713 lines
97 KiB
XML
713 lines
97 KiB
XML
<document id="F76815E4D8C45DEFF687550C4148FB61" ID-DOI="10.1093/zoolinnean/zlad049" ID-ISSN="0024-4082" ID-Zenodo-Dep="10469936" ID-ZooBank="14A0E12E-1A69-4725-8F99-4747F284C738" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1704717040264" checkinUser="plazi" docAuthor="Gündüz, İslam, Demirtaş, Sadık, Silsüpür, Metin, Özmen, Medine, Polly, P. David & Bilton, David T." docDate="2023" docId="03F787F4FFCF61102F14FC72FBBD0068" docLanguage="en" docName="zlad049.pdf" docOrigin="Zoological Journal of the Linnean Society 199 (3)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad049" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Talpa hakkariensis Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton 2023, sp.nov." docType="treatment" docVersion="4" lastPageNumber="579" masterDocId="FFCEFF8CFFC4611C2E3AFFB4FFB80600" masterDocTitle="Notes from the Anatolian underground: two new mole taxa from Eastern Turkey, together with a revised phylogeny of the genus Talpa (Mammalia: Eulipotyphla: Talpidae)" masterLastPageNumber="593" masterPageNumber="567" pageNumber="578" updateTime="1704893671711" updateUser="ExternalLinkService">
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<mods:title id="3860790B11653B02B0EC8EFAB134CFAC">Notes from the Anatolian underground: two new mole taxa from Eastern Turkey, together with a revised phylogeny of the genus Talpa (Mammalia: Eulipotyphla: Talpidae)</mods:title>
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<mods:namePart id="84C2AB579C309D4B6E343ED1EFF24AA8">Gündüz, İslam</mods:namePart>
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<mods:affiliation id="3F052C957DF60CD795D90A858ED3623B">Department of Biology, Faculty of Sciences, Ondokuz Mayıs University, Samsun, Turkey</mods:affiliation>
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<mods:nameIdentifier id="D658ECE717A7CF2DCDB9DDFDE620F1CE" type="email">gunduzi@omu.edu.tr</mods:nameIdentifier>
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<mods:namePart id="2E14B5D2CE37EAEDEE6CE3218936231C">Demirtaş, Sadık</mods:namePart>
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<mods:affiliation id="573FF0149BF1AA935D74EB1FE01D510D">Department of Biology, Faculty of Sciences, Ondokuz Mayıs University, Samsun, Turkey</mods:affiliation>
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<mods:namePart id="6E35410754162A57647B77EEF07427A6">Silsüpür, Metin</mods:namePart>
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<mods:affiliation id="54E65F9F4515A1C68F58342CF2AD2659">Department of Biology, Faculty of Sciences, Ondokuz Mayıs University, Samsun, Turkey</mods:affiliation>
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<mods:namePart id="BFE76C2818D7ACF8F9BD3FA7581F7737">Özmen, Medine</mods:namePart>
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<mods:affiliation id="A289BE429985CB7E2DF28D360DB29717">Department of Biology, Faculty of Sciences, Ondokuz Mayıs University, Samsun, Turkey</mods:affiliation>
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<mods:namePart id="D25CBB27C0605EB12146324B620AEA75">Polly, P. David</mods:namePart>
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<mods:affiliation id="F647CDAC902254615557E79690608AA3">Department of Earth & Atmospheric Sciences, Indiana University, Bloomington, IN, USA</mods:affiliation>
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<mods:namePart id="F2F6D6FB14E20B4B6A97CEBFE2FA4413">Bilton, David T.</mods:namePart>
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<mods:affiliation id="B4805A372462D4EC798607050F63B4C5">School of Biological and Marine Sciences, University of Plymouth, Drake Circus, Plymouth PL 4 8 AA, Devon, UK & Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, Johannesburg 2006, South Africa</mods:affiliation>
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<mods:date id="08BF95E3E7FA1228C69565A191704F9E">2023</mods:date>
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<emphasis id="B92AEAF0FFCF61172F14FC72FE5405E1" bold="true" box="[302,492,966,993]" italics="true" pageId="11" pageNumber="578">Talpa hakkariensis</emphasis>
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<emphasis id="B92AEAF0FFCF61172E4BFBD4FF4E0277" box="[113,246,1119,1143]" italics="true" pageId="11" pageNumber="578">Type locality:</emphasis>
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,
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,
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yaylası
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NNE
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of
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, 37˚38ʹN, 43˚37ʹE,
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(
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<figureCitation id="13652A67FFCF61172CEAFBCBFFC602B6" captionStart="Figure 1" captionStartId="2.[129,194,1931,1955]" captionTargetBox="[164,1439,1345,1902]" captionTargetId="figure-668@2.[161,1441,1342,1904]" captionTargetPageId="2" captionText="Figure 1. Map of locations (see Table 1) of Talpa moles from Turkey and Iran studied here." figureDoi="http://doi.org/10.5281/zenodo.10469938" httpUri="https://zenodo.org/record/10469938/files/figure.png" pageId="11" pageNumber="578">Figs 1</figureCitation>
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,
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<figureCitation id="13652A67FFCF61172EB3FB2AFF1F02B6" box="[137,167,1182,1206]" captionStart="Figure 5" captionStartId="13.[113,178,1773,1797]" captionTargetBox="[254,1318,145,1745]" captionTargetId="figure-7@13.[254,1319,145,1745]" captionTargetPageId="13" captionText="Figure 5. Freshly trapped Anatolian Talpa in situ, with overviews of habitats at collecting localities.A, T. hakkariensis sp. nov., type locality, Turkey, Hakkari Province, Cemşililan yaylası c. 8 km NNE of Durankaya; B, T. hakkariensis sp. nov., holotype, specimen 1533; C, T. davidiana davidiana, topotypical locality, Turkey, Hatay Province, Hassa, Çardak yaylası; D, T. davidiana davidiana, specimen 1565; E, T.davidiana tatvanensis ssp.nov., type locality, Turkey, Bitlis Province, Uslu, Tatvan c. 1 km E of Kaynarca; F, T.davidiana tatvanensis ssp. nov., holotype, specimen 1530." figureDoi="http://doi.org/10.5281/zenodo.10469946" httpUri="https://zenodo.org/record/10469946/files/figure.png" pageId="11" pageNumber="578">5A</figureCitation>
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;
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<tableCitation id="C6DC0359FFCF61172E8EFB2AFF4702B6" box="[180,255,1182,1206]" captionStart="Table 1" captionStartId="3.[111,166,394,419]" captionTargetPageId="3" captionText="Table 1. Geographic origin of Talpa sampled in this and previous studies, with details on the distribution of haplotypes. The frequencies of haplotypes retrieved from GenBank are stated in parentheses. Cytb haplotypes (Hap.1–20, GenBank accession numbers:OQ695509–OQ695528) and BRCA2 haplotypes (Hap.1–8, GenBank accession numbers: OQ695501–OQ695508) were new to this study. *When we aligned our BRCA2 haplotypes with the shorter (656 bp) published haplotypes, our T. davidiana haplotypes (Hap.4–6) and that (KP717132) in Bannikova et al. (2015) became identical, giving us a single BRCA2 haplotype in T. davidiana in the final dataset used for phylogenetic analyses.Locality codes correspond to Figure 1. See Supporting Information, Table S1 for sequences from other taxa used in phylogenetic analyses.Abbreviations used for collections: OMU-IG/SD, personal collection of İ. Gündüz and S. Demirtaş, Department of Biology, Faculty of Sciences, Ondokuz Mayıs University, Samsun, Turkey; FMNH, Field Museum of Natural History, Chicago, USA. † The last four (OMU-IG/SD collection) or five/six (FMNH collection) digits of the museum collection numbers were used to identify individual specimens in geometric morphometric analysis (e.g. see Supporting Information, Figs S1– S3; Table S3; Files S1, S2)" httpUri="http://table.plazi.org/id/DF21666AFFC7611F2E55FE3EFEDE0466" pageId="11" pageNumber="578" tableUuid="DF21666AFFC7611F2E55FE3EFEDE0466">Table 1</tableCitation>
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)
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.
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<typeStatus id="54E58840FFCF61172E4BFB63FF7402EF" box="[113,204,1239,1263]" pageId="11" pageNumber="578" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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Adult female, specimen no.
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, stuffed skin labelled ‘13/08/21-1533
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IG/SD//
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<emphasis id="B92AEAF0FFCF61172C36FB43FD76030F" box="[524,718,1271,1295]" italics="true" pageId="11" pageNumber="578">Talpa hakkariensis</emphasis>
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<taxonomicNameLabel id="A219578BFFCF61172CE5FB43FF1A032E" pageId="11" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
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// Cemşililan yaylası, Hakkari// 160-31-21 =
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.// 13, August, 2021’. Body frozen at –50˚C, internal organs in ethanol, skull extracted and labelled with specimen number (
|
||
<figureCitation id="13652A67FFCF61172E46FAC0FF7F038C" box="[124,199,1396,1420]" captionStart="Figure 6" captionStartId="14.[130,195,1251,1275]" captionTargetBox="[164,1400,155,1178]" captionTargetId="figure-390@14.[161,1441,144,1223]" captionTargetPageId="14" captionText="Figure 6. Dorsal view of Anatolian Talpa skins. A, T. hakkariensis sp. nov., holotype, specimen 1533; B, topotypical T. davidiana davidiana, specimen 1565; C, T. davidiana tatvanensis ssp.nov., holotype, specimen 1530. Scale bar = 50 mm." figureDoi="http://doi.org/10.5281/zenodo.10469948" httpUri="https://zenodo.org/record/10469948/files/figure.png" pageId="11" pageNumber="578">Fig. 6A</figureCitation>
|
||
). Collected by
|
||
<collectorName id="26AB5334FFCF61172F50FAC1FE6C038C" box="[362,468,1396,1424]" pageId="11" pageNumber="578">İ. Gündüz</collectorName>
|
||
and
|
||
<collectorName id="26AB5334FFCF61172C32FAC0FD3C038C" box="[520,644,1396,1420]" pageId="11" pageNumber="578">S. Demirtaş</collectorName>
|
||
. Sequences available from GenBank: (
|
||
<emphasis id="B92AEAF0FFCF61172FA5FA27FE7603AB" box="[415,462,1427,1451]" italics="true" pageId="11" pageNumber="578">Cytb</emphasis>
|
||
haplotype: Hap.15, accession number
|
||
<accessionNumber id="940DAB01FFCF61172EC7FA07FECF03CB" box="[253,375,1459,1483]" pageId="11" pageNumber="578">OQ695523</accessionNumber>
|
||
;
|
||
<emphasis id="B92AEAF0FFCF61172FB9FA07FE6903CB" box="[387,465,1459,1483]" italics="true" pageId="11" pageNumber="578">BRCA2</emphasis>
|
||
haplotype: Hap.7, accession number
|
||
<accessionNumber id="940DAB01FFCF61172EFDFA66FEFC03EA" box="[199,324,1490,1514]" pageId="11" pageNumber="578">OQ695507</accessionNumber>
|
||
); standard voucher specimens (skin, frozen body, skull, and various tissues in ethanol) are deposited in the Department of Biology, Faculty of Sciences, Ondokuz Mayıs University (
|
||
<collectionCode id="ED4FAE27FFCF61172ED2F984FEC60048" box="[232,382,1584,1608]" pageId="11" pageNumber="578">OMU-IG/SD</collectionCode>
|
||
), Samsun, Turkey.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BE136E2FFCF61172E4BF9DEFA8C047E" blockId="11.[113,763,1641,1979]" lastBlockId="11.[810,1460,144,638]" pageId="11" pageNumber="578">
|
||
<emphasis id="B92AEAF0FFCF61172E4BF9DEFEBC0081" box="[113,260,1641,1665]" italics="true" pageId="11" pageNumber="578">
|
||
<typeStatus id="54E58840FFCF61172E4BF9DEFF680081" box="[113,208,1642,1665]" pageId="11" pageNumber="578" type="paratype">Paratypes</typeStatus>
|
||
(6):
|
||
</emphasis>
|
||
<materialsCitation id="3B363CBFFFCF61172F34F9DDFE5300DF" collectingDate="2021-06-10" collectionCode="IG" location="Bilmizit" municipality="June" pageId="11" pageNumber="578" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D58FD6BFFCF61172F34F9DDFEC10081" box="[270,377,1641,1665]" count="1" pageId="11" pageNumber="578" type="male">One male</specimenCount>
|
||
, specimen no. 1487, stuffed skin labelled ‘10/06/21-1487
|
||
<specimenCount id="9D58FD6BFFCF61172F43F938FE3300A0" box="[377,395,1676,1696]" count="1" pageId="11" pageNumber="578" type="male">♂</specimenCount>
|
||
IG/SD//
|
||
<taxonomicName id="4C5E4D61FFCF61172C37F93DFD7700A1" authority="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton, 2023" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" box="[525,719,1673,1697]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="hakkariensis" status="sp. nov.">
|
||
<emphasis id="B92AEAF0FFCF61172C37F93DFD7700A1" box="[525,719,1673,1697]" italics="true" pageId="11" pageNumber="578">Talpa hakkariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219578BFFCF61172CE5F93DFF1A00C0" pageId="11" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
//
|
||
<location id="8E816039FFCF61172EFDF91CFEA000C0" LSID="urn:lsid:plazi:treatment:03F787F4FFCF61102F14FC72FBBD0068:8E816039FFCF61172EFDF91CFEA000C0" box="[199,280,1704,1728]" municipality="June" name="Bilmizit" pageId="11" pageNumber="578">Bilmizit</location>
|
||
yaylası-Hakkari, 37˚36ʹN 43˚38ʹE// 164- 28-20 =
|
||
<quantity id="4CA69B07FFCF61172EEBF973FE9D00DF" box="[209,293,1735,1759]" metricMagnitude="-2" metricUnit="kg" metricValue="7.305" pageId="11" pageNumber="578" unit="g" value="73.05">73.05 g</quantity>
|
||
.//
|
||
<collectingDate id="EFA4E9CAFFCF61172F70F973FE5300DF" box="[330,491,1735,1759]" pageId="11" pageNumber="578">
|
||
<date id="FFE01022FFCF61172F70F973FE5300DF" box="[330,491,1735,1759]" pageId="11" pageNumber="578" value="2021-06-10">10, June, 2021</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
’;
|
||
<materialsCitation id="3B363CBFFFCF61172FC7F97CFE53013D" collectingDate="2021-06-10" collectionCode="IG" location="Cemsililan" municipality="June" pageId="11" pageNumber="578" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D58FD6BFFCF61172FC7F97CFDD900DF" box="[509,609,1735,1759]" count="1" pageId="11" pageNumber="578" type="male">one male</specimenCount>
|
||
, specimen no. 1488, stuffed skin labelled ‘10/06/21-1488
|
||
<specimenCount id="9D58FD6BFFCF61172C07F95EFDF700FE" box="[573,591,1770,1790]" count="1" pageId="11" pageNumber="578" type="male">♂</specimenCount>
|
||
IG/SD//
|
||
<taxonomicName id="4C5E4D61FFCF61172CF8F953FF52011E" authority="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton, 2023" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="hakkariensis" status="sp. nov.">
|
||
<emphasis id="B92AEAF0FFCF61172CF8F953FF52011E" italics="true" pageId="11" pageNumber="578">Talpa hakkariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219578BFFCF61172EC7F8B2FEE4011E" box="[253,348,1798,1822]" pageId="11" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
//
|
||
<location id="8E816039FFCF61172FBBF8B2FE57011E" LSID="urn:lsid:plazi:treatment:03F787F4FFCF61102F14FC72FBBD0068:8E816039FFCF61172FBBF8B2FE57011E" box="[385,495,1798,1822]" municipality="June" name="Cemsililan" pageId="11" pageNumber="578">Cemşililan</location>
|
||
yaylası-Hakkari// 166- 30-20 =
|
||
<quantity id="4CA69B07FFCF61172EEBF891FE9D013D" box="[209,293,1829,1853]" metricMagnitude="-2" metricUnit="kg" metricValue="7.7379999999999995" pageId="11" pageNumber="578" unit="g" value="77.38">77.38 g</quantity>
|
||
.//
|
||
<collectingDate id="EFA4E9CAFFCF61172F70F891FE53013D" box="[330,491,1829,1853]" pageId="11" pageNumber="578">
|
||
<date id="FFE01022FFCF61172F70F891FE53013D" box="[330,491,1829,1853]" pageId="11" pageNumber="578" value="2021-06-10">10, June, 2021</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
’;
|
||
<materialsCitation id="3B363CBFFFCF61172FC7F892FE53019B" collectingDate="2021-06-10" collectionCode="IG" location="Cemsililan" municipality="June" pageId="11" pageNumber="578" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D58FD6BFFCF61172FC7F892FDD9013D" box="[509,609,1829,1853]" count="1" pageId="11" pageNumber="578" type="male">one male</specimenCount>
|
||
, specimen no. 1489, stuffed skin labelled ‘10/06/21-1489
|
||
<specimenCount id="9D58FD6BFFCF61172C07F8FCFDF7015C" box="[573,591,1864,1884]" count="1" pageId="11" pageNumber="578" type="male">♂</specimenCount>
|
||
IG/SD//
|
||
<taxonomicName id="4C5E4D61FFCF61172CF8F8F1FF52017C" authority="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton, 2023" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="hakkariensis" status="sp. nov.">
|
||
<emphasis id="B92AEAF0FFCF61172CF8F8F1FF52017C" italics="true" pageId="11" pageNumber="578">Talpa hakkariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219578BFFCF61172EC7F8D0FEE4017C" box="[253,348,1892,1916]" pageId="11" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
//
|
||
<location id="8E816039FFCF61172FBBF8D0FE57017C" LSID="urn:lsid:plazi:treatment:03F787F4FFCF61102F14FC72FBBD0068:8E816039FFCF61172FBBF8D0FE57017C" box="[385,495,1892,1916]" municipality="June" name="Cemsililan" pageId="11" pageNumber="578">Cemşililan</location>
|
||
yaylası-Hakkari// 170- 31-21 =
|
||
<quantity id="4CA69B07FFCF61172EEBF837FE9D019B" box="[209,293,1923,1947]" metricMagnitude="-2" metricUnit="kg" metricValue="7.125" pageId="11" pageNumber="578" unit="g" value="71.25">71.25 g</quantity>
|
||
.//
|
||
<collectingDate id="EFA4E9CAFFCF61172F70F837FE53019B" box="[330,491,1923,1947]" pageId="11" pageNumber="578">
|
||
<date id="FFE01022FFCF61172F70F837FE53019B" box="[330,491,1923,1947]" pageId="11" pageNumber="578" value="2021-06-10">10, June, 2021</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
’;
|
||
<materialsCitation id="3B363CBFFFCF61172FC7F830FB3306C7" collectingDate="2021-06-10" collectionCode="IG" location="Cemsililan" municipality="June" pageId="11" pageNumber="578" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D58FD6BFFCF61172FC7F830FDD9019B" box="[509,609,1923,1947]" count="1" pageId="11" pageNumber="578" type="male">one male</specimenCount>
|
||
, specimen no. 1490, stuffed skin labelled ‘10/6/21-1490
|
||
<specimenCount id="9D58FD6BFFCF61172C03F812FDF301BA" box="[569,587,1958,1978]" count="1" pageId="11" pageNumber="578" type="male">♂</specimenCount>
|
||
IG/SD//
|
||
<taxonomicName id="4C5E4D61FFCF61172CF8F817FC1B06A8" authority="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton, 2023" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" box="[706,931,144,1979]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="hakkariensis" status="sp. nov.">
|
||
<emphasis id="B92AEAF0FFCF61172CF8F817FC1B06A8" box="[706,931,144,1979]" italics="true" pageId="11" pageNumber="578">Talpa hakkariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219578BFFCF61172D95FF24FBB006A8" box="[943,1032,144,168]" pageId="11" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
//
|
||
<location id="8E816039FFCF61172A1CFF24FB2C06A8" LSID="urn:lsid:plazi:treatment:03F787F4FFCF61102F14FC72FBBD0068:8E816039FFCF61172A1CFF24FB2C06A8" box="[1062,1172,144,168]" municipality="June" name="Cemsililan" pageId="11" pageNumber="578">Cemşililan</location>
|
||
yaylası-Hakkari// 165-29- 21 =
|
||
<quantity id="4CA69B07FFCF61172D51FF1BFC7806C7" box="[875,960,175,199]" metricMagnitude="-2" metricUnit="kg" metricValue="6.795999999999999" pageId="11" pageNumber="578" unit="g" value="67.96">67.96 g</quantity>
|
||
.//
|
||
<collectingDate id="EFA4E9CAFFCF61172DDDFF1BFB3306C7" box="[999,1163,175,199]" pageId="11" pageNumber="578">
|
||
<date id="FFE01022FFCF61172DDDFF1BFB3306C7" box="[999,1163,175,199]" pageId="11" pageNumber="578" value="2021-06-10">10, June, 2021</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
’;
|
||
<materialsCitation id="3B363CBFFFCF61172AA5FF04FB2D0725" collectingDate="2021-08-11" collectionCode="IG" location="Cemsililan" municipality="August" pageId="11" pageNumber="578" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D58FD6BFFCF61172AA5FF04FAAE06C7" box="[1183,1302,175,199]" count="1" pageId="11" pageNumber="578" type="female">one female</specimenCount>
|
||
, specimen no. 1531, stuffed skin labelled ‘11/08/21-1531
|
||
<specimenCount id="9D58FD6BFFCF61172ACDFF66FABF06E6" box="[1271,1287,210,230]" count="1" pageId="11" pageNumber="578" type="female">♀</specimenCount>
|
||
IG/SD//
|
||
<taxonomicName id="4C5E4D61FFCF61172B40FF7AFC1B0706" authority="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton, 2023" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="hakkariensis" status="sp. nov.">
|
||
<emphasis id="B92AEAF0FFCF61172B40FF7AFC1B0706" italics="true" pageId="11" pageNumber="578">Talpa hakkariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219578BFFCF61172D95FF5AFBB00706" box="[943,1032,238,262]" pageId="11" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
//
|
||
<location id="8E816039FFCF61172A1CFF5AFB2C0706" LSID="urn:lsid:plazi:treatment:03F787F4FFCF61102F14FC72FBBD0068:8E816039FFCF61172A1CFF5AFB2C0706" box="[1062,1172,238,262]" municipality="August" name="Cemsililan" pageId="11" pageNumber="578">Cemşililan</location>
|
||
yaylası-Hakkari// 147-33- 21 =
|
||
<quantity id="4CA69B07FFCF61172D5FFEB9FC000725" box="[869,952,269,293]" metricMagnitude="-2" metricUnit="kg" metricValue="6.034" pageId="11" pageNumber="578" unit="g" value="60.34">60.34 g</quantity>
|
||
.//
|
||
<collectingDate id="EFA4E9CAFFCF61172DE6FEB9FB2D0725" box="[988,1173,269,293]" pageId="11" pageNumber="578">
|
||
<date id="FFE01022FFCF61172DE6FEB9FB2D0725" box="[988,1173,269,293]" pageId="11" pageNumber="578" value="2021-08-11">11, August, 2021</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
’;
|
||
<materialsCitation id="3B363CBFFFCF61172A9DFEBAFC4C07C2" collectingDate="2021-08-12" collectionCode="OMU-IG/SD" country="Turkey" county="Faculty of Sciences" location="Cemsililan" municipality="Department of Biology" pageId="11" pageNumber="578" specimenCode="BRCA2" specimenCount="1" specimenCount-female="1" stateProvince="Samsun" typeStatus="paratype">
|
||
<specimenCount id="9D58FD6BFFCF61172A9DFEBAFAA40725" box="[1191,1308,269,293]" count="1" pageId="11" pageNumber="578" type="female">one female</specimenCount>
|
||
, specimen no. 1532, stuffed skin labelled ‘12/08/21-1532
|
||
<specimenCount id="9D58FD6BFFCF61172ACDFE84FABF0744" box="[1271,1287,304,324]" count="1" pageId="11" pageNumber="578" type="female">♀</specimenCount>
|
||
IG/SD//
|
||
<taxonomicName id="4C5E4D61FFCF61172B40FE98FC1B0764" authority="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton, 2023" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="hakkariensis" status="sp. nov.">
|
||
<emphasis id="B92AEAF0FFCF61172B40FE98FC1B0764" italics="true" pageId="11" pageNumber="578">Talpa hakkariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219578BFFCF61172D95FEF8FBB00764" box="[943,1032,332,356]" pageId="11" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
//
|
||
<location id="8E816039FFCF61172A1CFEF8FB2C0764" LSID="urn:lsid:plazi:treatment:03F787F4FFCF61102F14FC72FBBD0068:8E816039FFCF61172A1CFEF8FB2C0764" box="[1062,1172,332,356]" country="Turkey" county="Faculty of Sciences" municipality="Department of Biology" name="Cemsililan" pageId="11" pageNumber="578" stateProvince="Samsun">Cemşililan</location>
|
||
yaylası-Hakkari// 154-30- 21 =
|
||
<quantity id="4CA69B07FFCF61172D58FEDFFC0B0783" box="[866,947,363,387]" metricMagnitude="-2" metricUnit="kg" metricValue="6.739" pageId="11" pageNumber="578" unit="g" value="67.39">67.39 g</quantity>
|
||
.//
|
||
<date id="FFE01022FFCF61172DEFFEDFFB3E0783" box="[981,1158,363,387]" pageId="11" pageNumber="578" value="2021-08-12">
|
||
<collectingDate id="EFA4E9CAFFCF61172DEFFEDFFB3E0783" box="[981,1158,363,387]" pageId="11" pageNumber="578">12. August, 2021</collectingDate>
|
||
</date>
|
||
’. Bodies frozen at –50˚C, internal organs in ethanol, skulls extracted and labelled with specimen numbers.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B363CBFFFCF61172DC1FE1EFA88047E" acessionNumber="OQ365519, OQ365520, OQ365521, OQ365522, OQ695507, OQ695508" country="Turkey" county="Faculty of Sciences" location="Cemsililan" municipality="Department of Biology" pageId="11" pageNumber="578" specimenCode="BRCA2" specimenCount="6" stateProvince="Samsun" typeStatus="paratype">
|
||
Sequences available from GenBank: (
|
||
<emphasis id="B92AEAF0FFCF61172BBEFE1EFA0B07C2" box="[1412,1459,426,450]" italics="true" pageId="11" pageNumber="578">Cytb</emphasis>
|
||
haplotypes: Hap.11–14, accession numbers
|
||
<accessionNumber id="940DAB01FFCF61172B14FE7DFA1C07E1" box="[1326,1444,457,481]" pageId="11" pageNumber="578">OQ695519</accessionNumber>
|
||
–
|
||
<accessionNumber id="940DAB01FFCF61172D10FE5CFC1C0400" box="[810,932,488,512]" pageId="11" pageNumber="578">OQ695522</accessionNumber>
|
||
;
|
||
<emphasis id="B92AEAF0FFCF61172D8BFE5CFC470400" box="[945,1023,488,512]" italics="true" pageId="11" pageNumber="578">BRCA2</emphasis>
|
||
haplotypes: Hap.7–8, accession numbers
|
||
<accessionNumber id="940DAB01FFCF61172D10FDBCFC1C0420" box="[810,932,520,544]" pageId="11" pageNumber="578">OQ695507</accessionNumber>
|
||
–
|
||
<accessionNumber id="940DAB01FFCF61172D89FDBCFB940420" box="[947,1068,520,544]" pageId="11" pageNumber="578">OQ695508</accessionNumber>
|
||
); standard voucher specimens (skins, frozen body, skulls, and various tissues in ethanol) are deposited in the Department of Biology, Faculty of Sciences, Ondokuz Mayıs University (
|
||
<collectionCode id="ED4FAE27FFCF61172DDEFDD2FBC2047E" box="[996,1146,614,638]" pageId="11" pageNumber="578">OMU-IG/SD</collectionCode>
|
||
), Samsun, Turkey
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3446569FFCF61172DCFFD12FA1D0320" pageId="11" pageNumber="578" type="description">
|
||
<paragraph id="8BE136E2FFCF61172DCFFD12FB5F04C0" blockId="11.[1013,1255,678,704]" box="[1013,1255,678,704]" pageId="11" pageNumber="578">
|
||
<emphasis id="B92AEAF0FFCF61172DCFFD12FB5F04C0" box="[1013,1255,678,704]" italics="true" pageId="11" pageNumber="578">
|
||
Measurements of
|
||
<typeStatus id="54E58840FFCF61172AADFD12FB5F04C0" box="[1175,1255,678,704]" pageId="11" pageNumber="578" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE136E2FFCF61172D10FD78FB260581" blockId="11.[810,1459,716,897]" pageId="11" pageNumber="578">
|
||
Body mass
|
||
<quantity id="4CA69B07FFCF61172D98FD78FC4804E5" box="[930,1008,716,741]" metricMagnitude="-2" metricUnit="kg" metricValue="7.384" pageId="11" pageNumber="578" unit="g" value="73.84">73.84 g</quantity>
|
||
, head and body length
|
||
<quantity id="4CA69B07FFCF61172ADCFD78FA8704E4" box="[1254,1343,716,740]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="578" unit="mm" value="160.0">160 mm</quantity>
|
||
, tail length
|
||
<quantity id="4CA69B07FFCF61172D10FD58FCC00503" box="[810,888,748,772]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="11" pageNumber="578" unit="mm" value="31.0">31 mm</quantity>
|
||
, hindfoot length
|
||
<quantity id="4CA69B07FFCF61172A0BFD58FBC70503" box="[1073,1151,748,772]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="578" unit="mm" value="21.0">21 mm</quantity>
|
||
, condylobasal length of skull
|
||
<quantity id="4CA69B07FFCF61172D10FCBFFC2D0522" box="[810,917,779,803]" metricMagnitude="-2" metricUnit="m" metricValue="3.34" pageId="11" pageNumber="578" unit="mm" value="33.4">33.40 mm</quantity>
|
||
, maxillary tooth row
|
||
<quantity id="4CA69B07FFCF61172A55FCBFFB620522" box="[1135,1242,779,803]" metricMagnitude="-2" metricUnit="m" metricValue="1.4929999999999999" pageId="11" pageNumber="578" unit="mm" value="14.93">14.93 mm</quantity>
|
||
, breadth of braincase
|
||
<quantity id="4CA69B07FFCF61172D10FC9EFC210542" box="[810,921,810,834]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="11" pageNumber="578" unit="mm" value="16.9">16.90 mm</quantity>
|
||
, height of braincase
|
||
<quantity id="4CA69B07FFCF61172A40FC9EFB630542" box="[1146,1243,810,834]" metricMagnitude="-3" metricUnit="m" metricValue="9.29" pageId="11" pageNumber="578" unit="mm" value="9.29">9.29 mm</quantity>
|
||
, breadth of rostrum over canines
|
||
<quantity id="4CA69B07FFCF61172D94FCFDFBB30561" box="[942,1035,841,865]" metricMagnitude="-3" metricUnit="m" metricValue="5.02" pageId="11" pageNumber="578" unit="mm" value="5.02">5.02 mm</quantity>
|
||
, breadth of rostrum over molars
|
||
<quantity id="4CA69B07FFCF61172B62FCFDFA0B0561" box="[1368,1459,841,865]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="11" pageNumber="578" unit="mm" value="9.3">9.30 mm</quantity>
|
||
(Supporting Information, Table S3).
|
||
</paragraph>
|
||
<paragraph id="8BE136E2FFCF61172DCAFC1DFB5605C2" blockId="11.[1008,1262,936,962]" box="[1008,1262,936,962]" pageId="11" pageNumber="578">
|
||
<emphasis id="B92AEAF0FFCF61172DCAFC1DFB5605C2" box="[1008,1262,936,962]" italics="true" pageId="11" pageNumber="578">
|
||
Measurements of
|
||
<typeStatus id="54E58840FFCF61172AA8FC1DFB5605C2" box="[1170,1262,937,962]" pageId="11" pageNumber="578" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE136E2FFCF61172D10FC7BFA1D0320" blockId="11.[810,1459,975,1312]" pageId="11" pageNumber="578">Means ± standard deviation (minimum–maximum). Linear measurements are in millimetres; body mass in grams. Sample size = 6 for all measurements. Body mass 69.56 ± 5.30 (60.34– 77.38), head and body length 161.00 ± 7.920 (147.0–170.0), tail length 30.17 ± 1.570 (28.0–33.0), hindfoot 20.67 ± 0.471 (20.0–21.0), condylobasal length of skull 32.88 ± 0.421 (32.10– 33.24), maxillary tooth row 14.15 ± 0.142 (13.88–14.31), breadth of braincase 16.30 ± 0.348 (15.8–16.88), height of braincase 9.31 ± 0.102 (9.20–9.48), breadth of rostrum over canines 5.24 ± 0.279 (4.88–5.59), breadth of rostrum over molars 9.86 ± 0.442 (9.24–10.4) (Supporting Information, Table S3).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3446569FFCF61102A7AFAFDFD060579" lastPageId="12" lastPageNumber="579" pageId="11" pageNumber="578" type="diagnosis">
|
||
<paragraph id="8BE136E2FFCF61172A7AFAFDFB250362" blockId="11.[1088,1181,1353,1378]" box="[1088,1181,1353,1378]" pageId="11" pageNumber="578">
|
||
<emphasis id="B92AEAF0FFCF61172A7AFAFDFB250362" box="[1088,1181,1353,1378]" italics="true" pageId="11" pageNumber="578">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE136E2FFCF61102D10FADBFD060579" blockId="11.[810,1462,1390,1978]" lastBlockId="12.[129,780,144,889]" lastPageId="12" lastPageNumber="579" pageId="11" pageNumber="578">
|
||
Very large mole; skull large with a stout, broad rostrum; dental formula of C3/3 I1/1 P4/4 M3/3. A member of the subgenus
|
||
<taxonomicName id="4C5E4D61FFCF61172D10FA19FCDB03C5" box="[810,867,1453,1477]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92AEAF0FFCF61172D10FA19FCDB03C5" box="[810,867,1453,1477]" italics="true" pageId="11" pageNumber="578">Talpa</emphasis>
|
||
</taxonomicName>
|
||
, closest to
|
||
<taxonomicName id="4C5E4D61FFCF61172DDAFA1AFBE603C5" box="[992,1118,1453,1477]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="davidiana">
|
||
<emphasis id="B92AEAF0FFCF61172DDAFA1AFBE603C5" box="[992,1118,1453,1477]" italics="true" pageId="11" pageNumber="578">T. davidiana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5E4D61FFCF61172AA2FA1AFAA503C5" baseAuthorityName="Krystufek and Vohralik" baseAuthorityYear="2005" box="[1176,1309,1454,1477]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="streetorum">
|
||
<emphasis id="B92AEAF0FFCF61172AA2FA1AFAA503C5" box="[1176,1309,1454,1477]" italics="true" pageId="11" pageNumber="578">T. streetorum</emphasis>
|
||
</taxonomicName>
|
||
morphologically and
|
||
<taxonomicName id="4C5E4D61FFCF61172DBCFA79FBBC03E4" box="[902,1028,1484,1508]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="davidiana">
|
||
<emphasis id="B92AEAF0FFCF61172DBCFA79FBBC03E4" box="[902,1028,1484,1508]" italics="true" pageId="11" pageNumber="578">T. davidiana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5E4D61FFCF61172A06FA79FB7003E4" authorityName="Vereschagin" authorityYear="1945" box="[1084,1224,1484,1508]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="talyschensis">
|
||
<emphasis id="B92AEAF0FFCF61172A06FA79FB7003E4" box="[1084,1224,1484,1508]" italics="true" pageId="11" pageNumber="578">T. talyschensis</emphasis>
|
||
</taxonomicName>
|
||
genetically (
|
||
<figureCitation id="13652A67FFCF61172B68FA78FA2A03E4" box="[1362,1426,1484,1508]" captionStart="Figure2" captionStartId="8.[129,192,1856,1880]" captionTargetBox="[428,1140,145,1827]" captionTargetId="graphics-10@8.[428,995,182,1813]" captionTargetPageId="8" captionText="Figure2. Results of BI, ML,and MP analyses combined on a BI tree based on the complete Cytb gene sequences ofTalpa species.ML and MP bootstrap values greater than 70% and posterior probabilities (PPs)greater than 0.80 are shown at nodes.Haplotypes Hap.1–20 are from this study. Species assignments from one distance-based (ASAP) and two tree-based (GMYC, mPTP) delimitation methods are represented by vertical bars onthe right side of the phylogeny.The black colour on the bars signifies that thereis agreement across delimitation approaches,whereas grey colour indicatesdisagreement.For geographical origins of the published sequences, see Table 1 and Supporting Information,Table S1." figureDoi="http://doi.org/10.5281/zenodo.10469940" httpUri="https://zenodo.org/record/10469940/files/figure.png" pageId="11" pageNumber="578">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="13652A67FFCF61172B9AFA78FA1503E4" box="[1440,1453,1484,1508]" captionStart="Figure 3" captionStartId="9.[113,178,1772,1796]" captionTargetBox="[317,1256,144,1743]" captionTargetId="graphics-89@9.[317,1108,144,1725]" captionTargetPageId="9" captionText="Figure 3. Results of BI, ML, and MP analyses combined on a ML tree based on BRCA2 sequences of Talpa and outgroup species.ML and MP bootstrap values greater than 70% and posterior probabilities greater than 0.80 are shown at nodes. Haplotypes Hap.1–8 are from this study. For the geographical origins of the published sequences, see Table 1 and Supporting Information, Table S1." figureDoi="http://doi.org/10.5281/zenodo.10469942" httpUri="https://zenodo.org/record/10469942/files/figure.png" pageId="11" pageNumber="578">3</figureCitation>
|
||
,
|
||
<figureCitation id="13652A67FFCF61172D10FA5FFC8F0003" box="[810,823,1515,1539]" captionStart="Figure 6" captionStartId="14.[130,195,1251,1275]" captionTargetBox="[164,1400,155,1178]" captionTargetId="figure-390@14.[161,1441,144,1223]" captionTargetPageId="14" captionText="Figure 6. Dorsal view of Anatolian Talpa skins. A, T. hakkariensis sp. nov., holotype, specimen 1533; B, topotypical T. davidiana davidiana, specimen 1565; C, T. davidiana tatvanensis ssp.nov., holotype, specimen 1530. Scale bar = 50 mm." figureDoi="http://doi.org/10.5281/zenodo.10469948" httpUri="https://zenodo.org/record/10469948/files/figure.png" pageId="11" pageNumber="578">6</figureCitation>
|
||
). Distinguished from both
|
||
<taxonomicName id="4C5E4D61FFCF61172A6DFA58FB6B0003" box="[1111,1235,1515,1539]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="davidiana">
|
||
<emphasis id="B92AEAF0FFCF61172A6DFA58FB6B0003" box="[1111,1235,1515,1539]" italics="true" pageId="11" pageNumber="578">T. davidiana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5E4D61FFCF61172B33FA58FA2B0003" authorityName="Vereschagin" authorityYear="1945" box="[1289,1427,1515,1539]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="talyschensis">
|
||
<emphasis id="B92AEAF0FFCF61172B33FA58FA2B0003" box="[1289,1427,1515,1539]" italics="true" pageId="11" pageNumber="578">T. talyschensis</emphasis>
|
||
</taxonomicName>
|
||
by the larger body size [
|
||
<taxonomicName id="4C5E4D61FFCF61172A3DF9B8FB230023" authority="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton, 2023" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" box="[1031,1179,1547,1571]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="hakkariensis" status="sp. nov.">
|
||
<emphasis id="B92AEAF0FFCF61172A3DF9B8FB230023" box="[1031,1179,1547,1571]" italics="true" pageId="11" pageNumber="578">T. hakkariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219578BFFCF61172A98F9BFFB480022" box="[1186,1264,1547,1570]" pageId="11" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
body mass
|
||
<quantity id="4CA69B07FFCF61172B56F9BFFCC60043" metricMagnitude="-2" metricUnit="kg" metricValue="6.886" metricValueMax="7.7379999999999995" metricValueMin="6.034" pageId="11" pageNumber="578" unit="g" value="68.86" valueMax="77.38" valueMin="60.34">60.34– 77.38 g</quantity>
|
||
, head and body length
|
||
<quantity id="4CA69B07FFCF61172AB7F99EFA990042" box="[1165,1313,1578,1602]" metricMagnitude="-1" metricUnit="m" metricValue="1.585" metricValueMax="1.7" metricValueMin="1.47" pageId="11" pageNumber="578" unit="mm" value="158.5" valueMax="170.0" valueMin="147.0">147–170 mm</quantity>
|
||
;
|
||
<taxonomicName id="4C5E4D61FFCF61172B08F99FFA0A0042" box="[1330,1458,1578,1602]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="davidiana">
|
||
<emphasis id="B92AEAF0FFCF61172B08F99FFA0A0042" box="[1330,1458,1578,1602]" italics="true" pageId="11" pageNumber="578">T. davidiana</emphasis>
|
||
</taxonomicName>
|
||
body mass
|
||
<quantity id="4CA69B07FFCF61172DA7F9FDFB8A0062" box="[925,1074,1609,1634]" metricMagnitude="-2" metricUnit="kg" metricValue="4.3495" metricValueMax="5.287" metricValueMin="3.412" pageId="11" pageNumber="578" unit="g" value="43.495" valueMax="52.87" valueMin="34.12">34.12–52.87 g</quantity>
|
||
, head and body length
|
||
<quantity id="4CA69B07FFCF61172B1BF9FDFA160061" box="[1313,1454,1609,1633]" metricMagnitude="-1" metricUnit="m" metricValue="1.405" metricValueMax="1.53" metricValueMin="1.28" pageId="11" pageNumber="578" unit="mm" value="140.5" valueMax="153.0" valueMin="128.0">128–153 mm</quantity>
|
||
;
|
||
<taxonomicName id="4C5E4D61FFCF61172D10F9DEFC030081" authorityName="Vereschagin" authorityYear="1945" box="[810,955,1641,1665]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="talyschensis">
|
||
<emphasis id="B92AEAF0FFCF61172D10F9DEFC030081" box="[810,955,1641,1665]" italics="true" pageId="11" pageNumber="578">T. talyschensis</emphasis>
|
||
</taxonomicName>
|
||
body mass
|
||
<quantity id="4CA69B07FFCF61172A76F9DDFB110081" box="[1100,1193,1641,1665]" metricMagnitude="-2" metricUnit="kg" metricValue="4.0" metricValueMax="4.9" metricValueMin="3.1" pageId="11" pageNumber="578" unit="g" value="40.0" valueMax="49.0" valueMin="31.0">31–49 g</quantity>
|
||
, head and body length
|
||
<quantity id="4CA69B07FFCF61172D10F93CFC0400A0" box="[810,956,1672,1696]" metricMagnitude="-1" metricUnit="m" metricValue="1.09" metricValueMax="1.14" metricValueMin="1.04" pageId="11" pageNumber="578" unit="mm" value="109.0" valueMax="114.0" valueMin="104.0">104–114 mm</quantity>
|
||
; (
|
||
<figureCitation id="13652A67FFCF61172DEFF93CFB9A00A0" box="[981,1058,1672,1696]" captionStart="Figure 6" captionStartId="14.[130,195,1251,1275]" captionTargetBox="[164,1400,155,1178]" captionTargetId="figure-390@14.[161,1441,144,1223]" captionTargetPageId="14" captionText="Figure 6. Dorsal view of Anatolian Talpa skins. A, T. hakkariensis sp. nov., holotype, specimen 1533; B, topotypical T. davidiana davidiana, specimen 1565; C, T. davidiana tatvanensis ssp.nov., holotype, specimen 1530. Scale bar = 50 mm." figureDoi="http://doi.org/10.5281/zenodo.10469948" httpUri="https://zenodo.org/record/10469948/files/figure.png" pageId="11" pageNumber="578">Fig. 6A</figureCitation>
|
||
)]. Condylobasal length of skull in
|
||
<taxonomicName id="4C5E4D61FFCF61172BA5F93DFC1B00BF" authority="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton, 2023" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="hakkariensis" status="sp. nov.">
|
||
<emphasis id="B92AEAF0FFCF61172BA5F93DFC1B00BF" italics="true" pageId="11" pageNumber="578">T. hakkariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219578BFFCF61172D90F91CFC4000BF" box="[938,1016,1704,1727]" pageId="11" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
also consistently greater,
|
||
<quantity id="4CA69B07FFCF61172B38F91CFA0B00BF" box="[1282,1459,1704,1728]" metricMagnitude="-2" metricUnit="m" metricValue="3.275" metricValueMax="3.34" metricValueMin="3.21" pageId="11" pageNumber="578" unit="mm" value="32.75" valueMax="33.4" valueMin="32.1">32.10–33.40 mm</quantity>
|
||
vs.
|
||
<quantity id="4CA69B07FFCF61172D75F973FC5600DF" box="[847,1006,1735,1759]" metricMagnitude="-1" metricUnit="m" metricValue="7.65937" metricValueMax="7.9425799999999995" metricValueMin="7.37616" pageId="11" pageNumber="578" unit="in" value="30.155" valueMax="31.27" valueMin="29.04">29.04–31.27 in</quantity>
|
||
<taxonomicName id="4C5E4D61FFCF61172DC3F973FBC000DF" box="[1017,1144,1735,1759]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="davidiana">
|
||
<emphasis id="B92AEAF0FFCF61172DC3F973FBC000DF" box="[1017,1144,1735,1759]" italics="true" pageId="11" pageNumber="578">T. davidiana</emphasis>
|
||
</taxonomicName>
|
||
studied here, and a mean of
|
||
<quantity id="4CA69B07FFCF61172D10F952FC2900FE" box="[810,913,1766,1790]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" pageId="11" pageNumber="578" unit="mm" value="31.1">31.10 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C5E4D61FFCF61172D95F953FABE00FE" authority="(Zaitsev et al. 2014)" baseAuthorityName="Zaitsev" baseAuthorityYear="2014" box="[943,1286,1766,1790]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="talyschensis">
|
||
<emphasis id="B92AEAF0FFCF61172D95F953FB8E00FE" box="[943,1078,1766,1790]" italics="true" pageId="11" pageNumber="578">T. talyschensis</emphasis>
|
||
(Zaitsev
|
||
<emphasis id="B92AEAF0FFCF61172AAEF953FB0400FE" box="[1172,1212,1766,1790]" italics="true" pageId="11" pageNumber="578">et al</emphasis>
|
||
. 2014)
|
||
</taxonomicName>
|
||
. The new species also differs from
|
||
<taxonomicName id="4C5E4D61FFCF61172DECF8B2FB00011D" baseAuthorityName="Milne-Edwards" baseAuthorityYear="1884" box="[982,1208,1797,1821]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="subSpecies" species="davidiana" subSpecies="davidiana">
|
||
<emphasis id="B92AEAF0FFCF61172DECF8B2FB00011D" box="[982,1208,1797,1821]" italics="true" pageId="11" pageNumber="578">T. davidiana davidiana</emphasis>
|
||
</taxonomicName>
|
||
on: the broader, blunter rostrum (
|
||
<quantity id="4CA69B07FFCF61172DB1F891FB8B013C" box="[907,1075,1829,1853]" metricMagnitude="-3" metricUnit="m" metricValue="9.82" metricValueMax="10.4" metricValueMin="9.24" pageId="11" pageNumber="578" unit="mm" value="9.82" valueMax="10.4" valueMin="9.24">9.24–10.40 mm</quantity>
|
||
over the molars in
|
||
<taxonomicName id="4C5E4D61FFCF61172AC7F891FA29013D" authority="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton, 2023" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" box="[1277,1425,1829,1853]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="hakkariensis" status="sp. nov.">
|
||
<emphasis id="B92AEAF0FFCF61172AC7F891FA29013D" box="[1277,1425,1829,1853]" italics="true" pageId="11" pageNumber="578">T. hakkariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219578BFFCF61172BA2F891FCED015C" pageId="11" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
vs.
|
||
<quantity id="4CA69B07FFCF61172D43F8F0FBB5015C" box="[889,1037,1860,1884]" metricMagnitude="-3" metricUnit="m" metricValue="9.11" metricValueMax="9.87" metricValueMin="8.35" pageId="11" pageNumber="578" unit="mm" value="9.11" valueMax="9.87" valueMin="8.35">8.35–9.87 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C5E4D61FFCF61172A11F8F1FB1C015C" box="[1067,1188,1860,1884]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="species" species="davidiana">
|
||
<emphasis id="B92AEAF0FFCF61172A11F8F1FB1C015C" box="[1067,1188,1860,1884]" italics="true" pageId="11" pageNumber="578">T. davidiana</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="13652A67FFCF61172A8BF8F0FAAC015C" box="[1201,1300,1860,1884]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStart-2="Figure 9" captionStart-3="Figure 10" captionStartId-0="15.[113,178,1554,1578]" captionStartId-1="16.[130,195,1568,1592]" captionStartId-2="17.[113,178,1592,1616]" captionStartId-3="18.[129,194,1594,1618]" captionTargetBox-0="[308,1263,146,1523]" captionTargetBox-1="[323,1278,146,1537]" captionTargetBox-2="[308,1263,146,1561]" captionTargetBox-3="[324,1278,147,1564]" captionTargetId-0="figure-231@15.[306,1266,144,1526]" captionTargetId-1="figure-219@16.[321,1281,144,1541]" captionTargetId-2="figure-141@17.[306,1266,144,1564]" captionTargetId-3="figure-122@18.[321,1281,144,1567]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionText-0="Figure 7. Talpa hakkariensis sp. nov., holotype, specimen 1533, cranium in (A) lateral, (B) ventral, and (C) dorsal views.Scale bar = 10 mm." captionText-1="Figure 8. Topotypical Talpa davidiana davidiana, specimen 1565, cranium in (A) lateral, (B) ventral, and (C) dorsal views.Scale bar = 10 mm." captionText-2="Figure 9. Talpa davidiana tatvanensis ssp. nov., holotype, specimen 1530, cranium in (A) lateral, (B) ventral, and (C) dorsal views.Scale bar = 10 mm." captionText-3="Figure 10. Talpa streetorum.Holotype, Field Museum of Natural History, Chicago, specimen 96424, cranium in (A) lateral, (B) ventral, and (C) dorsal views.Scale bar = 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10469950" figureDoi-1="http://doi.org/10.5281/zenodo.10469952" figureDoi-2="http://doi.org/10.5281/zenodo.10469954" figureDoi-3="http://doi.org/10.5281/zenodo.10469956" httpUri-0="https://zenodo.org/record/10469950/files/figure.png" httpUri-1="https://zenodo.org/record/10469952/files/figure.png" httpUri-2="https://zenodo.org/record/10469954/files/figure.png" httpUri-3="https://zenodo.org/record/10469956/files/figure.png" pageId="11" pageNumber="578">Figs 7–10</figureCitation>
|
||
); typically four rather than three premolars in each maxilla (
|
||
<figureCitation id="13652A67FFCF61172AD1F8D7FAF6017B" box="[1259,1358,1891,1915]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStart-2="Figure 9" captionStart-3="Figure 10" captionStartId-0="15.[113,178,1554,1578]" captionStartId-1="16.[130,195,1568,1592]" captionStartId-2="17.[113,178,1592,1616]" captionStartId-3="18.[129,194,1594,1618]" captionTargetBox-0="[308,1263,146,1523]" captionTargetBox-1="[323,1278,146,1537]" captionTargetBox-2="[308,1263,146,1561]" captionTargetBox-3="[324,1278,147,1564]" captionTargetId-0="figure-231@15.[306,1266,144,1526]" captionTargetId-1="figure-219@16.[321,1281,144,1541]" captionTargetId-2="figure-141@17.[306,1266,144,1564]" captionTargetId-3="figure-122@18.[321,1281,144,1567]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionText-0="Figure 7. Talpa hakkariensis sp. nov., holotype, specimen 1533, cranium in (A) lateral, (B) ventral, and (C) dorsal views.Scale bar = 10 mm." captionText-1="Figure 8. Topotypical Talpa davidiana davidiana, specimen 1565, cranium in (A) lateral, (B) ventral, and (C) dorsal views.Scale bar = 10 mm." captionText-2="Figure 9. Talpa davidiana tatvanensis ssp. nov., holotype, specimen 1530, cranium in (A) lateral, (B) ventral, and (C) dorsal views.Scale bar = 10 mm." captionText-3="Figure 10. Talpa streetorum.Holotype, Field Museum of Natural History, Chicago, specimen 96424, cranium in (A) lateral, (B) ventral, and (C) dorsal views.Scale bar = 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10469950" figureDoi-1="http://doi.org/10.5281/zenodo.10469952" figureDoi-2="http://doi.org/10.5281/zenodo.10469954" figureDoi-3="http://doi.org/10.5281/zenodo.10469956" httpUri-0="https://zenodo.org/record/10469950/files/figure.png" httpUri-1="https://zenodo.org/record/10469952/files/figure.png" httpUri-2="https://zenodo.org/record/10469954/files/figure.png" httpUri-3="https://zenodo.org/record/10469956/files/figure.png" pageId="11" pageNumber="578">Figs 7–10</figureCitation>
|
||
; only one of the nine
|
||
<taxonomicName id="4C5E4D61FFCF61172D98F837FB3E019B" baseAuthorityName="Milne-Edwards" baseAuthorityYear="1884" box="[930,1158,1923,1947]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="578" phylum="Chordata" rank="subSpecies" species="davidiana" subSpecies="davidiana">
|
||
<emphasis id="B92AEAF0FFCF61172D98F837FB3E019B" box="[930,1158,1923,1947]" italics="true" pageId="11" pageNumber="578">T. davidiana davidiana</emphasis>
|
||
</taxonomicName>
|
||
specimens we examined had four premolars; see also: Kryštufek
|
||
<emphasis id="B92AEAF0FFCF61172AA8F817FB0301BA" box="[1170,1211,1954,1978]" italics="true" pageId="11" pageNumber="578">et al</emphasis>
|
||
. 2001); 1st upper molar with small, reduced parastyle, always much more prominent in
|
||
<taxonomicName id="4C5E4D61FFC861102EBBFF04FED806C7" baseAuthorityName="Milne-Edwards" baseAuthorityYear="1884" box="[129,352,175,199]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="subSpecies" species="davidiana" subSpecies="davidiana">
|
||
<emphasis id="B92AEAF0FFC861102EBBFF04FED806C7" box="[129,352,175,199]" italics="true" pageId="12" pageNumber="579">T. davidiana davidiana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13652A67FFC861102F4AFF1BFE7E06C7" box="[368,454,175,199]" captionStart="Figure 11" captionStartId="19.[113,178,1138,1162]" captionTargetBox="[149,1423,147,1108]" captionTargetId="figure-344@19.[146,1426,144,1110]" captionTargetPageId="19" captionText="Figure 11. Lateral view of less upper molar in Anatolian–Iranian Talpa, anterior to the less. A, T. hakkariensis sp. nov., holotype, specimen 1533; B, topotypical T. davidiana davidiana, specimen 1565; C, T. davidiana tatvanensis ssp. nov., holotype, specimen 1530; D, T.streetorum Holotype, Field Museum of Natural History, Chicago, specimen 96424. Arrows indicate parastyle." figureDoi="http://doi.org/10.5281/zenodo.10469958" httpUri="https://zenodo.org/record/10469958/files/figure.png" pageId="12" pageNumber="579">Fig. 11A</figureCitation>
|
||
); larger hands and feet (see
|
||
<figureCitation id="13652A67FFC861102CD9FF1BFF2306E7" captionStart="Figure 12" captionStartId="20.[129,194,1182,1206]" captionTargetBox="[163,1441,150,1097]" captionTargetId="figure-422@20.[161,1441,144,1154]" captionTargetPageId="20" captionText="Figure 12. Feet and tails of Anatolian Talpa. A, D, T. hakkariensis sp. nov., holotype, specimen 1533; B, E, topotypical T. davidiana davidiana, specimen 1565; C, F, T. davidiana tatvanensis ssp. nov., holotype, specimen 1530. Scale bars: A–C = 25 mm; D–F = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.10469962" httpUri="https://zenodo.org/record/10469962/files/figure.png" pageId="12" pageNumber="579">Fig. 12</figureCitation>
|
||
); and longer (28–33 vs.
|
||
<quantity id="4CA69B07FFC861102FACFF7BFDBF06E7" box="[406,519,207,231]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="12" pageNumber="579" unit="mm" value="17.0" valueMax="18.0" valueMin="16.0">16–18 mm</quantity>
|
||
) and paler tail, with most tail hairs of more uniform length (compare
|
||
<figureCitation id="13652A67FFC861102C6BFF5AFD700705" box="[593,712,238,262]" captionStart="Figure 12" captionStartId="20.[129,194,1182,1206]" captionTargetBox="[163,1441,150,1097]" captionTargetId="figure-422@20.[161,1441,144,1154]" captionTargetPageId="20" captionText="Figure 12. Feet and tails of Anatolian Talpa. A, D, T. hakkariensis sp. nov., holotype, specimen 1533; B, E, topotypical T. davidiana davidiana, specimen 1565; C, F, T. davidiana tatvanensis ssp. nov., holotype, specimen 1530. Scale bars: A–C = 25 mm; D–F = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.10469962" httpUri="https://zenodo.org/record/10469962/files/figure.png" pageId="12" pageNumber="579">Fig. 12D, E</figureCitation>
|
||
). The broad snout of the new species resembles that of
|
||
<taxonomicName id="4C5E4D61FFC861102CB6FEBAFF540744" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="subSpecies" species="davidiana" subSpecies="tatvanensis">
|
||
<emphasis id="B92AEAF0FFC861102CB6FEBAFF540744" italics="true" pageId="12" pageNumber="579">T. davidiana tatvanensis</emphasis>
|
||
</taxonomicName>
|
||
ssp. nov., but this taxon is much smaller than
|
||
<taxonomicName id="4C5E4D61FFC861102CCCFE99FF420764" authority="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton, 2023" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="hakkariensis" status="sp. nov.">
|
||
<emphasis id="B92AEAF0FFC861102CCCFE99FF420764" italics="true" pageId="12" pageNumber="579">T. hakkariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219578BFFC861102F3AFEF8FEF50763" box="[256,333,332,355]" pageId="12" pageNumber="579" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(see below) and has a relatively prominent parastyle on the first upper molar, as in
|
||
<taxonomicName id="4C5E4D61FFC861102C25FED8FCBB0783" baseAuthorityName="Milne-Edwards" baseAuthorityYear="1884" box="[543,771,363,387]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="subSpecies" species="davidiana" subSpecies="davidiana">
|
||
<emphasis id="B92AEAF0FFC861102C25FED8FCBB0783" box="[543,771,363,387]" italics="true" pageId="12" pageNumber="579">T. davidiana davidiana</emphasis>
|
||
</taxonomicName>
|
||
. Distinguished from
|
||
<taxonomicName id="4C5E4D61FFC861102F69FE3FFE6E07A2" baseAuthorityName="Krystufek and Vohralik" baseAuthorityYear="2005" box="[339,470,395,418]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="streetorum">
|
||
<emphasis id="B92AEAF0FFC861102F69FE3FFE6E07A2" box="[339,470,395,418]" italics="true" pageId="12" pageNumber="579">T. streetorum</emphasis>
|
||
</taxonomicName>
|
||
by the latter’s larger cranium (see above), in having the posteriormost mandibular foramen positioned more anteriorly, and having a coronoid process that is narrower in anterior–posterior breadth and inclined more anteriorly (Supporting Information,
|
||
<figureCitation id="13652A67FFC861102FE5FDBCFD8C0420" box="[479,564,520,544]" captionStart="Figure 6" captionStartId="14.[130,195,1251,1275]" captionTargetBox="[164,1400,155,1178]" captionTargetId="figure-390@14.[161,1441,144,1223]" captionTargetPageId="14" captionText="Figure 6. Dorsal view of Anatolian Talpa skins. A, T. hakkariensis sp. nov., holotype, specimen 1533; B, topotypical T. davidiana davidiana, specimen 1565; C, T. davidiana tatvanensis ssp.nov., holotype, specimen 1530. Scale bar = 50 mm." figureDoi="http://doi.org/10.5281/zenodo.10469948" httpUri="https://zenodo.org/record/10469948/files/figure.png" pageId="12" pageNumber="579">Fig. S6B</figureCitation>
|
||
). The net
|
||
<emphasis id="B92AEAF0FFC861102C9AFDBDFD150420" box="[672,685,521,544]" italics="true" pageId="12" pageNumber="579">p</emphasis>
|
||
-distance in
|
||
<emphasis id="B92AEAF0FFC861102EA7FD93FF74043F" box="[157,204,551,575]" italics="true" pageId="12" pageNumber="579">Cytb</emphasis>
|
||
sequence between
|
||
<taxonomicName id="4C5E4D61FFC861102FA6FD9CFDA1043F" box="[412,537,551,575]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="davidiana">
|
||
<emphasis id="B92AEAF0FFC861102FA6FD9CFDA1043F" box="[412,537,551,575]" italics="true" pageId="12" pageNumber="579">T. davidiana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5E4D61FFC861102C6AFD9CFD5D043F" authority="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton, 2023" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" box="[592,741,551,575]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="hakkariensis" status="sp. nov.">
|
||
<emphasis id="B92AEAF0FFC861102C6AFD9CFD5D043F" box="[592,741,551,575]" italics="true" pageId="12" pageNumber="579">T. hakkariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219578BFFC861102CD4FD9CFF14045E" pageId="12" pageNumber="579" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is 10.50%; raw distance 11.33%. The net and raw K2P distances of 11.79% and 12.64%, respectively, between these taxa is in the range observed between other species of moles (e.g. within
|
||
<taxonomicName id="4C5E4D61FFC861102EF6FD10FEBD04BC" box="[204,261,676,700]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92AEAF0FFC861102EF6FD10FEBD04BC" box="[204,261,676,700]" italics="true" pageId="12" pageNumber="579">Talpa</emphasis>
|
||
</taxonomicName>
|
||
, interspecific net and raw K2P distances average
|
||
<emphasis id="B92AEAF0FFC861102EBBFD71FF3204DC" box="[129,138,709,732]" italics="true" pageId="12" pageNumber="579">c</emphasis>
|
||
. 11.47% and 12.61%, respectively). In fact, the net genetic distance in
|
||
<emphasis id="B92AEAF0FFC861102EC0FD57FE9104FB" box="[250,297,739,763]" italics="true" pageId="12" pageNumber="579">Cytb</emphasis>
|
||
sequence
|
||
<taxonomicName id="4C5E4D61FFC861102FA2FD50FDAF04FB" box="[408,535,739,763]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="davidiana">
|
||
<emphasis id="B92AEAF0FFC861102FA2FD50FE1104FB" box="[408,425,740,763]" italics="true" pageId="12" pageNumber="579">T</emphasis>
|
||
.
|
||
<emphasis id="B92AEAF0FFC861102F8CFD57FDAF04FB" box="[438,535,739,763]" italics="true" pageId="12" pageNumber="579">davidiana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5E4D61FFC861102C74FD50FD5E04FB" authority="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton, 2023" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" box="[590,742,739,763]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="hakkariensis" status="sp. nov.">
|
||
<emphasis id="B92AEAF0FFC861102C74FD50FDE704FB" box="[590,607,740,763]" italics="true" pageId="12" pageNumber="579">T</emphasis>
|
||
.
|
||
<emphasis id="B92AEAF0FFC861102C57FD57FD5E04FB" box="[621,742,739,763]" italics="true" pageId="12" pageNumber="579">hakkariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219578BFFC861102CD4FD50FF14051A" pageId="12" pageNumber="579" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is much higher than the raw (total) genetic distance between several recognized sister-species in
|
||
<taxonomicName id="4C5E4D61FFC861102C1EFC96FDE5053A" box="[548,605,802,826]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92AEAF0FFC861102C1EFC96FDE5053A" box="[548,605,802,826]" italics="true" pageId="12" pageNumber="579">Talpa</emphasis>
|
||
</taxonomicName>
|
||
(e.g.
|
||
<taxonomicName id="4C5E4D61FFC861102CABFC97FCB1053A" box="[657,777,802,826]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="aquitania">
|
||
<emphasis id="B92AEAF0FFC861102CABFC97FD1A053A" box="[657,674,803,826]" italics="true" pageId="12" pageNumber="579">T</emphasis>
|
||
.
|
||
<emphasis id="B92AEAF0FFC861102C91FC97FCB1053A" box="[683,777,803,826]" italics="true" pageId="12" pageNumber="579">aquitania</emphasis>
|
||
</taxonomicName>
|
||
vs.
|
||
<taxonomicName id="4C5E4D61FFC861102E9BFCF6FE940559" authorityName="Cabrera" authorityYear="1907" box="[161,300,833,857]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="occidentalis">
|
||
<emphasis id="B92AEAF0FFC861102E9BFCF6FF0A0559" box="[161,178,834,857]" italics="true" pageId="12" pageNumber="579">T</emphasis>
|
||
.
|
||
<emphasis id="B92AEAF0FFC861102E87FCF5FE940559" box="[189,300,833,857]" italics="true" pageId="12" pageNumber="579">occidentalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5E4D61FFC861102F02FCF6FE170559" authorityName="Satunin" authorityYear="1908" box="[312,431,834,857]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="caucasica">
|
||
<emphasis id="B92AEAF0FFC861102F02FCF6FEF10559" box="[312,329,834,857]" italics="true" pageId="12" pageNumber="579">T</emphasis>
|
||
.
|
||
<emphasis id="B92AEAF0FFC861102F6EFCF6FE170559" box="[340,431,834,857]" italics="true" pageId="12" pageNumber="579">caucasica</emphasis>
|
||
</taxonomicName>
|
||
vs.
|
||
<taxonomicName id="4C5E4D61FFC861102FECFCF6FD880559" authorityName="Stroganov" authorityYear="1944" box="[470,560,834,857]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="ognevi">
|
||
<emphasis id="B92AEAF0FFC861102FECFCF6FE5F0559" box="[470,487,834,857]" italics="true" pageId="12" pageNumber="579">T</emphasis>
|
||
.
|
||
<emphasis id="B92AEAF0FFC861102FC8FCF6FD880559" box="[498,560,834,857]" italics="true" pageId="12" pageNumber="579">ognevi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5E4D61FFC861102C5DFCF6FD750559" authorityName="Thomas" authorityYear="1906" box="[615,717,833,857]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="levantis">
|
||
<emphasis id="B92AEAF0FFC861102C5DFCF6FDC00559" box="[615,632,834,857]" italics="true" pageId="12" pageNumber="579">T</emphasis>
|
||
.
|
||
<emphasis id="B92AEAF0FFC861102CBEFCF5FD750559" box="[644,717,833,857]" italics="true" pageId="12" pageNumber="579">levantis</emphasis>
|
||
</taxonomicName>
|
||
vs.
|
||
<taxonomicName id="4C5E4D61FFC861102CC9FCF6FEB50578" authorityName="Dahl" authorityYear="1945" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="transcaucasica">
|
||
<emphasis id="B92AEAF0FFC861102CC9FCF6FCBC0559" box="[755,772,834,857]" italics="true" pageId="12" pageNumber="579">T</emphasis>
|
||
.
|
||
<emphasis id="B92AEAF0FFC861102EBBFCD5FEB50578" box="[129,269,865,888]" italics="true" pageId="12" pageNumber="579">transcaucasica</emphasis>
|
||
</taxonomicName>
|
||
) (Supporting Information, Tables S4, S5).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3446569FFC861102FB5FC16FBA40400" pageId="12" pageNumber="579" type="description">
|
||
<paragraph id="8BE136E2FFC861102FB5FC16FE4305BB" blockId="12.[399,507,930,955]" box="[399,507,930,955]" pageId="12" pageNumber="579">
|
||
<emphasis id="B92AEAF0FFC861102FB5FC16FE4305BB" box="[399,507,930,955]" italics="true" pageId="12" pageNumber="579">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE136E2FFC861102EBBFC73FDE603B5" blockId="12.[128,778,967,1962]" pageId="12" pageNumber="579">
|
||
A relatively large, heavy mole (
|
||
<figureCitation id="13652A67FFC861102F82FC73FDBB05DF" box="[440,515,967,991]" captionStart="Figure 5" captionStartId="13.[113,178,1773,1797]" captionTargetBox="[254,1318,145,1745]" captionTargetId="figure-7@13.[254,1319,145,1745]" captionTargetPageId="13" captionText="Figure 5. Freshly trapped Anatolian Talpa in situ, with overviews of habitats at collecting localities.A, T. hakkariensis sp. nov., type locality, Turkey, Hakkari Province, Cemşililan yaylası c. 8 km NNE of Durankaya; B, T. hakkariensis sp. nov., holotype, specimen 1533; C, T. davidiana davidiana, topotypical locality, Turkey, Hatay Province, Hassa, Çardak yaylası; D, T. davidiana davidiana, specimen 1565; E, T.davidiana tatvanensis ssp.nov., type locality, Turkey, Bitlis Province, Uslu, Tatvan c. 1 km E of Kaynarca; F, T.davidiana tatvanensis ssp. nov., holotype, specimen 1530." figureDoi="http://doi.org/10.5281/zenodo.10469946" httpUri="https://zenodo.org/record/10469946/files/figure.png" pageId="12" pageNumber="579">Figs 5B</figureCitation>
|
||
,
|
||
<figureCitation id="13652A67FFC861102C34FC73FD9405DF" box="[526,556,967,991]" captionStart="Figure 6" captionStartId="14.[130,195,1251,1275]" captionTargetBox="[164,1400,155,1178]" captionTargetId="figure-390@14.[161,1441,144,1223]" captionTargetPageId="14" captionText="Figure 6. Dorsal view of Anatolian Talpa skins. A, T. hakkariensis sp. nov., holotype, specimen 1533; B, topotypical T. davidiana davidiana, specimen 1565; C, T. davidiana tatvanensis ssp.nov., holotype, specimen 1530. Scale bar = 50 mm." figureDoi="http://doi.org/10.5281/zenodo.10469948" httpUri="https://zenodo.org/record/10469948/files/figure.png" pageId="12" pageNumber="579">6A</figureCitation>
|
||
), with a medium tail; 17.07–22.45% of head and body length. Tail (
|
||
<figureCitation id="13652A67FFC861102C6AFC52FD1F05FF" box="[592,679,998,1023]" captionStart="Figure 12" captionStartId="20.[129,194,1182,1206]" captionTargetBox="[163,1441,150,1097]" captionTargetId="figure-422@20.[161,1441,144,1154]" captionTargetPageId="20" captionText="Figure 12. Feet and tails of Anatolian Talpa. A, D, T. hakkariensis sp. nov., holotype, specimen 1533; B, E, topotypical T. davidiana davidiana, specimen 1565; C, F, T. davidiana tatvanensis ssp. nov., holotype, specimen 1530. Scale bars: A–C = 25 mm; D–F = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.10469962" httpUri="https://zenodo.org/record/10469962/files/figure.png" pageId="12" pageNumber="579">Fig. 12D</figureCitation>
|
||
) parallel-sided, with sparse, stiff, sub-erect, overlapping bristles; individual bristles mostly pale greyish, almost translucent, up to 25% blackish or with black cores; bristles relatively even in length; tail tip tapered and rounded, visible when bristles are moved. Overall appearance of tail paler than body. Dorsal and ventral pelage greyish brown throughout, with marked silvery sheen; individual hairs typically brownish at tips, greyish below. Body elongated. Palm of manus (
|
||
<figureCitation id="13652A67FFC861102FADFB55FE5502F9" box="[407,493,1249,1273]" captionStart="Figure 12" captionStartId="20.[129,194,1182,1206]" captionTargetBox="[163,1441,150,1097]" captionTargetId="figure-422@20.[161,1441,144,1154]" captionTargetPageId="20" captionText="Figure 12. Feet and tails of Anatolian Talpa. A, D, T. hakkariensis sp. nov., holotype, specimen 1533; B, E, topotypical T. davidiana davidiana, specimen 1565; C, F, T. davidiana tatvanensis ssp. nov., holotype, specimen 1530. Scale bars: A–C = 25 mm; D–F = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.10469962" httpUri="https://zenodo.org/record/10469962/files/figure.png" pageId="12" pageNumber="579">Fig. 12A</figureCitation>
|
||
) large, broad, relatively flat; claws at ends of all five phalanges, relatively broad and flattened dorsoventrally, with convex upper and concave lower margins, flattened oval in cross-section. Feet (
|
||
<figureCitation id="13652A67FFC861102C3CFA8BFDD90357" box="[518,609,1343,1367]" captionStart="Figure 12" captionStartId="20.[129,194,1182,1206]" captionTargetBox="[163,1441,150,1097]" captionTargetId="figure-422@20.[161,1441,144,1154]" captionTargetPageId="20" captionText="Figure 12. Feet and tails of Anatolian Talpa. A, D, T. hakkariensis sp. nov., holotype, specimen 1533; B, E, topotypical T. davidiana davidiana, specimen 1565; C, F, T. davidiana tatvanensis ssp. nov., holotype, specimen 1530. Scale bars: A–C = 25 mm; D–F = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.10469962" httpUri="https://zenodo.org/record/10469962/files/figure.png" pageId="12" pageNumber="579">Fig. 12D</figureCitation>
|
||
) large, approximately two-thirds tail length. Muzzle (
|
||
<figureCitation id="13652A67FFC861102C2EFAEAFDD90376" box="[532,609,1374,1398]" captionStart="Figure 5" captionStartId="13.[113,178,1773,1797]" captionTargetBox="[254,1318,145,1745]" captionTargetId="figure-7@13.[254,1319,145,1745]" captionTargetPageId="13" captionText="Figure 5. Freshly trapped Anatolian Talpa in situ, with overviews of habitats at collecting localities.A, T. hakkariensis sp. nov., type locality, Turkey, Hakkari Province, Cemşililan yaylası c. 8 km NNE of Durankaya; B, T. hakkariensis sp. nov., holotype, specimen 1533; C, T. davidiana davidiana, topotypical locality, Turkey, Hatay Province, Hassa, Çardak yaylası; D, T. davidiana davidiana, specimen 1565; E, T.davidiana tatvanensis ssp.nov., type locality, Turkey, Bitlis Province, Uslu, Tatvan c. 1 km E of Kaynarca; F, T.davidiana tatvanensis ssp. nov., holotype, specimen 1530." figureDoi="http://doi.org/10.5281/zenodo.10469946" httpUri="https://zenodo.org/record/10469946/files/figure.png" pageId="12" pageNumber="579">Figs 5B</figureCitation>
|
||
,
|
||
<figureCitation id="13652A67FFC861102C55FAEAFD350376" box="[623,653,1374,1398]" captionStart="Figure 6" captionStartId="14.[130,195,1251,1275]" captionTargetBox="[164,1400,155,1178]" captionTargetId="figure-390@14.[161,1441,144,1223]" captionTargetPageId="14" captionText="Figure 6. Dorsal view of Anatolian Talpa skins. A, T. hakkariensis sp. nov., holotype, specimen 1533; B, topotypical T. davidiana davidiana, specimen 1565; C, T. davidiana tatvanensis ssp.nov., holotype, specimen 1530. Scale bar = 50 mm." figureDoi="http://doi.org/10.5281/zenodo.10469948" httpUri="https://zenodo.org/record/10469948/files/figure.png" pageId="12" pageNumber="579">6A</figureCitation>
|
||
) elongated, with hairs, including longer sensory bristles, paler than on head. Snout pinkish with close, fine, small, pale hairs.
|
||
</paragraph>
|
||
<paragraph id="8BE136E2FFC861102EA6FA08FD1F00CF" blockId="12.[128,778,967,1962]" pageId="12" pageNumber="579">
|
||
Skull (
|
||
<figureCitation id="13652A67FFC861102EE5FA08FEAF03D4" box="[223,279,1468,1492]" captionStart="Figure 7" captionStartId="15.[113,178,1554,1578]" captionTargetBox="[308,1263,146,1523]" captionTargetId="figure-231@15.[306,1266,144,1526]" captionTargetPageId="15" captionText="Figure 7. Talpa hakkariensis sp. nov., holotype, specimen 1533, cranium in (A) lateral, (B) ventral, and (C) dorsal views.Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10469950" httpUri="https://zenodo.org/record/10469950/files/figure.png" pageId="12" pageNumber="579">Fig. 7</figureCitation>
|
||
) large, with a stout, broad rostrum. Width across canines 15.0–17.05%, and width across molars 27.84–31.92% condylobasal length. Maxillary tooth row equals 42.36–44.7% of the condylobasal length. The braincase is broad and deep, flattened above; height of neurocranium 27.71–29.12% of condylobasal length. The posterior margin of palatine is usually slightly posterior to the imaginary line connecting the posterior alveolar margins of 3rd upper molars, and the anterior border of the infraorbital foramen is above the 2nd upper molar.
|
||
</paragraph>
|
||
<paragraph id="8BE136E2FFC861102EA6F963FBA40400" blockId="12.[128,778,967,1962]" lastBlockId="12.[825,1474,144,512]" pageId="12" pageNumber="579">
|
||
Upper incisors of decreasing size; 3rd incisor approximately half the size of the 1st. Upper molars robust, decreasing in size 1–3. Fourth upper premolar largest; 1st intermediate and 2nd and 3rd small (
|
||
<figureCitation id="13652A67FFC861102F1FF880FED9014C" box="[293,353,1844,1868]" captionStart="Figure 7" captionStartId="15.[113,178,1554,1578]" captionTargetBox="[308,1263,146,1523]" captionTargetId="figure-231@15.[306,1266,144,1526]" captionTargetPageId="15" captionText="Figure 7. Talpa hakkariensis sp. nov., holotype, specimen 1533, cranium in (A) lateral, (B) ventral, and (C) dorsal views.Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10469950" httpUri="https://zenodo.org/record/10469950/files/figure.png" pageId="12" pageNumber="579">Fig. 7</figureCitation>
|
||
). First upper molar with very small, almost imperceptible parastyle (
|
||
<figureCitation id="13652A67FFC861102FF0F8E0FD9D016C" box="[458,549,1876,1900]" captionStart="Figure 11" captionStartId="19.[113,178,1138,1162]" captionTargetBox="[149,1423,147,1108]" captionTargetId="figure-344@19.[146,1426,144,1110]" captionTargetPageId="19" captionText="Figure 11. Lateral view of less upper molar in Anatolian–Iranian Talpa, anterior to the less. A, T. hakkariensis sp. nov., holotype, specimen 1533; B, topotypical T. davidiana davidiana, specimen 1565; C, T. davidiana tatvanensis ssp. nov., holotype, specimen 1530; D, T.streetorum Holotype, Field Museum of Natural History, Chicago, specimen 96424. Arrows indicate parastyle." figureDoi="http://doi.org/10.5281/zenodo.10469958" httpUri="https://zenodo.org/record/10469958/files/figure.png" pageId="12" pageNumber="579">Fig. 11A</figureCitation>
|
||
), visible in
|
||
<typeStatus id="54E58840FFC861102C91F8E0FCB1016C" box="[683,777,1876,1900]" pageId="12" pageNumber="579" type="holotype">holotype</typeStatus>
|
||
and some
|
||
<typeStatus id="54E58840FFC861102ECEF8C7FEE3018B" box="[244,347,1907,1931]" pageId="12" pageNumber="579" type="paratype">paratypes</typeStatus>
|
||
, but lost in specimens with substantial tooth wear; protocone much larger than paracone; mesostyle relatively flattened at apex, with clear mesal furrow. Second upper molar with well-developed parastyle, somewhat bilobed at apex in
|
||
<typeStatus id="54E58840FFC861102DDEFF7AFBFA06E6" box="[996,1090,206,230]" pageId="12" pageNumber="579" type="holotype">holotype</typeStatus>
|
||
and most
|
||
<typeStatus id="54E58840FFC861102A95FF7BFAAF06E7" box="[1199,1303,207,231]" pageId="12" pageNumber="579" type="paratype">paratypes</typeStatus>
|
||
; protocone only slightly larger than paracone; mesostyle well developed and clearly bilobed at apex. Third upper molar with well-developed parastyle, slightly more prominent than mesostyle; mesostyle clearly bilobed at apex; protocone only slightly larger than paracone. Mandible
|
||
<figureCitation id="13652A67FFC861102A2FFEDFFBC80783" box="[1045,1136,363,387]" captionStart="Figure 13" captionStartId="21.[113,178,1087,1111]" captionTargetBox="[149,1423,147,1057]" captionTargetId="figure-388@21.[146,1426,144,1060]" captionTargetPageId="21" captionText="Figure 13. Mandibles of Anatolian–Iranian Talpa. A, T. hakkariensis sp. nov., holotype, specimen 1533; B, topotypical T. davidiana davidiana, specimen 1565; C, T. davidiana tatvanensis ssp. nov., holotype, specimen 1530; D, T.streetorum Holotype, Field Museum of Natural History, Chicago, specimen 96424. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10469964" httpUri="https://zenodo.org/record/10469964/files/figure.png" pageId="12" pageNumber="579">Fig. 13A</figureCitation>
|
||
. Lower incisors subequal. First lower premolar large, with prominent distal cusp; 4th lower premolar subequal to 1st; 2nd and 3rd lower premolars much smaller. First and 2nd lower molars large; 3rd smaller; all with prominent parastyles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3446569FFC861102A71FD82FAED0494" pageId="12" pageNumber="579" type="etymology">
|
||
<paragraph id="8BE136E2FFC861102A71FD82FB090450" blockId="12.[1099,1201,566,592]" box="[1099,1201,566,592]" pageId="12" pageNumber="579">
|
||
<emphasis id="B92AEAF0FFC861102A71FD82FB090450" box="[1099,1201,566,592]" italics="true" pageId="12" pageNumber="579">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE136E2FFC861102D03FDE9FAED0494" blockId="12.[825,1474,605,660]" pageId="12" pageNumber="579">
|
||
<materialsCitation id="3B363CBFFFC861102D03FDE9FAE90494" country="Turkey" location="The" pageId="12" pageNumber="579" specimenCount="1" stateProvince="Hakkari">
|
||
<location id="8E816039FFC861102D03FDE9FCD80475" LSID="urn:lsid:plazi:treatment:03F787F4FFCF61102F14FC72FBBD0068:8E816039FFC861102D03FDE9FCD80475" box="[825,864,605,629]" country="Turkey" name="The" pageId="12" pageNumber="579" stateProvince="Hakkari">The</location>
|
||
new species is named in reference to
|
||
<collectingRegion id="499AF800FFC861102B3FFDE9FA060475" box="[1285,1470,605,629]" country="Turkey" name="Hakkari" pageId="12" pageNumber="579">Hakkari Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3497672FFC861102D03FDC8FC3A0494" box="[825,898,636,660]" name="Turkey" pageId="12" pageNumber="579">Turkey</collectingCountry>
|
||
where the
|
||
<typeStatus id="54E58840FFC861102DCBFDC8FBA70494" box="[1009,1055,636,660]" pageId="12" pageNumber="579">type</typeStatus>
|
||
locality is situated (adjective)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3446569FFC861102A31FD07FB3705CD" pageId="12" pageNumber="579" type="distribution">
|
||
<paragraph id="8BE136E2FFC861102A31FD07FB4804CD" blockId="12.[1035,1264,691,717]" box="[1035,1264,691,717]" pageId="12" pageNumber="579">
|
||
<emphasis id="B92AEAF0FFC861102A31FD07FB4804CD" box="[1035,1264,691,717]" italics="true" pageId="12" pageNumber="579">Distribution and ecology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE136E2FFC861102D03FD6EFB3705CD" blockId="12.[825,1475,730,973]" pageId="12" pageNumber="579">
|
||
Known from a handful of geographically close localities in the mountains of
|
||
<collectingRegion id="499AF800FFC861102DC1FD4DFB0B0511" box="[1019,1203,761,785]" country="Turkey" name="Hakkari" pageId="12" pageNumber="579">Hakkari province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3497672FFC861102AFBFD4DFAB30511" box="[1217,1291,761,785]" name="Turkey" pageId="12" pageNumber="579">Turkey</collectingCountry>
|
||
. Moles were collected from moist soils beside a mountain stream at Cemşililan yaylası (
|
||
<figureCitation id="13652A67FFC861102DB4FC8CFC610550" box="[910,985,824,848]" captionStart="Figure 5" captionStartId="13.[113,178,1773,1797]" captionTargetBox="[254,1318,145,1745]" captionTargetId="figure-7@13.[254,1319,145,1745]" captionTargetPageId="13" captionText="Figure 5. Freshly trapped Anatolian Talpa in situ, with overviews of habitats at collecting localities.A, T. hakkariensis sp. nov., type locality, Turkey, Hakkari Province, Cemşililan yaylası c. 8 km NNE of Durankaya; B, T. hakkariensis sp. nov., holotype, specimen 1533; C, T. davidiana davidiana, topotypical locality, Turkey, Hatay Province, Hassa, Çardak yaylası; D, T. davidiana davidiana, specimen 1565; E, T.davidiana tatvanensis ssp.nov., type locality, Turkey, Bitlis Province, Uslu, Tatvan c. 1 km E of Kaynarca; F, T.davidiana tatvanensis ssp. nov., holotype, specimen 1530." figureDoi="http://doi.org/10.5281/zenodo.10469946" httpUri="https://zenodo.org/record/10469946/files/figure.png" pageId="12" pageNumber="579">Fig. 5A</figureCitation>
|
||
), in Eastern Anatolian Montane Steppe subject to nomadic grazing, but otherwise with little human influence. Most of the precipitation in this region falls as winter snows. At Bilmizit yaylası, specimen was collected in a similar habitat, but in this case without surface water.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3446569FFC861102A74FC42FBBD0068" pageId="12" pageNumber="579" type="discussion">
|
||
<paragraph id="8BE136E2FFC861102A74FC42FB140210" blockId="12.[1102,1196,1014,1040]" box="[1102,1196,1014,1040]" pageId="12" pageNumber="579">
|
||
<emphasis id="B92AEAF0FFC861102A74FC42FB140210" box="[1102,1196,1014,1040]" italics="true" pageId="12" pageNumber="579">Comment</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE136E2FFC861102D03FBA8FBBD0068" blockId="12.[825,1474,1052,1640]" pageId="12" pageNumber="579">
|
||
This new mole probably corresponds to the specimens from Hakkari referred to
|
||
<taxonomicName id="4C5E4D61FFC861102A29FB88FB2B0253" box="[1043,1171,1083,1107]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="davidiana">
|
||
<emphasis id="B92AEAF0FFC861102A29FB88FB2B0253" box="[1043,1171,1083,1107]" italics="true" pageId="12" pageNumber="579">T. davidiana</emphasis>
|
||
</taxonomicName>
|
||
by Kryštufek
|
||
<emphasis id="B92AEAF0FFC861102B09FB88FAD90253" box="[1331,1377,1083,1107]" italics="true" pageId="12" pageNumber="579">et al</emphasis>
|
||
. (2001),
|
||
<taxonomicName id="4C5E4D61FFC861102D03FBEFFC070272" baseAuthorityName="Krystufek and Vohralik" baseAuthorityYear="2005" box="[825,959,1115,1138]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="streetorum">
|
||
<emphasis id="B92AEAF0FFC861102D03FBEFFC070272" box="[825,959,1115,1138]" italics="true" pageId="12" pageNumber="579">T. streetorum</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFCF4B13FFC861102DD5FBEFFB000273" author="Dogramaci S" box="[1007,1208,1115,1139]" pageId="12" pageNumber="579" pagination="204 - 219" refId="ref19474" refString="Dogramaci S. Taxonomy and distribution of moles (genus: Talpa) in Turkey. Doga TU Zooloji 1989; 13: 204 - 219 [in Turkish]." type="journal article" year="1989">Doğramacı (1989)</bibRefCitation>
|
||
, and
|
||
<emphasis id="B92AEAF0FFC861102AC1FBEFFA260272" box="[1275,1438,1115,1139]" italics="true" pageId="12" pageNumber="579">
|
||
<collectionCode id="ED4FAE27FFC861102AC1FBEFFAB30272" box="[1275,1291,1115,1138]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="579">T</collectionCode>
|
||
. d.
|
||
<taxonomicName id="4C5E4D61FFC861102B0DFBEFFA260272" baseAuthorityName="Krystufek and Vohralik" baseAuthorityYear="2005" box="[1335,1438,1115,1138]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="streetorum">streetorum</taxonomicName>
|
||
</emphasis>
|
||
by
|
||
<bibRefCitation id="EFCF4B13FFC861102D03FBCEFB3E0292" author="Krystufek B & Vohralik V & Mammals of Turkey and Cyprus & Rodentia I" box="[825,1158,1146,1170]" pageId="12" pageNumber="579" refId="ref20318" refString="Krystufek B, Vohralik V. Mammals of Turkey and Cyprus. Rodentia I. Sciuridae, Dipodidae, Gliridae, Arvicolinae. Koper, Slovenia: Knjiznica Annales Majora. 2005." type="book" year="2005">Kryštufek and Vohralik (2005)</bibRefCitation>
|
||
. Our genetic, morphological, and morphometric data indicate that Hakkari populations represent a distinct species of broad-rostrum mole, rather than a subspecies of
|
||
<taxonomicName id="4C5E4D61FFC861102D9AFB6DFBA202F0" box="[928,1050,1240,1264]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="davidiana">
|
||
<emphasis id="B92AEAF0FFC861102D9AFB6DFBA202F0" box="[928,1050,1240,1264]" italics="true" pageId="12" pageNumber="579">T. davidiana</emphasis>
|
||
</taxonomicName>
|
||
, and our morphometric analysis indicates that it is also distinct from
|
||
<taxonomicName id="4C5E4D61FFC861102A72FB4CFB70030F" baseAuthorityName="Krystufek and Vohralik" baseAuthorityYear="2005" box="[1096,1224,1272,1295]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="streetorum">
|
||
<emphasis id="B92AEAF0FFC861102A72FB4CFB70030F" box="[1096,1224,1272,1295]" italics="true" pageId="12" pageNumber="579">T. streetorum</emphasis>
|
||
</taxonomicName>
|
||
(see above). At the time of writing, we have been unable to sequence the
|
||
<typeStatus id="54E58840FFC861102B03FAA3FADF032F" box="[1337,1383,1303,1327]" pageId="12" pageNumber="579">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C5E4D61FFC861102D6CFA83FC61034E" baseAuthorityName="Krystufek and Vohralik" baseAuthorityYear="2005" box="[854,985,1335,1358]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="streetorum">
|
||
<emphasis id="B92AEAF0FFC861102D6CFA83FC61034E" box="[854,985,1335,1358]" italics="true" pageId="12" pageNumber="579">T. streetorum</emphasis>
|
||
</taxonomicName>
|
||
or to obtain new material from
|
||
<collectingCountry id="F3497672FFC861102B0BFA82FAE4034D" box="[1329,1372,1334,1357]" name="Iran" pageId="12" pageNumber="579">Iran</collectingCountry>
|
||
, but consider the morphometric results from our comparison of skulls conclusive.
|
||
<bibRefCitation id="EFCF4B13FFC861102D94FAC1FB50038C" author="Krystufek B & Vohralik V & Mammals of Turkey and Cyprus & Rodentia I" box="[942,1256,1396,1421]" pageId="12" pageNumber="579" refId="ref20318" refString="Krystufek B, Vohralik V. Mammals of Turkey and Cyprus. Rodentia I. Sciuridae, Dipodidae, Gliridae, Arvicolinae. Koper, Slovenia: Knjiznica Annales Majora. 2005." type="book" year="2005">Kryštufek and Vohralik (2005)</bibRefCitation>
|
||
noted the similarities between the moles from Hakkari and a fossil mole from the Late Pleistocene of
|
||
<collectingCountry id="F3497672FFC861102DEBFA07FBB203CB" box="[977,1034,1459,1483]" name="Israel" pageId="12" pageNumber="579">Israel</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C5E4D61FFC861102A2FFA00FB4303CB" authority="Bate, 1937" authorityName="Bate" authorityYear="1937" box="[1045,1275,1459,1483]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="chthonia">
|
||
<emphasis id="B92AEAF0FFC861102A2FFA00FB3C03CB" box="[1045,1156,1459,1483]" italics="true" pageId="12" pageNumber="579">T. chthonia</emphasis>
|
||
Bate, 1937
|
||
</taxonomicName>
|
||
, which they speculated could be conspecific. Because we consider
|
||
<taxonomicName id="4C5E4D61FFC861102B14FA67FA7A03EA" authority="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton, 2023" authorityName="Gündüz & Demirtaş & Silsüpür & Özmen & Polly & Bilton" authorityYear="2023" box="[1326,1474,1490,1514]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="hakkariensis" status="sp. nov.">
|
||
<emphasis id="B92AEAF0FFC861102B14FA67FA7A03EA" box="[1326,1474,1490,1514]" italics="true" pageId="12" pageNumber="579">T. hakkariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219578BFFC861102D03FA46FC300009" box="[825,904,1522,1545]" pageId="12" pageNumber="579" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C5E4D61FFC861102D87FA47FB87000A" baseAuthorityName="Krystufek and Vohralik" baseAuthorityYear="2005" box="[957,1087,1523,1546]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="streetorum">
|
||
<emphasis id="B92AEAF0FFC861102D87FA47FB87000A" box="[957,1087,1523,1546]" italics="true" pageId="12" pageNumber="579">T. streetorum</emphasis>
|
||
</taxonomicName>
|
||
to be distinct, it is not clear without further analysis which, if either, would be associated with this fossil but, if conspecific with either, the earlier name
|
||
<taxonomicName id="4C5E4D61FFC861102B6EF985FA7A0048" authorityName="Bate" authorityYear="1937" box="[1364,1474,1584,1608]" class="Mammalia" family="Talpidae" genus="Talpa" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="579" phylum="Chordata" rank="species" species="chthonia">
|
||
<emphasis id="B92AEAF0FFC861102B6EF985FA7A0048" box="[1364,1474,1584,1608]" italics="true" pageId="12" pageNumber="579">T. chthonia</emphasis>
|
||
</taxonomicName>
|
||
would have priority.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |