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<document ID-CLB-Dataset="21459" ID-DOI="http://dx.doi.org/10.3897/phytokeys.153.49388" ID-GBIF-Dataset="322737dd-e6d7-4f3f-8493-b466ffb6dcec" ID-PMC="PMC7381717" ID-Pensoft-Pub="1314-2003-153-1" ID-Pensoft-UUID="8BF3C31AA36950BE8BBC287988998FF6" ID-PubMed="32765177" ModsDocID="1314-2003-153-1" checkinTime="1594919040269" checkinUser="pensoft" docAuthor="Nguy ễn 1, Danh-Hung, Lương 2, Văn-Dũng, Le 3, Thi-Hương, Tr ần 4, Qu ốc-Thanh, Đ ỗ 1,5, Ng ọc-Đai &amp; Ly 1,6, Ng ọc-Sam" docDate="2020" docId="44ED8C82E5AF56829C357A3549E0CA24" docLanguage="en" docName="PhytoKeys 153: 1-11" docOrigin="PhytoKeys 153" docSource="http://dx.doi.org/10.3897/phytokeys.153.49388" docTitle="Camellia puhoatensis N. S. Ly, V. D. Luong, T. H. Le, D. H. Nguyen &amp; N. D. Do 2020, sp. nov." docType="treatment" docVersion="6" id="8BF3C31AA36950BE8BBC287988998FF6" lastPageNumber="1" masterDocId="8BF3C31AA36950BE8BBC287988998FF6" masterDocTitle="Camellia puhoatensis (Sect. Archecamellia - Theaceae), a new species from Vietnam" masterLastPageNumber="11" masterPageNumber="1" pageNumber="1" updateTime="1732871053670" updateUser="ExternalLinkService">
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<mods:title id="E08CD4660A2857DF5A9F06F89AAAE415">Camellia puhoatensis (Sect. Archecamellia - Theaceae), a new species from Vietnam</mods:title>
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<mods:namePart id="3F9D8EDCC2150AD516B8460F8040DA3A">Nguy ễn 1, Danh-Hung</mods:namePart>
<mods:affiliation id="80E94CC1C1DBB13BC36C9E77CEFF4595">Graduate University of Science and Technology, VAST, 18 Hoang Quoc Viet, Cau Giay District, Ha Noi, Vietnam</mods:affiliation>
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<mods:namePart id="2983F7D324B9F690BA6972535F1D907B">Lương 2, Văn-Dũng</mods:namePart>
<mods:affiliation id="BA78EEA4A4E3DE4B3D5FFF1D36FA6B2E">Graduate University of Science and Technology, VAST, 18 Hoang Quoc Viet, Cau Giay District, Ha Noi, Vietnam</mods:affiliation>
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<mods:namePart id="BA4C9D141E832A1189C63A7623E8D311">Le 3, Thi-Hương</mods:namePart>
<mods:affiliation id="311289A6AA2D7A2F98FEEC2D68F8B761">Graduate University of Science and Technology, VAST, 18 Hoang Quoc Viet, Cau Giay District, Ha Noi, Vietnam</mods:affiliation>
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<mods:namePart id="033D44E683318043D980DC4876EEBAF6">Tr ần 4, Qu ốc-Thanh</mods:namePart>
<mods:affiliation id="280C722F0E0C43A7A36E85F0838ED308">Graduate University of Science and Technology, VAST, 18 Hoang Quoc Viet, Cau Giay District, Ha Noi, Vietnam</mods:affiliation>
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<mods:namePart id="485223F5EE6F27C27D04E192CB99F6D9">Đ ỗ 1,5, Ng ọc-Đai</mods:namePart>
<mods:affiliation id="2965EA0844A39E90BDA59CECF091D085">Graduate University of Science and Technology, VAST, 18 Hoang Quoc Viet, Cau Giay District, Ha Noi, Vietnam</mods:affiliation>
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<mods:namePart id="42D51ABF867E56045F5BEB258DA27E17">Ly 1,6, Ng ọc-Sam</mods:namePart>
<mods:affiliation id="677B79D50A33D5EFC5DAC53EEAAB9B26">Graduate University of Science and Technology, VAST, 18 Hoang Quoc Viet, Cau Giay District, Ha Noi, Vietnam</mods:affiliation>
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<mods:date id="4CC54290492A922D80B8B9915A869AEA">2020</mods:date>
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<mods:classification id="BDC042A02E52166343AEA9586D8DEBA5">journal article</mods:classification>
<mods:identifier id="87303ABA7B567ED642463B40280B186B" type="DOI">http://dx.doi.org/10.3897/phytokeys.153.49388</mods:identifier>
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<treatment id="44ED8C82E5AF56829C357A3549E0CA24" ID-GBIF-Taxon="165119416" LSID="urn:lsid:plazi:treatment:44ED8C82E5AF56829C357A3549E0CA24" httpUri="http://treatment.plazi.org/id/44ED8C82E5AF56829C357A3549E0CA24" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Theaceae">
<subSubSection id="221F006EFB896199A4F4AB53819BFD3D" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="90F6FD05A64B7A7399F6CD93FD2B1E0E" pageId="0" pageNumber="1">
<taxonomicName id="4F67B0EC1098AB86485829FD9DB77F0E" ID-CoL="7T89Y" LSID="44ED8C82-E5AF-5682-9C35-7A3549E0CA24" authority="N. S. Lý, V. D. Luong, T. H. Le, D. H. Nguyen &amp; N. D. Do" authorityName="N. S. Ly, V. D. Luong, T. H. Le, D. H. Nguyen &amp; N. D. Do" authorityYear="2020" class="Magnoliopsida" family="Theaceae" genus="Camellia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Camellia puhoatensis" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="puhoatensis" status="sp. nov.">
Camellia puhoatensis N.S.
<normalizedToken id="4B242E23F20D3E6C1E34D7D0E07263C8" originalValue="Lý">Ly</normalizedToken>
, V.D. Luong, T.H. Le, D.H. Nguyen &amp; N.D. Do
</taxonomicName>
<taxonomicNameLabel id="179DCF42B1406C73116469F750AECBF0" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="C8C5B102FAF13BC3E850A9ADD6B05C4F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Camellia puhoatensis. A Leaf, adaxial view B venation detail of leaf (abaxial surface) C flower (lateral view) D flower (top view) E bracteoles (inner surfaces shown) F sepals (adaxial surfaces) G petals (adaxial surfaces) H androecium (one part) I stamen J gynoecium (with sepals and petals). Drawn from the holotype by Van-Dung Luong." figureDoi="10.3897/phytokeys.153.49388.figure1" httpUri="https://binary.pensoft.net/fig/432011" pageId="0" pageNumber="1">Figures 1</figureCitation>
<figureCitation id="FA7F9C3074AF4C8B2C0F6684C55BACAF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Camellia puhoatensis. A young shoot B terminal buds C solitary bud and axillary flower (side view) D flower and pollinated flower (side view) E close-up of flower (front view) F immature fruit G a part of branch showing leaves abaxial and opening flower H leaves adaxially I bracteoles J sepals K petals L androecium with stamens M gynoecium (with sepals and styles). Photos by Ngoc-Dai Do, the colour plate prepared by Ngoc-Sam Ly." figureDoi="10.3897/phytokeys.153.49388.figure2" httpUri="https://binary.pensoft.net/fig/432012" pageId="0" pageNumber="1">, 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="EE5B795F327CA72456D469B26FBEA900" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="255026F549D5BD14FA0FD4A3FDF90415" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="7C6A24098318F51A2E7058FE7F9BCAD5" pageId="0" pageNumber="1">
<taxonomicName id="B5D3937A462CE89AE26D49BDD0E9C5BA" authorityName="N. S. Ly, V. D. Luong, T. H. Le, D. H. Nguyen &amp; N. D. Do" authorityYear="2020" class="Magnoliopsida" family="Theaceae" genus="Camellia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Camellia puhoatensis" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="puhoatensis">
<emphasis id="F631F0F841E056E135DA15931351EC50" italics="true" pageId="0" pageNumber="1">Camellia puhoatensis</emphasis>
</taxonomicName>
is morphologically similar to
<taxonomicName id="B872725656362B9704F85D2B1383BA7D" lsidName="C. chrysanthoides" pageId="0" pageNumber="1" rank="species" species="chrysanthoides">
<emphasis id="01F0989990EC646358A66061A6D866A6" italics="true" pageId="0" pageNumber="1">C. chrysanthoides</emphasis>
</taxonomicName>
,
<taxonomicName id="39DA3031CA2BE9C5D604CB55EB1DF8AA" lsidName="C. flavida" pageId="0" pageNumber="1" rank="species" species="flavida">
<emphasis id="F156DF753BDD1A26FE24AD1FF1C2DE40" italics="true" pageId="0" pageNumber="1">C. flavida</emphasis>
</taxonomicName>
and
<taxonomicName id="26B70CE73726D6A4234F71B445AA5356" lsidName="C. petelotii" pageId="0" pageNumber="1" rank="species" species="petelotii">
<emphasis id="12E02D51C6ABB2782E0B3877CF5E3B19" italics="true" pageId="0" pageNumber="1">C. petelotii</emphasis>
</taxonomicName>
, but differs in having young puberulous shoots, mature leaves sparsely puberulous abaxially with leaf bases rounded or broadly obtuse, petioles and pedicels puberulous, tepals 12-13 and the ovary and styles pubescent.
</paragraph>
<paragraph id="BBEE6E2EE01282469832F203006CBCA7" pageId="0" pageNumber="1">
Type. VIETNAM. Nghe An Province: Que Phong District, Dong Van Commune, Pu Hoat NR,
<geoCoordinate id="ACF1F611BDE422D4C4B95D5CFFE00825" degrees="19" direction="north" minutes="43" orientation="latitude" precision="15" seconds="31" value="19.725279">19°43'31&quot;N</geoCoordinate>
,
<geoCoordinate id="D9EEAF56ACB0D38D8FBA274467A7B1A5" degrees="105" direction="east" minutes="05" orientation="longitude" precision="15" seconds="43" value="105.09528">105°05'43&quot;E</geoCoordinate>
, 270 m elev., 30 December 2018,
<emphasis id="A0FB8FC914D57EE63876320FE5E87AF0" italics="true" pageId="0" pageNumber="1">Do Ngoc Dai, Le Thi Huong, Nguyen Danh Hung, DHH-682</emphasis>
(holotype VNM; isotypes P, DLU).
</paragraph>
<caption id="FDC4F026B3647B8A57BA1DCC880CF8DF" doi="10.3897/phytokeys.153.49388.figure1" httpUri="https://binary.pensoft.net/fig/432011" pageId="0" pageNumber="1" start="Figure 1" startId="F1">
<paragraph id="301B6C0B84EDACD70BBAD1219E4F4414" pageId="0" pageNumber="1">
<emphasis id="493370225BCEFCB75BF6FAE6117EF7E3" bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
<taxonomicName id="E0DF33E344E08FF9F6CBEC979A6C1FD4" authorityName="N. S. Ly, V. D. Luong, T. H. Le, D. H. Nguyen &amp; N. D. Do" authorityYear="2020" class="Magnoliopsida" family="Theaceae" genus="Camellia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Camellia puhoatensis" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="puhoatensis">
<emphasis id="70DAFC1273F8D301B522EBDC0B656951" italics="true" pageId="0" pageNumber="1">Camellia puhoatensis</emphasis>
</taxonomicName>
.
<emphasis id="270D26404F67625F2BA580C350226A66" bold="true" pageId="0" pageNumber="1">A</emphasis>
Leaf, adaxial view
<emphasis id="F9E32A4A589109E5F06E1563ABE74C9D" bold="true" pageId="0" pageNumber="1">B</emphasis>
venation detail of leaf (abaxial surface)
<emphasis id="9955405145E1BD04FD99AE2548163EEB" bold="true" pageId="0" pageNumber="1">C</emphasis>
flower (lateral view)
<emphasis id="E0BD5866DDCB5D11A07E5D7F46A4A944" bold="true" pageId="0" pageNumber="1">D</emphasis>
flower (top view)
<emphasis id="628AA30BB6702D75CA12AE1FD365D884" bold="true" pageId="0" pageNumber="1">E</emphasis>
bracteoles (inner surfaces shown)
<emphasis id="A85BBFA4E4C7B7552778CD120E4888E7" bold="true" pageId="0" pageNumber="1">F</emphasis>
sepals (adaxial surfaces)
<emphasis id="5C1FC7D118F9661B2CC4B42965295290" bold="true" pageId="0" pageNumber="1">G</emphasis>
petals (adaxial surfaces)
<emphasis id="272B4D8B51D0572D6D21ADF812356F42" bold="true" pageId="0" pageNumber="1">H</emphasis>
androecium (one part)
<emphasis id="F2948320F933FA921D9E824060066788" bold="true" pageId="0" pageNumber="1">I</emphasis>
stamen
<emphasis id="9A50CD65AC196D642A69F6B188E7F4DC" bold="true" pageId="0" pageNumber="1">J</emphasis>
gynoecium (with sepals and petals). Drawn from the holotype by Van-Dung Luong.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="12D3F1B5287BEA3DD83B15448303CD93" pageId="0" pageNumber="1" type="description">
<paragraph id="773BEAA6DC114704862AD6A6CF601E9E" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="743C2F7856F63ACFC63A646A6DDC2E5C" pageId="0" pageNumber="1">
Shrub to tree, 2-7 m tall;
<emphasis id="0A5FE3AC758411B7AD5424046CA6031C" italics="true" pageId="0" pageNumber="1">young shoots</emphasis>
puberulous then glabrous when aging, purple towards terminals; semi-mature branches brown, smooth, glabrous, leaf scars prominent;
<emphasis id="B048F54D6CEF87CDED7C20BF25DBEF0B" italics="true" pageId="0" pageNumber="1">adult branches</emphasis>
and trunk light grey, smooth with lighter-coloured patches and covered by lichens;
<emphasis id="740FA9D40680BFA10778382D223C9291" italics="true" pageId="0" pageNumber="1">axillary leaf buds</emphasis>
rudimentary, roughly triangular, flat, with rounded apex, pubescent, brown, bud scales small but prominent, 1-3 mm long.
<emphasis id="D8E07AAF3D79ABF8A33CA1C8C4353D83" italics="true" pageId="0" pageNumber="1">Leaves</emphasis>
: juvenile leaves forming a narrow tube when young, soft, somewhat pendulous, purple in colour; young leaves slightly serrate, shiny, purple, adaxially glabrous, abaxially puberulous; developing leaves descending, narrow, shiny, purple to green-purple tinted, abaxial surface puberulous; mature leaves serrate, irregularly towards the apex, 17-23
<normalizedToken id="364475215A2B376EB2E1110096C4F1EC" originalValue="×">x</normalizedToken>
5.0-6.5 cm; petiole 8-16
<normalizedToken id="BA9D643308771719280F4BB7837D788D" originalValue="×">x</normalizedToken>
4-5 mm, puberulous; lamina thin, coriaceous, oblong ovate or oblong, leaf apex acuminate or narrowly acuminate, base rounded or broadly obtuse, adaxially dark green and glabrous, abaxially pale green and sparsely puberulous; primary vein continues as a shallow channel on the adaxial side of the petiole, 2.0-2.5 mm wide proximally, less than 1.0 mm distally, proximally light green and shiny on both sides; secondary venation pinnate, indistinctly brochidodromous, partially eucamptodromous on some leaves, with 10-13 pairs; midribs and lateral veins sunken adaxially; veins distinct proximally, less so towards the apex and the margins; tertiary venation very indistinct, sometimes lacking, more prominent at the leaf margins.
<emphasis id="D47D6BDC23F6CFA7E8BC07BE529F968F" italics="true" pageId="0" pageNumber="1">Flowers</emphasis>
usually solitary, sometimes together in groups of 2 flowers borne on a short bracteate shoot, terminal, rarely axillary, lacking scent, 4.5-6.0 cm in diameter;
<emphasis id="7E46381C6B5DE9B785196C2AB6F017CA" italics="true" pageId="0" pageNumber="1">pedicel</emphasis>
stout, covered by purplish-red perulae, 7-10 mm long, puberulous; flower buds unevenly globose in shape, 2.2-2.6
<normalizedToken id="5E5F022E6DD060FD66986075743D34EA" originalValue="×">x</normalizedToken>
2.0-2.3 cm, yellowish-red tinted, open flowers somewhat circular;
<emphasis id="B7E0C7A32D2DFF36E22E832246D13E60" italics="true" pageId="0" pageNumber="1">bracteoles</emphasis>
(sensu
<bibRefCitation id="55B2676199433770F8FAFF6D3D4442A5" author="Sealy, JR" journalOrPublisher="Adansonia" pageId="0" pageNumber="1" refId="B37" refString="Sealy, JR, 1958. A revision of the genus Camellia. Royal Horticultural Society London, 1-239." title="A revision of the genus Camellia. Royal Horticultural Society London, 1 - 239." year="1958">Sealy 1958</bibRefCitation>
) 3-4, opposite, orbicular, 1.5-2.5
<normalizedToken id="D9B8640971234736F950C44415CE5814" originalValue="×">x</normalizedToken>
1.5-3.0 mm, abaxially red to yellow-red tinted, adaxially paler, glabrous, margins ciliate, persistent;
<emphasis id="13BA2036FEE62A6C117B8CC11BD7CD25" italics="true" pageId="0" pageNumber="1">sepals</emphasis>
5, persistent, orbicular or subglobose, 0.6-1.5
<normalizedToken id="37292306117C1E2CF5F05F0834F8B35D" originalValue="×">x</normalizedToken>
0.8-1.8 cm, abaxially dull red and pubescent, adaxially pale yellowish and glabrous, margins ciliate;
<emphasis id="F1875EAF3E2394D6B4D9D7D60886646D" italics="true" pageId="0" pageNumber="1">petals</emphasis>
12-13, arranged in 3 whorls, bright yellow, sometimes with large red patch on the outer ones; outermost whorl comprising 3 or 4 petals, orbicular to broadly obovate, 2.2-2.8
<normalizedToken id="90334FA2C7B2DBFD61A91FE28751E7C2" originalValue="×">x</normalizedToken>
1.6-2.3 cm, abaxially pubescent, adaxially glabrous; middle whorl comprising 4 or 5 petals, broadly obovate, 2.4-3.3
<normalizedToken id="3F641329BD7B9A0F386085225CFBCB65" originalValue="×">x</normalizedToken>
1.8-2.5 cm, abaxially pubescent, adaxially glabrous; innermost whorl of 3 or 4 petals, orbicular to broadly obovate, 2.3-2.5
<normalizedToken id="BF02D30C17F6B25E4006C26D08784F4F" originalValue="×">x</normalizedToken>
1.7-2.2 cm, abaxially pubescent, adaxially glabrous, basally united with outermost filaments 5-7 mm.
<emphasis id="6645D52F0F58DFCA111111C0614D3E9F" italics="true" pageId="0" pageNumber="1">Androecium</emphasis>
numerous stamens, in 4-5 whorls, light yellow, 2.5-2.8 cm long, glabrous; outer filaments basally united for 1.5-1.8 cm forming a cup, inner ones basally united for 3-5 mm, free above union;
<emphasis id="3C352386B6F98E250F73D14A2A95B915" italics="true" pageId="0" pageNumber="1">anthers</emphasis>
yellow, 2.2-2.8
<normalizedToken id="51DF3241840EDD7DDBB30D2F7E0D6F1C" originalValue="×">x</normalizedToken>
1-1.5 mm, with two longitudinal striations, dorsifixed.
<emphasis id="C9025871ABE16C45699D1252B1EFF7AB" italics="true" pageId="0" pageNumber="1">Gynoecium</emphasis>
superior, 3-(4)-loculed, ovoid terminating into 3-(4) styles, 2.5-3.0
<normalizedToken id="0209BE68EB2CA9732459DC76A86D9B41" originalValue="×">x</normalizedToken>
3.0-3.5 mm, slightly longitudinal striations, pubescent, 2 ovules per locule;
<emphasis id="A6BAD27B6EC152601AD95852BACE1462" italics="true" pageId="0" pageNumber="1">styles</emphasis>
free to the base, 1.8-2.3 cm long, pubescent.
<emphasis id="076F5F34518364E0A5F0A6A4CF32D77F" italics="true" pageId="0" pageNumber="1">Capsule</emphasis>
not seen.
</paragraph>
<caption id="0ADEAB5B5920F2A56A4A4D5508D83E8B" doi="10.3897/phytokeys.153.49388.figure2" httpUri="https://binary.pensoft.net/fig/432012" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph id="3BC5B8C835AE0172C9EB1628DA7E275D" pageId="0" pageNumber="1">
<emphasis id="08833CB620B01750A1AF04E101DE3DF3" bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
<taxonomicName id="8C10F1DD8A34190BB79218A3BE90E9CA" authorityName="N. S. Ly, V. D. Luong, T. H. Le, D. H. Nguyen &amp; N. D. Do" authorityYear="2020" class="Magnoliopsida" family="Theaceae" genus="Camellia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Camellia puhoatensis" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="puhoatensis">
<emphasis id="2C702A307082662330031088F98BE01A" italics="true" pageId="0" pageNumber="1">Camellia puhoatensis</emphasis>
</taxonomicName>
.
<emphasis id="5C78837BA590D70FFB4C32FE281C9AB8" bold="true" pageId="0" pageNumber="1">A</emphasis>
young shoot
<emphasis id="80A2F4E5937065E91D211525B30C09F0" bold="true" pageId="0" pageNumber="1">B</emphasis>
terminal buds
<emphasis id="D81754576F1176B407A22DBDCD9090EF" bold="true" pageId="0" pageNumber="1">C</emphasis>
solitary bud and axillary flower (side view)
<emphasis id="054C6C2DD0203DA3625A26B04CC8DC94" bold="true" pageId="0" pageNumber="1">D</emphasis>
flower and pollinated flower (side view)
<emphasis id="CB1E275F3B5F2B9BA8FAF426D5947A1C" bold="true" pageId="0" pageNumber="1">E</emphasis>
close-up of flower (front view)
<emphasis id="056212C1BFAA1233778264DDE752A33C" bold="true" pageId="0" pageNumber="1">F</emphasis>
immature fruit
<emphasis id="286F67C40CAA52BC8104DA13F40C648A" bold="true" pageId="0" pageNumber="1">G</emphasis>
a part of branch showing leaves abaxial and opening flower
<emphasis id="8F767A4105C85C8CF2E8C04FDFC03ADF" bold="true" pageId="0" pageNumber="1">H</emphasis>
leaves adaxially
<emphasis id="579A3C32D42A59E3A0CC8F6A04C153CD" bold="true" pageId="0" pageNumber="1">I</emphasis>
bracteoles
<emphasis id="84104B6D42E1AE27F76CADBCE7565478" bold="true" pageId="0" pageNumber="1">J</emphasis>
sepals
<emphasis id="ADD3A6C8C07DD3505F248F301715E946" bold="true" pageId="0" pageNumber="1">K</emphasis>
petals
<emphasis id="8F94B928B9AF37BE6901CEB158A0684B" bold="true" pageId="0" pageNumber="1">L</emphasis>
androecium with stamens
<emphasis id="550DE27220C9F8D3DB72CD3C16C17F63" bold="true" pageId="0" pageNumber="1">M</emphasis>
gynoecium (with sepals and styles). Photos by Ngoc-Dai Do, the colour plate prepared by Ngoc-Sam Ly.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="B2D2075BDE1DBA047608976E460F3E57" pageId="0" pageNumber="1" type="phenology">
<paragraph id="723FFA8C95A463520DDF6F7B5A93CC8C" pageId="0" pageNumber="1">Phenology.</paragraph>
<paragraph id="4B38C2A0A671454276945ECFAB1BC104" pageId="0" pageNumber="1">Flowering from November to January of the next year.</paragraph>
</subSubSection>
<subSubSection id="9BC1D9DE88BFFBCF85EE9FC8A23362CB" pageId="0" pageNumber="1" type="distribution">
<paragraph id="CBAA85CBB7BC6CF76293855368EA0F00" pageId="0" pageNumber="1">Distribution and habitat.</paragraph>
<paragraph id="F0DA4DEA24526AD43875C3574BE8A121" pageId="0" pageNumber="1">
<taxonomicName id="636ABB9143518B5FFDF9A4B4CB69EB96" authorityName="N. S. Ly, V. D. Luong, T. H. Le, D. H. Nguyen &amp; N. D. Do" authorityYear="2020" class="Magnoliopsida" family="Theaceae" genus="Camellia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Camellia puhoatensis" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="puhoatensis">
<emphasis id="96F86A43C688D4615FD5A030F775014F" italics="true" pageId="0" pageNumber="1">Camellia puhoatensis</emphasis>
</taxonomicName>
is currently found only from the type locality. It grows on moist fertile and sandy soils along mountain streams and hill slopes in evergreen broad-leaved forests in Pu Hoat Nature Reserve, Vietnam, at elevations of 270-450 m.
</paragraph>
</subSubSection>
<subSubSection id="1B6CEB64B627600D4E5ED669F773148F" pageId="0" pageNumber="1" type="provisional conservation assessment">
<paragraph id="6668491A61947AD9A7620BDA9D9D7542" pageId="0" pageNumber="1">Provisional conservation assessment.</paragraph>
<paragraph id="631F826743598BE19199249E58B780C1" pageId="0" pageNumber="1">
At present, only a single population of about 300 scattered mature individuals of
<taxonomicName id="43370BE566065E51DD528F94CC4B3157" authorityName="N. S. Ly, V. D. Luong, T. H. Le, D. H. Nguyen &amp; N. D. Do" authorityYear="2020" class="Magnoliopsida" family="Theaceae" genus="Camellia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Camellia puhoatensis" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="puhoatensis">
<emphasis id="6097B7DFCAE65A7B197CA3907238CFE6" italics="true" pageId="0" pageNumber="1">Camellia puhoatensis</emphasis>
</taxonomicName>
was observed in the type locality, with an estimated extent of occurrence (EOO) less than 100 km2 and an area of occupancy (AOO) less than 1 km2. The population is highly threatened due to loss of habitat within its range and high market demands for wild, yellow-flowered camellias which are intensively collected for sale by local people. Therefore,
<taxonomicName id="51FFB4129F370CB35E7F9FCBAD9CC26E" lsidName="C. puhoatensis" pageId="0" pageNumber="1" rank="species" species="puhoatensis">
<emphasis id="0FA96EF8D7C2E6250055BDA6A49F098F" italics="true" pageId="0" pageNumber="1">C. puhoatensis</emphasis>
</taxonomicName>
is preliminarily categorised as Critically Endangered [B1ab (i, ii, iii) + 2ab (ii, iii), D], according to the IUCN Categories and Criteria (
<bibRefCitation id="7F4FF8E680372FA8F2A16831561C1410" author="IUCN" journalOrPublisher="International Camellia Journal" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org/resources/redlistguidelines" refId="B11" refString="IUCN, 2017. Guidelines for Using the IUCN Red List Categories and Criteria. Version 13. Prepared by the Standards and Petitions Committee. https://www.iucnredlist.org/resources/redlistguidelines" title="Guidelines for Using the IUCN Red List Categories and Criteria. Version 13. Prepared by the Standards and Petitions Committee." url="https://www.iucnredlist.org/resources/redlistguidelines" year="2017">IUCN 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="24797B7DD7F5B9A3587E61A3F95A9CDB" pageId="0" pageNumber="1" type="etymology">
<paragraph id="A3C1B04EB01B9F094A4EF74B6EF1EB60" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="715990B1FFEC3687E8C2F67D33987CAC" pageId="0" pageNumber="1">
The specify epithet '
<taxonomicName id="1963AAD5D97B0AE4BDD8BBA324C202EF" lsidName="puhoatensis" pageId="0" pageNumber="1" rank="species" species="puhoatensis">
<emphasis id="91E0C1EF0973707201D3ECE279E79DDA" italics="true" pageId="0" pageNumber="1">puhoatensis</emphasis>
</taxonomicName>
' refers to the type locality.
</paragraph>
</subSubSection>
<subSubSection id="50554848D5E051476EF3A91BE9346A42" pageId="0" pageNumber="1" type="uses">
<paragraph id="3A68EEA22E2BDD1C0B10BFC3EFDC9C76" pageId="0" pageNumber="1">Uses.</paragraph>
<paragraph id="A38C32C3B6D1C0F55D286AC00E4DF44F" pageId="0" pageNumber="1">Leaves and flowers were harvested and used for tea by the local people.</paragraph>
</subSubSection>
<subSubSection id="627BCDB25C5A134F32D6BED503AA8607" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="74D2AB69F99EE5DD130CBF5D0148ED86" pageId="0" pageNumber="1">Additional specimens examined.</paragraph>
<paragraph id="216BCE744626AA93A60D416D53323A50" pageId="0" pageNumber="1">
<emphasis id="04D117C194A0777D0A3811D930A14B00" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes.</emphasis>
Vietnam. Nghe An Province: Que Phong District, Dong Van Commune,
<normalizedToken id="5DE75CCCB1EA18671222235E4030644C" originalValue="Pù">Pu</normalizedToken>
Hoạt NR,
<geoCoordinate id="A93975CD3AFB3BCDD0DC5938643EB4D7" degrees="19" direction="north" minutes="48" orientation="latitude" precision="15" seconds="45" value="19.8125">19°48'45&quot;N</geoCoordinate>
,
<geoCoordinate id="0328E9D1AAB30C6F3E21D3CD82815097" degrees="105" direction="east" minutes="5" orientation="longitude" precision="15" seconds="39" value="105.09417">105°5'39&quot;E</geoCoordinate>
, 320 m elev., 2 September 2018,
<emphasis id="D2A6A3559ED1F0402CF626450A2A7182" italics="true" pageId="0" pageNumber="1">
<normalizedToken id="470DC7AE55A10C6F7C179EBDC5D554AD" originalValue="Đ">Đ</normalizedToken>
ỗ Ngọc
<normalizedToken id="059AA89F3EE7F3E8CEBEF9A5C2103471" originalValue="Đài">Đai</normalizedToken>
, Nguyễn Danh
<normalizedToken id="BE5B2BD9352394DB738491DA9A311456" originalValue="Hùng">Hung</normalizedToken>
,
<normalizedToken id="7572F85E54272ADD2A7E3C61E85BB10B" originalValue="Lê">Le</normalizedToken>
Thị
<normalizedToken id="617B4A479E68A0A101CC833D35318520" originalValue="Hương">Hương</normalizedToken>
</emphasis>
,
<emphasis id="B223D2F9C205E2B1002A8E2B3685256A" italics="true" pageId="0" pageNumber="1">DHH 120</emphasis>
(VNM); the same locality,
<geoCoordinate id="5E5330877AD95AC59526AFBA5A1F0CFA" degrees="19" direction="north" minutes="48" orientation="latitude" precision="15" seconds="31" value="19.80861">19°48'31&quot;N</geoCoordinate>
,
<geoCoordinate id="B04AD7AF31F76D6B31C9B6B53A5564C2" degrees="105" direction="east" minutes="05" orientation="longitude" precision="15" seconds="43" value="105.09528">105°05'43&quot;E</geoCoordinate>
, 280 m elev., 16 January 2019,
<emphasis id="781FC6D87A966875F545BE10D3EDB803" italics="true" pageId="0" pageNumber="1">
<normalizedToken id="879C636C1DF9C61D64D241E9914B3AF1" originalValue="Đ">Đ</normalizedToken>
ỗ Ngọc
<normalizedToken id="B2DA6481A55DF3F0F7A27A4DDE910FD8" originalValue="Đài">Đai</normalizedToken>
, Nguyễn Danh
<normalizedToken id="1F240A02917937E85D37F92DAE0DD93B" originalValue="Hùng">Hung</normalizedToken>
,
<normalizedToken id="FED5198B0641F9E57969863BF2C42335" originalValue="Lê">Le</normalizedToken>
Thị
<normalizedToken id="57C38570FFE9036E39B6E95B6EAA4FEA" originalValue="Hương">Hương</normalizedToken>
</emphasis>
,
<emphasis id="F3D33E5284D4E7375D4A02B616C1D39F" italics="true" pageId="0" pageNumber="1">DHH 790</emphasis>
(DLU),
<emphasis id="E0872817A26209C4FC9510D36FCE1D6D" italics="true" pageId="0" pageNumber="1">DHH</emphasis>
791 (HN).
</paragraph>
</subSubSection>
<subSubSection id="0F1B3DF4BE6D19CE4209CAD6805390CF" pageId="0" pageNumber="1" type="vernacular name">
<paragraph id="2C7A3F39BB47C8B4819D865311A7E9F7" pageId="0" pageNumber="1">Vernacular name.</paragraph>
<paragraph id="046AF1257EA0065D11C27C6A2864FE78" pageId="0" pageNumber="1">
Vietnamese language:
<normalizedToken id="92FB6DB33AB9F7B1724603C9AAC2B5C8" originalValue="Trà">Tra</normalizedToken>
hoa
<normalizedToken id="80D12EB563C594CFE6C899117FF97A4E" originalValue="vàng">vang</normalizedToken>
<normalizedToken id="80EC9F303D2A8AA82910292BB2574E0F" originalValue="pù">pu</normalizedToken>
hoạt.
</paragraph>
</subSubSection>
<subSubSection id="BB18F02AE82FFE4C93AA7D41A756CC60" pageId="0" pageNumber="1" type="taxonomic notes">
<paragraph id="D045F5EC528AF36A2795AE200D2DD4EC" pageId="0" pageNumber="1">Taxonomic notes.</paragraph>
<paragraph id="5D6E18FEA65596189090CEDA84112F28" pageId="0" pageNumber="1">
The current infrageneric classification of
<taxonomicName id="C20A597DA0CD90AE3D528CB7D9ED1C22" class="Magnoliopsida" family="Theaceae" genus="Camellia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Camellia" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="266254D738037B939532A0856EA03DEE" italics="true" pageId="0" pageNumber="1">Camellia</emphasis>
</taxonomicName>
is derived from three previous publications (
<bibRefCitation id="292D208EB75472258DA30DABD6DF572C" author="Sealy, JR" journalOrPublisher="Adansonia" pageId="0" pageNumber="1" refId="B37" refString="Sealy, JR, 1958. A revision of the genus Camellia. Royal Horticultural Society London, 1-239." title="A revision of the genus Camellia. Royal Horticultural Society London, 1 - 239." year="1958">Sealy 1958</bibRefCitation>
;
<bibRefCitation id="6FF465569EB781E00E67DEDEACD5DAB2" author="Chang, HT" journalOrPublisher="Acta Scientiarum Naturalium Universitatis Sunyatseni Monograph Series" pageId="0" pageNumber="1" refId="B3" refString="Chang, HT, Bartholomew, B, 1984. Camellias. Timber Press Portland London, 1-211." title="Camellias. Timber Press Portland London, 1 - 211." year="1984">Chang and Bartholomew 1984</bibRefCitation>
;
<bibRefCitation id="D2AB1C6C915A81879B5536BD4E49E523" author="Ming, TL" journalOrPublisher="Academia Journal of Biology" pageId="0" pageNumber="1" refId="B19" refString="Ming, TL, 2000. Monograph of the genus Camellia. Yunnan Science and Technology Press Kunming, 1-120." title="Monograph of the genus Camellia. Yunnan Science and Technology Press Kunming, 1 - 120." year="2000">Ming 2000</bibRefCitation>
) and was based on the assessment of morphological characteristics. The taxonomic system of
<bibRefCitation id="2169E8EF1588DDDA17B4D7AD2E79AF67" author="Sealy, JR" journalOrPublisher="Adansonia" pageId="0" pageNumber="1" refId="B37" refString="Sealy, JR, 1958. A revision of the genus Camellia. Royal Horticultural Society London, 1-239." title="A revision of the genus Camellia. Royal Horticultural Society London, 1 - 239." year="1958">Sealy (1958)</bibRefCitation>
and
<bibRefCitation id="D6636AB4565EFE2C8EA72D321806740C" author="Chang, HT" journalOrPublisher="Acta Scientiarum Naturalium Universitatis Sunyatseni Monograph Series" pageId="0" pageNumber="1" refId="B3" refString="Chang, HT, Bartholomew, B, 1984. Camellias. Timber Press Portland London, 1-211." title="Camellias. Timber Press Portland London, 1 - 211." year="1984">Chang and Bartholomew (1984)</bibRefCitation>
are used to describe and determine the placement of new taxa within
<taxonomicName id="58662544647FB0177879DEA48DFB596C" class="Magnoliopsida" family="Theaceae" genus="Camellia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Camellia" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="465E7211AFFB149D85102A370EE8961A" italics="true" pageId="0" pageNumber="1">Camellia</emphasis>
</taxonomicName>
. These systems are the most detailed and comprehensive study of the genus and also provide the basis for our current understanding of the genus. The taxonomic system of Ming (sensu
<bibRefCitation id="49B928F41E6078375F600C3DD12D1BD3" author="Ming, TL" editor="Wu, ZY" journalOrPublisher="12. Science Press Beijing and Missouri Botanical Garden Press St Louis" pageId="0" pageNumber="1" pagination="366 - 478" refId="B20" refString="Ming, TL, Bartholomew, B, 2007. Theaceae. In: Wu, ZY, Raven, PH, Eds., Flora of China, Vol. 12. Science Press Beijing and Missouri Botanical Garden Press St Louis: 366 - 478" title="Theaceae." volumeTitle="Flora of China, Vol." year="2007">Ming and Bartholomew 2007</bibRefCitation>
) was used for supplementary data only as it appears to be superficially similar to the system of Sealy (
<bibRefCitation id="41A2638F771E7F9BFF99BD7D3EF4D2E1" DOI="https://doi.org/10.1111/j.1756-1051.2010.00652.x" author="Orel, G" journalOrPublisher="Nordic Journal of Botany" pageId="0" pageNumber="1" pagination="280 - 284" refId="B24" refString="Orel, G, Wilson, PG, 2010a. Camellia luteocerata sp nov and a new section of Camellia Dalatia from Vietnam. Nordic Journal of Botany 28 (3): 280 - 284, DOI: https://doi.org/10.1111/j.1756-1051.2010.00652.x" title="Camellia luteocerata sp nov and a new section of Camellia Dalatia from Vietnam." url="https://doi.org/10.1111/j.1756-1051.2010.00652.x" volume="28" year="2010 a">Orel and Wilson 2010a</bibRefCitation>
). In this paper, we have followed the
<bibRefCitation id="26C3331B2944C5AC00A5C5F7BF5CBCA8" author="Sealy, JR" journalOrPublisher="Adansonia" pageId="0" pageNumber="1" refId="B37" refString="Sealy, JR, 1958. A revision of the genus Camellia. Royal Horticultural Society London, 1-239." title="A revision of the genus Camellia. Royal Horticultural Society London, 1 - 239." year="1958">
<normalizedToken id="F4577869609C4E8491946A21E48E3E13" originalValue="Sealys">Sealy's</normalizedToken>
(1958)
</bibRefCitation>
taxonomic system to consider the placement of the new species within Section
<taxonomicName id="D03D107EF4C8C04B40CBC68F603494A7" class="Magnoliopsida" family="Theaceae" genus="Camellia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Archecamellia">Archecamellia</taxonomicName>
Sealy of
<taxonomicName id="7C6CEBA7A2E4DE09B20143FAE085E933" class="Magnoliopsida" family="Theaceae" genus="Camellia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Camellia" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="194CD60FBD124C513A1205FC3BFEEAF9" italics="true" pageId="0" pageNumber="1">Camellia</emphasis>
</taxonomicName>
. Morphologically,
<taxonomicName id="1A30891AE9030C01760E329C71A4324E" lsidName="C. puhoatensis" pageId="0" pageNumber="1" rank="species" species="puhoatensis">
<emphasis id="CB6A44A3B028969141706CDBBC3EDC72" italics="true" pageId="0" pageNumber="1">C. puhoatensis</emphasis>
</taxonomicName>
possesses a solitary or paired flowers at terminal (sometimes axillary), stout, thick and erected pedicel, 3-4 persistent bracteoles, 5 persistent sepals (undifferentiated bracteoles and sepals), large yellow flowers with 12 tepals that are inner ones basally connate and adnate to androecium, androecium free above the union with the petals or unified for some distance to form a fleshy cup, filaments glabrous, gynoecium 3(-4)-loculi, styles 3(-4) and free to the base. These characteristics are not only identical to the species of sect.
<taxonomicName id="989CF1EF14DAA0C27EAB37B71EF4B932" lsidName="" pageId="0" pageNumber="1" rank="section" section="Archaecamellia">Archaecamellia</taxonomicName>
Sealy (sensu
<bibRefCitation id="00F41F09DC7A9FF7DA0A7D94A9F1B242" author="Sealy, JR" journalOrPublisher="Adansonia" pageId="0" pageNumber="1" refId="B37" refString="Sealy, JR, 1958. A revision of the genus Camellia. Royal Horticultural Society London, 1-239." title="A revision of the genus Camellia. Royal Horticultural Society London, 1 - 239." year="1958">Sealy 1958</bibRefCitation>
; sensu
<bibRefCitation id="B8C5DF23F2BA302D09E4E9241728F105" author="Chang, HT" journalOrPublisher="Acta Scientiarum Naturalium Universitatis Sunyatseni Monograph Series" pageId="0" pageNumber="1" refId="B3" refString="Chang, HT, Bartholomew, B, 1984. Camellias. Timber Press Portland London, 1-211." title="Camellias. Timber Press Portland London, 1 - 211." year="1984">Chang and Bartholomew 1984</bibRefCitation>
), but also share with species of sect.
<taxonomicName id="73A34EC9DC054993C9A777A1752586BC" lsidName="" pageId="0" pageNumber="1" rank="section" section="Stereocarpus">Stereocarpus</taxonomicName>
which possesses 2 or 4 bracteoles (sensu
<bibRefCitation id="1DD97F25E43A7F1C80890D94E598D83C" author="Sealy, JR" journalOrPublisher="Adansonia" pageId="0" pageNumber="1" refId="B37" refString="Sealy, JR, 1958. A revision of the genus Camellia. Royal Horticultural Society London, 1-239." title="A revision of the genus Camellia. Royal Horticultural Society London, 1 - 239." year="1958">Sealy 1958</bibRefCitation>
), terminal flowers (sensu
<bibRefCitation id="1E1B77B823515A1C93B54E25E7116B1E" author="Chang, HT" journalOrPublisher="Acta Scientiarum Naturalium Universitatis Sunyatseni Monograph Series" pageId="0" pageNumber="1" refId="B3" refString="Chang, HT, Bartholomew, B, 1984. Camellias. Timber Press Portland London, 1-211." title="Camellias. Timber Press Portland London, 1 - 211." year="1984">Chang and Bartholomew 1984</bibRefCitation>
), stamens in 3-4 whorls, ovary with 3-5 locules (sensu
<bibRefCitation id="94B74CA5109467573552FFEFB3A20D2A" author="Ming, TL" journalOrPublisher="Academia Journal of Biology" pageId="0" pageNumber="1" refId="B19" refString="Ming, TL, 2000. Monograph of the genus Camellia. Yunnan Science and Technology Press Kunming, 1-120." title="Monograph of the genus Camellia. Yunnan Science and Technology Press Kunming, 1 - 120." year="2000">Ming 2000</bibRefCitation>
), as well as sharing with species of sect.
<taxonomicName id="78CC261EE592E02B5F210A2F05263777" lsidName="" pageId="0" pageNumber="1" rank="section" section="Chrysantha">Chrysantha</taxonomicName>
Chang, such as distinct peduncle, small floral bracteolates, yellow flowers, three carpels of gynoecium and separate styles (sensu
<bibRefCitation id="CE02933193374D0AD653D37EF9169064" author="Chang, HT" journalOrPublisher="Acta Scientiarum Naturalium Universitatis Sunyatseni" pageId="0" pageNumber="1" pagination="69 - 74" refId="B1" refString="Chang, HT, 1979. Chrysantha, a section of golden camellias from Cathaysian flora. Acta Scientiarum Naturalium Universitatis Sunyatseni 18: 69 - 74" title="Chrysantha, a section of golden camellias from Cathaysian flora." volume="18" year="1979">Chang 1979</bibRefCitation>
). As characterized by
<bibRefCitation id="301ECCD0FB331B993D550139989A32D3" author="Sealy, JR" journalOrPublisher="Adansonia" pageId="0" pageNumber="1" refId="B37" refString="Sealy, JR, 1958. A revision of the genus Camellia. Royal Horticultural Society London, 1-239." title="A revision of the genus Camellia. Royal Horticultural Society London, 1 - 239." year="1958">Sealy (1958)</bibRefCitation>
, sect.
<taxonomicName id="DA37F2E97952B04F8327697442E34479" genus="Archaecamellia" lsidName="Archaecamellia" pageId="0" pageNumber="1" rank="genus">Archaecamellia</taxonomicName>
shares several traits with sect.
<taxonomicName id="22AC431B896A07B4EA01FAD0609999CC" genus="Archaecamellia" lsidName="" pageId="0" pageNumber="1" rank="section" section="Stereocarpus">Stereocarpus</taxonomicName>
. These include a solitary and erected flower at the end of the branches, persistent bracteoles and sepals, stamens united with the petals and glabrous filaments. However, traits that are distinctive to sect.
<taxonomicName id="33DF4F95560AD0970535A26A33BF79D5" genus="Archaecamellia" lsidName="" pageId="0" pageNumber="1" rank="section" section="Archaecamellia">Archaecamellia</taxonomicName>
include (6-)11-16 indistinct bracteoles and sepals, 8-14 petals, glabrous or pubescent gynoecium and 3 or 5 free styles (vs. 2 or 4 bracteoles and 5 or 6 sepals, ca. 12 petals, glabrous gynoecium and a single style in sect.
<taxonomicName id="9C1F297FA2232EE89AC2B875E1CA9602" genus="Archaecamellia" lsidName="" pageId="0" pageNumber="1" rank="section" section="Stereocarpus">Stereocarpus</taxonomicName>
). The sect.
<taxonomicName id="F33F8F24BC25EE8ECD82455EF89519FD" genus="Chrysantha" lsidName="Chrysantha" pageId="0" pageNumber="1" rank="genus">Chrysantha</taxonomicName>
also shares several traits with sect.
<taxonomicName id="807900FC1DE5E43A8AB7233CDFDAC0AD" genus="Chrysantha" lsidName="" pageId="0" pageNumber="1" rank="section" section="Archaecamellia">Archaecamellia</taxonomicName>
in having yellow and pedicellate flowers, persistent bracteoles and sepals, glabrous or hairy filaments and gynoecium, but it can be distinguished from sect.
<taxonomicName id="4F69811A0A4272B5B4F79AD1E0B660E3" genus="Chrysantha" lsidName="" pageId="0" pageNumber="1" rank="section" section="Archaecamellia">Archaecamellia</taxonomicName>
by the axillary flowers, distinct bracts and sepals and 3-5 cleft styles (
<bibRefCitation id="C53239E3219039A12D8889244C2946BF" author="Chang, HT" journalOrPublisher="Acta Scientiarum Naturalium Universitatis Sunyatseni" pageId="0" pageNumber="1" pagination="69 - 74" refId="B1" refString="Chang, HT, 1979. Chrysantha, a section of golden camellias from Cathaysian flora. Acta Scientiarum Naturalium Universitatis Sunyatseni 18: 69 - 74" title="Chrysantha, a section of golden camellias from Cathaysian flora." volume="18" year="1979">Chang 1979</bibRefCitation>
). Section
<taxonomicName id="E7F0B0C327551B9183347DCBC5BACCA1" genus="Chrysantha" lsidName="" pageId="0" pageNumber="1" rank="section" section="Archaecamellia">Archaecamellia</taxonomicName>
is currently comprised of 19 species (
<bibRefCitation id="6076F23F668A6B0E702FB9B24F6E080E" author="Sealy, JR" journalOrPublisher="Adansonia" pageId="0" pageNumber="1" refId="B37" refString="Sealy, JR, 1958. A revision of the genus Camellia. Royal Horticultural Society London, 1-239." title="A revision of the genus Camellia. Royal Horticultural Society London, 1 - 239." year="1958">Sealy 1958</bibRefCitation>
;
<bibRefCitation id="C15B2C506775C22A1CBF6913167DF7BD" author="Ming, TL" journalOrPublisher="Academia Journal of Biology" pageId="0" pageNumber="1" refId="B19" refString="Ming, TL, 2000. Monograph of the genus Camellia. Yunnan Science and Technology Press Kunming, 1-120." title="Monograph of the genus Camellia. Yunnan Science and Technology Press Kunming, 1 - 120." year="2000">Ming 2000</bibRefCitation>
;
<bibRefCitation id="7FBDD513D05E016769D91C60EEE15535" DOI="https://doi.org/10.1007/s12225-012-9317-0" author="Orel, G" journalOrPublisher="Kew Bulletin" pageId="0" pageNumber="1" pagination="565 - 569" refId="B26" refString="Orel, G, Wilson, PG, 2012a. Camellia cattienensis: a new species of Camellia sect. Archaecamellia: Theaceae from Vietnam. Kew Bulletin 66 (4): 565 - 569, DOI: https://doi.org/10.1007/s12225-012-9317-0" title="Camellia cattienensis: a new species of Camellia sect. Archaecamellia: Theaceae from Vietnam." url="https://doi.org/10.1007/s12225-012-9317-0" volume="66" year="2012 a">Orel and Wilson 2012a</bibRefCitation>
;
<bibRefCitation id="25E58B859474805020C28882BFAC600A" DOI="https://doi.org/10.11110/kjpt.2019.49.1.90" author="Do, ND" journalOrPublisher="Korean Journal of Plant Taxonomy" pageId="0" pageNumber="1" pagination="90 - 95" refId="B5" refString="Do, ND, Luong, VD, Nguyen, DC, Hoang, TS, Le, TH, Le Han, JE, Park, HS, 2019a. A new yellow Camellia species from Central Vietnam. Korean Journal of Plant Taxonomy 49 (1): 90 - 95, DOI: https://doi.org/10.11110/kjpt.2019.49.1.90" title="A new yellow Camellia species from Central Vietnam." url="https://doi.org/10.11110/kjpt.2019.49.1.90" volume="49" year="2019 a">Do et al. 2019a</bibRefCitation>
). The new species is most similar to
<taxonomicName id="28E25F45E166BFA04530D1909E316407" lsidName="C. chrysanthoides" pageId="0" pageNumber="1" rank="species" species="chrysanthoides">
<emphasis id="0DDF0FF09DB95E59458A8FEE802FB597" italics="true" pageId="0" pageNumber="1">C. chrysanthoides</emphasis>
</taxonomicName>
H.T.Chang,
<taxonomicName id="D0D664C2CBC96D346D57A481A468E822" lsidName="C. flavida" pageId="0" pageNumber="1" rank="species" species="flavida">
<emphasis id="BF6F97B1477AF3DC2A041D774DEB1D5A" italics="true" pageId="0" pageNumber="1">C. flavida</emphasis>
</taxonomicName>
H.T.Chang and
<taxonomicName id="BCA4CADFB1A7E72894B93FE03F3F2507" lsidName="C. petelotii" pageId="0" pageNumber="1" rank="species" species="petelotii">
<emphasis id="6FF0E745975131367B45415B3556AEE6" italics="true" pageId="0" pageNumber="1">C. petelotii</emphasis>
</taxonomicName>
(Merr.) Sealy in having the same plant habit, somewhat oblong leaves, yellow flowers, glabrous 3-loculed gynoecium with 3 styles free to the base. A detailed morphological comparison between
<taxonomicName id="F45EA1EAE1C42F79AC4C0B794DFD6568" lsidName="C. puhoatensis" pageId="0" pageNumber="1" rank="species" species="puhoatensis">
<emphasis id="BF685C224D9A8AE06AEC4892042A90F7" italics="true" pageId="0" pageNumber="1">C. puhoatensis</emphasis>
</taxonomicName>
and these three species is provided in the above diagnosis and in Table
<tableCitation id="B4164D50F55FE2968DBA6B78E21CDC87" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparison of C. puhoatensis with its most closely-related taxa (based on Sealy 1958; Chang and Bartholomew 1984; Tran and Hakoda 1998; Pham et al. 2019)." httpUri="http://table.plazi.org/id/9832B9EFCABB2302834C710943B08ACE" pageId="0" pageNumber="1" tableUuid="9832B9EFCABB2302834C710943B08ACE">1</tableCitation>
. Moreover,
<taxonomicName id="F5DA7FA42ADB49DF986C62E9BF87EEFE" lsidName="C. puhoatensis" pageId="0" pageNumber="1" rank="species" species="puhoatensis">
<emphasis id="A3F9177CD4ABCF8B27D26BA2EE4A6FFA" italics="true" pageId="0" pageNumber="1">C. puhoatensis</emphasis>
</taxonomicName>
also resembles
<taxonomicName id="F10F1AD35B2FDAC3E11B24DF1AE05597" lsidName="C. dormoyana" pageId="0" pageNumber="1" rank="species" species="dormoyana">
<emphasis id="0394A69403C596D0E84098F4725A46C1" italics="true" pageId="0" pageNumber="1">C. dormoyana</emphasis>
</taxonomicName>
(Pierre) Sealy of sect.
<taxonomicName id="88B35CB6B165F672FF1DDD075826535C" lsidName="" pageId="0" pageNumber="1" rank="section" section="Stereocarpus">Stereocarpus</taxonomicName>
(
<bibRefCitation id="B6004E80203DFC06AD90060945A5C97A" author="Sealy, JR" journalOrPublisher="Adansonia" pageId="0" pageNumber="1" refId="B37" refString="Sealy, JR, 1958. A revision of the genus Camellia. Royal Horticultural Society London, 1-239." title="A revision of the genus Camellia. Royal Horticultural Society London, 1 - 239." year="1958">Sealy 1958</bibRefCitation>
) and
<taxonomicName id="E088B3FD21E07A00CAE5B42DC1988049" lsidName="C. velutina" pageId="0" pageNumber="1" rank="species" species="velutina">
<emphasis id="FEA45B2BD256A0EEC1274FB3011DA714" italics="true" pageId="0" pageNumber="1">C. velutina</emphasis>
</taxonomicName>
V.T. Pham et al. of sect.
<taxonomicName id="1EB2E3C2AFA86BDB481FB864211B6933" lsidName="" pageId="0" pageNumber="1" rank="section" section="Chrysantha">Chrysantha</taxonomicName>
(
<bibRefCitation id="D9DC4287B3678649C3CDFC9B7065561C" DOI="https://doi.org/10.30848/PJB2019-4(33)" author="Pham, VT" journalOrPublisher="Pakistan Journal of Botany" pageId="0" pageNumber="1" pagination="1441 - 1446" refId="B33" refString="Pham, VT, Luong, VD, Averyanov, LV, Trinh, NB, Nguyen, TL, Nguyen, TLT, 2019. Camellia velutina (Theaceae: Sect. Chrysanthae), a new species from northern Vietnam. Pakistan Journal of Botany 51 (4): 1441 - 1446, DOI: https://doi.org/10.30848/PJB2019-4(33)" title="Camellia velutina (Theaceae: Sect. Chrysanthae), a new species from northern Vietnam." url="https://doi.org/10.30848/PJB2019-4(33)" volume="51" year="2019">Pham et al. 2019</bibRefCitation>
) by somewhat oblong leaves, yellow flowers and glabrous stamens. However,
<taxonomicName id="9A0B61EDAF81FAE4E07027141B6B636B" authorityName="Sealy" authorityYear="1958" baseAuthorityName="Pierre" class="Magnoliopsida" family="Theaceae" genus="Camellia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Camellia dormoyana" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="dormoyana">
<emphasis id="E2104941ADA9E7C798A5B8B9E1E90019" italics="true" pageId="0" pageNumber="1">Camellia dormoyana</emphasis>
</taxonomicName>
is easily distinguished from
<taxonomicName id="A9821CEDE732F607A75DBB7A27740BE4" lsidName="C. puhoatensis" pageId="0" pageNumber="1" rank="species" species="puhoatensis">
<emphasis id="E195CD9B00BB493A6524E8432B81E941" italics="true" pageId="0" pageNumber="1">C. puhoatensis</emphasis>
</taxonomicName>
by having the young shoots, mature leaves and petioles all glabrous, the sessile pedicel and 5-6 bracteoles abaxially velutinous, the abaxial petals silky velutinous, the ovary being glabrous and with five locules and the styles united for their entire length and glabrous. Similarly,
<taxonomicName id="AF39F685C378136B43347B03B131D442" lsidName="C. velutina" pageId="0" pageNumber="1" rank="species" species="velutina">
<emphasis id="B39BFCC9E716DDCD239963600EE0744C" italics="true" pageId="0" pageNumber="1">C. velutina</emphasis>
</taxonomicName>
is readily distinguished from
<taxonomicName id="3F9A0CD7CE4EFDC73A73F6516A29CCB5" lsidName="C. puhoatensis" pageId="0" pageNumber="1" rank="species" species="puhoatensis">
<emphasis id="E91536B4611C2595915EEF0DECF4CA37" italics="true" pageId="0" pageNumber="1">C. puhoatensis</emphasis>
</taxonomicName>
by its glabrous young shoots, mature leaves and petioles, sepals that are silky velutinous abaxially and velutinous adaxially, the 10 (occasionally 11) petals that are silky velutinous and glabrous ovary and style (see Table
<tableCitation id="2BEC96CFC6EE710F83BC71DDD995445B" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparison of C. puhoatensis with its most closely-related taxa (based on Sealy 1958; Chang and Bartholomew 1984; Tran and Hakoda 1998; Pham et al. 2019)." httpUri="http://table.plazi.org/id/9832B9EFCABB2302834C710943B08ACE" pageId="0" pageNumber="1" tableUuid="9832B9EFCABB2302834C710943B08ACE">1</tableCitation>
).
</paragraph>
<caption id="9832B9EFCABB2302834C710943B08ACE" ID-Table-UUID="9832B9EFCABB2302834C710943B08ACE" httpUri="http://table.plazi.org/id/9832B9EFCABB2302834C710943B08ACE" pageId="0" pageNumber="1" start="Table 1" startId="T1">
<paragraph id="D361513FC5BDB7D483E833B6BB5E9401" pageId="0" pageNumber="1">
<emphasis id="2870119D5140049DA3F1782BE0870ED3" bold="true" pageId="0" pageNumber="1">Table 1.</emphasis>
Morphological comparison of
<taxonomicName id="A297942983AEFCACCFD25EA8183D1487" lsidName="C. puhoatensis" pageId="0" pageNumber="1" rank="species" species="puhoatensis">
<emphasis id="03BF06AF10078651F6CC56338A2FFA32" italics="true" pageId="0" pageNumber="1">C. puhoatensis</emphasis>
</taxonomicName>
with its most closely-related taxa (based on
<bibRefCitation id="23A9A5A94BC52329355D9181857DFC25" author="Sealy, JR" journalOrPublisher="Adansonia" pageId="0" pageNumber="1" refId="B37" refString="Sealy, JR, 1958. A revision of the genus Camellia. Royal Horticultural Society London, 1-239." title="A revision of the genus Camellia. Royal Horticultural Society London, 1 - 239." year="1958">Sealy 1958</bibRefCitation>
;
<bibRefCitation id="289985F036BF1D3D0A09BC41B2968E52" author="Chang, HT" journalOrPublisher="Acta Scientiarum Naturalium Universitatis Sunyatseni Monograph Series" pageId="0" pageNumber="1" refId="B3" refString="Chang, HT, Bartholomew, B, 1984. Camellias. Timber Press Portland London, 1-211." title="Camellias. Timber Press Portland London, 1 - 211." year="1984">Chang and Bartholomew 1984</bibRefCitation>
;
<bibRefCitation id="C21DD948AB41C4317BE11ECD5DF363EA" author="Tran, N" journalOrPublisher="International Camellia Journal" pageId="0" pageNumber="1" pagination="81 - 83" refId="B41" refString="Tran, N, Hakoda, N, 1998. Camellia petelotii: A species of yellow Camellia from Vietnam. International Camellia Journal 30: 81 - 83" title="Camellia petelotii: A species of yellow Camellia from Vietnam." volume="30" year="1998">Tran and Hakoda 1998</bibRefCitation>
;
<bibRefCitation id="B64950479F7FC1EDFB7070B258E88647" DOI="https://doi.org/10.30848/PJB2019-4(33)" author="Pham, VT" journalOrPublisher="Pakistan Journal of Botany" pageId="0" pageNumber="1" pagination="1441 - 1446" refId="B33" refString="Pham, VT, Luong, VD, Averyanov, LV, Trinh, NB, Nguyen, TL, Nguyen, TLT, 2019. Camellia velutina (Theaceae: Sect. Chrysanthae), a new species from northern Vietnam. Pakistan Journal of Botany 51 (4): 1441 - 1446, DOI: https://doi.org/10.30848/PJB2019-4(33)" title="Camellia velutina (Theaceae: Sect. Chrysanthae), a new species from northern Vietnam." url="https://doi.org/10.30848/PJB2019-4(33)" volume="51" year="2019">Pham et al. 2019</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="C100056ACAFDDC5299D6D07B1435BB55" pageId="0" pageNumber="1">
<table id="D11A2CB56C11DAB0951664B80F876E1C" pageId="0" pageNumber="1">
<tr id="9FCB440A88678DAD78890B365D143C6F" pageId="0" pageNumber="1">
<th id="7319BB907A93D9986812485672BC1E63" colspan="1" pageId="0" pageNumber="1" rowspan="1">Characters</th>
<th id="4E204F386A1FBDD0BE6041D7FD72715D" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="29CE8B0A3ACE635FFA0251C5E1C5A4E1" lsidName="C. puhoatensis" pageId="0" pageNumber="1" rank="species" species="puhoatensis">
<emphasis id="B14CB1F13CDA9125BE7914513D23FF06" italics="true" pageId="0" pageNumber="1">C. puhoatensis</emphasis>
</taxonomicName>
</th>
<th id="4D836D249863151BA4901B436E4D02B5" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="5A5F51DEA0FE92FEAAF1CC584460EDE2" lsidName="C. chrysanthoides" pageId="0" pageNumber="1" rank="species" species="chrysanthoides">
<emphasis id="8DCF4C4361ABC39C1EF41B9F879BB931" italics="true" pageId="0" pageNumber="1">C. chrysanthoides</emphasis>
</taxonomicName>
</th>
<th id="4E0732514EB5FC0D150663AC6FB216AA" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="5DE03A1204B42B9D7B0CEAE250B8D26C" lsidName="C. flavida" pageId="0" pageNumber="1" rank="species" species="flavida">
<emphasis id="03FC3568DAED318A3D974E703FF3A438" italics="true" pageId="0" pageNumber="1">C. flavida</emphasis>
</taxonomicName>
</th>
<th id="29BDEC4ED7905C5FD8D01ABB46A576C8" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="2B39A4F38F5C0704D8017A11BC1817FC" lsidName="C. petelotii" pageId="0" pageNumber="1" rank="species" species="petelotii">
<emphasis id="5A534F7783EAE1D0D4C5FEF823BEDE41" italics="true" pageId="0" pageNumber="1">C. petelotii</emphasis>
</taxonomicName>
</th>
<th id="BCC8A59899063DD58908C92400B27744" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="33D0A06F8D5E6D296419D75C25827DC8" lsidName="C. velutina" pageId="0" pageNumber="1" rank="species" species="velutina">
<emphasis id="116D0194982BB3AFC28EADFF63D9B38E" italics="true" pageId="0" pageNumber="1">C. velutina</emphasis>
</taxonomicName>
</th>
<th id="268D7A69863BF3C0286AE0D70AD97F98" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="ECD80E459C042D20D824583FC4C7C1C9" lsidName="C. dormoyana" pageId="0" pageNumber="1" rank="species" species="dormoyana">
<emphasis id="3781CDDB2DE6690BEE677679FB5FD5BC" italics="true" pageId="0" pageNumber="1">C. dormoyana</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3145D2F142ED70368A8B9F449E2ABDBC" pageId="0" pageNumber="1">
<td id="C3E203E49B601A96253F3AF68157CB90" colspan="1" pageId="0" pageNumber="1" rowspan="1">Young shoot</td>
<td id="22631A2E7566F17BA8A8226A179B0D60" colspan="1" pageId="0" pageNumber="1" rowspan="1">puberulous</td>
<td id="F8D125E57E9BDE676E450AEE2900EF38" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="F662BA77BE7EF4D55CC671106B5CF0A4" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="DA4CF74C2BFCA6958EAC992939B5A5D7" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="260C7C65AD70D1D1F71778AE53EEF14C" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="591B2BDB1BE5662AE6DC5CBEBABB5FE1" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
</tr>
<tr id="58681A4E6AA8E189B65777AC880D08C5" pageId="0" pageNumber="1">
<td id="EA0479A6647F90A542893C7DB85871B3" colspan="1" pageId="0" pageNumber="1" rowspan="1">Leaf blade</td>
<td id="FBD09ABF8819524F5E7294953580BF53" colspan="1" pageId="0" pageNumber="1" rowspan="1">
oblong ovate or oblong, 17-23
<normalizedToken id="1DE3A4981353164E382784FA35FD85C5" originalValue="×">x</normalizedToken>
5-6.5 cm, base rounded or broadly obtuse, abaxially sparsely puberulous
</td>
<td id="C2225BD26D869801D3BA890A1841E8BE" colspan="1" pageId="0" pageNumber="1" rowspan="1">
oblong, 10-18
<normalizedToken id="1B47D8FD2EB211467D9E36F331B8AC10" originalValue="×">x</normalizedToken>
3.0-6.5 cm, base cuneate, glabrous
</td>
<td id="99B37AC6C6CB8150101AEE5F0163BC79" colspan="1" pageId="0" pageNumber="1" rowspan="1">
elliptic to oblong, 6.0-10
<normalizedToken id="AFDD72267930075369325D3351303D65" originalValue="×">x</normalizedToken>
2.1-4.5 cm, base broadly cuneate, glabrous
</td>
<td id="FE4B4BAD7DE33B2E2720FE94A00B446A" colspan="1" pageId="0" pageNumber="1" rowspan="1">
broadly oblong or oblong-oval, 14.5-18
<normalizedToken id="2FDA024CEB60C365D6D33E72397723BF" originalValue="×">x</normalizedToken>
4.5-7.5 cm, base broadly cuneate, glabrous
</td>
<td id="D5F24517AABB3F28623192CEF6FB7D94" colspan="1" pageId="0" pageNumber="1" rowspan="1">
oblong to elliptic, 15-22
<normalizedToken id="F96862DB451E108635BC1B590650F4E2" originalValue="×">x</normalizedToken>
5-11 cm, base broadly cuneate to rounded, glabrous
</td>
<td id="C6AAAD161C06AA0432E5DB18B69B67A7" colspan="1" pageId="0" pageNumber="1" rowspan="1">
oval or oblong or ovate, 11-18(-25)
<normalizedToken id="8F0B6384073D9C9360F82B8E6A7E8E70" originalValue="×">x</normalizedToken>
5.5-8.5 cm, base cuneate to rounded, glabrous
</td>
</tr>
<tr id="4AEC72FDF5F0A46FFE2A07E00E0927A8" pageId="0" pageNumber="1">
<td id="BF76EB437A04C3A3E0CC5644A9A0583A" colspan="1" pageId="0" pageNumber="1" rowspan="1">Petiole</td>
<td id="B323B42363E05B2F403D555731A5A962" colspan="1" pageId="0" pageNumber="1" rowspan="1">puberulous</td>
<td id="57CE82CA667AF44BB4C607E18010A0B5" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="118CFA333E15825D9C86BB51F09CA252" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="C7B6F6107B3377A870DA09D9A2A26A27" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="C983E548A313903ACA0F82F54051C44E" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="CE40E90082930494D786E2CEE35AA286" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
</tr>
<tr id="D75C6E2C9885D8AEF4AE9638FB22A41A" pageId="0" pageNumber="1">
<td id="80D1E11A4CC62744ED6B8647AD07C4C0" colspan="1" pageId="0" pageNumber="1" rowspan="1">Flower</td>
<td id="9AA5EB7A41AF9C9376D7CBB003522E31" colspan="1" pageId="0" pageNumber="1" rowspan="1">solitary (2 flowers), terminal, rarely axillary</td>
<td id="BC8F66D76DB1CEF45B76F76F63D7178E" colspan="1" pageId="0" pageNumber="1" rowspan="1">solitary, mostly axillary</td>
<td id="E2FACE83D324C6F9DC48137C337634D2" colspan="1" pageId="0" pageNumber="1" rowspan="1">solitary, terminal and axillary</td>
<td id="0AED855F9CACD401D2575B693FE177ED" colspan="1" pageId="0" pageNumber="1" rowspan="1">solitary, terminal</td>
<td id="3091B5131CB9BD5C2E9CF2498569CC36" colspan="1" pageId="0" pageNumber="1" rowspan="1">solitary, terminal or axillary</td>
<td id="0A65E593AEF87B7A906D88382ABB045A" colspan="1" pageId="0" pageNumber="1" rowspan="1">solitary, terminal</td>
</tr>
<tr id="5959AD06C2C7E7C8CCC3F75B18BF6A01" pageId="0" pageNumber="1">
<td id="AC4C016F66003D206B0C5D3969113DB6" colspan="1" pageId="0" pageNumber="1" rowspan="1">Pedicel</td>
<td id="03C6C6AE7FA8D5FE4E5C1AB901BFFCE3" colspan="1" pageId="0" pageNumber="1" rowspan="1">7-10 mm long</td>
<td id="D8CE2CB78A5C50DFFC2B7236194172E1" colspan="1" pageId="0" pageNumber="1" rowspan="1">3-4 mm long</td>
<td id="A84153120680267D33F6306509E1716F" colspan="1" pageId="0" pageNumber="1" rowspan="1">1-2 mm long</td>
<td id="4A1070050C97C8BBD574EA09CFFA8C10" colspan="1" pageId="0" pageNumber="1" rowspan="1">10-12 mm long</td>
<td id="0A396AFF6040948E6531DDAACACF5925" colspan="1" pageId="0" pageNumber="1" rowspan="1">10-13 mm</td>
<td id="AF74731F24BA3C2CD9BB75D1DBDF8AC9" colspan="1" pageId="0" pageNumber="1" rowspan="1">sessile</td>
</tr>
<tr id="59D89BECDBF6D23CBDC5BD7ACCEDA8B0" pageId="0" pageNumber="1">
<td id="69FEF3A5A99849F31DD6D5C0F93053A3" colspan="1" pageId="0" pageNumber="1" rowspan="1">Bracteoles</td>
<td id="6F5CD7B1EB5E133D2A70A0EC925DC259" colspan="1" pageId="0" pageNumber="1" rowspan="1">3-4, glabrous</td>
<td id="72E4851818E47564F1D7647C7E7596B0" colspan="1" pageId="0" pageNumber="1" rowspan="1">4-6, abaxially pubescent</td>
<td id="D11B041884B36DDA16C218BFB79341AC" colspan="1" pageId="0" pageNumber="1" rowspan="1">4-5, glabrous</td>
<td id="3D3C972F0906241AF3B47A124B2CEB9B" colspan="1" pageId="0" pageNumber="1" rowspan="1">(6-)8-10, abaxially puberulous</td>
<td id="751CC73BE2CB76BD2CCFB4F892643D54" colspan="1" pageId="0" pageNumber="1" rowspan="1">2(-3), abaxially velutinous</td>
<td id="A3D1FF80FDB6C1304C45B37A0181767D" colspan="1" pageId="0" pageNumber="1" rowspan="1">5-6, abaxially silky velutinous</td>
</tr>
<tr id="6F666DB9BC75BCC1A7CCA798F89AF42A" pageId="0" pageNumber="1">
<td id="341A8381D85B4ADC77B36BB95400AA68" colspan="1" pageId="0" pageNumber="1" rowspan="1">Sepals</td>
<td id="FDE507BE4C1309ACBA83B760170E1A73" colspan="1" pageId="0" pageNumber="1" rowspan="1">5, abaxially pubescent</td>
<td id="67551FD95D2ACF8A3A3FF38E7053FEA1" colspan="1" pageId="0" pageNumber="1" rowspan="1">5, abaxially puberulent</td>
<td id="A2F619E4C0848CDFCAB6AFA782483C5B" colspan="1" pageId="0" pageNumber="1" rowspan="1">4-6, glabrous</td>
<td id="9ACC0A0A86129F68A27954B3D003AB92" colspan="1" pageId="0" pageNumber="1" rowspan="1">5, abaxially puberulous</td>
<td id="3FA17EDFF04E54FF8263BA4FE99EEA9C" colspan="1" pageId="0" pageNumber="1" rowspan="1">5, adaxially velutinous</td>
<td id="D832A9689C63376239D0ADBBF83AC46A" colspan="1" pageId="0" pageNumber="1" rowspan="1">abaxially silky velutinous</td>
</tr>
<tr id="5E6857016803BD8CF7E750C115DBA661" pageId="0" pageNumber="1">
<td id="BF320469523E2A9CC4907377F18A427B" colspan="1" pageId="0" pageNumber="1" rowspan="1">Petals</td>
<td id="1FC1229B2CFAB56E6E0CB1051C69C152" colspan="1" pageId="0" pageNumber="1" rowspan="1">12-13, abaxially pubescent</td>
<td id="21165F4B54C88F4F323A24FC80BAD743" colspan="1" pageId="0" pageNumber="1" rowspan="1">8-9, abaxially puberulent</td>
<td id="793B6420CD3E03766CF903EBEFFCA6DE" colspan="1" pageId="0" pageNumber="1" rowspan="1">8, glabrous</td>
<td id="1F990905101CB42E9F493F3269959A2F" colspan="1" pageId="0" pageNumber="1" rowspan="1">ca. 14, abaxially puberulous</td>
<td id="B837672B95CAEDD83502A14AB7C543B5" colspan="1" pageId="0" pageNumber="1" rowspan="1">10(-11), velutinous</td>
<td id="DFDB93979E0E233189557CFF44BDE709" colspan="1" pageId="0" pageNumber="1" rowspan="1">12, silky velutinous</td>
</tr>
<tr id="795509540802E98CCC611F5196298078" pageId="0" pageNumber="1">
<td id="2279F4A618A136D8969CB100B1840229" colspan="1" pageId="0" pageNumber="1" rowspan="1">Stamen</td>
<td id="857BA1D5E77736B328D119EEC7FA0495" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="51C94B6FC950C6E391F1C5DDECC5574C" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="4584650E78EA8EE4360BDF444532B666" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="D73B4910EE0FB17D7C0CB22815C76823" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="BD91E0CCCB944A74BA2E6E3F4C37AD76" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="C8DB781F0D1C45DED33D6CD7AA1DBF9D" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
</tr>
<tr id="40FE6E7625B4A1AB5900743080A18BFE" pageId="0" pageNumber="1">
<td id="3FFBE09E1088D55972872F5101907ED9" colspan="1" pageId="0" pageNumber="1" rowspan="1">Ovary</td>
<td id="5002B804ED5EEEF7371E36878681749B" colspan="1" pageId="0" pageNumber="1" rowspan="1">3-(4) loculi, ovoid, pubescent</td>
<td id="E9FEC538A28CEFE8A920920DD30B58D8" colspan="1" pageId="0" pageNumber="1" rowspan="1">ovoid, 3 loculi slightly tomentose</td>
<td id="4FE6ACC9A6911709FE06959943321135" colspan="1" pageId="0" pageNumber="1" rowspan="1">globose, 3 loculi, glabrous</td>
<td id="A44976DC839D11266B4B50C30B8811D8" colspan="1" pageId="0" pageNumber="1" rowspan="1">globose, 3 loculi, glabrous</td>
<td id="0EE98314DF907BEC421195822CA76014" colspan="1" pageId="0" pageNumber="1" rowspan="1">3 loculi, ovoid, glabrous</td>
<td id="34547BA5D34DC2BDBDA5C33A8005D76D" colspan="1" pageId="0" pageNumber="1" rowspan="1">globose, 5 loculi, glabrous</td>
</tr>
<tr id="44C9A3AA247A200F8B431A1350433C34" pageId="0" pageNumber="1">
<td id="6CE8861D84E3794ABBC5579C437C0646" colspan="1" pageId="0" pageNumber="1" rowspan="1">Style</td>
<td id="80EB9C095332DE40820BC2DBFA32D5C2" colspan="1" pageId="0" pageNumber="1" rowspan="1">3-(4), free to base, pubescent</td>
<td id="5175F9CB78BEB0B6DB549B6CA9DE32AC" colspan="1" pageId="0" pageNumber="1" rowspan="1">3, free to base, glabrous or sparsely pubescent</td>
<td id="37287486E87EDBA000DF143E26468FD9" colspan="1" pageId="0" pageNumber="1" rowspan="1">3, free to base, glabrous</td>
<td id="0895FA1BF3810CFF4C74A009411488AE" colspan="1" pageId="0" pageNumber="1" rowspan="1">3, free to base, glabrous</td>
<td id="DC4007421F7198860BC05AA5235AF866" colspan="1" pageId="0" pageNumber="1" rowspan="1">3, free to base, glabrous</td>
<td id="AEB2409D6BF78477989CC7C248FED90A" colspan="1" pageId="0" pageNumber="1" rowspan="1">5, united, glabrous</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>