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<document id="CF51C8CBF66C291FBC24A5DB89EF0056" ID-CLB-Dataset="9527" ID-DOI="10.11646/zootaxa.4881.2.5" ID-GBIF-Dataset="ddbf20bd-a496-402e-9048-804711808b44" ID-ISSN="1175-5326" ID-Zenodo-Dep="4283704" ID-ZooBank="7E570EDA-1A01-45AB-8ED2-79F8B8A6B9FE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1605976222792" checkinUser="plazi" docAuthor="Komai, Tomoyuki" docDate="2020" docId="03898797FFEFFF9CFF14FF7FFB72FE04" docLanguage="en" docName="zootaxa.4881.2.5.pdf" docOrigin="Zootaxa 4881 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gebiacantha fortispinata Komai 2020, n. sp." docType="treatment" docUuid="005FFE26-CBEC-4A8C-A4CB-584B39D3480E" docUuidSource="ZooBank" docVersion="8" lastPageNumber="320" masterDocId="FFB0FFEFFFE7FF91FF83FFE8FFD3FFD5" masterDocTitle="Redescription of a poorly known upogebiid mud shrimp Gebiacantha acanthochela (Sakai, 1967) and description of a new species of the genus from the Ryukyu Islands, Japan (Decapoda: Gebiidea)" masterLastPageNumber="322" masterPageNumber="307" pageNumber="315" updateTime="1698846437294" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="1867B3629DCDB75E35C0704B212A2E77" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A4BF86C09BB9908ABEAE3227BFE2FB95">
<mods:title id="05649268A4DC17F9371010CB9CA83E05">Redescription of a poorly known upogebiid mud shrimp Gebiacantha acanthochela (Sakai, 1967) and description of a new species of the genus from the Ryukyu Islands, Japan (Decapoda: Gebiidea)</mods:title>
</mods:titleInfo>
<mods:name id="454533FEB27BBE7E1EB154626686ED58" type="personal">
<mods:role id="20EB1665DD773E2BFEF8106571F46BA0">
<mods:roleTerm id="50E29AB8F59B9F22EC182D315B55782D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9AA27426770CCB5A03D429D0C47CDCB5">Komai, Tomoyuki</mods:namePart>
</mods:name>
<mods:typeOfResource id="4B00DDE720F49F567391EF6547FB5212">text</mods:typeOfResource>
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<mods:title id="5C755D8F157F6DB373F33BC89A094656">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FB91C9971741F51826FAD507C80691CD">
<mods:date id="1A3CDFB4F89520810C6ED9EE0493440F">2020</mods:date>
<mods:detail id="449E39711E47EDCD6A7F865DB8D1BC95" type="pubDate">
<mods:number id="6418EFEFECA7805AD348D18D726C714D">2020-11-19</mods:number>
</mods:detail>
<mods:detail id="2960A6287DCC4226D6EFBE054D22B943" type="volume">
<mods:number id="F1C207EB3535DBC08E4D6CE926D1A4E8">4881</mods:number>
</mods:detail>
<mods:detail id="3B150B4598777BB8803A1C64A55013B0" type="issue">
<mods:number id="E11D78666DC1A63FBEEE9373028ABAE8">2</mods:number>
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<mods:classification id="DF753E23ACEC50F501EDF7AE84C4138F">journal article</mods:classification>
<mods:identifier id="A9B83883BF48A34A1BE884D7B9259F03" type="CLB-Dataset">9527</mods:identifier>
<mods:identifier id="B7088C486C30BEC5C7D6D570176FF9EC" type="DOI">10.11646/zootaxa.4881.2.5</mods:identifier>
<mods:identifier id="B60FCD8624BC5A8F333395045B3FC446" type="GBIF-Dataset">ddbf20bd-a496-402e-9048-804711808b44</mods:identifier>
<mods:identifier id="DA7B19AF14425ECC0EE0CBAD43BB2BFA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A9593D3738291CB6B919A4D8539A85DB" type="Zenodo-Dep">4283704</mods:identifier>
<mods:identifier id="A981CF8A12CA74EC4064FC773CFA2910" type="ZooBank">7E570EDA-1A01-45AB-8ED2-79F8B8A6B9FE</mods:identifier>
</mods:mods>
<treatment id="03898797FFEFFF9CFF14FF7FFB72FE04" ID-DOI="http://doi.org/10.5281/zenodo.4328019" ID-GBIF-Taxon="170510577" ID-Zenodo-Dep="4328019" LSID="urn:lsid:zoobank.org:act:005FFE26-CBEC-4A8C-A4CB-584B39D3480E" httpUri="http://treatment.plazi.org/id/03898797FFEFFF9CFF14FF7FFB72FE04" lastPageId="13" lastPageNumber="320" pageId="8" pageNumber="315">
<subSubSection id="C33A650AFFEFFF99FF14FF7FFDA8FF00" pageId="8" pageNumber="315" type="nomenclature">
<paragraph id="8B9F3681FFEFFF99FF14FF7FFDDCFF64" blockId="8.[151,635,151,250]" box="[151,527,151,178]" pageId="8" pageNumber="315">
<heading id="D0D781EDFFEFFF99FF14FF7FFDDCFF64" bold="true" box="[151,527,151,178]" fontSize="11" level="1" pageId="8" pageNumber="315" reason="1">
<emphasis id="B954EA93FFEFFF99FF14FF7FFDDCFF64" bold="true" box="[151,527,151,178]" pageId="8" pageNumber="315">
<taxonomicName id="4C204D02FFEFFF99FF14FF7FFE16FF64" ID-CoL="e0444d81-043b-4649-b711-e1ad49f3a7eb" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[151,453,151,178]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="315" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEFFF99FF14FF7FFE16FF64" bold="true" box="[151,453,151,178]" italics="true" pageId="8" pageNumber="315">Gebiacantha fortispinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26757E8FFEFFF99FE4EFF7FFDDCFF64" box="[461,527,151,177]" pageId="8" pageNumber="315" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9F3681FFEFFF99FF14FF54FDA8FF00" blockId="8.[151,635,151,250]" box="[151,635,186,214]" pageId="8" pageNumber="315">[New Japanese name: Toge-toge-anajyako]</paragraph>
</subSubSection>
<subSubSection id="C33A650AFFEFFF99FF14FF37FED8FF2F" box="[151,267,223,250]" pageId="8" pageNumber="315" type="description">
<paragraph id="8B9F3681FFEFFF99FF14FF37FED8FF2F" blockId="8.[151,635,151,250]" box="[151,267,223,250]" pageId="8" pageNumber="315">
(
<figureCitation id="131B2A04FFEFFF99FF1CFF37FED0FF2F" box="[159,259,223,250]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStartId-0="8.[151,250,1723,1747]" captionStartId-1="10.[151,250,1908,1932]" captionStartId-2="11.[151,250,1906,1930]" captionStartId-3="13.[151,250,1668,1692]" captionTargetBox-0="[161,1426,391,1690]" captionTargetBox-1="[166,1422,350,1875]" captionTargetBox-2="[167,1422,606,1872]" captionTargetBox-3="[201,1385,537,1629]" captionTargetId-0="figure-64@8.[151,1436,381,1700]" captionTargetId-1="figure-115@10.[158,1430,345,1885]" captionTargetId-2="figure-205@11.[157,1431,597,1882]" captionTargetId-3="figure-144@13.[181,1406,489,1645]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="13" captionText-0="FIGURE 6. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, carapace and cephalic appendages, lateral view; B, thoracomere 8 (including coxa of pereopod 5), pleon, telson and left pleopod 1, lateral view (setae omitted)." captionText-1="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." captionText-2="FIGURE 8. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, maxilliped 3, lateral view (setae omitted); B, left coxae of pereopods 1–3 (P1–P3), ventromesial view; C, ischium to carpus of left pereopod 3, lateral view (propodus and dactylus missing; setae omitted); D, left pereopod 4, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 5, lateral view; G, same, propodus and dactylus, upper view." captionText-3="FIGURE 9. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176, right pereopod 1. A, ischium and merus, lateral view (proximal part of merus damaged; setae omitted); B, carpus to dactylus, lateral view; C, same, mesial view (setae omitted); D, dactylus, lateral view (setae omitted)." figureDoi-0="http://doi.org/10.5281/zenodo.4283716" figureDoi-1="http://doi.org/10.5281/zenodo.4283718" figureDoi-2="http://doi.org/10.5281/zenodo.4283720" figureDoi-3="http://doi.org/10.5281/zenodo.4283722" httpUri-0="https://zenodo.org/record/4283716/files/figure.png" httpUri-1="https://zenodo.org/record/4283718/files/figure.png" httpUri-2="https://zenodo.org/record/4283720/files/figure.png" httpUri-3="https://zenodo.org/record/4283722/files/figure.png" pageId="8" pageNumber="315">Figs 6–9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33A650AFFEFFF99FF14FECFFB59FEB0" pageId="8" pageNumber="315" type="materials_examined">
<paragraph id="8B9F3681FFEFFF99FF14FECFFB59FEB0" blockId="8.[151,1436,295,358]" pageId="8" pageNumber="315">
<emphasis id="B954EA93FFEFFF99FF14FECFFEA8FE94" bold="true" box="[151,379,295,321]" pageId="8" pageNumber="315">Material examined</emphasis>
.
<materialsCitation id="3B483CDCFFEFFF99FE09FECFFB59FEB0" ID-GBIF-Occurrence="2985241302" collectingDate="2010-01-10" collectorName="Naoki Shirakawa." county="Apo-gama" elevation="30" location="Onna Village" municipality="Manza" pageId="8" pageNumber="315" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<typeStatus id="549B8823FFEFFF99FE09FECFFE24FE94" box="[394,503,295,321]" pageId="8" pageNumber="315" type="holotype">Holotype</typeStatus>
: CBM-ZC 16176, female (cl
<quantity id="4CD89B64FFEFFF99FCC6FECFFC73FE97" box="[837,928,295,322]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="8" pageNumber="315" unit="mm" value="7.4">7.4 mm</quantity>
),
<collectingCounty id="62FE4E0DFFEFFF99FC31FECFFBF8FE94" box="[946,1067,295,321]" pageId="8" pageNumber="315">Apo-gama</collectingCounty>
,
<collectingMunicipality id="6BFBACFBFFEFFF99FBBBFEC0FB54FE97" box="[1080,1159,296,322]" pageId="8" pageNumber="315">Manza</collectingMunicipality>
,
<location id="8EFF605AFFEFFF99FB10FECFFAF8FE94" LSID="urn:lsid:plazi:treatment:03898797FFEFFF9CFF14FF7FFB72FE04:8EFF605AFFEFFF99FB10FECFFAF8FE94" box="[1171,1323,295,321]" county="Apo-gama" municipality="Manza" name="Onna Village" pageId="8" pageNumber="315" stateProvince="Okinawa">Onna Village</location>
,
<collectingRegion id="49E4F863FFEFFF99FAB4FECFFA4FFE94" box="[1335,1436,295,321]" country="Japan" name="Okinawa" pageId="8" pageNumber="315">Okinawa</collectingRegion>
<location id="8EFF605AFFEFFF99FF14FEA3FF0DFEB0" LSID="urn:lsid:plazi:treatment:03898797FFEFFF9CFF14FF7FFB72FE04:8EFF605AFFEFFF99FF14FEA3FF0DFEB0" box="[151,222,331,357]" county="Apo-gama" municipality="Manza" name="Island" pageId="8" pageNumber="315" stateProvince="Okinawa">Island</location>
,
<location id="8EFF605AFFEFFF99FF69FEA3FE49FEB0" LSID="urn:lsid:plazi:treatment:03898797FFEFFF9CFF14FF7FFB72FE04:8EFF605AFFEFFF99FF69FEA3FE49FEB0" box="[234,410,331,357]" county="Apo-gama" municipality="Manza" name="Ryukyu Islands" pageId="8" pageNumber="315" stateProvince="Okinawa">Ryukyu Islands</location>
,
<quantity id="4CD89B64FFEFFF99FE26FEA3FE0DFEB3" box="[421,478,331,358]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="315" unit="m" value="30.0">
<elevation id="000DD1B2FFEFFF99FE26FEA3FE0DFEB3" box="[421,478,331,358]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="315" unit="m" value="30.0">30 m</elevation>
</quantity>
deep,
<date id="FF9E1041FFEFFF99FDABFEA3FD37FEB0" box="[552,740,331,358]" pageId="8" pageNumber="315" value="2010-01-10">
<collectingDate id="EFDAE9A9FFEFFF99FDABFEA3FD37FEB0" box="[552,740,331,358]" pageId="8" pageNumber="315" value="2010-01-10">10 January 2010</collectingDate>
</date>
, yabby pump, coll.
<collectorName id="26D55357FFEFFF99FC43FEA3FB59FEB0" box="[960,1162,331,357]" pageId="8" pageNumber="315">Naoki Shirakawa.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF5F6609FFEFFF99FF14F953FF27F8CE" ID-DOI="http://doi.org/10.5281/zenodo.4283716" ID-Zenodo-Dep="4283716" httpUri="https://zenodo.org/record/4283716/files/figure.png" pageId="8" pageNumber="315" startId="8.[151,250,1723,1747]" targetBox="[161,1426,391,1690]" targetPageId="8">
<paragraph id="8B9F3681FFEFFF99FF14F953FF27F8CE" blockId="8.[151,1436,1723,1819]" pageId="8" pageNumber="315">
<emphasis id="B954EA93FFEFFF99FF14F953FEC6F906" bold="true" box="[151,277,1723,1747]" pageId="8" pageNumber="315">FIGURE 6.</emphasis>
<taxonomicName id="4C204D02FFEFFF99FE9CF953FDF2F901" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[287,545,1723,1748]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="315" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEFFF99FE9CF953FDF2F901" box="[287,545,1723,1748]" italics="true" pageId="8" pageNumber="315">Gebiacantha fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEFFF99FDA9F954FDB5F901" bold="true" box="[554,614,1724,1748]" pageId="8" pageNumber="315">
<taxonomicNameLabel id="A26757E8FFEFFF99FDA9F954FDB5F901" box="[554,614,1724,1748]" pageId="8" pageNumber="315" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, female (cl 7.4 mm), CBM-ZC 16176. A, carapace and cephalic appendages, lateral view; B, thoracomere 8 (including coxa of pereopod 5), pleon, telson and left pleopod 1, lateral view (setae omitted).
</paragraph>
</caption>
<subSubSection id="C33A650AFFEFFF9DFF44F8A1FDE6FDCC" lastPageId="12" lastPageNumber="319" pageId="8" pageNumber="315" type="description">
<paragraph id="8B9F3681FFEFFF98FF44F8A1FAE4FE20" blockId="8.[151,1437,1865,2036]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="316" pageId="8" pageNumber="315">
<emphasis id="B954EA93FFEFFF99FF44F8A1FE8AF8B6" bold="true" box="[199,345,1865,1891]" pageId="8" pageNumber="315">Description.</emphasis>
Rostrum (
<figureCitation id="131B2A04FFEFFF99FE57F8A2FDE6F8B1" box="[468,565,1866,1892]" captionStart="FIGURE 6" captionStartId="8.[151,250,1723,1747]" captionTargetBox="[161,1426,391,1690]" captionTargetId="figure-64@8.[151,1436,381,1700]" captionTargetPageId="8" captionText="FIGURE 6. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, carapace and cephalic appendages, lateral view; B, thoracomere 8 (including coxa of pereopod 5), pleon, telson and left pleopod 1, lateral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283716" httpUri="https://zenodo.org/record/4283716/files/figure.png" pageId="8" pageNumber="315">Figs 6A</figureCitation>
,
<figureCitation id="131B2A04FFEFFF99FDC1F8A2FD5BF8B1" box="[578,648,1866,1892]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="8" pageNumber="315">7A, B</figureCitation>
) long, reaching beyond midlength of article 4 of antennal peduncle, subtriangular in dorsal view, 1.3 times as long as basal width, directed forward in lateral view, somewhat flattened dorsoventrally; dorsolateral margins with 7 (left) and 8 (right) narrowly spaced, relatively large spiniform tubercles, each tubercle directed anterodorsally, anteriormost pair terminal in position; dorsal surface setose, with shallow median sulcus proximally; ventral surface nearly straight, armed with 4 strong, moderately spaced spines in distal half, distalmost spine most slender, directed anteriorly; rostral apex roundly truncate in lateral view, overreached by dorsal terminal spines and distalmost ventral spine. Carapace (
<figureCitation id="131B2A04FFEEFF98FCF1FF53FC19FF03" box="[882,970,187,214]" captionStart="FIGURE 6" captionStartId="8.[151,250,1723,1747]" captionTargetBox="[161,1426,391,1690]" captionTargetId="figure-64@8.[151,1436,381,1700]" captionTargetPageId="8" captionText="FIGURE 6. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, carapace and cephalic appendages, lateral view; B, thoracomere 8 (including coxa of pereopod 5), pleon, telson and left pleopod 1, lateral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283716" httpUri="https://zenodo.org/record/4283716/files/figure.png" pageId="9" pageNumber="316">Fig. 6A</figureCitation>
) somewhat compressed laterally; anterior part (
<figureCitation id="131B2A04FFEEFF98FF51FF37FE9AFF2C" box="[210,329,223,249]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="9" pageNumber="316">Fig. 7A, B</figureCitation>
) with lateral gastric ridges slightly diverging posteriorly, each terminating anteriorly in small spine and bearing row of 11 spiniform tubercles decreasing in size toward posterior; gastric region flanked by shallow longitudinal grooves running along gastric lateral ridges and having 2 irregular, longitudinal rows of small spiniform tubercles, extending anteriorly to rostral base on either side of midline; median part almost flat, unarmed; anterolateral margin with 7 spinules, including postocular spine; postorbital region with 2 spinules; pterygostomial margin rounded, unarmed; linea thalassinica extending from anterolateral notch to posterodorsal end of carapace, interrupted at midway between cervical groove and carapace posterior margin; posterior part with shoulder along cervical groove bearing row of spinules extending to anterolateral notch; branchiostegite poorly calcified.
</paragraph>
<paragraph id="8B9F3681FFEEFF98FF44FE17FB09FCEC" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="316">
Pleon (
<figureCitation id="131B2A04FFEEFF98FE94FE17FEBDFDCF" box="[279,366,511,538]" captionStart="FIGURE 6" captionStartId="8.[151,250,1723,1747]" captionTargetBox="[161,1426,391,1690]" captionTargetId="figure-64@8.[151,1436,381,1700]" captionTargetPageId="8" captionText="FIGURE 6. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, carapace and cephalic appendages, lateral view; B, thoracomere 8 (including coxa of pereopod 5), pleon, telson and left pleopod 1, lateral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283716" httpUri="https://zenodo.org/record/4283716/files/figure.png" pageId="9" pageNumber="316">Fig. 6B</figureCitation>
) fairly flattened dorsoventrally. Pleuron 1 narrow, ventrally rounded, demarcated from tergum by distinct longitudinal groove; pleuron 2 also narrow, anterior part demarcated from tergum by distinct longitudinal groove reaching to midlength of somite, ventral margin slightly upturned. Pleomeres 3–5 each with short curved groove on anterior part; pleural margins with row of seta directed laterally, each posterolateral margin rounded. Pleomere 6 (
<figureCitation id="131B2A04FFEEFF98FEA0FD67FEA1FD7F" box="[291,370,655,682]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="9" pageNumber="316">Fig.7C</figureCitation>
) longest, 1.1 times as long as greatest width at 0.6 length of somite; lateral margins slightly sinuous, with low convexity at 0.6 length; posterior margin nearly straight, with small notch on each lateral part. Telson (
<figureCitation id="131B2A04FFEEFF98FF23FD3FFF25FD24" box="[160,246,727,753]" captionStart="FIGURE 1" captionStartId="2.[151,250,1580,1604]" captionTargetBox="[156,1431,533,1551]" captionTargetId="figure-185@2.[151,1436,525,1557]" captionTargetPageId="2" captionText="FIGURE 1. Gebiacantha acanthochela (Sakai, 1967), ovigerous female (cl 8.2 mm), CBM-ZC 16175.A, carapace and cephalic appendages, lateral view (setae omitted); B, thoracomere 8 (including coxa of pereopod 5), pleon, telson and left pleopod 1, lateral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283706" httpUri="https://zenodo.org/record/4283706/files/figure.png" pageId="9" pageNumber="316">Fig. 1C</figureCitation>
) subrectangular, 1.2 times as long as greatest width at anterior 0.2; dorsal surface with clearly delimited longitudinal ridges laterally, proximal part slightly elevated, though not forming distinct transverse carina; lateral margins unarmed; posterior margin bilobed with deep median notch, fringed with fine long setae.
</paragraph>
<paragraph id="8B9F3681FFEEFF98FF44FCABFAD9FC54" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="316">
Ocular peduncle (
<figureCitation id="131B2A04FFEEFF98FE1BFCABFE26FC88" box="[408,501,835,861]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="9" pageNumber="316">Fig. 7A</figureCitation>
) moderately stout, subcylindrical, unarmed, not reaching midlength of rostrum, slightly widened basally; cornea terminal, pigmented, corneal width subequal to diameter of eyestalk.
</paragraph>
<paragraph id="8B9F3681FFEEFF98FF44FC63FEC0FBE0" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="316">
Antennular peduncle (
<figureCitation id="131B2A04FFEEFF98FE42FC63FDE8FC70" box="[449,571,907,933]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="9" pageNumber="316">Fig. 7A, D</figureCitation>
) not reaching distal margin of antennal peduncle article 4. Article 1 subequal in length to distal 2 articles combined, armed with prominent ventrodistal spine directed anteriorly; statocyst lobe slightly inflated. Article 2 shortest, cup-like. Article 3 slightly widened distally, unarmed. Upper flagellum shorter than peduncle consisting of about 16 articles; lower flagellum much slenderer than upper flagellum, consisting of 12 articles.
</paragraph>
<paragraph id="8B9F3681FFEEFF98FF44FBD7FE4FFB3C" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="316">
Antennal peduncle (
<figureCitation id="131B2A04FFEEFF98FE2EFBD7FDD4FB8C" box="[429,519,1087,1113]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="9" pageNumber="316">Fig. 7A</figureCitation>
) article 1 with conspicuous, ventrally directed spine on ventrodistal margin. Articles 2 and 3 fused, unarmed on dorsolateral margin proximally; ventral margin armed with 4 evenly spaced spines. Article 4 with 3 subequal spines on ventral margin.Article 5 less than half-length of article 4, unarmed. Scaphocerite clearly demarcated basally, with 1 small spine dorsodistally. Flagellum longer than carapace, each article with thin setae of various length.
</paragraph>
<paragraph id="8B9F3681FFEEFF98FF44FB1BFE27FA30" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="316">
Mouthparts not dissected, though external observation made. Mandible without prominent tooth on occlusal margin of incisor process. Maxillule and maxilla without distinctive features. Maxilliped 1 without epipod. Maxilliped 2 without distinctive features. Maxilliped 3 (
<figureCitation id="131B2A04FFEEFF98FD6BFAD3FC91FA80" box="[744,834,1339,1365]" captionStart="FIGURE 8" captionStartId="11.[151,250,1906,1930]" captionTargetBox="[167,1422,606,1872]" captionTargetId="figure-205@11.[157,1431,597,1882]" captionTargetPageId="11" captionText="FIGURE 8. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, maxilliped 3, lateral view (setae omitted); B, left coxae of pereopods 1–3 (P1–P3), ventromesial view; C, ischium to carpus of left pereopod 3, lateral view (propodus and dactylus missing; setae omitted); D, left pereopod 4, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 5, lateral view; G, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283720" httpUri="https://zenodo.org/record/4283720/files/figure.png" pageId="9" pageNumber="316">Fig. 8A</figureCitation>
) with endopod moderately stout, extending as far as distal end of antennal peduncle; ischium to dactylus with thick long setae on lower (flexor) margins; ischium with 2 spinules on mesial face proximally; merus to propodus unarmed; dactylus nearly straight, subequal in length to propodus, terminating in blunt tip; exopod slender, not reaching dorsodistal margin of ischium, flagellum slender, simple; epipod small, leaf-like.
</paragraph>
<paragraph id="8B9F3681FFEEFF9BFF44FA07FA84FE94" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1437,151,321]" lastPageId="10" lastPageNumber="317" pageId="9" pageNumber="316">
Pereopod 1 (
<figureCitation id="131B2A04FFEEFF98FED6FA07FE44F9DC" box="[341,407,1519,1545]" captionStart="FIGURE 9" captionStartId="13.[151,250,1668,1692]" captionTargetBox="[201,1385,537,1629]" captionTargetId="figure-144@13.[181,1406,489,1645]" captionTargetPageId="13" captionText="FIGURE 9. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176, right pereopod 1. A, ischium and merus, lateral view (proximal part of merus damaged; setae omitted); B, carpus to dactylus, lateral view; C, same, mesial view (setae omitted); D, dactylus, lateral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283722" httpUri="https://zenodo.org/record/4283722/files/figure.png" pageId="9" pageNumber="316">Fig. 9</figureCitation>
A–D) relatively slender, subchelate; setation of merus to carpus typical of upogebiids. Coxa (
<figureCitation id="131B2A04FFEEFF98FF23F9FBFF25F9F8" box="[160,246,1555,1581]" captionStart="FIGURE 8" captionStartId="11.[151,250,1906,1930]" captionTargetBox="[167,1422,606,1872]" captionTargetId="figure-205@11.[157,1431,597,1882]" captionTargetPageId="11" captionText="FIGURE 8. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, maxilliped 3, lateral view (setae omitted); B, left coxae of pereopods 1–3 (P1–P3), ventromesial view; C, ischium to carpus of left pereopod 3, lateral view (propodus and dactylus missing; setae omitted); D, left pereopod 4, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 5, lateral view; G, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283720" httpUri="https://zenodo.org/record/4283720/files/figure.png" pageId="9" pageNumber="316">Fig. 8B</figureCitation>
) with 1 conspicuous procurved spine on ventral surface. Ischium with 2 small hook-like spines on lower margin. Merus with row of 6 evenly spaced, erect spines in proximal 0.6 of lower margin and 2 additional spinules subdistally; upper margin nearly straight, armed with 1 small subdistal spine. Carpus cup-shaped, widened distally, slightly more than half length of palm; upper surface with spines of various size, arranged in 3 irregular rows (distomesial spine strongest); upper lateral surface with shallow longitudinal sulcus, lower lateral margin sharply carinate; mesial face with few small spines, distomesial margin with 2 prominent, elongate spines (lower spine extending proximal 0.25 of palm); lower margin with small subterminal spine. Carpus-propodus articulation with clockwise torsion of about 20° from perpendicular in right side. Palm gently arcuate, about 2.9 times as long as maximal height at proximal end; upper margin with row of 12 small to strong, slender spines and sparse moderately long setae over entire length (distalmost spine terminal); dactylar condyle on lateral face little developed; mesial face with about 30 small to moderately strong spines arranged in 4 irregular longitudinal rows (uppermost row adjacent to palm upper margin); lower margin gently concave, with 1 prominent spine proximal to base of fixed finger and row of 6 small spines in proximal half; setose ridge extending from lower proximal angle to about 0.3 length of palm; fixed finger less than half-length of dactylus, gently curved, occlusal margin with 3 small, acute teeth proximally. Dactylus 0.6 length of palm, slightly curved, terminating in minute corneous claw; upper (extensor) margin bearing about 10 minute denticles becoming obsolete distally and row of numerous setae increasing in length proximally; lateral face with longitudinal carina adjacent to upper margin and 2 rows of setae along midline; mesial face with double row of setae superior to midline; occlusal margin carinate in proximal 0.6, rounded in distal 0.4.
</paragraph>
<caption id="DF5F6609FFEDFF9BFF14F89CFAADF82C" ID-DOI="http://doi.org/10.5281/zenodo.4283718" ID-Zenodo-Dep="4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="10" pageNumber="317" startId="10.[151,250,1908,1932]" targetBox="[166,1422,350,1875]" targetPageId="10">
<paragraph id="8B9F3681FFEDFF9BFF14F89CFAADF82C" blockId="10.[151,1437,1908,2041]" pageId="10" pageNumber="317">
<emphasis id="B954EA93FFEDFF9BFF14F89CFEC0F858" bold="true" box="[151,275,1908,1933]" pageId="10" pageNumber="317">FIGURE 7.</emphasis>
<taxonomicName id="4C204D02FFEDFF9BFE99F89DFDC9F858" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[282,538,1909,1933]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="317" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEDFF9BFE99F89DFDC9F858" box="[282,538,1909,1933]" italics="true" pageId="10" pageNumber="317">Gebiacantha fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEDFF9BFDA3F89DFD89F858" bold="true" box="[544,602,1909,1933]" pageId="10" pageNumber="317">
<taxonomicNameLabel id="A26757E8FFEDFF9BFDA3F89DFD89F858" box="[544,602,1909,1933]" pageId="10" pageNumber="317" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted).
</paragraph>
</caption>
<paragraph id="8B9F3681FFECFF9AFF44FF7FFAFCFF64" blockId="11.[151,1437,150,573]" box="[199,1327,150,177]" pageId="11" pageNumber="318">
Pereopod 2 (
<figureCitation id="131B2A04FFECFF9AFED6FF7FFE78FF64" box="[341,427,151,177]" captionStart="FIGURE 8" captionStartId="11.[151,250,1906,1930]" captionTargetBox="[167,1422,606,1872]" captionTargetId="figure-205@11.[157,1431,597,1882]" captionTargetPageId="11" captionText="FIGURE 8. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, maxilliped 3, lateral view (setae omitted); B, left coxae of pereopods 1–3 (P1–P3), ventromesial view; C, ischium to carpus of left pereopod 3, lateral view (propodus and dactylus missing; setae omitted); D, left pereopod 4, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 5, lateral view; G, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283720" httpUri="https://zenodo.org/record/4283720/files/figure.png" pageId="11" pageNumber="318">Fig. 8B</figureCitation>
) coxa bearing 5 spinules on ventromesial face; otherwise missing on both sides.
</paragraph>
<paragraph id="8B9F3681FFECFF9AFF44FF53FD69FE94" blockId="11.[151,1437,150,573]" pageId="11" pageNumber="318">
Pereopod 3 (only left preserved) (
<figureCitation id="131B2A04FFECFF9AFDBCFF53FD47FF00" box="[575,660,187,213]" captionStart="FIGURE 8" captionStartId="11.[151,250,1906,1930]" captionTargetBox="[167,1422,606,1872]" captionTargetId="figure-205@11.[157,1431,597,1882]" captionTargetPageId="11" captionText="FIGURE 8. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, maxilliped 3, lateral view (setae omitted); B, left coxae of pereopods 1–3 (P1–P3), ventromesial view; C, ischium to carpus of left pereopod 3, lateral view (propodus and dactylus missing; setae omitted); D, left pereopod 4, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 5, lateral view; G, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283720" httpUri="https://zenodo.org/record/4283720/files/figure.png" pageId="11" pageNumber="318">Fig. 8C</figureCitation>
) with coxa having scattered spinules on mesial face, including 1 conspicuous spine anterolateral to gonopore (
<figureCitation id="131B2A04FFECFF9AFDE6FF37FD6FFF2C" box="[613,700,223,249]" captionStart="FIGURE 8" captionStartId="11.[151,250,1906,1930]" captionTargetBox="[167,1422,606,1872]" captionTargetId="figure-205@11.[157,1431,597,1882]" captionTargetPageId="11" captionText="FIGURE 8. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, maxilliped 3, lateral view (setae omitted); B, left coxae of pereopods 1–3 (P1–P3), ventromesial view; C, ischium to carpus of left pereopod 3, lateral view (propodus and dactylus missing; setae omitted); D, left pereopod 4, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 5, lateral view; G, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283720" httpUri="https://zenodo.org/record/4283720/files/figure.png" pageId="11" pageNumber="318">Fig. 8B</figureCitation>
). Ischium unarmed. Merus with 1 small upper distal spine and row of 6 spines, widely spaced, on lower margin. Carpus widened distally; upper margin unarmed; lower margin with 1 tiny distal spine. Propodus and dactylus missing.
</paragraph>
<paragraph id="8B9F3681FFECFF9AFF44FEA3FA40FE04" blockId="11.[151,1437,150,573]" pageId="11" pageNumber="318">
Pereopod 4 (
<figureCitation id="131B2A04FFECFF9AFED6FEA3FE7EFEB0" box="[341,429,331,357]" captionStart="FIGURE 8" captionStartId="11.[151,250,1906,1930]" captionTargetBox="[167,1422,606,1872]" captionTargetId="figure-205@11.[157,1431,597,1882]" captionTargetPageId="11" captionText="FIGURE 8. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, maxilliped 3, lateral view (setae omitted); B, left coxae of pereopods 1–3 (P1–P3), ventromesial view; C, ischium to carpus of left pereopod 3, lateral view (propodus and dactylus missing; setae omitted); D, left pereopod 4, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 5, lateral view; G, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283720" httpUri="https://zenodo.org/record/4283720/files/figure.png" pageId="11" pageNumber="318">Fig. 8D</figureCitation>
) with coxa bearing 1 small spine on ventrodistal margin. Ischium to carpus unarmed; carpus cup-shaped. Propodus subequal in length to carpus, with gently arcuate upper margin; lower margin with row of minute denticles in proximal half. Dactylus (
<figureCitation id="131B2A04FFECFF9AFD37FE7BFCDAFE78" box="[692,777,403,429]" captionStart="FIGURE 8" captionStartId="11.[151,250,1906,1930]" captionTargetBox="[167,1422,606,1872]" captionTargetId="figure-205@11.[157,1431,597,1882]" captionTargetPageId="11" captionText="FIGURE 8. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, maxilliped 3, lateral view (setae omitted); B, left coxae of pereopods 1–3 (P1–P3), ventromesial view; C, ischium to carpus of left pereopod 3, lateral view (propodus and dactylus missing; setae omitted); D, left pereopod 4, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 5, lateral view; G, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283720" httpUri="https://zenodo.org/record/4283720/files/figure.png" pageId="11" pageNumber="318">Fig. 8E</figureCitation>
) subequal in length to propodus, faintly recurved distally, terminating in acute tip; lower (flexor) margin slightly sinuous, with row of 4 minute spiniform setae in distal 0.3.
</paragraph>
<paragraph id="8B9F3681FFECFF9AFF44FE33FDB5FDE8" blockId="11.[151,1437,150,573]" pageId="11" pageNumber="318">
Pereopod 5 (
<figureCitation id="131B2A04FFECFF9AFED6FE33FE1BFE20" box="[341,456,475,501]" captionStart="FIGURE 8" captionStartId="11.[151,250,1906,1930]" captionTargetBox="[167,1422,606,1872]" captionTargetId="figure-205@11.[157,1431,597,1882]" captionTargetPageId="11" captionText="FIGURE 8. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, maxilliped 3, lateral view (setae omitted); B, left coxae of pereopods 1–3 (P1–P3), ventromesial view; C, ischium to carpus of left pereopod 3, lateral view (propodus and dactylus missing; setae omitted); D, left pereopod 4, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 5, lateral view; G, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283720" httpUri="https://zenodo.org/record/4283720/files/figure.png" pageId="11" pageNumber="318">Fig. 8F, G</figureCitation>
) subchelate, articles unarmed; no trace of gonopore on coxa. Carpus subequal in length to merus and propodus. Propodus slightly arcuate, with short fixed finger reaching beyond midlength of dactylus. Dactylus curved, terminating in blunt tip.
</paragraph>
<caption id="DF5F6609FFECFF9AFF14F89AFB95F823" ID-DOI="http://doi.org/10.5281/zenodo.4283720" ID-Zenodo-Dep="4283720" httpUri="https://zenodo.org/record/4283720/files/figure.png" pageId="11" pageNumber="318" startId="11.[151,250,1906,1930]" targetBox="[167,1422,606,1872]" targetPageId="11">
<paragraph id="8B9F3681FFECFF9AFF14F89AFB95F823" blockId="11.[151,1437,1906,2038]" pageId="11" pageNumber="318">
<emphasis id="B954EA93FFECFF9AFF14F89AFEC0F85F" bold="true" box="[151,275,1906,1930]" pageId="11" pageNumber="318">FIGURE 8.</emphasis>
<taxonomicName id="4C204D02FFECFF9AFE99F89AFDCAF85F" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[282,537,1906,1930]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="318" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFECFF9AFE99F89AFDCAF85F" box="[282,537,1906,1930]" italics="true" pageId="11" pageNumber="318">Gebiacantha fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFECFF9AFDA3F89AFD8AF85F" bold="true" box="[544,601,1906,1930]" pageId="11" pageNumber="318">
<taxonomicNameLabel id="A26757E8FFECFF9AFDA3F89AFD8AF85F" box="[544,601,1906,1930]" pageId="11" pageNumber="318" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, female (cl 7.4 mm), CBM-ZC 16176. A, maxilliped 3, lateral view (setae omitted); B, left coxae of pereopods 1–3 (P1–P3), ventromesial view; C, ischium to carpus of left pereopod 3, lateral view (propodus and dactylus missing; setae omitted); D, left pereopod 4, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 5, lateral view; G, same, propodus and dactylus, upper view.
</paragraph>
</caption>
<paragraph id="8B9F3681FFEBFF9DFF44FF7FFF31FF00" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="319">
Arthrobranchs of
<typeStatus id="549B8823FFEBFF9DFE12FF7FFE12FF64" box="[401,449,151,177]" pageId="12" pageNumber="319">type</typeStatus>
C (cf.
<bibRefCitation id="EFB14B70FFEBFF9DFD90FF7FFD6CFF64" author="Ngoc-Ho, N." box="[531,703,151,177]" pageId="12" pageNumber="319" pagination="237 - 273" refId="ref8226" refString="Ngoc-Ho, N. (1981) A taxonomic study of the larvae of four thalassinid species (Decapoda, Thalassinidea) from the Gulf of Mexico. Bulletin of the British Museum (Natural History), London, Zoology, 40, 237 - 273." type="journal article" year="1981">Ngoc-Ho 1981</bibRefCitation>
), composed of deeply divided, narrow lamellae on either side of rachis.
</paragraph>
<paragraph id="8B9F3681FFEBFF9DFF44FF37FCCCFEC8" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="319">
Pleopods 1 (
<figureCitation id="131B2A04FFEBFF9DFED7FF37FE78FF2F" box="[340,427,223,250]" captionStart="FIGURE 6" captionStartId="8.[151,250,1723,1747]" captionTargetBox="[161,1426,391,1690]" captionTargetId="figure-64@8.[151,1436,381,1700]" captionTargetPageId="8" captionText="FIGURE 6. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, carapace and cephalic appendages, lateral view; B, thoracomere 8 (including coxa of pereopod 5), pleon, telson and left pleopod 1, lateral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283716" httpUri="https://zenodo.org/record/4283716/files/figure.png" pageId="12" pageNumber="319">Fig. 6B</figureCitation>
) of 2 articles. Pleopods 2–5 similar; endopods small and slender, about half-length of respective exopod; exopods relatively slender (cf.
<figureCitation id="131B2A04FFEBFF9DFD36FEEBFCDAFEC8" box="[693,777,259,285]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="12" pageNumber="319">Fig. 7E</figureCitation>
).
</paragraph>
<paragraph id="8B9F3681FFEBFF9DFF44FECFFD14FE20" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="319">
Uropod (
<figureCitation id="131B2A04FFEBFF9DFEA8FECFFEAEFE94" box="[299,381,295,321]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="12" pageNumber="319">Fig. 7F</figureCitation>
) with both endopod and exopod reaching posterior margin of telson. Protopod with small spine on posterior margin. Endopod subtrapezoidal with rounded distomesial angle and somewhat produced, rounded distolateral angle, 1.3 times as long as wide; inner margin nearly straight, outer margin slightly concave, with small knob-like tubercle proximally; dorsal surface with median carina, outer margin also bluntly carinate. Exopod widened distally, subtriangular, 1.4 times as long as wide, slightly exceeding beyond endopod, distal margin roundly truncate; upper surface with 2 longitudinal carina.
</paragraph>
<paragraph id="8B9F3681FFEBFF9DFF44FE16FDE6FDCC" blockId="12.[151,1437,151,2013]" box="[199,565,510,537]" pageId="12" pageNumber="319">
<emphasis id="B954EA93FFEBFF9DFF44FE16FE79FDCD" bold="true" box="[199,426,510,536]" pageId="12" pageNumber="319">Colouration in life.</emphasis>
Not known.
</paragraph>
</subSubSection>
<subSubSection id="C33A650AFFEBFF9CFF44FDCBFC97FE5C" lastPageId="13" lastPageNumber="320" pageId="12" pageNumber="319" type="discussion">
<paragraph id="8B9F3681FFEBFF9DFF44FDCBFEE8FDB4" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="319">
<emphasis id="B954EA93FFEBFF9DFF44FDCBFEE8FDE8" bold="true" box="[199,315,547,573]" pageId="12" pageNumber="319">Remarks.</emphasis>
The
<typeStatus id="549B8823FFEBFF9DFEF7FDCBFE05FDE8" box="[372,470,547,573]" pageId="12" pageNumber="319" type="holotype">holotype</typeStatus>
of the new species is parasitized by an unidentified bopyrid, inhabiting the right branchial chamber.
</paragraph>
<paragraph id="8B9F3681FFEBFF9DFF48FD83FB55FAE4" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="319">
As mentioned above,
<taxonomicName id="4C204D02FFEBFF9DFE38FD83FD06FD50" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[443,725,619,645]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFE38FD83FD06FD50" box="[443,725,619,645]" italics="true" pageId="12" pageNumber="319">Gebiacantha fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFD59FD83FCC4FD50" bold="true" box="[730,791,619,645]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFD59FD83FCC4FD50" box="[730,791,619,645]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is assigned to the species group characterised by a distally widened, subtriangular uropodal exopod (with a subtruncate distal margin). The new species appears closest to
<taxonomicName id="4C204D02FFEBFF9DFA02FD67FEF5FD18" authorityName="Ngoc-Ho" authorityYear="1994" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="multispinosa">
<emphasis id="B954EA93FFEBFF9DFA02FD67FEF5FD18" italics="true" pageId="12" pageNumber="319">G. multispinosa</emphasis>
</taxonomicName>
in having a few spinules on the postocular region of the carapace, six or more spinules on the anterolateral margin of the carapace, many spinules on the shoulder along the cervical groove, 11 or more spines on the upper margin of the pereopod 1 propodus, more than 20 spines, arranged in four longitudinal rows, on the mesial face of the pereopod 1 palm and the presence of denticles on the occlusal margin of the pereopod 1 fixed finger (cf.
<bibRefCitation id="EFB14B70FFEBFF9DFF14FCABFE83FC88" author="Ngoc-Ho, N." box="[151,336,835,861]" pageId="12" pageNumber="319" pagination="193 - 216" refId="ref8328" refString="Ngoc-Ho, N. (1994 a) Notes on some Indo-Pacific Upogebiidae with descriptions of four new species (Crustacea: Thalassinidea). Memoirs of the Queensland Museum, 35, 193 - 216." type="journal article" year="1994">Ngoc-Ho 1994a</bibRefCitation>
). Nevertheless, comparison with the original description of
<taxonomicName id="4C204D02FFEBFF9DFC77FCABFB75FC88" authorityName="Ngoc-Ho" authorityYear="1994" box="[1012,1190,835,861]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="multispinosa">
<emphasis id="B954EA93FFEBFF9DFC77FCABFB75FC88" box="[1012,1190,835,861]" italics="true" pageId="12" pageNumber="319">G. multispinosa</emphasis>
</taxonomicName>
reveals the following differences warranting the recognition of the new taxon: (1) four rostral ventral spines are rather widely spaced in the distal 0.7 length of the rostrum in
<taxonomicName id="4C204D02FFEBFF9DFDBCFC63FD37FC70" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[575,740,907,933]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFDBCFC63FD37FC70" box="[575,740,907,933]" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFD68FC63FCF9FC70" bold="true" box="[747,810,907,933]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFD68FC63FCF9FC70" box="[747,810,907,933]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B2A04FFEBFF9DFCB9FC63FC47FC70" box="[826,916,907,933]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="12" pageNumber="319">Fig. 7A</figureCitation>
), whereas two or three ventral spines are clustered in the distal
<quantity id="4CD89B64FFEBFF9DFEE0FC47FE77FC1C" box="[355,420,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="319" unit="in" value="0.3">0.3 in</quantity>
<taxonomicName id="4C204D02FFEBFF9DFE2FFC47FD8DFC1C" authorityName="Ngoc-Ho" authorityYear="1994" box="[428,606,943,969]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="multispinosa">
<emphasis id="B954EA93FFEBFF9DFE2FFC47FD8DFC1C" box="[428,606,943,969]" italics="true" pageId="12" pageNumber="319">G. multispinosa</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFB14B70FFEBFF9DFD17FC47FC9EFC1C" author="Ngoc-Ho, N." box="[660,845,943,969]" pageId="12" pageNumber="319" pagination="193 - 216" refId="ref8328" refString="Ngoc-Ho, N. (1994 a) Notes on some Indo-Pacific Upogebiidae with descriptions of four new species (Crustacea: Thalassinidea). Memoirs of the Queensland Museum, 35, 193 - 216." type="journal article" year="1994">Ngoc-Ho 1994a</bibRefCitation>
: fig. 2B); (2) the telson is relatively narrower with a more deeply notched posterior margin in
<taxonomicName id="4C204D02FFEBFF9DFDEEFC3BFCC0FC38" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[621,787,979,1005]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFDEEFC3BFCC0FC38" box="[621,787,979,1005]" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFC98FC3BFC88FC38" bold="true" box="[795,859,979,1005]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFC98FC3BFC88FC38" box="[795,859,979,1005]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
than in
<taxonomicName id="4C204D02FFEBFF9DFC3AFC3BFBBFFC38" authorityName="Ngoc-Ho" authorityYear="1994" box="[953,1132,979,1005]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="multispinosa">
<emphasis id="B954EA93FFEBFF9DFC3AFC3BFBBFFC38" box="[953,1132,979,1005]" italics="true" pageId="12" pageNumber="319">G. multispinosa</emphasis>
</taxonomicName>
(1.2 times as wide as long versus 1.5 times: cf.
<figureCitation id="131B2A04FFEBFF9DFE03FC1FFE07FBC4" box="[384,468,1015,1041]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="12" pageNumber="319">Fig. 7C</figureCitation>
and
<bibRefCitation id="EFB14B70FFEBFF9DFD88FC1FFD17FBC4" author="Ngoc-Ho, N." box="[523,708,1015,1041]" pageId="12" pageNumber="319" pagination="193 - 216" refId="ref8328" refString="Ngoc-Ho, N. (1994 a) Notes on some Indo-Pacific Upogebiidae with descriptions of four new species (Crustacea: Thalassinidea). Memoirs of the Queensland Museum, 35, 193 - 216." type="journal article" year="1994">Ngoc-Ho 1994a</bibRefCitation>
: fig. 2G); (3) the cornea of the eye is relatively larger in
<taxonomicName id="4C204D02FFEBFF9DFAC3FC1FFF3AFBE1" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFAC3FC1FFF3AFBE1" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFF72FBF3FEE3FBE0" bold="true" box="[241,304,1051,1077]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFF72FBF3FEE3FBE0" box="[241,304,1051,1077]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
than in
<taxonomicName id="4C204D02FFEBFF9DFE0FFBF3FDEDFBE0" authorityName="Ngoc-Ho" authorityYear="1994" box="[396,574,1051,1077]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="multispinosa">
<emphasis id="B954EA93FFEBFF9DFE0FFBF3FDEDFBE0" box="[396,574,1051,1077]" italics="true" pageId="12" pageNumber="319">G. multispinosa</emphasis>
</taxonomicName>
(cf.
<figureCitation id="131B2A04FFEBFF9DFDF0FBF3FD14FBE0" box="[627,711,1051,1077]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="12" pageNumber="319">Fig. 7A</figureCitation>
and Ngoc-Ho 1994: fig. 2B); (4) the pereopod 1 propodus bears six small spines on the proximal half of the lower margin in
<taxonomicName id="4C204D02FFEBFF9DFCB0FBD7FC04FB8C" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[819,983,1087,1113]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFCB0FBD7FC04FB8C" box="[819,983,1087,1113]" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFC5EFBD7FBC8FB8C" bold="true" box="[989,1051,1087,1113]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFC5EFBD7FBC8FB8C" box="[989,1051,1087,1113]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B2A04FFEBFF9DFBA9FBD7FB52FB8C" box="[1066,1153,1087,1113]" captionStart="FIGURE 9" captionStartId="13.[151,250,1668,1692]" captionTargetBox="[201,1385,537,1629]" captionTargetId="figure-144@13.[181,1406,489,1645]" captionTargetPageId="13" captionText="FIGURE 9. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176, right pereopod 1. A, ischium and merus, lateral view (proximal part of merus damaged; setae omitted); B, carpus to dactylus, lateral view; C, same, mesial view (setae omitted); D, dactylus, lateral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283722" httpUri="https://zenodo.org/record/4283722/files/figure.png" pageId="12" pageNumber="319">Fig. 9B</figureCitation>
), whereas there are about 10 spinules in the proximal two-thirds in
<taxonomicName id="4C204D02FFEBFF9DFDE1FB8BFCC0FBA8" authorityName="Ngoc-Ho" authorityYear="1994" box="[610,787,1123,1149]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="multispinosa">
<emphasis id="B954EA93FFEBFF9DFDE1FB8BFCC0FBA8" box="[610,787,1123,1149]" italics="true" pageId="12" pageNumber="319">G. multispinosa</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFB14B70FFEBFF9DFCC5FB8BFC2EFBA8" author="Ngoc-Ho, N." box="[838,1021,1123,1149]" pageId="12" pageNumber="319" pagination="193 - 216" refId="ref8328" refString="Ngoc-Ho, N. (1994 a) Notes on some Indo-Pacific Upogebiidae with descriptions of four new species (Crustacea: Thalassinidea). Memoirs of the Queensland Museum, 35, 193 - 216." type="journal article" year="1994">Ngoc-Ho 1994a</bibRefCitation>
: fig. 2C); (5) the pereopod 1 dactylus bears 10 more widely spaced small denticles on the upper margin in
<taxonomicName id="4C204D02FFEBFF9DFC23FB6FFB95FB74" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[928,1094,1159,1185]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFC23FB6FFB95FB74" box="[928,1094,1159,1185]" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFBCEFB6FFB5EFB74" bold="true" box="[1101,1165,1159,1185]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFBCEFB6FFB5EFB74" box="[1101,1165,1159,1185]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B2A04FFEBFF9DFB1EFB6FFB25FB74" box="[1181,1270,1159,1185]" captionStart="FIGURE 9" captionStartId="13.[151,250,1668,1692]" captionTargetBox="[201,1385,537,1629]" captionTargetId="figure-144@13.[181,1406,489,1645]" captionTargetPageId="13" captionText="FIGURE 9. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176, right pereopod 1. A, ischium and merus, lateral view (proximal part of merus damaged; setae omitted); B, carpus to dactylus, lateral view; C, same, mesial view (setae omitted); D, dactylus, lateral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283722" httpUri="https://zenodo.org/record/4283722/files/figure.png" pageId="12" pageNumber="319">Fig. 9D</figureCitation>
) (versus about 15 denticles more narrowly spaced in
<taxonomicName id="4C204D02FFEBFF9DFDBEFB43FD3EFB10" authorityName="Ngoc-Ho" authorityYear="1994" box="[573,749,1195,1221]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="multispinosa">
<emphasis id="B954EA93FFEBFF9DFDBEFB43FD3EFB10" box="[573,749,1195,1221]" italics="true" pageId="12" pageNumber="319">G. multispinosa</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFB14B70FFEBFF9DFD78FB43FC61FB10" author="Ngoc-Ho, N." box="[763,946,1195,1221]" pageId="12" pageNumber="319" pagination="193 - 216" refId="ref8328" refString="Ngoc-Ho, N. (1994 a) Notes on some Indo-Pacific Upogebiidae with descriptions of four new species (Crustacea: Thalassinidea). Memoirs of the Queensland Museum, 35, 193 - 216." type="journal article" year="1994">Ngoc-Ho 1994a</bibRefCitation>
: fig. 2C); (6) the pereopod 3 merus is armed with a small dorsodistal spine in
<taxonomicName id="4C204D02FFEBFF9DFD87FB27FD7AFB3C" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[516,681,1231,1257]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFD87FB27FD7AFB3C" box="[516,681,1231,1257]" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFD2CFB27FD3EFB3C" bold="true" box="[687,749,1231,1257]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFD2CFB27FD3EFB3C" box="[687,749,1231,1257]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B2A04FFEBFF9DFD7FFB27FC86FB3C" box="[764,853,1231,1257]" captionStart="FIGURE 9" captionStartId="13.[151,250,1668,1692]" captionTargetBox="[201,1385,537,1629]" captionTargetId="figure-144@13.[181,1406,489,1645]" captionTargetPageId="13" captionText="FIGURE 9. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176, right pereopod 1. A, ischium and merus, lateral view (proximal part of merus damaged; setae omitted); B, carpus to dactylus, lateral view; C, same, mesial view (setae omitted); D, dactylus, lateral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283722" httpUri="https://zenodo.org/record/4283722/files/figure.png" pageId="12" pageNumber="319">Fig. 9A</figureCitation>
), while such a spine is absent in
<taxonomicName id="4C204D02FFEBFF9DFB3CFB27FAA3FB3C" authorityName="Ngoc-Ho" authorityYear="1994" box="[1215,1392,1231,1257]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="multispinosa">
<emphasis id="B954EA93FFEBFF9DFB3CFB27FAA3FB3C" box="[1215,1392,1231,1257]" italics="true" pageId="12" pageNumber="319">G. multispinosa</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFB14B70FFEBFF9DFF14FB1BFE83FAD8" author="Ngoc-Ho, N." box="[151,336,1267,1293]" pageId="12" pageNumber="319" pagination="193 - 216" refId="ref8328" refString="Ngoc-Ho, N. (1994 a) Notes on some Indo-Pacific Upogebiidae with descriptions of four new species (Crustacea: Thalassinidea). Memoirs of the Queensland Museum, 35, 193 - 216." type="journal article" year="1994">Ngoc-Ho 1994a</bibRefCitation>
: fig. 2F); (8) the posterolateral angle of the uropodal endopod is rounded in
<taxonomicName id="4C204D02FFEBFF9DFB2CFB1BFA86FAD8" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[1199,1365,1267,1293]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFB2CFB1BFA86FAD8" box="[1199,1365,1267,1293]" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFADFFB1BFA4FFAD8" bold="true" box="[1372,1436,1267,1293]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFADFFB1BFA4FFAD8" box="[1372,1436,1267,1293]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B2A04FFEBFF9DFF23FAFFFF27FAE4" box="[160,244,1303,1329]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="12" pageNumber="319">Fig. 7F</figureCitation>
), rather than subacutely pointed in
<taxonomicName id="4C204D02FFEBFF9DFDFCFAFFFCE3FAE4" authorityName="Ngoc-Ho" authorityYear="1994" box="[639,816,1303,1329]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="multispinosa">
<emphasis id="B954EA93FFEBFF9DFDFCFAFFFCE3FAE4" box="[639,816,1303,1329]" italics="true" pageId="12" pageNumber="319">G. multispinosa</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFB14B70FFEBFF9DFCE7FAFFFBCFFAE4" author="Ngoc-Ho, N." box="[868,1052,1303,1329]" pageId="12" pageNumber="319" pagination="193 - 216" refId="ref8328" refString="Ngoc-Ho, N. (1994 a) Notes on some Indo-Pacific Upogebiidae with descriptions of four new species (Crustacea: Thalassinidea). Memoirs of the Queensland Museum, 35, 193 - 216." type="journal article" year="1994">Ngoc-Ho 1994a</bibRefCitation>
: fig. 2G).
</paragraph>
<paragraph id="8B9F3681FFEBFF9DFF44FAD3FB13F8A4" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="319">
<taxonomicName id="4C204D02FFEBFF9DFF44FAD3FE26FA80" baseAuthorityName="Sakai" baseAuthorityYear="1967" box="[199,501,1339,1365]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="acanthochela">
<emphasis id="B954EA93FFEBFF9DFF44FAD3FE26FA80" box="[199,501,1339,1365]" italics="true" pageId="12" pageNumber="319">Gebiacantha acanthochela</emphasis>
</taxonomicName>
is also similar to the new species in having relatively numerous spines on the upper margin and mesial face of the pereopod 1 propodus.
<taxonomicName id="4C204D02FFEBFF9DFD77FAB7FBC3FAAC" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[756,1040,1375,1401]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFD77FAB7FBC3FAAC" box="[756,1040,1375,1401]" italics="true" pageId="12" pageNumber="319">Gebiacantha fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFB9AFAB7FB8AFAAC" bold="true" box="[1049,1113,1375,1401]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFB9AFAB7FB8AFAAC" box="[1049,1113,1375,1401]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is easily distinguished from
<taxonomicName id="4C204D02FFEBFF9DFF14FA6BFE81FA48" baseAuthorityName="Sakai" baseAuthorityYear="1967" box="[151,338,1411,1437]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="acanthochela">
<emphasis id="B954EA93FFEBFF9DFF14FA6BFE81FA48" box="[151,338,1411,1437]" italics="true" pageId="12" pageNumber="319">G. acanthochela</emphasis>
</taxonomicName>
by the following particulars: (1) dorsolateral tubercles on the rostrum are more acuminate in
<taxonomicName id="4C204D02FFEBFF9DFA02FA6BFEC9FA14" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFA02FA6BFEC9FA14" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFEA1FA4FFEB1FA14" bold="true" box="[290,354,1447,1473]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFEA1FA4FFEB1FA14" box="[290,354,1447,1473]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
than in
<taxonomicName id="4C204D02FFEBFF9DFE41FA4FFDAEFA14" baseAuthorityName="Sakai" baseAuthorityYear="1967" box="[450,637,1447,1473]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="acanthochela">
<emphasis id="B954EA93FFEBFF9DFE41FA4FFDAEFA14" box="[450,637,1447,1473]" italics="true" pageId="12" pageNumber="319">G. acanthochela</emphasis>
</taxonomicName>
(cf.
<figureCitation id="131B2A04FFEBFF9DFD37FA4FFCDAFA14" box="[692,777,1447,1473]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="12" pageNumber="319">Fig. 7A</figureCitation>
and
<figureCitation id="131B2A04FFEBFF9DFCC1FA4FFC4EFA14" box="[834,925,1447,1473]" captionStart="FIGURE 3" captionStartId="5.[151,250,1864,1888]" captionTargetBox="[218,1365,193,1838]" captionTargetId="figure-15@5.[209,1378,181,1840]" captionTargetPageId="5" captionText="FIGURE 3. Gebiacantha acanthochela (Sakai, 1967), ovigerous female (cl 8.2 mm), CBM-ZC 16175. A, maxilliped 3, lateral view (setae omitted); B, right coxae of pereopods 1–3 (P1–P3), ventromesial view; C, left pereopod 2, lateral view (setae omitted); D, left pereopod 3, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 4, lateral view (setae omitted); G, same, dactylus, mesial view (setae omitted); H, left pereopod 5, lateral view; I, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283710" httpUri="https://zenodo.org/record/4283710/files/figure.png" pageId="12" pageNumber="319">Fig. 3A</figureCitation>
); (2) four rostral ventral spines are relatively stronger, of which the posteriormost spine is located proximal to the midlength of the rostrum in
<taxonomicName id="4C204D02FFEBFF9DFB76FA23FA4EFA30" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[1269,1437,1483,1509]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFB76FA23FA4EFA30" box="[1269,1437,1483,1509]" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFF14FA07FF0BF9DC" bold="true" box="[151,216,1519,1545]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFF14FA07FF0BF9DC" box="[151,216,1519,1545]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B2A04FFEBFF9DFF69FA07FE97F9DC" box="[234,324,1519,1545]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="12" pageNumber="319">Fig. 7A</figureCitation>
) (versus two or three ventral spines are clustered in the subterminal portion of the rostrum in
<taxonomicName id="4C204D02FFEBFF9DFA02FA07FEFDF9F8" baseAuthorityName="Sakai" baseAuthorityYear="1967" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="acanthochela">
<emphasis id="B954EA93FFEBFF9DFA02FA07FEFDF9F8" italics="true" pageId="12" pageNumber="319">G. acanthochela</emphasis>
</taxonomicName>
;
<figureCitation id="131B2A04FFEBFF9DFEBFF9FBFE46F9F8" box="[316,405,1555,1581]" captionStart="FIGURE 3" captionStartId="5.[151,250,1864,1888]" captionTargetBox="[218,1365,193,1838]" captionTargetId="figure-15@5.[209,1378,181,1840]" captionTargetPageId="5" captionText="FIGURE 3. Gebiacantha acanthochela (Sakai, 1967), ovigerous female (cl 8.2 mm), CBM-ZC 16175. A, maxilliped 3, lateral view (setae omitted); B, right coxae of pereopods 1–3 (P1–P3), ventromesial view; C, left pereopod 2, lateral view (setae omitted); D, left pereopod 3, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 4, lateral view (setae omitted); G, same, dactylus, mesial view (setae omitted); H, left pereopod 5, lateral view; I, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283710" httpUri="https://zenodo.org/record/4283710/files/figure.png" pageId="12" pageNumber="319">Fig. 3A</figureCitation>
); (3) the carapace postocular region is armed with two spinules in
<taxonomicName id="4C204D02FFEBFF9DFBF4F9FBFAC8F9F8" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[1143,1307,1555,1581]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFBF4F9FBFAC8F9F8" box="[1143,1307,1555,1581]" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFAA1F9FBFAB3F9F8" bold="true" box="[1314,1376,1555,1581]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFAA1F9FBFAB3F9F8" box="[1314,1376,1555,1581]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B2A04FFEBFF9DFAECF9FBFF6FF984" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="12" pageNumber="319">Fig. 7A</figureCitation>
), rather than unarmed or at most with a few minute granules in
<taxonomicName id="4C204D02FFEBFF9DFC02F9DFFBE9F984" baseAuthorityName="Sakai" baseAuthorityYear="1967" box="[897,1082,1591,1617]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="acanthochela">
<emphasis id="B954EA93FFEBFF9DFC02F9DFFBE9F984" box="[897,1082,1591,1617]" italics="true" pageId="12" pageNumber="319">G. acanthochela</emphasis>
</taxonomicName>
(
<figureCitation id="131B2A04FFEBFF9DFBCAF9DFFB71F984" box="[1097,1186,1591,1617]" captionStart="FIGURE 3" captionStartId="5.[151,250,1864,1888]" captionTargetBox="[218,1365,193,1838]" captionTargetId="figure-15@5.[209,1378,181,1840]" captionTargetPageId="5" captionText="FIGURE 3. Gebiacantha acanthochela (Sakai, 1967), ovigerous female (cl 8.2 mm), CBM-ZC 16175. A, maxilliped 3, lateral view (setae omitted); B, right coxae of pereopods 1–3 (P1–P3), ventromesial view; C, left pereopod 2, lateral view (setae omitted); D, left pereopod 3, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 4, lateral view (setae omitted); G, same, dactylus, mesial view (setae omitted); H, left pereopod 5, lateral view; I, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283710" httpUri="https://zenodo.org/record/4283710/files/figure.png" pageId="12" pageNumber="319">Fig. 3A</figureCitation>
); (4) the shoulder bordering the cervical groove is armed with eight spinules in
<taxonomicName id="4C204D02FFEBFF9DFCAEF9B3FC00F9A0" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[813,979,1627,1653]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFCAEF9B3FC00F9A0" box="[813,979,1627,1653]" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFC58F9B3FBC8F9A0" bold="true" box="[987,1051,1627,1653]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFC58F9B3FBC8F9A0" box="[987,1051,1627,1653]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B2A04FFEBFF9DFBAFF9B3FB54F9A0" box="[1068,1159,1627,1653]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="12" pageNumber="319">Fig. 7A</figureCitation>
), whereas with only one small spine at the crossing point with the linea thalassinica in
<taxonomicName id="4C204D02FFEBFF9DFCD4F997FBC2F94C" baseAuthorityName="Sakai" baseAuthorityYear="1967" box="[855,1041,1663,1689]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="acanthochela">
<emphasis id="B954EA93FFEBFF9DFCD4F997FBC2F94C" box="[855,1041,1663,1689]" italics="true" pageId="12" pageNumber="319">G. acanthochela</emphasis>
</taxonomicName>
(
<figureCitation id="131B2A04FFEBFF9DFBA2F997FBAFF94C" box="[1057,1148,1663,1689]" captionStart="FIGURE 3" captionStartId="5.[151,250,1864,1888]" captionTargetBox="[218,1365,193,1838]" captionTargetId="figure-15@5.[209,1378,181,1840]" captionTargetPageId="5" captionText="FIGURE 3. Gebiacantha acanthochela (Sakai, 1967), ovigerous female (cl 8.2 mm), CBM-ZC 16175. A, maxilliped 3, lateral view (setae omitted); B, right coxae of pereopods 1–3 (P1–P3), ventromesial view; C, left pereopod 2, lateral view (setae omitted); D, left pereopod 3, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 4, lateral view (setae omitted); G, same, dactylus, mesial view (setae omitted); H, left pereopod 5, lateral view; I, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283710" httpUri="https://zenodo.org/record/4283710/files/figure.png" pageId="12" pageNumber="319">Fig. 3A</figureCitation>
); (5) the median notch of the telson posterior margin is deeper in
<taxonomicName id="4C204D02FFEBFF9DFDD0F94BFD2AF968" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[595,761,1699,1725]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFDD0F94BFD2AF968" box="[595,761,1699,1725]" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFC83F94BFCEDF968" bold="true" box="[768,830,1699,1725]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFC83F94BFCEDF968" box="[768,830,1699,1725]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
than in
<taxonomicName id="4C204D02FFEBFF9DFC19F94BFB80F968" baseAuthorityName="Sakai" baseAuthorityYear="1967" box="[922,1107,1699,1725]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="acanthochela">
<emphasis id="B954EA93FFEBFF9DFC19F94BFB80F968" box="[922,1107,1699,1725]" italics="true" pageId="12" pageNumber="319">G. acanthochela</emphasis>
</taxonomicName>
(
<figureCitation id="131B2A04FFEBFF9DFBE0F94BFB6BF968" box="[1123,1208,1699,1725]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="12" pageNumber="319">Fig. 7C</figureCitation>
versus
<figureCitation id="131B2A04FFEBFF9DFA8DF94BFAB4F968" box="[1294,1383,1699,1725]" captionStart="FIGURE 3" captionStartId="5.[151,250,1864,1888]" captionTargetBox="[218,1365,193,1838]" captionTargetId="figure-15@5.[209,1378,181,1840]" captionTargetPageId="5" captionText="FIGURE 3. Gebiacantha acanthochela (Sakai, 1967), ovigerous female (cl 8.2 mm), CBM-ZC 16175. A, maxilliped 3, lateral view (setae omitted); B, right coxae of pereopods 1–3 (P1–P3), ventromesial view; C, left pereopod 2, lateral view (setae omitted); D, left pereopod 3, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 4, lateral view (setae omitted); G, same, dactylus, mesial view (setae omitted); H, left pereopod 5, lateral view; I, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283710" httpUri="https://zenodo.org/record/4283710/files/figure.png" pageId="12" pageNumber="319">Fig. 3D</figureCitation>
); (6) the antennal peduncle article 1 is armed with one prominent spine on the ventrodistal margin in
<taxonomicName id="4C204D02FFEBFF9DFB5BF92FFAADF934" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[1240,1406,1735,1761]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFB5BF92FFAADF934" box="[1240,1406,1735,1761]" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFA05F92FFF6BF8D0" bold="true" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFA05F92FFF6BF8D0" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B2A04FFEBFF9DFF4BF903FEDAF8D0" box="[200,265,1771,1797]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1932]" captionTargetBox="[166,1422,350,1875]" captionTargetId="figure-115@10.[158,1430,345,1885]" captionTargetPageId="10" captionText="FIGURE 7. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176. A, rostrum, anterior part of carapace and cephalic appendages, lateral view (setae omitted); B, rostrum and anterior part of carapace, dorsal view (setae omitted); C, pleomere 6 and telson, dorsal view, perpendicular to horizontal plane (setae omitted); D, left antennule, ventrolateral view; F, left pleopod 2, posterior view (setae omitted); G, left uropod, dorsal view, perpendicular to horizontal plane (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283718" httpUri="https://zenodo.org/record/4283718/files/figure.png" pageId="12" pageNumber="319">Fig. 7</figureCitation>
AO, while unarmed in
<taxonomicName id="4C204D02FFEBFF9DFD8FF903FD16F8D0" baseAuthorityName="Sakai" baseAuthorityYear="1967" box="[524,709,1771,1797]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="acanthochela">
<emphasis id="B954EA93FFEBFF9DFD8FF903FD16F8D0" box="[524,709,1771,1797]" italics="true" pageId="12" pageNumber="319">G. acanthochela</emphasis>
</taxonomicName>
(
<figureCitation id="131B2A04FFEBFF9DFD56F903FCFDF8D0" box="[725,814,1771,1797]" captionStart="FIGURE 3" captionStartId="5.[151,250,1864,1888]" captionTargetBox="[218,1365,193,1838]" captionTargetId="figure-15@5.[209,1378,181,1840]" captionTargetPageId="5" captionText="FIGURE 3. Gebiacantha acanthochela (Sakai, 1967), ovigerous female (cl 8.2 mm), CBM-ZC 16175. A, maxilliped 3, lateral view (setae omitted); B, right coxae of pereopods 1–3 (P1–P3), ventromesial view; C, left pereopod 2, lateral view (setae omitted); D, left pereopod 3, lateral view (setae omitted); E, same, dactylus, mesial view (setae omitted); F, left pereopod 4, lateral view (setae omitted); G, same, dactylus, mesial view (setae omitted); H, left pereopod 5, lateral view; I, same, propodus and dactylus, upper view." figureDoi="http://doi.org/10.5281/zenodo.4283710" httpUri="https://zenodo.org/record/4283710/files/figure.png" pageId="12" pageNumber="319">Fig. 3A</figureCitation>
); (7) the occlusal margin of the pereopod 1 fixed finger bears three small acute denticles proximally in
<taxonomicName id="4C204D02FFEBFF9DFD2EF8E7FC83F8FC" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[685,848,1807,1833]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortitpinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFD2EF8E7FC83F8FC" box="[685,848,1807,1833]" italics="true" pageId="12" pageNumber="319">G. fortitpinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFCD4F8E7FC44F8FC" bold="true" box="[855,919,1807,1833]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFCD4F8E7FC44F8FC" box="[855,919,1807,1833]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B2A04FFEBFF9DFC24F8E7FBD3F8FC" box="[935,1024,1807,1833]" captionStart="FIGURE 9" captionStartId="13.[151,250,1668,1692]" captionTargetBox="[201,1385,537,1629]" captionTargetId="figure-144@13.[181,1406,489,1645]" captionTargetPageId="13" captionText="FIGURE 9. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176, right pereopod 1. A, ischium and merus, lateral view (proximal part of merus damaged; setae omitted); B, carpus to dactylus, lateral view; C, same, mesial view (setae omitted); D, dactylus, lateral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283722" httpUri="https://zenodo.org/record/4283722/files/figure.png" pageId="12" pageNumber="319">Fig. 9C</figureCitation>
), while unarmed in
<taxonomicName id="4C204D02FFEBFF9DFB61F8E7FA4FF8FC" baseAuthorityName="Sakai" baseAuthorityYear="1967" box="[1250,1436,1807,1833]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="acanthochela">
<emphasis id="B954EA93FFEBFF9DFB61F8E7FA4FF8FC" box="[1250,1436,1807,1833]" italics="true" pageId="12" pageNumber="319">G. acanthochela</emphasis>
</taxonomicName>
(
<figureCitation id="131B2A04FFEBFF9DFF23F8DBFF25F898" box="[160,246,1842,1869]" captionStart="FIGURE 5" captionStartId="7.[151,250,1892,1916]" captionTargetBox="[151,1435,1245,1869]" captionTargetId="figure-549@7.[151,1435,1245,1869]" captionTargetPageId="7" captionText="FIGURE 5. Gebiacantha acanthochela (Sakai, 1967), ovigerous female (cl 8.2 mm), CBM-ZC 16175, habitus in dorsal view, showing living colouration." figureDoi="http://doi.org/10.5281/zenodo.4283714" httpUri="https://zenodo.org/record/4283714/files/figure.png" pageId="12" pageNumber="319">Fig. 5C</figureCitation>
); (8) the pereopod 1 dactylus bears 10 spiniform denticles on the upper (extensor) margin in
<taxonomicName id="4C204D02FFEBFF9DFB7BF8DBFA4EF898" authority="Komai, 2020" authorityName="Komai" authorityYear="2020" box="[1272,1437,1843,1869]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="fortispinata" status="sp. nov.">
<emphasis id="B954EA93FFEBFF9DFB7BF8DBFA4EF898" box="[1272,1437,1843,1869]" italics="true" pageId="12" pageNumber="319">G. fortispinata</emphasis>
</taxonomicName>
<emphasis id="B954EA93FFEBFF9DFF14F8BFFF05F8A4" bold="true" box="[151,214,1879,1905]" pageId="12" pageNumber="319">
<taxonomicNameLabel id="A26757E8FFEBFF9DFF14F8BFFF05F8A4" box="[151,214,1879,1905]" pageId="12" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B2A04FFEBFF9DFF65F8BFFEECF8A4" box="[230,319,1879,1905]" captionStart="FIGURE 9" captionStartId="13.[151,250,1668,1692]" captionTargetBox="[201,1385,537,1629]" captionTargetId="figure-144@13.[181,1406,489,1645]" captionTargetPageId="13" captionText="FIGURE 9. Gebiacantha fortispinata n. sp., holotype, female (cl 7.4 mm), CBM-ZC 16176, right pereopod 1. A, ischium and merus, lateral view (proximal part of merus damaged; setae omitted); B, carpus to dactylus, lateral view; C, same, mesial view (setae omitted); D, dactylus, lateral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4283722" httpUri="https://zenodo.org/record/4283722/files/figure.png" pageId="12" pageNumber="319">Fig. 9D</figureCitation>
), whereas having only four or five blunt denticles in
<taxonomicName id="4C204D02FFEBFF9DFC12F8BFFB99F8A4" baseAuthorityName="Sakai" baseAuthorityYear="1967" box="[913,1098,1879,1905]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="acanthochela">
<emphasis id="B954EA93FFEBFF9DFC12F8BFFB99F8A4" box="[913,1098,1879,1905]" italics="true" pageId="12" pageNumber="319">G. acanthochela</emphasis>
</taxonomicName>
(
<figureCitation id="131B2A04FFEBFF9DFBDAF8BFFB61F8A4" box="[1113,1202,1878,1905]" captionStart="FIGURE 5" captionStartId="7.[151,250,1892,1916]" captionTargetBox="[151,1435,1245,1869]" captionTargetId="figure-549@7.[151,1435,1245,1869]" captionTargetPageId="7" captionText="FIGURE 5. Gebiacantha acanthochela (Sakai, 1967), ovigerous female (cl 8.2 mm), CBM-ZC 16175, habitus in dorsal view, showing living colouration." figureDoi="http://doi.org/10.5281/zenodo.4283714" httpUri="https://zenodo.org/record/4283714/files/figure.png" pageId="12" pageNumber="319">Fig. 5D</figureCitation>
).
</paragraph>
<paragraph id="8B9F3681FFEBFF9CFF44F893FEFEFEC8" blockId="12.[151,1437,151,2013]" lastBlockId="13.[151,1437,151,465]" lastPageId="13" lastPageNumber="320" pageId="12" pageNumber="319">
<bibRefCitation id="EFB14B70FFEBFF9DFF44F893FE48F840" author="Dworschak, P. C." box="[199,411,1915,1941]" pageId="12" pageNumber="319" pagination="19 - 32" refId="ref8009" refString="Dworschak, P. C. (2019) Axiidea (Micheleidae) and Gebiidea (Laomediidae and Upogebiidae) of Panglao, the Philippines. Annalen des Naturhistorischen Museums in Wien, Serie B, 121, 19 - 32." type="journal article" year="2019">Dworschak (2019)</bibRefCitation>
reported
<taxonomicName id="4C204D02FFEBFF9DFD89F893FD6DF840" authorityName="Ngoc-Ho" authorityYear="1994" box="[522,702,1915,1941]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="multispinosa">
<emphasis id="B954EA93FFEBFF9DFD89F893FD6DF840" box="[522,702,1915,1941]" italics="true" pageId="12" pageNumber="319">G. multispinosa</emphasis>
</taxonomicName>
from the
<collectingCountry id="F3377611FFEBFF9DFCB1F893FC63F840" box="[818,944,1915,1941]" name="Philippines" pageId="12" pageNumber="319">Philippines</collectingCountry>
with notes on variation in the number and shape of the rostral ventral spines, the number of the anterolateral and cervical spinules on the carapace and the armature of the article 5 of the antennal peduncle. Specimens illustrated by
<bibRefCitation id="EFB14B70FFEBFF9DFC5FF82BFB7EF808" author="Dworschak, P. C." box="[988,1197,1987,2013]" pageId="12" pageNumber="319" pagination="19 - 32" refId="ref8009" refString="Dworschak, P. C. (2019) Axiidea (Micheleidae) and Gebiidea (Laomediidae and Upogebiidae) of Panglao, the Philippines. Annalen des Naturhistorischen Museums in Wien, Serie B, 121, 19 - 32." type="journal article" year="2019">Dworschak (2019)</bibRefCitation>
do not agree with the original description and illustration by
<bibRefCitation id="EFB14B70FFEAFF9CFDC6FF7FFCD9FF64" author="Ngoc-Ho, N." box="[581,778,151,177]" pageId="13" pageNumber="320" pagination="193 - 216" refId="ref8328" refString="Ngoc-Ho, N. (1994 a) Notes on some Indo-Pacific Upogebiidae with descriptions of four new species (Crustacea: Thalassinidea). Memoirs of the Queensland Museum, 35, 193 - 216." type="journal article" year="1994">Ngoc-Ho (1994a)</bibRefCitation>
in the rounded posterolateral angle of the uropodal exopod. The above comparison between the new species and
<taxonomicName id="4C204D02FFEAFF9CFD74FF53FC79FF00" authorityName="Ngoc-Ho" authorityYear="1994" box="[759,938,187,213]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="320" phylum="Arthropoda" rank="species" species="multispinosa">
<emphasis id="B954EA93FFEAFF9CFD74FF53FC79FF00" box="[759,938,187,213]" italics="true" pageId="13" pageNumber="320">G. multispinosa</emphasis>
</taxonomicName>
relies only upon the original description of the latter species, considering the possibility that the Philippine specimens might be specifically distinct from
<taxonomicName id="4C204D02FFEAFF9CFA02FF37FEF5FEC8" authorityName="Ngoc-Ho" authorityYear="1994" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="320" phylum="Arthropoda" rank="species" species="multispinosa">
<emphasis id="B954EA93FFEAFF9CFA02FF37FEF5FEC8" italics="true" pageId="13" pageNumber="320">G. multispinosa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9F3681FFEAFF9CFF44FECFFC97FE5C" blockId="13.[151,1437,151,465]" pageId="13" pageNumber="320">
This study increases representatives of
<taxonomicName id="4C204D02FFEAFF9CFDFCFECFFCC3FE94" authorityName="Ngoc-Ho" authorityYear="1989" box="[639,784,295,321]" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B954EA93FFEAFF9CFDFCFECFFCC3FE94" box="[639,784,295,321]" italics="true" pageId="13" pageNumber="320">Gebiacantha</emphasis>
</taxonomicName>
known from Japanese waters to three.
<taxonomicName id="4C204D02FFEAFF9CFB45FECFFF38FEB1" authorityName="Komai" authorityYear="2017" class="Malacostraca" family="Upogebiidae" genus="Gebiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="320" phylum="Arthropoda" rank="species" species="sagamiensis">
<emphasis id="B954EA93FFEAFF9CFB45FECFFF38FEB1" italics="true" pageId="13" pageNumber="320">Gebiacantha sagamiensis</emphasis>
</taxonomicName>
is referred to the species group characterised by the oval-shaped uropodal exopod, and is notable in the genus for the absence of rostral ventral spines (
<bibRefCitation id="EFB14B70FFEAFF9CFD25FE87FCEBFE5C" author="Komai, T." box="[678,824,367,393]" pageId="13" pageNumber="320" pagination="578 - 586" refId="ref8135" refString="Komai, T. (2017) Gebiacantha sagamiensis, a new species of upogebiid shrimp (Crustacea: Decapoda: Gebiidea) from Sagami Bay, central Japan. Zootaxa, 4263 (3), 578 - 586. https: // doi. org / 10.11646 / zootaxa. 4263.3.9" type="journal article" year="2017">Komai 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A650AFFEAFF9CFF44FE7BFB72FE04" pageId="13" pageNumber="320" type="etymology">
<paragraph id="8B9F3681FFEAFF9CFF44FE7BFB72FE04" blockId="13.[151,1437,151,465]" pageId="13" pageNumber="320">
<emphasis id="B954EA93FFEAFF9CFF44FE7BFE9DFE78" bold="true" box="[199,334,403,429]" pageId="13" pageNumber="320">Etymology.</emphasis>
From the combination of the Latin,
<emphasis id="B954EA93FFEAFF9CFD60FE7BFCCDFE78" box="[739,798,403,429]" italics="true" pageId="13" pageNumber="320">fortis</emphasis>
(= strong) and
<emphasis id="B954EA93FFEAFF9CFC49FE7BFBF4FE79" box="[970,1063,403,428]" italics="true" pageId="13" pageNumber="320">spinatus</emphasis>
(spined), in reference to the relatively strong armature on the carapace and cheliped in the new species, used as an adjective.
</paragraph>
</subSubSection>
</treatment>
</document>
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