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<document id="BF6518EA7AAA18090587EC643E646B45" ID-CLB-Dataset="205013" ID-DOI="10.11646/zootaxa.5219.3.1" ID-GBIF-Dataset="cddff8b1-3eb5-45bc-8262-147b2fb3b7c8" ID-ISSN="1175-5326" ID-Zenodo-Dep="7417441" ID-ZooBank="4601FC23-BC31-438C-AF32-6EEAE5ACE7CF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1670570701893" checkinUser="plazi" docAuthor="Landestoy, Miguel A., Schools, Molly &amp; Hedges, S. Blair" docDate="2022" docId="03F787C2E43DFF89FF10323F25A0FA18" docLanguage="en" docName="zootaxa.5219.3.1.pdf" docOrigin="Zootaxa 5219 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guarocuyus Landestoy &amp; Schools &amp; Hedges 2022, gen. nov." docType="treatment" docUuid="1BA79AB3-AAB2-4F7C-BDC8-D449A622F228" docUuidSource="ZooBank" docVersion="7" lastPageNumber="208" masterDocId="FFCEFFBAE43BFF8EFF8735652037FF85" masterDocTitle="A new genus and species of Caribbean forest lizard (Diploglossidae; Celestinae) from southern Hispaniola" masterLastPageNumber="226" masterPageNumber="201" pageNumber="207" updateTime="1699413251006" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="60AE8FD14D71B04635692511110C6EFF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="286B14325465B111C9313E46B7E27663">A new genus and species of Caribbean forest lizard (Diploglossidae; Celestinae) from southern Hispaniola</mods:title>
</mods:titleInfo>
<mods:name id="B8D902A61A7BDAECF02B2E243A04CA16" type="personal">
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<mods:roleTerm id="DABF9BF2FE583542C235794693FA325F">Author</mods:roleTerm>
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<mods:namePart id="6D347162FB330150C0BC6F278DD6CBD7">Landestoy, Miguel A.</mods:namePart>
<mods:nameIdentifier id="C261A08DD3E645D30D8A9F58FA9AB974" type="ORCID">0000-0002-5072-5769</mods:nameIdentifier>
<mods:affiliation id="A4422A8261837AD7B8F52948981E6B18">Escuela de Biología, Facultad de Ciencias de la Universidad Autónoma de Santo Domingo, República Dominicana. hispanioland @ gmail. com; https: // orcid. org / 0000 - 0002 - 5072 - 5769</mods:affiliation>
<mods:nameIdentifier id="C5443A1135796ABC05F288ADCEA0ADC6" type="email">hispanioland@gmail.com</mods:nameIdentifier>
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<mods:namePart id="2CD617B53F3F83283CD75A8485DA1A4D">Schools, Molly</mods:namePart>
<mods:affiliation id="875764D648CA09D3960F1EEAEB39695D">Center for Biodiversity, 502 SERC Building, Temple University, 1925 N. 12 th St., Philadelphia, PA 19122, USA.</mods:affiliation>
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<mods:namePart id="0B60924AC4C91112E27CD81210B99BC0">Hedges, S. Blair</mods:namePart>
<mods:affiliation id="AEC1ECE630D85E1D921343ECDDCE2D13">Center for Biodiversity, 502 SERC Building, Temple University, 1925 N. 12 th St., Philadelphia, PA 19122, USA.</mods:affiliation>
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<mods:title id="80DCF88F1BBD1CCE078B43F6376A2656">Zootaxa</mods:title>
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<mods:date id="7FCE67A5102D654E96216171F88AB3D8">2022</mods:date>
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<mods:number id="6A06CBBDEF7E25EE08CE34C78CC1A7FD">2022-12-09</mods:number>
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<mods:number id="C8CCAA5BA8A42ECD88B4BD67D0F28C15">5219</mods:number>
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<treatment id="03F787C2E43DFF89FF10323F25A0FA18" ID-DOI="http://doi.org/10.5281/zenodo.7419906" ID-GBIF-Taxon="206168295" ID-Zenodo-Dep="7419906" LSID="urn:lsid:zoobank.org:act:1BA79AB3-AAB2-4F7C-BDC8-D449A622F228" httpUri="http://treatment.plazi.org/id/03F787C2E43DFF89FF10323F25A0FA18" lastPageId="7" lastPageNumber="208" pageId="6" pageNumber="207">
<subSubSection id="C344655FE43DFF88FF10323F2196F8F1" box="[151,417,1882,1908]" pageId="6" pageNumber="207" type="nomenclature">
<paragraph id="8BE136D4E43DFF88FF10323F2196F8F1" blockId="6.[151,991,1882,1944]" box="[151,417,1882,1908]" pageId="6" pageNumber="207">
<heading id="D0A981B8E43DFF88FF10323F2196F8F1" bold="true" box="[151,417,1882,1908]" fontSize="11" level="1" pageId="6" pageNumber="207" reason="1">
<emphasis id="B92AEAC6E43DFF88FF10323F2196F8F1" bold="true" box="[151,417,1882,1908]" pageId="6" pageNumber="207">
<taxonomicName id="4C5E4D57E43DFF88FF10323F211CF8F1" authority="Landestoy &amp; Schools &amp; Hedges, 2022" authorityName="Landestoy &amp; Schools &amp; Hedges" authorityYear="2022" box="[151,299,1882,1908]" class="Reptilia" family="Diploglossidae" genus="Guarocuyus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="207" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B92AEAC6E43DFF88FF10323F211CF8F1" bold="true" box="[151,299,1882,1908]" italics="true" pageId="6" pageNumber="207">Guarocuyus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21957BDE43DFF88FEB4323F2196F8F1" box="[307,417,1882,1908]" pageId="6" pageNumber="207" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C344655FE43DFF88FF10321B23E8F81D" box="[151,991,1918,1944]" pageId="6" pageNumber="207" type="description">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C344655FE43DFF88FF1032A323C0F865" box="[151,1015,1989,2016]" pageId="6" pageNumber="207" type="type_taxon">
<paragraph id="8BE136D4E43DFF88FF1032A323C0F865" blockId="6.[151,1015,1989,2016]" box="[151,1015,1989,2016]" pageId="6" pageNumber="207">
<emphasis id="B92AEAC6E43DFF88FF1032A32105F85A" bold="true" box="[151,306,1989,2016]" pageId="6" pageNumber="207">
<typeStatus id="54E58876E43DFF88FF1032A320E5F865" box="[151,210,1990,2016]" pageId="6" pageNumber="207">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C5E4D57E43DFF88FEBE32A3227FF865" authority="Landestoy &amp; Schools &amp; Hedges, 2022" authorityName="Landestoy &amp; Schools &amp; Hedges" authorityYear="2022" box="[313,584,1990,2016]" class="Reptilia" family="Diploglossidae" genus="Guarocuyus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="jaraguanus" status="sp. nov.">
<emphasis id="B92AEAC6E43DFF88FEBE32A3227FF865" box="[313,584,1990,2016]" italics="true" pageId="6" pageNumber="207">Guarocuyus jaraguanus</emphasis>
</taxonomicName>
<emphasis id="B92AEAC6E43DFF88FDC832A3229FF865" bold="true" box="[591,680,1990,2016]" pageId="6" pageNumber="207">
<taxonomicNameLabel id="A21957BDE43DFF88FDC832A3229FF865" box="[591,680,1990,2016]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, herein described (
<figureCitation id="13652A51E43DFF88FCF832A323DCF865" box="[895,1003,1990,2016]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStartId-0="10.[151,250,1485,1509]" captionStartId-1="11.[151,250,1800,1824]" captionStartId-2="12.[151,250,1880,1904]" captionStartId-3="13.[151,250,725,749]" captionTargetBox-0="[160,1406,642,1442]" captionTargetBox-1="[151,1425,821,1770]" captionTargetBox-2="[155,1436,185,1852]" captionTargetBox-3="[271,1320,186,694]" captionTargetId-0="figure-226@10.[151,1436,633,1462]" captionTargetId-1="figure-285@11.[151,1436,813,1777]" captionTargetId-2="figure-22@12.[151,1436,181,1856]" captionTargetId-3="figure-19@13.[264,1323,181,701]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionText-0="FIGURE 6. Living individual (MNHNSD 23.3935) of Guarocuyus jaraguanus sp. nov." captionText-1="FIGURE 7. Dorsal view of a series of Guarocuyus jaraguanus sp. nov. showing variation. Specimens: males, MNHNSD 23.3934 (A), MNHNSD 23.3936 (B), 1005 (C), MNHNSD 23.3937 (D), MNHNSD 23.3935 (E), and IIBZ-HER00006 (F), and females, IIBZ-HER00007–00008 (G–H)." captionText-2="FIGURE 8. Holotype MNHNSD 23.3937 (MALT1004) Guarocuyus jaraguanus sp. nov. in dorsal (A), lateral (B), and ventral (C) views. Scale bar = 10 mm." captionText-3="FIGURE 9. Lateral view of the head of Guarocuyus jaraguanus sp. nov. (MNHNSD 23.3912) showing two (red shading) postnasal scales." figureDoi-0="http://doi.org/10.5281/zenodo.7417453" figureDoi-1="http://doi.org/10.5281/zenodo.7417455" figureDoi-2="http://doi.org/10.5281/zenodo.7417457" figureDoi-3="http://doi.org/10.5281/zenodo.7417459" httpUri-0="https://zenodo.org/record/7417453/files/figure.png" httpUri-1="https://zenodo.org/record/7417455/files/figure.png" httpUri-2="https://zenodo.org/record/7417457/files/figure.png" httpUri-3="https://zenodo.org/record/7417459/files/figure.png" pageId="6" pageNumber="207">Figs. 6–9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C344655FE43CFF89FF4035F323B6FF50" pageId="7" pageNumber="208" type="distribution">
<paragraph id="8BE136D4E43CFF89FF4035F323B6FF50" blockId="7.[151,1437,150,1437]" pageId="7" pageNumber="208">
<emphasis id="B92AEAC6E43CFF89FF4035F32156FF35" bold="true" box="[199,353,150,176]" pageId="7" pageNumber="208">Distribution.</emphasis>
Cayo de las Iguanas and adjacent Cayo Pei, Laguna de Oviedo, in the
<collectingRegion id="499AF836E43CFF89FB3535F2252AFF34" box="[1202,1309,151,177]" country="Dominican Republic" name="Barahona" pageId="7" pageNumber="208">Barahona</collectingRegion>
Peninsula,
<collectingCountry id="F3497644E43CFF89FF1035DE21B4FF50" box="[151,387,187,213]" name="Dominican Republic" pageId="7" pageNumber="208">Dominican Republic</collectingCountry>
, the southernmost part of Hispaniola (
<figureCitation id="13652A51E43CFF89FCA935DE2344FF50" box="[814,883,186,213]" captionStart="FIGURE 5" captionStartId="9.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-19@9.[151,1436,181,1089]" captionTargetPageId="9" captionText="FIGURE 5. Map showing known locality for Guarocuyus jaraguanus sp. nov. marked with a red circle. Inset shows all of Hispaniola, with a red box indicating the mapped area." figureDoi="http://doi.org/10.5281/zenodo.7417451" httpUri="https://zenodo.org/record/7417451/files/figure.png" pageId="7" pageNumber="208">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C344655FE43CFF89FF4035BA2437FEC4" pageId="7" pageNumber="208" type="etymology">
<paragraph id="8BE136D4E43CFF89FF4035BA2437FEC4" blockId="7.[151,1437,150,1437]" pageId="7" pageNumber="208">
<emphasis id="B92AEAC6E43CFF89FF4035BA2179FF7C" bold="true" box="[199,334,223,249]" pageId="7" pageNumber="208">Etymology.</emphasis>
The generic name is a latinized masculine patronym honoring the taíno Guarocuya (a.k.a Enriquillo), the last cacique of the Jaragua region and a hero of the last standing free community of this native Antillean people, who died around the first half of the 16
<superScript id="7C2B9B9CE43CFF89FDC93443226CFEB1" attach="left" box="[590,603,294,308]" fontSize="6" pageId="7" pageNumber="208">th</superScript>
century (
<bibRefCitation id="EFCF4B25E43CFF89FD433442236BFEC4" author="Altman, I." box="[708,860,295,321]" pageId="7" pageNumber="208" pagination="587 - 614" refId="ref11572" refString="Altman, I. (2007) The Revolt of Enriquillo and the Historiography of Early Spanish America. The Americas, 63 (4), 587 - 614. https: // doi. org / 10.1353 / tam. 2007.0052" type="journal article" year="2007">Altman 2007</bibRefCitation>
;
<bibRefCitation id="EFCF4B25E43CFF89FCEF344223C2FEC4" author="Ozuna, A." box="[872,1013,295,321]" pageId="7" pageNumber="208" pagination="77 - 96" refId="ref12418" refString="Ozuna, A. (2018) Rebellion and Anti-colonial Struggle in Hispaniola: From Indigenous Agitators to African Rebels. Africology: The Journal of Pan African Studies, 11, 77 - 96." type="journal article" year="2018">Ozuna 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C344655FE43CFF89FF40342F2483FAFD" pageId="7" pageNumber="208" type="diagnosis">
<paragraph id="8BE136D4E43CFF89FF40342F2426FDB8" blockId="7.[151,1437,150,1437]" pageId="7" pageNumber="208">
<emphasis id="B92AEAC6E43CFF89FF40342F2175FEE1" bold="true" box="[199,322,330,356]" pageId="7" pageNumber="208">Diagnosis.</emphasis>
The new genus can be distinguished by the following combination of characters (see
<bibRefCitation id="EFCF4B25E43CFF89FAA3342E2118FE0C" author="Schools, M. &amp; Hedges, S. B." pageId="7" pageNumber="208" pagination="201 - 257" refId="ref12671" refString="Schools, M. &amp; Hedges, S. B. (2021) Phylogenetics, classification, and biogeography of the Neotropical forest lizards (Squamata, Diploglossidae). Zootaxa, 4974 (2), 201 - 257. https: // doi. org / 10.11646 / zootaxa. 4974.2.1" type="journal article" year="2021">Schools &amp; Hedges 2021</bibRefCitation>
): (1) claw sheath, absent or present, (2) contact between the nasal and rostral scales, absent, (3) scales in contact with the nasal scale, four or five (mode 5), (4) postnasal scales, one or two (mode 1), (5) position of the nostril in the nasal scale, central, (6) keels on dorsal body scales, present, (7) digits per limb, five, (8) longest toe lamellae, 21–25, (9) dorsal scale rows, 88–98, (10) relative head width, 12.0–13.9, (11) relative rostral height, 54.1–64.5, (12) relative frontonasal length, 3.07–4.48, (13) relative interparietal distance,
<geoCoordinate id="EE6A5013E43CFF89FBFC349B248CFD9C" box="[1147,1211,510,537]" degrees="0.179" direction="north" orientation="latitude" pageId="7" pageNumber="208" precision="55" value="0.179">0.179</geoCoordinate>
<geoCoordinate id="EE6A5013E43CFF89FB3C349B253DFD9C" box="[1211,1290,510,537]" degrees="0.755" direction="west" orientation="longitude" pageId="7" pageNumber="208" precision="55" value="-0.755">–0.755</geoCoordinate>
, (14) relative axilla-groin distance, 53.2–58.7, (15) relative axilla-cloaca distance, 62.5–67.7.
</paragraph>
<paragraph id="8BE136D4E43CFF89FF4037222483FAFD" blockId="7.[151,1437,150,1437]" pageId="7" pageNumber="208">
Within
<taxonomicName id="4C5E4D57E43CFF89FE90372221BFFDE4" box="[279,392,583,609]" pageId="7" pageNumber="204" rank="subFamily" subFamily="Celestinae">Celestinae</taxonomicName>
,
<taxonomicName id="4C5E4D57E43CFF89FE1437222223FDE4" authority="Landestoy &amp; Schools &amp; Hedges, 2022" authorityName="Landestoy &amp; Schools &amp; Hedges" authorityYear="2022" box="[403,532,583,609]" class="Reptilia" family="Diploglossidae" genus="Guarocuyus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B92AEAC6E43CFF89FE1437222223FDE4" box="[403,532,583,609]" italics="true" pageId="7" pageNumber="208">Guarocuyus</emphasis>
</taxonomicName>
<emphasis id="B92AEAC6E43CFF89FD9C37222248FDE4" bold="true" box="[539,639,583,609]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="A21957BDE43CFF89FD9C37222248FDE4" box="[539,639,583,609]" pageId="7" pageNumber="208" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from
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<emphasis id="B92AEAC6E43CFF89FCEA372223F0FDE4" box="[877,967,583,609]" italics="true" pageId="7" pageNumber="208">Advenus</emphasis>
Schools &amp; Hedges
</taxonomicName>
by the presence of keels on the dorsal scales (vs. absent in
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<emphasis id="B92AEAC6E43CFF89FD8F370E2255FD00" box="[520,610,619,645]" italics="true" pageId="7" pageNumber="208">Advenus</emphasis>
</taxonomicName>
), longest toe lamellae count (21–25 vs.
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), relative width of the ear opening (EW/SVL 2.33–2.90 vs.
<quantity id="4CA69B31E43CFF89FDB537EA22B7FD2C" box="[562,640,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="2.5146" pageId="7" pageNumber="208" unit="in" value="0.99">0.99 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FD0E37EA22D4FD2C" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[649,739,655,681]" class="Squamata" family="Diploglossidae" genus="Advenus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FD0E37EA22D4FD2C" box="[649,739,655,681]" italics="true" pageId="7" pageNumber="208">Advenus</emphasis>
</taxonomicName>
), RH/RW (54.1–64.5 vs.
<quantity id="4CA69B31E43CFF89FB8637EB2478FD2C" box="[1025,1103,654,681]" metricMagnitude="0" metricUnit="m" metricValue="1.27254" pageId="7" pageNumber="208" unit="in" value="50.1">50.1 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FBDF37EA2485FD2C" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[1112,1202,655,681]" class="Squamata" family="Diploglossidae" genus="Advenus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FBDF37EA2485FD2C" box="[1112,1202,655,681]" italics="true" pageId="7" pageNumber="208">Advenus</emphasis>
</taxonomicName>
), relative frontonasal length (FN/SVL 3.07–4.48 vs.
<quantity id="4CA69B31E43CFF89FE6537D6221BFD48" box="[482,556,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="6.2484" pageId="7" pageNumber="208" unit="in" value="2.46">2.46 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FDB437D622BAFD48" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[563,653,691,717]" class="Squamata" family="Diploglossidae" genus="Advenus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FDB437D622BAFD48" box="[563,653,691,717]" italics="true" pageId="7" pageNumber="208">Advenus</emphasis>
</taxonomicName>
), and by the AGD/SVL (53.2–58.7 vs. 60.0 in
<taxonomicName id="4C5E4D57E43CFF89FB0537D624EBFD48" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[1154,1244,691,717]" class="Squamata" family="Diploglossidae" genus="Advenus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FB0537D624EBFD48" box="[1154,1244,691,717]" italics="true" pageId="7" pageNumber="208">Advenus</emphasis>
</taxonomicName>
). From
<taxonomicName id="4C5E4D57E43CFF89FAB437D62156FD74" authority="Schools &amp; Hedges" authorityName="Schools &amp; Hedges" authorityYear="2021" class="Squamata" family="Diploglossidae" genus="Caribicus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FAB437D625ABFD48" box="[1331,1436,691,717]" italics="true" pageId="7" pageNumber="208">Caribicus</emphasis>
Schools &amp; Hedges
</taxonomicName>
,
<taxonomicName id="4C5E4D57E43CFF89FEEA37B221D9FD74" authority="Landestoy &amp; Schools &amp; Hedges, 2022" authorityName="Landestoy &amp; Schools &amp; Hedges" authorityYear="2022" box="[365,494,727,753]" class="Reptilia" family="Diploglossidae" genus="Guarocuyus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B92AEAC6E43CFF89FEEA37B221D9FD74" box="[365,494,727,753]" italics="true" pageId="7" pageNumber="208">Guarocuyus</emphasis>
</taxonomicName>
<emphasis id="B92AEAC6E43CFF89FE7137B2226DFD74" bold="true" box="[502,602,727,753]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="A21957BDE43CFF89FE7137B2226DFD74" box="[502,602,727,753]" pageId="7" pageNumber="208" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished by the presence of an interdigital scaleless web between toes II–IV (vs. absent in
<taxonomicName id="4C5E4D57E43CFF89FE27379E223EFC90" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[416,521,763,789]" class="Squamata" family="Diploglossidae" genus="Caribicus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FE27379E223EFC90" box="[416,521,763,789]" italics="true" pageId="7" pageNumber="208">Caribicus</emphasis>
</taxonomicName>
), longest toe lamellae count (21–25 vs.
<quantity id="4CA69B31E43CFF89FC32379E2423FC90" box="[949,1044,763,789]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="5.08" metricValueMin="3.302" pageId="7" pageNumber="208" unit="in" value="16.5" valueMax="20.0" valueMin="13.0">13–20 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FB9C379E24B3FC90" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[1051,1156,763,789]" class="Squamata" family="Diploglossidae" genus="Caribicus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FB9C379E24B3FC90" box="[1051,1156,763,789]" italics="true" pageId="7" pageNumber="208">Caribicus</emphasis>
</taxonomicName>
), EW/SVL (2.33–2.90 vs.
<quantity id="4CA69B31E43CFF89FF10367A212AFCBC" box="[151,285,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="3.8735" metricValueMax="4.7752" metricValueMin="2.9718" pageId="7" pageNumber="208" unit="in" value="1.525" valueMax="1.88" valueMin="1.17">1.17–1.88 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FEA4367A21BBFCBC" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[291,396,799,825]" class="Squamata" family="Diploglossidae" genus="Caribicus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FEA4367A21BBFCBC" box="[291,396,799,825]" italics="true" pageId="7" pageNumber="208">Caribicus</emphasis>
</taxonomicName>
), and by the DTC, FSSS (vs. ES in
<taxonomicName id="4C5E4D57E43CFF89FC80367A2347FCBC" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[775,880,799,825]" class="Squamata" family="Diploglossidae" genus="Caribicus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FC80367A2347FCBC" box="[775,880,799,825]" italics="true" pageId="7" pageNumber="208">Caribicus</emphasis>
</taxonomicName>
). From
<taxonomicName id="4C5E4D57E43CFF89FC41367A246BFCBC" authority="Gray" authorityName="Gray" box="[966,1116,799,825]" class="Reptilia" family="Diploglossidae" genus="Celestus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FC41367A2417FCBC" box="[966,1056,799,825]" italics="true" pageId="7" pageNumber="208">Celestus</emphasis>
Gray
</taxonomicName>
,
<taxonomicName id="4C5E4D57E43CFF89FBE1367A24D0FCBC" authority="Landestoy &amp; Schools &amp; Hedges, 2022" authorityName="Landestoy &amp; Schools &amp; Hedges" authorityYear="2022" box="[1126,1255,799,825]" class="Reptilia" family="Diploglossidae" genus="Guarocuyus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B92AEAC6E43CFF89FBE1367A24D0FCBC" box="[1126,1255,799,825]" italics="true" pageId="7" pageNumber="208">Guarocuyus</emphasis>
</taxonomicName>
<emphasis id="B92AEAC6E43CFF89FB69367A2565FCBC" bold="true" box="[1262,1362,799,825]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="A21957BDE43CFF89FB69367A2565FCBC" box="[1262,1362,799,825]" pageId="7" pageNumber="208" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
differs by the presence of an interdigital scaleless web between toes II–IV (vs. absent or vestigial scaly web only between toes III–IV in
<taxonomicName id="4C5E4D57E43CFF89FF7836022230FC04" authority="Grant" authorityName="Grant" authorityYear="1940" box="[255,519,871,897]" class="Reptilia" family="Diploglossidae" genus="Celestus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="species" species="barbouri">
<emphasis id="B92AEAC6E43CFF89FF7836022189FC04" box="[255,446,871,897]" italics="true" pageId="7" pageNumber="208">Celestus barbouri</emphasis>
Grant
</taxonomicName>
;
<tableCitation id="C6DC036FE43CFF89FD9536022256FC04" box="[530,609,871,897]" captionStart="TABLE 1" captionStartId="3.[151,239,460,486]" captionTargetId="graphics-130@3.[151,2040,633,1687]" captionTargetPageId="3" captionText="TABLE 1. Diagnostic characters used to distinguish genera of the subfamily Celestinae in Schools and Hedges (2021), with additional characters (see Methods): (1) claw sheath: absent (a), present (p), (2) scales in contact with the nasal scale, (3) postnasal scales, (4) medial keel on dorsal body scales, (5) digits per limb, (6) foot webbing: absent (a), present (p), vestigial (v), (7) longest toe lamellae, (8) dorsal scale rows, (9) DTC, (10) EW/SVL, (11) HW/SVL, (12) RH/RW, (13) FN/SVL, (14) IPD/SVL, (15) AGD/SVL, and (16) ACD/SVL. A dash indicates that no value was obtained for that trait. An asterisk (*) represents very rare values (1–2 cases). Parentheses for each character represent sample size." httpUri="http://table.plazi.org/id/DF21665CE438FF8DFF1034A920F9FDF3" pageId="7" pageNumber="208" tableUuid="DF21665CE438FF8DFF1034A920F9FDF3">Table 1</tableCitation>
), and by the EW/SVL (2.33–2.90 vs.
<quantity id="4CA69B31E43CFF89FC7B360224B4FC04" box="[1020,1155,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="4.064" metricValueMax="5.6134" metricValueMin="2.5146" pageId="7" pageNumber="208" unit="in" value="1.6" valueMax="2.21" valueMin="0.99">0.99–2.21 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FB0C360224D2FC04" box="[1163,1253,871,897]" class="Reptilia" family="Diploglossidae" genus="Celestus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FB0C360224D2FC04" box="[1163,1253,871,897]" italics="true" pageId="7" pageNumber="208">Celestus</emphasis>
</taxonomicName>
). From
<taxonomicName id="4C5E4D57E43CFF89FABA36022157FC20" authority="Schools &amp; Hedges" authorityName="Schools &amp; Hedges" authorityYear="2021" class="Reptilia" family="Diploglossidae" genus="Comptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FABA360225ABFC04" box="[1341,1436,871,897]" italics="true" pageId="7" pageNumber="208">Comptus</emphasis>
Schools &amp; Hedges
</taxonomicName>
,
<taxonomicName id="4C5E4D57E43CFF89FEEC36EE21DBFC20" authority="Landestoy &amp; Schools &amp; Hedges, 2022" authorityName="Landestoy &amp; Schools &amp; Hedges" authorityYear="2022" box="[363,492,907,933]" class="Reptilia" family="Diploglossidae" genus="Guarocuyus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B92AEAC6E43CFF89FEEC36EE21DBFC20" box="[363,492,907,933]" italics="true" pageId="7" pageNumber="208">Guarocuyus</emphasis>
</taxonomicName>
<emphasis id="B92AEAC6E43CFF89FE7436EE2260FC20" bold="true" box="[499,599,907,933]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="A21957BDE43CFF89FE7436EE2260FC20" box="[499,599,907,933]" pageId="7" pageNumber="208" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
differs by the presence of an interdigital scaleless web between toes II–IV (vs. absent in
<taxonomicName id="4C5E4D57E43CFF89FF7A36CA216BFC4C" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[253,348,943,969]" class="Reptilia" family="Diploglossidae" genus="Comptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FF7A36CA216BFC4C" box="[253,348,943,969]" italics="true" pageId="7" pageNumber="208">Comptus</emphasis>
</taxonomicName>
, except for vestigial scaly web only between toes III–IV in some
<taxonomicName id="4C5E4D57E43CFF89FB9F36CA252DFC4C" authority="Cope" authorityName="Cope" box="[1048,1306,943,969]" class="Reptilia" family="Diploglossidae" genus="Comptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="species" species="stenurus">
<emphasis id="B92AEAC6E43CFF89FB9F36CA24EFFC4D" box="[1048,1240,943,969]" italics="true" pageId="7" pageNumber="208">Comptus stenurus</emphasis>
Cope
</taxonomicName>
), and by the EW/SVL (2.33–2.90 vs.
<quantity id="4CA69B31E43CFF89FE1836B62213FC68" box="[415,548,979,1005]" metricMagnitude="-2" metricUnit="m" metricValue="3.5941" metricValueMax="5.2578" metricValueMin="1.9304" pageId="7" pageNumber="208" unit="in" value="1.415" valueMax="2.07" valueMin="0.76">0.76–2.07 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FDAC36B622BDFC68" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[555,650,979,1005]" class="Reptilia" family="Diploglossidae" genus="Comptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FDAC36B622BDFC68" box="[555,650,979,1005]" italics="true" pageId="7" pageNumber="208">Comptus</emphasis>
</taxonomicName>
). From
<taxonomicName id="4C5E4D57E43CFF89FD6736B623A4FC68" authority="Cope" authorityName="Cope" authorityYear="1861" box="[736,915,979,1005]" class="Reptilia" family="Diploglossidae" genus="Panolopus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FD6736B62365FC68" box="[736,850,979,1005]" italics="true" pageId="7" pageNumber="208">Panolopus</emphasis>
Cope
</taxonomicName>
,
<taxonomicName id="4C5E4D57E43CFF89FC1A36B62429FC68" authority="Landestoy &amp; Schools &amp; Hedges, 2022" authorityName="Landestoy &amp; Schools &amp; Hedges" authorityYear="2022" box="[925,1054,979,1005]" class="Reptilia" family="Diploglossidae" genus="Guarocuyus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B92AEAC6E43CFF89FC1A36B62429FC68" box="[925,1054,979,1005]" italics="true" pageId="7" pageNumber="208">Guarocuyus</emphasis>
</taxonomicName>
<emphasis id="B92AEAC6E43CFF89FBA236B624BEFC68" bold="true" box="[1061,1161,979,1005]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="A21957BDE43CFF89FBA236B624BEFC68" box="[1061,1161,979,1005]" pageId="7" pageNumber="208" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
differs by the presence of keels on the dorsal scales (vs. absent in adult
<taxonomicName id="4C5E4D57E43CFF89FDF5369222D3FB94" authorityName="Cope" authorityYear="1861" box="[626,740,1015,1041]" class="Reptilia" family="Diploglossidae" genus="Panolopus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FDF5369222D3FB94" box="[626,740,1015,1041]" italics="true" pageId="7" pageNumber="208">Panolopus</emphasis>
</taxonomicName>
), presence of an interdigital scaleless web between toes II–IV (vs. absent in
<taxonomicName id="4C5E4D57E43CFF89FF79317E2147FBB0" authorityName="Cope" authorityYear="1861" box="[254,368,1051,1077]" class="Reptilia" family="Diploglossidae" genus="Panolopus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FF79317E2147FBB0" box="[254,368,1051,1077]" italics="true" pageId="7" pageNumber="208">Panolopus</emphasis>
</taxonomicName>
), EW/SVL (2.33–2.90 vs.
<quantity id="4CA69B31E43CFF89FD09317F2322FBB0" box="[654,789,1050,1077]" metricMagnitude="-2" metricUnit="m" metricValue="3.5687" metricValueMax="5.6896" metricValueMin="1.4478" pageId="7" pageNumber="208" unit="in" value="1.405" valueMax="2.24" valueMin="0.57">0.57–2.24 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FC9C317E23BAFBB0" authorityName="Cope" authorityYear="1861" box="[795,909,1051,1077]" class="Reptilia" family="Diploglossidae" genus="Panolopus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FC9C317E23BAFBB0" box="[795,909,1051,1077]" italics="true" pageId="7" pageNumber="208">Panolopus</emphasis>
</taxonomicName>
), and by the RH/RW (54.1–64.5 vs.
<quantity id="4CA69B31E43CFF89FA91317E25ABFBB0" box="[1302,1436,1050,1077]" metricMagnitude="0" metricUnit="m" metricValue="1.14808" metricValueMax="1.3411199999999999" metricValueMin="0.95504" pageId="7" pageNumber="208" unit="in" value="45.2" valueMax="52.8" valueMin="37.6">37.6–52.8 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FF10315A213EFBDC" authorityName="Cope" authorityYear="1861" box="[151,265,1087,1113]" class="Reptilia" family="Diploglossidae" genus="Panolopus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FF10315A213EFBDC" box="[151,265,1087,1113]" italics="true" pageId="7" pageNumber="208">Panolopus</emphasis>
</taxonomicName>
). From
<taxonomicName id="4C5E4D57E43CFF89FED9315A21C1FBDC" authority="Gray" authorityName="Gray" authorityYear="1852" box="[350,502,1087,1113]" class="Reptilia" family="Diploglossidae" genus="Sauresia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FED9315A218DFBDC" box="[350,442,1087,1113]" italics="true" pageId="7" pageNumber="208">Sauresia</emphasis>
Gray
</taxonomicName>
,
<taxonomicName id="4C5E4D57E43CFF89FD87315A22B6FBDC" authority="Landestoy &amp; Schools &amp; Hedges, 2022" authorityName="Landestoy &amp; Schools &amp; Hedges" authorityYear="2022" box="[512,641,1087,1113]" class="Reptilia" family="Diploglossidae" genus="Guarocuyus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B92AEAC6E43CFF89FD87315A22B6FBDC" box="[512,641,1087,1113]" italics="true" pageId="7" pageNumber="208">Guarocuyus</emphasis>
</taxonomicName>
<emphasis id="B92AEAC6E43CFF89FD00315A22DDFBDC" bold="true" box="[647,746,1087,1113]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="A21957BDE43CFF89FD00315A22DDFBDC" box="[647,746,1087,1113]" pageId="7" pageNumber="208" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
differs by the presence of keels on the dorsal scales (vs. absent in
<taxonomicName id="4C5E4D57E43CFF89FF10310620C4FBF8" authorityName="Gray" authorityYear="1852" box="[151,243,1123,1149]" class="Reptilia" family="Diploglossidae" genus="Sauresia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FF10310620C4FBF8" box="[151,243,1123,1149]" italics="true" pageId="7" pageNumber="208">Sauresia</emphasis>
</taxonomicName>
), number of digits per limb (5 vs.
<quantity id="4CA69B31E43CFF89FDDD310622B4FBF8" box="[602,643,1123,1149]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="7" pageNumber="208" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FD0F310622D3FBF8" authorityName="Gray" authorityYear="1852" box="[648,740,1123,1149]" class="Reptilia" family="Diploglossidae" genus="Sauresia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FD0F310622D3FBF8" box="[648,740,1123,1149]" italics="true" pageId="7" pageNumber="208">Sauresia</emphasis>
</taxonomicName>
), presence of an interdigital scaleless web between toes II–IV (vs. absent in
<taxonomicName id="4C5E4D57E43CFF89FF7B31E2216FFB24" authorityName="Gray" authorityYear="1852" box="[252,344,1159,1185]" class="Reptilia" family="Diploglossidae" genus="Sauresia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FF7B31E2216FFB24" box="[252,344,1159,1185]" italics="true" pageId="7" pageNumber="208">Sauresia</emphasis>
</taxonomicName>
), longest toe lamellae count (21–25 vs.
<quantity id="4CA69B31E43CFF89FD7831E22367FB24" box="[767,848,1159,1185]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="3.048" metricValueMin="2.032" pageId="7" pageNumber="208" unit="in" value="10.0" valueMax="12.0" valueMin="8.0">8–12 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FCD131E22385FB24" authorityName="Gray" authorityYear="1852" box="[854,946,1159,1185]" class="Reptilia" family="Diploglossidae" genus="Sauresia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FCD131E22385FB24" box="[854,946,1159,1185]" italics="true" pageId="7" pageNumber="208">Sauresia</emphasis>
</taxonomicName>
), number of dorsal scales (88–98 vs.
<quantity id="4CA69B31E43CFF89FAB831E2209BFB40" metricMagnitude="0" metricUnit="m" metricValue="2.8956" metricValueMax="3.2257999999999996" metricValueMin="2.5654" pageId="7" pageNumber="208" unit="in" value="114.0" valueMax="127.0" valueMin="101.0">101–127 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FF3231CE2126FB40" authorityName="Gray" authorityYear="1852" box="[181,273,1195,1221]" class="Reptilia" family="Diploglossidae" genus="Sauresia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FF3231CE2126FB40" box="[181,273,1195,1221]" italics="true" pageId="7" pageNumber="208">Sauresia</emphasis>
</taxonomicName>
), EW/SVL (2.33–2.90 vs.
<quantity id="4CA69B31E43CFF89FDBC31CE22F3FB40" box="[571,708,1195,1221]" metricMagnitude="-2" metricUnit="m" metricValue="2.032" metricValueMax="2.9718" metricValueMin="1.0922" pageId="7" pageNumber="208" unit="in" value="0.7999999999999999" valueMax="1.17" valueMin="0.43">0.43–1.17 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FD4A31CE231EFB40" authorityName="Gray" authorityYear="1852" box="[717,809,1195,1221]" class="Reptilia" family="Diploglossidae" genus="Sauresia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FD4A31CE231EFB40" box="[717,809,1195,1221]" italics="true" pageId="7" pageNumber="208">Sauresia</emphasis>
</taxonomicName>
), FN/SVL (3.07–4.48 vs.
<quantity id="4CA69B31E43CFF89FBCD31CF24E4FB40" box="[1098,1235,1194,1221]" metricMagnitude="-2" metricUnit="m" metricValue="5.4102" metricValueMax="6.5024" metricValueMin="4.318" pageId="7" pageNumber="208" unit="in" value="2.13" valueMax="2.56" valueMin="1.7">1.70–2.56 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FB5B31CE250FFB40" authorityName="Gray" authorityYear="1852" box="[1244,1336,1195,1221]" class="Reptilia" family="Diploglossidae" genus="Sauresia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FB5B31CE250FFB40" box="[1244,1336,1195,1221]" italics="true" pageId="7" pageNumber="208">Sauresia</emphasis>
</taxonomicName>
), and by both AGD/SVL and ACD/SVL (53.2–58.7 vs.
<quantity id="4CA69B31E43CFF89FD1731AA232FFB6C" box="[656,792,1231,1257]" metricMagnitude="0" metricUnit="m" metricValue="1.6992600000000002" metricValueMax="1.7754600000000003" metricValueMin="1.6230600000000002" pageId="7" pageNumber="208" unit="in" value="66.9" valueMax="69.9" valueMin="63.9">63.9–69.9 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FCA731AA234BFB6C" authorityName="Gray" authorityYear="1852" box="[800,892,1231,1257]" class="Reptilia" family="Diploglossidae" genus="Sauresia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FCA731AA234BFB6C" box="[800,892,1231,1257]" italics="true" pageId="7" pageNumber="208">Sauresia</emphasis>
</taxonomicName>
, and 62.5–67.7 vs. 69.2–76.3, respectively). From
<taxonomicName id="4C5E4D57E43CFF89FF1031962146FA88" authority="Cochran" authorityName="Cochran" authorityYear="1927" box="[151,369,1267,1293]" class="Reptilia" family="Diploglossidae" genus="Wetmorena" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FF1031962139FA89" box="[151,270,1267,1292]" italics="true" pageId="7" pageNumber="208">Wetmorena</emphasis>
Cochran
</taxonomicName>
,
<taxonomicName id="4C5E4D57E43CFF89FEFD319621CCFA88" authority="Landestoy &amp; Schools &amp; Hedges, 2022" authorityName="Landestoy &amp; Schools &amp; Hedges" authorityYear="2022" box="[378,507,1267,1293]" class="Reptilia" family="Diploglossidae" genus="Guarocuyus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B92AEAC6E43CFF89FEFD319621CCFA88" box="[378,507,1267,1293]" italics="true" pageId="7" pageNumber="208">Guarocuyus</emphasis>
</taxonomicName>
<emphasis id="B92AEAC6E43CFF89FD8631962253FA88" bold="true" box="[513,612,1267,1293]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="A21957BDE43CFF89FD8631962253FA88" box="[513,612,1267,1293]" pageId="7" pageNumber="208" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
differs by the presence of keels on the dorsal scales (vs. absent in
<taxonomicName id="4C5E4D57E43CFF89FA92319625BBFA89" authorityName="Cochran" authorityYear="1927" box="[1301,1420,1267,1292]" class="Reptilia" family="Diploglossidae" genus="Wetmorena" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FA92319625BBFA89" box="[1301,1420,1267,1292]" italics="true" pageId="7" pageNumber="208">Wetmorena</emphasis>
</taxonomicName>
), number of digits per limb (5 vs.
<quantity id="4CA69B31E43CFF89FE6930722221FAB4" box="[494,534,1303,1329]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="7" pageNumber="208" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FD9A307222A3FAB5" authorityName="Cochran" authorityYear="1927" box="[541,660,1303,1328]" class="Reptilia" family="Diploglossidae" genus="Wetmorena" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FD9A307222A3FAB5" box="[541,660,1303,1328]" italics="true" pageId="7" pageNumber="208">Wetmorena</emphasis>
</taxonomicName>
), presence of an interdigital scaleless web between toes II–IV (vs. absent in
<taxonomicName id="4C5E4D57E43CFF89FF36305E211FFAD1" authorityName="Cochran" authorityYear="1927" box="[177,296,1339,1364]" class="Reptilia" family="Diploglossidae" genus="Wetmorena" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FF36305E211FFAD1" box="[177,296,1339,1364]" italics="true" pageId="7" pageNumber="208">Wetmorena</emphasis>
</taxonomicName>
), longest toe lamellae count (21–25 vs.
<quantity id="4CA69B31E43CFF89FD4E305E232FFAD0" box="[713,792,1339,1365]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="3.048" metricValueMin="2.032" pageId="7" pageNumber="208" unit="in" value="10.0" valueMax="12.0" valueMin="8.0">8–12 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FC9A305E23A3FAD1" authorityName="Cochran" authorityYear="1927" box="[797,916,1339,1364]" class="Reptilia" family="Diploglossidae" genus="Wetmorena" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FC9A305E23A3FAD1" box="[797,916,1339,1364]" italics="true" pageId="7" pageNumber="208">Wetmorena</emphasis>
</taxonomicName>
), EW/SVL (2.33–2.90 vs.
<quantity id="4CA69B31E43CFF89FB2A305E2527FAD0" box="[1197,1296,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="8.001" metricValueMax="16.002" metricValueMin="0.0" pageId="7" pageNumber="208" unit="in" value="0.315" valueMax="0.63" valueMin="0.0">0–0.63 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FA92305E25BBFAD1" authorityName="Cochran" authorityYear="1927" box="[1301,1420,1339,1364]" class="Reptilia" family="Diploglossidae" genus="Wetmorena" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FA92305E25BBFAD1" box="[1301,1420,1339,1364]" italics="true" pageId="7" pageNumber="208">Wetmorena</emphasis>
</taxonomicName>
), FN/SVL (3.07–4.48 vs.
<quantity id="4CA69B31E43CFF89FE10303A222BFAFC" box="[407,540,1375,1401]" metricMagnitude="-2" metricUnit="m" metricValue="5.2832" metricValueMax="6.8326" metricValueMin="3.7338" pageId="7" pageNumber="208" unit="in" value="2.08" valueMax="2.69" valueMin="1.47">1.47–2.69 in</quantity>
<taxonomicName id="4C5E4D57E43CFF89FDA4303A22ADFAFD" authorityName="Cochran" authorityYear="1927" box="[547,666,1375,1400]" class="Reptilia" family="Diploglossidae" genus="Wetmorena" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B92AEAC6E43CFF89FDA4303A22ADFAFD" box="[547,666,1375,1400]" italics="true" pageId="7" pageNumber="208">Wetmorena</emphasis>
</taxonomicName>
), and by the AGD/SVL (53.2–58.7 vs. 59.9–65.7).
</paragraph>
</subSubSection>
<subSubSection id="C344655FE43CFF89FF4030E725A0FA18" box="[199,1431,1410,1437]" pageId="7" pageNumber="208" type="description">
<paragraph id="8BE136D4E43CFF89FF4030E725A0FA18" blockId="7.[151,1437,150,1437]" box="[199,1431,1410,1437]" pageId="7" pageNumber="208">
<emphasis id="B92AEAC6E43CFF89FF4030E72118FA19" bold="true" box="[199,303,1410,1436]" pageId="7" pageNumber="208">Content.</emphasis>
The new genus currently bears a single species,
<taxonomicName id="4C5E4D57E43CFF89FCD730E62468FA19" authority="Landestoy &amp; Schools &amp; Hedges, 2022" authorityName="Landestoy &amp; Schools &amp; Hedges" authorityYear="2022" box="[848,1119,1411,1437]" class="Reptilia" family="Diploglossidae" genus="Guarocuyus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="species" species="jaraguanus" status="sp. nov.">
<emphasis id="B92AEAC6E43CFF89FCD730E62468FA19" box="[848,1119,1411,1437]" italics="true" pageId="7" pageNumber="208">Guarocuyus jaraguanus</emphasis>
</taxonomicName>
<emphasis id="B92AEAC6E43CFF89FBE230E62488FA18" bold="true" box="[1125,1215,1411,1437]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="A21957BDE43CFF89FBE230E62488FA18" box="[1125,1215,1411,1437]" pageId="7" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(described below).
</paragraph>
</subSubSection>
</treatment>
</document>
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