; red circles: A. pusillum Chirio, Legros &amp; Agnèse sp. nov.; yellow circles: A. rubrogaster Chirio, Legros &amp; Agnèse sp. nov.; red squares: A. aurantiacum Chirio, Legros &amp; Agnèse sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3825051" httpUri="https://zenodo.org/record/3825051/files/figure.png" pageId="20" pageNumber="21">Fig. 1</figureCitation>
) and never elsewhere. The species seems to be endemic to this hydrographic basin. It lives only in very small forest streams and water holes, often with a bottom consisting of roots and dead leaves, where it can be very abundant. In the southern-most locality, it was found hidden under dead leaves along the river banks in a 2 metre wide river with a sandy bottom. This species was found in syntopy with three undescribed species of
<taxonomicName id="4C254D53FFE0FFBAFF44F9A12266F9D3" authorityName="Myers" authorityYear="1924" box="[189,347,1544,1570]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B951EAC2FFE0FFBAFF44F9A12266F9D3" box="[189,347,1544,1570]" italics="true" pageId="20" pageNumber="21">Aphyosemion</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D53FFE0FFBAFE93F9A122E4F9D3" box="[362,473,1544,1570]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B951EAC2FFE0FFBAFE93F9A122E4F9D3" box="[362,473,1544,1570]" italics="true" pageId="20" pageNumber="21">Epiplatys</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D53FFE0FFBAFDEBF9A12180F9D3" authorityName="Ahl" authorityYear="1928" box="[530,701,1544,1570]" class="Actinopterygii" family="Poeciliidae" genus="Plataplochilus" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B951EAC2FFE0FFBAFDEBF9A12180F9D3" box="[530,701,1544,1570]" italics="true" pageId="20" pageNumber="21">Plataplochilus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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