262 lines
18 KiB
XML
262 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.90.20586" ID-PMC="PMC5784240" ID-Pensoft-Pub="1314-2003-90-1" ID-Pensoft-UUID="80067D29FFFB7D34FF80E95D553F4254" ID-PubMed="29391851" ID-Zenodo-Dep="1138341" ModsDocID="1314-2003-90-1" checkinTime="1510830429569" checkinUser="pensoft" docAuthor="Berry, Paul E., Kainulainen, Kent & Ee, Benjamin W. van" docDate="2017" docId="0C760245DE4A501C8BCC0ABDE9DE446D" docLanguage="en" docName="PhytoKeys 90: 1-87" docOrigin="PhytoKeys 90" docPubDate="2017-11-15" docSource="http://dx.doi.org/10.3897/phytokeys.90.20586" docTitle="Croton submetallicus Baill., Bull. Mens. Soc. Linn. Paris 2: 966. 1891, as ‘ submetallicum’" docType="treatment" docVersion="3" id="80067D29FFFB7D34FF80E95D553F4254" lastPageNumber="58" masterDocId="80067D29FFFB7D34FF80E95D553F4254" masterDocTitle="A Nomenclator of Croton (Euphorbiaceae) in Madagascar, the Comoros Archipelago, and the Mascarene Islands" masterLastPageNumber="87" masterPageNumber="1" pageNumber="57" updateTime="1668142129654" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A Nomenclator of Croton (Euphorbiaceae) in Madagascar, the Comoros Archipelago, and the Mascarene Islands</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Berry, Paul E.</mods:namePart>
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<mods:affiliation>Herbarium, Department of Ecology and Evolutionary Biology, University of Michigan, 3600 Varsity Drive, Ann Arbor, Michigan 48108, U. S. A.</mods:affiliation>
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<mods:nameIdentifier type="email">peberry@umich.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kainulainen, Kent</mods:namePart>
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<mods:affiliation>Herbarium, Department of Ecology and Evolutionary Biology, University of Michigan, 3600 Varsity Drive, Ann Arbor, Michigan 48108, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ee, Benjamin W. van</mods:namePart>
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<mods:affiliation>Department of Biology, Universidad de Puerto Rico, Recinto Universitario de Mayagueez, Mayagueez, PR 00680, Puerto Rico, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2017-11-15</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>90</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>87</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.90.20586</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.90.20586</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-90-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">80067D29FFFB7D34FF80E95D553F4254</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1138341</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182225632" LSID="urn:lsid:plazi:treatment:0C760245DE4A501C8BCC0ABDE9DE446D" httpUri="http://treatment.plazi.org/id/0C760245DE4A501C8BCC0ABDE9DE446D" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
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<subSubSection pageId="56" pageNumber="57" type="nomenclature">
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<paragraph pageId="56" pageNumber="57">
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<pageBreakToken pageId="56" pageNumber="57" start="start">114</pageBreakToken>
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.
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<taxonomicName LSID="0C760245-DE4A-501C-8BCC-0ABDE9DE446D" authority="Baill., Bull. Mens. Soc. Linn. Paris 2: 966. 1891, as ' submetallicum'" authorityName="Baill., Bull. Mens. Soc. Linn. Paris 2: 966. 1891, as ‘ submetallicum’" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton submetallicus" order="Malpighiales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="species" species="submetallicus">
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Croton submetallicus Baill., Bull. Mens. Soc. Linn. Paris 2: 966. 1891, as
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<normalizedToken originalValue="‘submetallicum’">'submetallicum'</normalizedToken>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="56" pageNumber="57" type="reference_group">
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<paragraph pageId="56" pageNumber="57">
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<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton chapelieri subsp. var. var. longepetiolata" order="Malpighiales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="variety" species="chapelieri" subSpecies="var." variety="longepetiolata">Croton chapelieri var. longepetiolata</taxonomicName>
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Radcl.-Sm., Gen. Croton Madag. Comoro 152. 2016,
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<emphasis bold="true" pageId="56" pageNumber="57">syn. nov.</emphasis>
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Type. Madagascar. Prov. Toamasina: W of
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<taxonomicName family="Euphorbiaceae" genus="Vavatenina" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="" order="Malpighiales" pageId="56" pageNumber="57" rank="genus">Vavatenina</taxonomicName>
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,
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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Naturelle
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<normalizedToken originalValue="Intégrale">Integrale</normalizedToken>
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Zahamena, Amboditamenaka Forest,
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<geoCoordinate degrees="17" direction="south" minutes="44" orientation="latitude" precision="925" value="-17.733334">17°44'S</geoCoordinate>
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,
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<geoCoordinate degrees="49" direction="east" minutes="00" orientation="longitude" precision="925" value="49.0">49°00'E</geoCoordinate>
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, 15-20 Sep 1993,
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<emphasis italics="true" pageId="56" pageNumber="57">S. Malcomber B. Rasolondraibe, L.M. Randrianjanaka & J.P. Abraham 2511</emphasis>
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(holotype: K!; isotypes: MO [MO-2990285]!, P [P00433109]!).
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</paragraph>
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<paragraph pageId="56" pageNumber="57">
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<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton fianarantsoae subsp. var. var. obovalifolius" order="Malpighiales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="variety" species="fianarantsoae" subSpecies="var." variety="obovalifolius">Croton fianarantsoae var. obovalifolius</taxonomicName>
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Radcl.-Sm., Gen. Croton Madag. Comoro 155. 2016,
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<emphasis bold="true" pageId="56" pageNumber="57">syn. nov.</emphasis>
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Type. Madagascar. Prov. Fianarantsoa: W of Ranomafana,
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<geoCoordinate degrees="21" direction="south" minutes="16" orientation="latitude" precision="925" value="-21.266666">21°16'S</geoCoordinate>
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,
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<geoCoordinate degrees="47" direction="east" minutes="28" orientation="longitude" precision="925" value="47.466667">47°28'E</geoCoordinate>
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, 8-10 Sep 1992,
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<emphasis italics="true" pageId="56" pageNumber="57">S. Malcomber</emphasis>
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&
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<emphasis italics="true" pageId="56" pageNumber="57">R. Rakoto 1550</emphasis>
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(holotype: K; isotypes: MO, P [P00422460]!).
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</paragraph>
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<paragraph pageId="56" pageNumber="57">
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<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton nitidulus subsp. var. var. acuminatus" order="Malpighiales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="variety" species="nitidulus" subSpecies="var." variety="acuminatus">Croton nitidulus var. acuminatus</taxonomicName>
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Radcl.-Sm., Gen. Croton Madag. Comoro 169. 2016,
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<emphasis bold="true" pageId="56" pageNumber="57">syn. nov.</emphasis>
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Type. Madagascar. Prov. Toamasina:
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<normalizedToken originalValue="Itinéraire">Itineraire</normalizedToken>
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de Didy
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<normalizedToken originalValue="à">a</normalizedToken>
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Brickaville (
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<normalizedToken originalValue="forêt">foret</normalizedToken>
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orientale),
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<normalizedToken originalValue="reçu">recu</normalizedToken>
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Apr 1954,
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<emphasis italics="true" pageId="56" pageNumber="57">G. Cours 4871</emphasis>
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(lectotype, designated here: P [P00133641]!; isolectotypes: P [P00133639]!; P [P00133640]!; P [P00133642]!).
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</paragraph>
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<paragraph pageId="56" pageNumber="57">
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<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton nitidulus subsp. var. var. cinereus" order="Malpighiales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="variety" species="nitidulus" subSpecies="var." variety="cinereus">Croton nitidulus var. cinereus</taxonomicName>
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Radcl.-Sm., Gen. Croton Madag. Comoro 170. 2016,
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<emphasis bold="true" pageId="56" pageNumber="57">syn. nov.</emphasis>
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Type. Madagascar. Prov. Toamasina: Ambatovy,
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<geoCoordinate degrees="18" direction="south" minutes="51" orientation="latitude" precision="15" seconds="34" value="-18.859446">18°51'34"S</geoCoordinate>
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,
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<geoCoordinate degrees="48" direction="east" minutes="18" orientation="longitude" precision="15" seconds="25" value="48.306942">48°18'25"E</geoCoordinate>
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, 3 Mar 1997,
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<emphasis italics="true" pageId="56" pageNumber="57">P. Rakotomalaza et al. 1217</emphasis>
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(holotype: K!; isotype: MO!).
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</paragraph>
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<paragraph pageId="56" pageNumber="57">
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<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton nitidulus subsp. var. var. hypopoliotes" order="Malpighiales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="variety" species="nitidulus" subSpecies="var." variety="hypopoliotes">Croton nitidulus var. hypopoliotes</taxonomicName>
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Radcl.-Sm., Gen. Croton Madag. Comoro 172. 2016,
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<emphasis bold="true" pageId="56" pageNumber="57">syn. nov.</emphasis>
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Type. Madagascar. "Central Madagascar", received Dec 1883,
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<emphasis italics="true" pageId="56" pageNumber="57">R. Baron s.n</emphasis>
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. (holotype: K!).
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</paragraph>
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<paragraph pageId="56" pageNumber="57">
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<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton nitidulus subsp. var. var. macrophyllus" order="Malpighiales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="variety" species="nitidulus" subSpecies="var." variety="macrophyllus">Croton nitidulus var. macrophyllus</taxonomicName>
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Radcl.-Sm., Gen. Croton Madag. Comoro 172. 2016,
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<emphasis bold="true" pageId="56" pageNumber="57">syn. nov.</emphasis>
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Type. Madagascar. Prov. Toamasina:
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<normalizedToken originalValue="Itinéraire">Itineraire</normalizedToken>
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de Didy
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<normalizedToken originalValue="à">a</normalizedToken>
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Brickaville (
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<normalizedToken originalValue="forêt">foret</normalizedToken>
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orientale), s.d.,
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<emphasis italics="true" pageId="56" pageNumber="57">G. Cours 4660</emphasis>
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(lectoype, designated here: P [P00133616]!; isolectotypes: P [P00133615]!, P [P00133617]!).
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</paragraph>
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<paragraph pageId="56" pageNumber="57">
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<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton submetallicus subsp. var. var. tomentosus" order="Malpighiales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="variety" species="submetallicus" subSpecies="var." variety="tomentosus">Croton submetallicus var. tomentosus</taxonomicName>
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Radcl.-Sm., Gen. Croton Madag. Comoro 58. 2016,
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<emphasis bold="true" pageId="56" pageNumber="57">syn. nov.</emphasis>
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Type. Madagascar. Prov. Toamasina:
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<normalizedToken originalValue="forêt">foret</normalizedToken>
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de
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<normalizedToken originalValue="l’Analamazaotra">l'Analamazaotra</normalizedToken>
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,
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<normalizedToken originalValue="reçu">recu</normalizedToken>
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3 Dec 1934,
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<emphasis italics="true" pageId="56" pageNumber="57">E. Ursch 21</emphasis>
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(holotype: P [P00133354]!).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="56" pageNumber="57" type="materials_examined">
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<paragraph pageId="56" pageNumber="57">
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<typeStatus>Type</typeStatus>
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.
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</paragraph>
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<paragraph pageId="56" pageNumber="57">
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<materialsCitation collectingDate="2017-01-01" collectingDateMax="2017-12-31" collectingDateMin="2017-01-01" collectorName="R. Baron" country="Madagascar" location="Madagascar" specimenCount="1" typeStatus="lectotype">
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<collectingCountry name="Madagascar">Madagascar</collectingCountry>
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. sin. loc., s.d.,
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<emphasis italics="true" pageId="56" pageNumber="57">
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<collectorName>R. Baron</collectorName>
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5286
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</emphasis>
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(
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<typeStatus>lectotype</typeStatus>
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, designated here: K [K000422582]!; isolectotypes: K [K000422581]!, P [P00133327]!)
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</materialsCitation>
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.
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<collectingCountry name="Madagascar">Madagascar</collectingCountry>
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.
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<materialsCitation collectingDate="2015-08-15" collectorName="B. van Ee, P. E. Berry, H. Razafindraibe" country="US" county="Andasibe" elevation="900" latitude="-18.985556" location="Alaotra-Mangoro Region. Forest" longLatPrecision="21" longitude="48.431946" municipality="Rn" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
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Prov.
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<collectingRegion country="Madagascar" name="Toamasina">Toamasina</collectingRegion>
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:
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<location LSID="urn:lsid:plazi:treatment:0C760245DE4A501C8BCC0ABDE9DE446D:429DB8FD509E87EA3EB67AFBFE77A37C" country="US" county="Andasibe" latitude="-18.985556" longLatPrecision="21" longitude="48.431946" municipality="Rn" name="Alaotra-Mangoro Region. Forest" stateProvince="Toamasina">Alaotra-Mangoro Region. Forest</location>
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<locationDeviation location="Eulophiella Hotel">W of Eulophiella Hotel</locationDeviation>
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, south of
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<collectingMunicipality>Rn</collectingMunicipality>
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2 and
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<collectingCounty>Andasibe</collectingCounty>
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,
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<geoCoordinate degrees="18" direction="south" minutes="59" orientation="latitude" precision="15" seconds="08" value="-18.985556">18°59'08"S</geoCoordinate>
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,
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<geoCoordinate degrees="48" direction="east" minutes="25" orientation="longitude" precision="15" seconds="55" value="48.431946">48°25'55"E</geoCoordinate>
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,
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<quantity metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</elevation>
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</quantity>
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,
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<collectingDate value="2015-08-15">15 Aug 2015</collectingDate>
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,
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<emphasis italics="true" pageId="56" pageNumber="57">
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<collectorName>B. van Ee</collectorName>
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,
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<collectorName>P.E. Berry</collectorName>
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&
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<collectorName>H. Razafindraibe</collectorName>
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2222
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</emphasis>
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(epitype, designated here: MICH [MICH1514786]!; additional duplicates: G, K, MO, P, TAN,
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<collectingCountry name="United States of America">US</collectingCountry>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="57" pageNumber="58" type="distribution">
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<paragraph pageId="57" pageNumber="58">
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<pageBreakToken pageId="57" pageNumber="58" start="start">Habit</pageBreakToken>
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and distribution.
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</paragraph>
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<paragraph pageId="57" pageNumber="58">Shrubs or trees; eastern montane forests of Madagascar (Fianarantsoa, Toamasina).</paragraph>
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</subSubSection>
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<subSubSection pageId="57" pageNumber="58" type="notes">
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<paragraph pageId="57" pageNumber="58">Notes.</paragraph>
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<paragraph pageId="57" pageNumber="58">
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The type of
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<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton submetallicus" order="Malpighiales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="species" species="submetallicus">
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<emphasis italics="true" pageId="57" pageNumber="58">Croton submetallicus</emphasis>
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</taxonomicName>
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is deficient for characterizing this species properly; it lacks pistillate flowers, which are very diagnostic. Therefore we are designating an epitype that is well distributed, has photographs in Tropicos®, and is also sampled molecularly. The only other described species name that could be applied here is
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<taxonomicName lsidName="C. macrochlamys" pageId="57" pageNumber="58" rank="species" species="macrochlamys">
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<emphasis italics="true" pageId="57" pageNumber="58">C. macrochlamys</emphasis>
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</taxonomicName>
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, but it has a very poor type specimen, and we are treating it here as a synonym of
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<taxonomicName lsidName="C. nitidulus" pageId="57" pageNumber="58" rank="species" species="nitidulus">
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<emphasis italics="true" pageId="57" pageNumber="58">C. nitidulus</emphasis>
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</taxonomicName>
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because of its smaller leaf size.
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</paragraph>
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<paragraph pageId="57" pageNumber="58">
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For
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<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Croton nitidulus subsp. var. var. acuminatus" order="Malpighiales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="variety" species="nitidulus" subSpecies="var." variety="acuminatus">Croton nitidulus var. acuminatus</taxonomicName>
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,
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<bibRefCitation author="Radcliffe-Smith, A" journalOrPublisher="Royal Botanic Gardens, Kew" pageId="73" pageNumber="74" refId="B55" refString="Radcliffe-Smith, A, 2016. The Genus Croton in Madagascar & the Comoro Is. Received at Kew, NHM and RHS Wisley libraries as a pre-print on 23 Dec 2016 via Martin Cheek, 1-213." title="The Genus Croton in Madagascar & the Comoro Is. Received at Kew, NHM and RHS Wisley libraries as a pre-print on 23 Dec 2016 via Martin Cheek, 1 - 213." year="2016">Radcliffe-Smith (2016)</bibRefCitation>
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designated
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<emphasis italics="true" pageId="57" pageNumber="58">Cours 4871</emphasis>
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at P as the holotype, but there are four nearly identical specimens of this collection at P, so we selected one of them here as lectotype. Likewise, for
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<taxonomicName lsidName="C. nitidulus subsp. var. var. macrophyllus" pageId="57" pageNumber="58" rank="variety" species="nitidulus" subSpecies="var." variety="macrophyllus">C. nitidulus var. macrophyllus</taxonomicName>
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,
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<bibRefCitation author="Radcliffe-Smith, A" journalOrPublisher="Royal Botanic Gardens, Kew" pageId="73" pageNumber="74" refId="B55" refString="Radcliffe-Smith, A, 2016. The Genus Croton in Madagascar & the Comoro Is. Received at Kew, NHM and RHS Wisley libraries as a pre-print on 23 Dec 2016 via Martin Cheek, 1-213." title="The Genus Croton in Madagascar & the Comoro Is. Received at Kew, NHM and RHS Wisley libraries as a pre-print on 23 Dec 2016 via Martin Cheek, 1 - 213." year="2016">Radcliffe-Smith (2016)</bibRefCitation>
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designated
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<emphasis italics="true" pageId="57" pageNumber="58">Cours 4660</emphasis>
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at P as the holotype, but since there are three duplicates of that collection there, we selected one of them as the lectotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |