264 lines
37 KiB
XML
264 lines
37 KiB
XML
<document id="6A8BDF0298EFB6957336C5EBD6F7905F" ID-DOI="10.1649/0010-065X-71.4.679" ID-ISSN="1938-4394" ID-Zenodo-Dep="5377913" ID-ZooBank="urn:lsid:zoobank.org:pub:379A516C-9DC2-41A8-9B60-B8AD506A968B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630283693544" checkinUser="diego" docAuthor="Liebherr, James K." docDate="2017" docId="03F787BFFF89FF9049DDFE757CB3FCE3" docLanguage="en" docName="0010-065X-71.4.679.pdf" docOrigin="The Coleopterists Bulletin 71 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-71.4.679" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Mecyclothorax medioconstrictus Liebherr 2017, new species" docType="treatment" docUuid="E08578DD-C91A-475F-AC8E-83D4DC8920A7" docUuidSource="ZooBank" docVersion="5" lastPageNumber="688" masterDocId="FFCEFFC7FF80FF9A4963FFC97E46FFB7" masterDocTitle="Review of Mecyclothorax Sharp (Coleoptera: Carabidae: Moriomorphini) from Papua New Guinea, with Descriptions of Five New Species" masterLastPageNumber="703" masterPageNumber="679" pageNumber="687" updateTime="1699213668230" updateUser="plazi">
|
||
<mods:mods id="11F7339FD0A69C7E246732760F82B3DE" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="3E1511E816E2F7050B383326E790B394">
|
||
<mods:title id="A1BF71B3B8C8178A1C25D26571F402A3">Review of Mecyclothorax Sharp (Coleoptera: Carabidae: Moriomorphini) from Papua New Guinea, with Descriptions of Five New Species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="143DC6814628E7C2C4497BD12983435F" type="personal">
|
||
<mods:role id="FFDE624098EBA702352CC3877103D702">
|
||
<mods:roleTerm id="E7D84649E9027545D984C3D074CE53FF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6F2882DD8E1CD467FE47C25F9A27D1BB">Liebherr, James K.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="075859822FBE16650CA2E6D67B526A6F">text</mods:typeOfResource>
|
||
<mods:relatedItem id="9380B2F5EB670590EABBAC4D6AFCCE65" type="host">
|
||
<mods:titleInfo id="D8EA1DAA67EBF2113A475FF59C7245F3">
|
||
<mods:title id="8C1B760CE43EB61C355F6F2759BD09CB">The Coleopterists Bulletin</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="26308DE20F3430B9DB227F7723A76B8E">
|
||
<mods:date id="73E2A49ADD152D05F7044B903332E2B7">2017</mods:date>
|
||
<mods:detail id="FC8446659EF5AA4C4EE307FD970EB455" type="pubDate">
|
||
<mods:number id="F4E334595730FF59ED36A31A13159617">2017-12-22</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="6ED54C02A58661E80E028BDDFAE05FAD" type="volume">
|
||
<mods:number id="DBD60125ACA5F115AE4FCB267BE9978A">71</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="C616E094023380261F5E23C82EE66A6B" type="issue">
|
||
<mods:number id="D8B8BF541191052547DA9D8A53436754">4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="ED44AD8058F2BD0C6EAD57B3CAE02BD2" unit="page">
|
||
<mods:start id="D978809B8F85CD8A8453C000C78D06B0">679</mods:start>
|
||
<mods:end id="25FCCD4012C36AA3E0C053021EDB797F">703</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="ABB77472F0224CA15534DB4E64386B8C">
|
||
<mods:url id="52A87CE37FC79131429CBCF93C7FD333">http://dx.doi.org/10.1649/0010-065x-71.4.679</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="D2A2CCFECBECF88E1E553B40D96094A4">journal article</mods:classification>
|
||
<mods:identifier id="633D7E9AF013CE187069CE712E525781" type="DOI">10.1649/0010-065X-71.4.679</mods:identifier>
|
||
<mods:identifier id="BDD8F1803250B1CCDFA890573BED1F01" type="ISSN">1938-4394</mods:identifier>
|
||
<mods:identifier id="B190764CD829E658512F7DD0C1EB8261" type="Zenodo-Dep">5377913</mods:identifier>
|
||
<mods:identifier id="CE6C67449FE0BDE37A3DCC4AC31F7DE4" type="ZooBank">urn:lsid:zoobank.org:pub:379A516C-9DC2-41A8-9B60-B8AD506A968B</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03F787BFFF89FF9049DDFE757CB3FCE3" ID-DOI="http://doi.org/10.5281/zenodo.5465040" ID-Zenodo-Dep="5465040" LSID="urn:lsid:zoobank.org:act:E08578DD-C91A-475F-AC8E-83D4DC8920A7" httpUri="http://treatment.plazi.org/id/03F787BFFF89FF9049DDFE757CB3FCE3" lastPageId="10" lastPageNumber="688" pageId="9" pageNumber="687">
|
||
<subSubSection id="C3446522FF89FF9349DDFE757F9AFDB1" pageId="9" pageNumber="687" type="nomenclature">
|
||
<paragraph id="8BE136A9FF89FF9349DDFE757F9AFDB1" blockId="9.[190,641,444,518]" pageId="9" pageNumber="687">
|
||
<heading id="D0A981C5FF89FF9349DDFE757F9CFE5B" centered="true" fontSize="9" level="2" pageId="9" pageNumber="687" reason="2">
|
||
<emphasis id="B92AEABBFF89FF9349DDFE757F9CFE5B" bold="true" pageId="9" pageNumber="687">
|
||
4.
|
||
<taxonomicName id="4C5E4D2AFF89FF9349BBFE757C3BFE66" ID-CoL="8MGZF" authority="Liebherr" authorityName="Liebherr" authorityYear="2017" box="[216,637,444,465]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="687" phylum="Arthropoda" rank="species" species="medioconstrictus" status="sp. nov.">
|
||
<emphasis id="B92AEABBFF89FF9349BBFE757C5CFE66" bold="true" box="[216,538,444,465]" italics="true" pageId="9" pageNumber="687">Mecyclothorax medioconstrictus</emphasis>
|
||
Liebherr
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A21957C0FF89FF934806FE1E7F9CFE5B" box="[357,474,471,492]" pageId="9" pageNumber="687" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
(
|
||
<figureCitation id="13652A2CFF89FF934808FE387FE9FDB1" box="[363,431,497,518]" captionStart="Figs" captionStartId="4.[175,214,1615,1634]" captionTargetBox="[154,1168,198,1580]" captionTargetId="figure-0@4.[151,1173,194,1599]" captionTargetPageId="4" captionText="Figs. 1–4. Mecyclothorax species, dorsal habitus. 1) M. kavanaughi; 2) M. baehri; 3) M. medioconstrictus; 4) M. andersoni. Range of standardized body lengths (in mm) and setal formulae (see text) for number of supraorbital setae each side, number of pronotal setae each side, number of dorsal elytral setae each elytron, and number of setae at the apex of each elytron are indicated." figureDoi="http://doi.org/10.5281/zenodo.5377915" httpUri="https://zenodo.org/record/5377915/files/figure.png" pageId="9" pageNumber="687">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="13652A2CFF89FF9348DFFE387F93FDB1" box="[444,469,497,518]" captionStart="Fig" captionStartId="17.[197,228,896,915]" captionTargetBox="[175,1191,199,878]" captionTargetId="figure-399@17.[173,1195,194,881]" captionTargetPageId="17" captionText="Fig. 32. Localities in Papua New Guinea from which Mecyclothorax spp. were collected and the particular species collected at each locality. 1 = Mt. Capella — M. baehri; 2 = Mt. Fugilil — M. brispex and M. medioconstrictus; 3 = Kubor Range — M. kubor; 4 = Mt. Wilhelm — M. sedlaceki; 5 = Finisterre Range — M. kavanaughi; 6 = Mt. Missim — M. gressitti; 7 = Mt. Kaindi, Edie Creek and Bulldog Road — M. andersoni, M. cuccodoroi, and M. loebli; 8 = Mt. Amungwiwa—M. amingwiwae." figureDoi="http://doi.org/10.5281/zenodo.5377933" httpUri="https://zenodo.org/record/5377933/files/figure.png" pageId="9" pageNumber="687">32</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3446522FF89FF9349A6FDD57C1FFC6D" pageId="9" pageNumber="687" type="diagnosis">
|
||
<paragraph id="8BE136A9FF89FF9349A6FDD57C1FFC6D" blockId="9.[173,660,540,1730]" pageId="9" pageNumber="687">
|
||
<emphasis id="B92AEABBFF89FF9349A6FDD57F6AFD86" bold="true" box="[197,300,540,561]" pageId="9" pageNumber="687">Diagnosis.</emphasis>
|
||
As noted above, this species and
|
||
<taxonomicName id="4C5E4D2AFF89FF934B1AFDD57EADFDFB" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="687" phylum="Arthropoda" rank="species" species="baehri">
|
||
<emphasis id="B92AEABBFF89FF934B1AFDD57EADFDFB" italics="true" pageId="9" pageNumber="687">M. baehri</emphasis>
|
||
</taxonomicName>
|
||
are the only two
|
||
<taxonomicName id="4C5E4D2AFF89FF9348E9FDFE7C26FDFB" box="[394,608,567,588]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="687" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B92AEABBFF89FF9348E9FDFE7C5EFDFB" box="[394,536,567,588]" italics="true" pageId="9" pageNumber="687">Mecyclothorax</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3497639FF89FF9349CEFD987F22FDD1" box="[173,356,593,614]" name="Papua New Guinea" pageId="9" pageNumber="687">Papua New Guinea</collectingCountry>
|
||
characterized by a quadrisetose pronotum, three dorsal elytral setae, and presence of both subapical and apical elytral setae. Both species comprise large-bodied beetles — the
|
||
<typeStatus id="54E5880BFF89FF934B60FD687C13FD01" box="[515,597,673,694]" pageId="9" pageNumber="687" type="holotype">holotype</typeStatus>
|
||
of this species has a standardized body length 5.0 mm — that exhibit a medially constricted body form. The pronotal base for
|
||
<taxonomicName id="4C5E4D2AFF89FF934839FD397C5CFCB2" authorityName="Liebherr" authorityYear="2017" box="[346,538,752,773]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="687" phylum="Arthropoda" rank="species" species="medioconstrictus">
|
||
<emphasis id="B92AEABBFF89FF934839FD397C5CFCB2" box="[346,538,752,773]" italics="true" pageId="9" pageNumber="687">M. medioconstrictus</emphasis>
|
||
</taxonomicName>
|
||
is relatively narrow, and the elytral lateral margins are narrowly curved posteriorly laterad and behind the humerus (
|
||
<figureCitation id="13652A2CFF89FF9349D7FC897EB0FCE2" box="[180,246,832,853]" captionStart="Figs" captionStartId="4.[175,214,1615,1634]" captionTargetBox="[154,1168,198,1580]" captionTargetId="figure-0@4.[151,1173,194,1599]" captionTargetPageId="4" captionText="Figs. 1–4. Mecyclothorax species, dorsal habitus. 1) M. kavanaughi; 2) M. baehri; 3) M. medioconstrictus; 4) M. andersoni. Range of standardized body lengths (in mm) and setal formulae (see text) for number of supraorbital setae each side, number of pronotal setae each side, number of dorsal elytral setae each elytron, and number of setae at the apex of each elytron are indicated." figureDoi="http://doi.org/10.5281/zenodo.5377915" httpUri="https://zenodo.org/record/5377915/files/figure.png" pageId="9" pageNumber="687">Fig. 3</figureCitation>
|
||
). This species can be diagnosed from
|
||
<taxonomicName id="4C5E4D2AFF89FF9349CEFC927F4DFCC7" box="[173,267,859,880]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="687" phylum="Arthropoda" rank="species" species="baehri">
|
||
<emphasis id="B92AEABBFF89FF9349CEFC927F4DFCC7" box="[173,267,859,880]" italics="true" pageId="9" pageNumber="687">M. baehri</emphasis>
|
||
</taxonomicName>
|
||
by the indistinctly punctate pronotal median base versus deeply and distinctly punctate base in
|
||
<taxonomicName id="4C5E4D2AFF89FF9349AFFC597F68FC12" box="[204,302,912,933]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="687" phylum="Arthropoda" rank="species" species="baehri">
|
||
<emphasis id="B92AEABBFF89FF9349AFFC597F68FC12" box="[204,302,912,933]" italics="true" pageId="9" pageNumber="687">M. baehri</emphasis>
|
||
</taxonomicName>
|
||
, and the isodiametric mesh microsculpture on the discal elytral intervals versus the transverse mesh microsculpture in
|
||
<taxonomicName id="4C5E4D2AFF89FF934895FC0C7C12FC6D" box="[502,596,965,986]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="687" phylum="Arthropoda" rank="species" species="baehri">
|
||
<emphasis id="B92AEABBFF89FF934895FC0C7C12FC6D" box="[502,596,965,986]" italics="true" pageId="9" pageNumber="687">M. baehri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3446522FF89FF9049A6FC297F2EFA2A" lastPageId="10" lastPageNumber="688" pageId="9" pageNumber="687" type="description">
|
||
<paragraph id="8BE136A9FF89FF9049A6FC297C07FCA8" blockId="9.[173,660,540,1730]" lastBlockId="10.[151,638,194,1729]" lastPageId="10" lastPageNumber="688" pageId="9" pageNumber="687">
|
||
<emphasis id="B92AEABBFF89FF9349A6FC297FC0FC42" bold="true" box="[197,390,992,1013]" pageId="9" pageNumber="687">Description. Head:</emphasis>
|
||
Frons broadly convex and bordered by shallow, moderately broad, subparallel frontal grooves that extend anteriorly from immediately mesad anterior supraorbital seta, are deepest at juncture with shallow frontoclypeal suture, and extend onto basolateral reaches of clypeus; eyes small in diameter, moderately protruding on an ocular lobe that obtusely meets gena behind the eye (
|
||
<figureCitation id="13652A2CFF89FF9349D7FB7D7EB6FB7E" box="[180,240,1204,1225]" captionStart="Figs" captionStartId="4.[175,214,1615,1634]" captionTargetBox="[154,1168,198,1580]" captionTargetId="figure-0@4.[151,1173,194,1599]" captionTargetPageId="4" captionText="Figs. 1–4. Mecyclothorax species, dorsal habitus. 1) M. kavanaughi; 2) M. baehri; 3) M. medioconstrictus; 4) M. andersoni. Range of standardized body lengths (in mm) and setal formulae (see text) for number of supraorbital setae each side, number of pronotal setae each side, number of dorsal elytral setae each elytron, and number of setae at the apex of each elytron are indicated." figureDoi="http://doi.org/10.5281/zenodo.5377915" httpUri="https://zenodo.org/record/5377915/files/figure.png" pageId="9" pageNumber="687">Fig. 3</figureCitation>
|
||
), ocular ratio = 1.50, ocular lobe ratio = 0.81; 3
|
||
<superScript id="7C2B9BE1FF89FF934994FB007F40FB60" attach="left" box="[247,262,1225,1239]" fontSize="6" pageId="9" pageNumber="687">rd</superScript>
|
||
antennomere largely glabrous, though several small setae present basad apical setal ring; antenna moderately elongate, antennomere 9 length 2.1X maximal breadth; anterior labral margin broadly, moderately concave; mandibles of moderate length, distance from dorsal condyle to apex 1.82X distance from condyle to anterolateral margin of labrum; vertex with broad, shallow, transverse depression, or neck, behind eyes; mentum tooth obtuse, apex broadly rounded; ligular apex truncate, broad, 2 ligular setae well separated, paraglossae thin and only slightly extending beyond ligular margin.
|
||
<emphasis id="B92AEABBFF89FF934861F9C47F36F995" bold="true" box="[258,368,1549,1570]" pageId="9" pageNumber="687">Pronotum:</emphasis>
|
||
Narrow, basally constricted, MPW/PL = 1.19, MPW/BPW = 1.48; pronotal lateral seta set on disc 2 setal diameters inside lateral marginal depression; hind angle a distinct, obtuse denticle with lateral margins on the denticle subparallel each side of pronotum, hind seta at or just before obtuse hind angle; median base slightly depressed relative to disc, sparsely, minutely punctate, with 14–16 small punctures each side; median longitudinal impression finely and narrowly impressed at front of median base, shallow and irregular on pronotal disc; anterior transverse impression narrow, finely impressed to border broadly convex anterior callosity, lateral impression deeper and more well defined in lateral 2/3 of length to front angle; front margin with indistinct, narrow convexity medially, though the marginal impression too shallow to call the convexity a marginal bead; front angles not protruding, tightly rounded; lateral marginal depression narrow at position of lateral seta, narrowly lined with microsculpture at depth, broader with irregular transverse wrinkles just anterad hind angle; laterobasal depression a widening of lateral marginal depression inside basal denticles, its medial surface roughened due to presence of 7 punctures; prosternum with distinct anteapical furrow, groove smooth, deep laterally, shallower and broader ventromedially; prosternal process convex anterad procoxal cavities, shallowly bisulcate ventrally between coxae; proepimeron bordered anteriorly and posteriorly by smooth grooves.
|
||
<emphasis id="B92AEABBFF89FF934A49FC897D34FCE2" bold="true" box="[810,882,832,853]" pageId="9" pageNumber="687">Elytra:</emphasis>
|
||
Ellipsoid, humeri narrowly rounded posterad the slightly recurved basal elytral groove, MEW/EL = 0.73, maximum width slightly behind midlength; parascutellar seta present; scutellum narrowly triangular, ratio of width at transverse groove/median length = 1.14; parascutellar striole a continuous shallow groove with 3–4 punctures; striae 1–4 deep, continuous on disc, 4
|
||
<superScript id="7C2B9BE1FF89FF934DF8FC3D7AEFFBB5" attach="none" box="[1179,1193,1012,1026]" fontSize="6" pageId="9" pageNumber="687">th</superScript>
|
||
striae distinctly punctate, inner striae nearly smooth; stria 5 a series of isolated punctures, stria 6 evidenced by shallow elongate impressions, stria 7 represented by sporadic, shallow, rounded impressions; striae 2–7 reduced on apex, only sutural stria deep and smooth, stria 2 shallow and irregularly punctate, stria 7 impressed in vicinity of subapical seta, striae 8 smoothly and deeply impressed at midlength; lateral marginal depression narrow, margin beaded throughout length; subapical sinuation broadly and shallowly impressed, elytral epipleuron narrowly visible in anterior half of sinuation; lateral elytral setae 7 + 6–7.
|
||
<emphasis id="B92AEABBFF89FF934D3AFA9A7D58FA34" bold="true" pageId="9" pageNumber="687">Mesepisternum:</emphasis>
|
||
With nine dimpled impressions in 1–3 dorsoventral rows; metepisternal maximum width/ lateral length = 0.65; metasternal process acute, apex tightly rounded, margin broadest and most convex medially; metathoracic wings vestigial, apex not extending beyond posterior margin of metanotum.
|
||
<emphasis id="B92AEABBFF89FF934A5AF9C47DE5F995" bold="true" box="[825,931,1549,1570]" pageId="9" pageNumber="687">Abdomen:</emphasis>
|
||
Ventrites 1 and 2 separated laterally by anteriorly sinuous suture, surface of ventrite 2 depressed within convex sinuosity; suture between ventrites 2 and 3 shallow but traceable laterally; broad shallow depressions present laterally on ventrites 3–6.
|
||
<emphasis id="B92AEABBFF89FF934AECF95B7A75F910" bold="true" box="[911,1075,1682,1703]" pageId="9" pageNumber="687">Microsculpture:</emphasis>
|
||
Reduced on head, frons glossy, vertex with shallow, transversely stretched sculpticells; pronotal disc covered with shallow elongate transverse mesh, sculpticell breadth 3–4X length, mixed with transverse lines; pronotal base with slightly transverse mesh between punctures; elytral disc with transversely stretched isodiametric mesh, apex with shallow isodiametric mesh, sculpticells flat; metasternum with welldeveloped transverse mesh; lateral portions of basal abdominal ventrites with swirling isodiametric and transverse sculpticells.
|
||
<emphasis id="B92AEABBFF8AFF904B6AFE787C3BFE71" bold="true" box="[521,637,433,454]" pageId="10" pageNumber="688">Coloration:</emphasis>
|
||
Head capsule piceous, clypeus and mandibles rufopiceous; antennae flavobrunneous, antennomeres 4–11 with piceous cast; pronotal disc piceous, anterior and posterior margins narrowly dark rufous; elytral disc piceous, sutural interval and lateral elytral margins concolorous, apical margin from posterior setae of lateral elytral setal series narrowly rufobrunneous; proepipleuron, proepisternum, elytral epipleuron, and metepisternum piceous; abdomen rufopiceous with ventrites 3–5 amber brunneous along posterior margins, apical ventrite 6 with apical third rufoflavous; femora dark rufous, tibiae dark rufous with piceous cast.
|
||
</paragraph>
|
||
<paragraph id="8BE136A9FF8AFF9049CCFCEC7F2EFA2A" blockId="10.[151,638,194,1729]" pageId="10" pageNumber="688">
|
||
<emphasis id="B92AEABBFF8AFF9049CCFCEC7F8DFC8D" bold="true" box="[175,459,805,826]" pageId="10" pageNumber="688">Female Reproductive Tract.</emphasis>
|
||
Based on external anatomical characters, this species exhibits a sistertaxon relationship with
|
||
<taxonomicName id="4C5E4D2AFF8AFF9048E9FC937FA8FCD8" box="[394,494,858,879]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="688" phylum="Arthropoda" rank="species" species="baehri">
|
||
<emphasis id="B92AEABBFF8AFF9048E9FC937FA8FCD8" box="[394,494,858,879]" italics="true" pageId="10" pageNumber="688">M. baehri</emphasis>
|
||
</taxonomicName>
|
||
in a cladistic analysis including eight of the 11 Papuan New
|
||
<collectingCountry id="F3497639FF8AFF9049F4FC467E9DFC13" box="[151,219,911,932]" name="Guinea-Bissau" pageId="10" pageNumber="688">Guinea</collectingCountry>
|
||
species reviewed here, plus 50 other taxa including
|
||
<taxonomicName id="4C5E4D2AFF8AFF904999FC637F93FC08" box="[250,469,938,959]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="688" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B92AEABBFF8AFF904999FC637FCEFC08" box="[250,392,938,959]" italics="true" pageId="10" pageNumber="688">Mecyclothorax</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
from throughout the generic distribution and moriomorphine outgroups (unpublished data). As such, the generic combination with
|
||
<taxonomicName id="4C5E4D2AFF8AFF90482DFC307F9AFBB9" box="[334,476,1017,1038]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="688" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92AEABBFF8AFF90482DFC307F9AFBB9" box="[334,476,1017,1038]" italics="true" pageId="10" pageNumber="688">Mecyclothorax</emphasis>
|
||
</taxonomicName>
|
||
is substantiated without the necessity to dissect the unique female
|
||
<typeStatus id="54E5880BFF8AFF9049F4FBE77EACFBF4" box="[151,234,1070,1091]" pageId="10" pageNumber="688" type="holotype">holotype</typeStatus>
|
||
for assessment of internal female reproductive tract characters. Nonetheless, the specimen was relaxed in warm water, allowing slight eversion of the gonocoxae, thereby permitting the following character assessments; basal gonocoxite 1 with 3 apical fringe setae and larger apicomedial seta present (see
|
||
<figureCitation id="13652A2CFF8AFF904833FB077F8FFB54" box="[336,457,1229,1251]" captionStart="Figs" captionStartId="15.[197,236,789,808]" captionTargetBox="[180,1191,201,764]" captionTargetId="figure-456@15.[173,1195,194,774]" captionTargetPageId="15" captionText="Figs. 29–31. Mecyclothorax species, female gonocoxae, right side, ventral view. 29) M. andersoni; 30) M. brispex; 31) M. gressitti. Abbreviations aredefined in Table 1; scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5377931" httpUri="https://zenodo.org/record/5377931/files/figure.png" pageId="10" pageNumber="688">Figs. 29–31</figureCitation>
|
||
for homologies); apical gonocoxite 2 broad basally, with base extended by lateral apodeme; apical gonocoxite 2 with 2 lateral ensiform setae present, the setae less than 0.4X the gonocoxite length and situated on the lateral margin of the gonocoxite; apical nematiform setae present in fossa situated less than 0.25X coxite length from the apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3446522FF8AFF9049CCFA6B7D91FE91" pageId="10" pageNumber="688" type="materials_examined">
|
||
<paragraph id="8BE136A9FF8AFF9049CCFA6B7D91FE91" blockId="10.[151,638,194,1729]" lastBlockId="10.[685,1173,194,852]" pageId="10" pageNumber="688">
|
||
<materialsCitation id="3B363CF4FF8AFF9049CCFA6B7FEEF9B0" collectionCode="EMEC, NMNHS, MT, BERON" country="Guinea-Bissau" county="New" elevation="3450" location="British Speleological" municipality="Papua" pageId="10" pageNumber="688" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B92AEABBFF8AFF9049CCFA6B7F57FA00" bold="true" box="[175,273,1442,1463]" pageId="10" pageNumber="688">
|
||
<typeStatus id="54E5880BFF8AFF9049CCFA6B7F4BFA00" box="[175,269,1442,1463]" pageId="10" pageNumber="688" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Female (
|
||
<collectionCode id="ED4FAE6CFF8AFF904819FA6B7F86FA00" box="[378,448,1442,1463]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="10" pageNumber="688" type="University or college">EMEC</collectionCode>
|
||
, for deposition in
|
||
<collectionCode id="ED4FAE6CFF8AFF9049F4FA747EA9FA65" box="[151,239,1469,1490]" pageId="10" pageNumber="688">NMNHS</collectionCode>
|
||
):
|
||
<collectionCode id="ED4FAE6CFF8AFF90499FFA747F59FA65" box="[252,287,1469,1490]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="10" pageNumber="688">MT</collectionCode>
|
||
FUGILIL / summit
|
||
<quantity id="4CA69B4CFF8AFF9048B2FA747C5EFA65" box="[465,536,1469,1490]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="10" pageNumber="688" unit="m" value="3450.0">
|
||
<elevation id="0073D19AFF8AFF9048B2FA747C5EFA65" box="[465,536,1469,1490]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="10" pageNumber="688" unit="m" value="3450.0">3450 m</elevation>
|
||
</quantity>
|
||
. / 29.ix.75 / P.
|
||
<collectionCode id="ED4FAE6CFF8AFF9049D1FA1E7F47FA5B" box="[178,257,1495,1516]" pageId="10" pageNumber="688">BERON</collectionCode>
|
||
//
|
||
<location id="8E816072FF8AFF904842FA1E7FA2FA5B" LSID="urn:lsid:plazi:treatment:03F787BFFF89FF9049DDFE757CB3FCE3:8E816072FF8AFF904842FA1E7FA2FA5B" box="[289,484,1495,1516]" country="Guinea-Bissau" county="New" municipality="Papua" name="British Speleological" pageId="10" pageNumber="688">British Speleological</location>
|
||
/
|
||
<location id="8E816072FF8AFF90489DFA1E7C24FA5B" LSID="urn:lsid:plazi:treatment:03F787BFFF89FF9049DDFE757CB3FCE3:8E816072FF8AFF90489DFA1E7C24FA5B" box="[510,610,1495,1516]" country="Guinea-Bissau" county="New" municipality="Papua" name="Expedition" pageId="10" pageNumber="688">Expedition</location>
|
||
to
|
||
<collectingMunicipality id="6B85ACD3FF8AFF9049F4FA3B7E89F9B0" box="[151,207,1522,1543]" pageId="10" pageNumber="688">Papua</collectingMunicipality>
|
||
/
|
||
<collectingCounty id="62804E25FF8AFF904984FA3B7F55F9B0" box="[231,275,1522,1543]" pageId="10" pageNumber="688">New</collectingCounty>
|
||
<collectingCountry id="F3497639FF8AFF904878FA3B7F1BF9B0" box="[283,349,1522,1543]" name="Guinea-Bissau" pageId="10" pageNumber="688">Guinea</collectingCountry>
|
||
1975 //
|
||
</materialsCitation>
|
||
<materialsCitation id="3B363CF4FF8AFF9048D2FA3B7D94FE91" collectionCode="EMEC, NMNHS" collectorName="J. K. Liebherr & This & Barry Moore" country="Bulgaria" location="Essig Museum of Entomology" municipality="National Museum of Natural History" pageId="10" pageNumber="688" specimenCount="1" stateProvince="Sofiya-Grad" typeStatus="holotype">
|
||
<typeStatus id="54E5880BFF8AFF9048D2FA3B7C6DF9B0" box="[433,555,1522,1543]" pageId="10" pageNumber="688" type="holotype">HOLOTYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="4C5E4D2AFF8AFF904B21FA3B7EAAF996" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="688" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
|
||
/ medioconstrictus /
|
||
<collectorName id="26AB537FFF8AFF9048A7F9C57C02F996" box="[452,580,1548,1569]" pageId="10" pageNumber="688">J.K. Liebherr</collectorName>
|
||
2017 [black-bordered red label].
|
||
<collectorName id="26AB537FFF8AFF9048EDF9EE7FF1F98B" box="[398,439,1575,1596]" pageId="10" pageNumber="688">This</collectorName>
|
||
specimen was part of the
|
||
<collectorName id="26AB537FFF8AFF9049DBF98B7F6BF9E0" box="[184,301,1602,1623]" pageId="10" pageNumber="688">Barry Moore</collectorName>
|
||
personal collection of carabid beetles transferred to the
|
||
<location id="8E816072FF8AFF90482CF9957C38F9C6" LSID="urn:lsid:plazi:treatment:03F787BFFF89FF9049DDFE757CB3FCE3:8E816072FF8AFF90482CF9957C38F9C6" box="[335,638,1628,1649]" country="Bulgaria" municipality="National Museum of Natural History" name="Essig Museum of Entomology" pageId="10" pageNumber="688" stateProvince="Sofiya-Grad">Essig Museum of Entomology</location>
|
||
(
|
||
<collectionCode id="ED4FAE6CFF8AFF9049FDF9BE7EA5F93B" box="[158,227,1655,1676]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="10" pageNumber="688" type="University or college">EMEC</collectionCode>
|
||
). The specimen no doubt comprised part of a loan that resulted in descriptions of three cave species based on the British Speleological Expedition (
|
||
<bibRefCitation id="EFCF4B58FF8AFF904B5CF9657C98FF60" author="Moore, B. P." box="[575,734,194,1729]" pageId="10" pageNumber="688" pagination="357 - 363" refId="ref18926" refString="Moore, B. P. 1977. A remarkable new genus and three new species of cavernicolous Carabidae (Coleoptera) (Zoological results of the British Speleological Expedition to Papua-New-Guinea 1975). International Journal of Speleology 78: 357 - 363." type="journal article" year="1977">Moore 1977</bibRefCitation>
|
||
). As such, it was returned to the
|
||
<collectionCode id="ED4FAE6CFF8AFF904D4DFF0B7A29FF60" box="[1070,1135,194,215]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="10" pageNumber="688" type="University or college">EMEC</collectionCode>
|
||
for accessioning and will be forwarded to the
|
||
<collectingMunicipality id="6B85ACD3FF8AFF904D25FF157DF1FEBB" pageId="10" pageNumber="688">National Museum of Natural History</collectingMunicipality>
|
||
(
|
||
<collectionCode id="ED4FAE6CFF8AFF904AA4FF3E7A58FEBB" box="[967,1054,247,268]" pageId="10" pageNumber="688">NMNHS</collectionCode>
|
||
,
|
||
<collectingRegion id="499AF84BFF8AFF904D4FFF3E7A18FEBB" box="[1068,1118,247,268]" country="Bulgaria" name="Sofiya-Grad" pageId="10" pageNumber="688">Sofia</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3497639FF8AFF904D08FF3E7C9DFE91" name="Bulgaria" pageId="10" pageNumber="688">Bulgaria</collectingCountry>
|
||
) for permanent deposition
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3446522FF8AFF904BA6FEE57A32FE27" pageId="10" pageNumber="688" type="etymology">
|
||
<paragraph id="8BE136A9FF8AFF904BA6FEE57A32FE27" blockId="10.[685,1173,194,852]" pageId="10" pageNumber="688">
|
||
<emphasis id="B92AEABBFF8AFF904BA6FEE57D7EFEF6" bold="true" box="[709,824,300,321]" pageId="10" pageNumber="688">Etymology.</emphasis>
|
||
The adjectival species epithet,
|
||
<taxonomicName id="4C5E4D2AFF8AFF904BCEFE8F7D09FEEC" authorityName="Liebherr" authorityYear="2017" box="[685,847,326,347]" class="Insecta" family="Carabidae" genus="Mecyclothorax" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="688" phylum="Arthropoda" rank="species" species="medioconstrictus">
|
||
<emphasis id="B92AEABBFF8AFF904BCEFE8F7D09FEEC" box="[685,847,326,347]" italics="true" pageId="10" pageNumber="688">medioconstrictus</emphasis>
|
||
</taxonomicName>
|
||
, denotes the hourglass body shape of the adults of this species, with their broad head, cordate pronotum, and ellipsoid elytra (
|
||
<figureCitation id="13652A2CFF8AFF904D4FFEB27A21FE27" box="[1068,1127,379,400]" captionStart="Figs" captionStartId="4.[175,214,1615,1634]" captionTargetBox="[154,1168,198,1580]" captionTargetId="figure-0@4.[151,1173,194,1599]" captionTargetPageId="4" captionText="Figs. 1–4. Mecyclothorax species, dorsal habitus. 1) M. kavanaughi; 2) M. baehri; 3) M. medioconstrictus; 4) M. andersoni. Range of standardized body lengths (in mm) and setal formulae (see text) for number of supraorbital setae each side, number of pronotal setae each side, number of dorsal elytral setae each elytron, and number of setae at the apex of each elytron are indicated." figureDoi="http://doi.org/10.5281/zenodo.5377915" httpUri="https://zenodo.org/record/5377915/files/figure.png" pageId="10" pageNumber="688">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3446522FF8AFF904BA6FE5F7CB3FCE3" pageId="10" pageNumber="688" type="distribution">
|
||
<paragraph id="8BE136A9FF8AFF904BA6FE5F7CB3FCE3" blockId="10.[685,1173,194,852]" pageId="10" pageNumber="688">
|
||
<emphasis id="B92AEABBFF8AFF904BA6FE5F7D8FFE1C" bold="true" box="[709,969,406,427]" pageId="10" pageNumber="688">Distribution and Habitat.</emphasis>
|
||
The elevation of the Mt. Fugilil summit (
|
||
<figureCitation id="13652A2CFF8AFF904AEBFE787D90FE71" box="[904,982,433,454]" captionStart="Fig" captionStartId="17.[197,228,896,915]" captionTargetBox="[175,1191,199,878]" captionTargetId="figure-399@17.[173,1195,194,881]" captionTargetPageId="17" captionText="Fig. 32. Localities in Papua New Guinea from which Mecyclothorax spp. were collected and the particular species collected at each locality. 1 = Mt. Capella — M. baehri; 2 = Mt. Fugilil — M. brispex and M. medioconstrictus; 3 = Kubor Range — M. kubor; 4 = Mt. Wilhelm — M. sedlaceki; 5 = Finisterre Range — M. kavanaughi; 6 = Mt. Missim — M. gressitti; 7 = Mt. Kaindi, Edie Creek and Bulldog Road — M. andersoni, M. cuccodoroi, and M. loebli; 8 = Mt. Amungwiwa—M. amingwiwae." figureDoi="http://doi.org/10.5281/zenodo.5377933" httpUri="https://zenodo.org/record/5377933/files/figure.png" pageId="10" pageNumber="688">Fig. 32</figureCitation>
|
||
) was recorded as
|
||
<quantity id="4CA69B4CFF8AFF904BCEFE027D46FE57" box="[685,768,459,480]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="10" pageNumber="688" unit="m" value="3450.0">3,450 m</quantity>
|
||
by P. Beron for the
|
||
<typeStatus id="54E5880BFF8AFF904ABFFE027A69FE57" box="[988,1071,459,480]" pageId="10" pageNumber="688" type="holotype">holotype</typeStatus>
|
||
specimen collected
|
||
<date id="FFE01069FF8AFF904A6FFE2F7D33FE4C" box="[780,885,486,507]" pageId="10" pageNumber="688" value="1975-09-29">29.ix.1975</date>
|
||
.
|
||
<bibRefCitation id="EFCF4B58FF8AFF904AE1FE2F7A46FE4C" author="Shear, W. A." box="[898,1024,486,507]" pageId="10" pageNumber="688" pagination="35 - 40" refId="ref19400" refString="Shear, W. A. 1980. Zoological Results of the British Speleological Expedition to Papua New Guinea 1975: Malayothrix papuana, n. sp., the first chordeumid milliped from New Guinea (Chordeumida, Metopidiotrichidae). Myriapodologica 1: 35 - 40." type="journal article" year="1980">Shear (1980)</bibRefCitation>
|
||
cites the summit’ s elevation as
|
||
<quantity id="4CA69B4CFF8AFF904A0EFDC97D87FDA2" box="[877,961,512,533]" metricMagnitude="3" metricUnit="m" metricValue="3.05" pageId="10" pageNumber="688" unit="m" value="3050.0">3,050 m</quantity>
|
||
for the
|
||
<typeStatus id="54E5880BFF8AFF904D43FDC97A35FDA2" box="[1056,1139,512,533]" pageId="10" pageNumber="688" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C5E4D2AFF8AFF904BCEFDD27D87FD87" authority="Shear" authorityName="Shear" authorityYear="1980" box="[685,961,539,560]" class="Diplopoda" family="Metopidiotrichidae" genus="Malayothrix" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="688" phylum="Arthropoda" rank="species" species="papuana">
|
||
<emphasis id="B92AEABBFF8AFF904BCEFDD27DC7FD87" box="[685,897,539,560]" italics="true" pageId="10" pageNumber="688">Malayothrix papuana</emphasis>
|
||
Shear
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C5E4D2AFF8AFF904AB2FDD27A7CFD87" authorityName="de Blainville in Gervais" authorityYear="1844" box="[977,1082,539,560]" class="Diplopoda" kingdom="Animalia" pageId="10" pageNumber="688" phylum="Arthropoda" rank="class">Diplopoda</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C5E4D2AFF8AFF904D24FDD27D67FDFD" authorityName="Attems" authorityYear="1907" class="Diplopoda" family="Metopidiotrichidae" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="688" phylum="Arthropoda" rank="family">Metopidiotrichidae</taxonomicName>
|
||
), also collected by Beron on
|
||
<date id="FFE01069FF8AFF904D4BFDFC7AD6FDFD" box="[1064,1168,565,586]" pageId="10" pageNumber="688" value="1975-09-29">29.ix.1975</date>
|
||
. The geographic coordinates for the summit (
|
||
<bibRefCitation id="EFCF4B58FF8AFF904D0CFD997A2EFD37" author="National Geospatial Intelligence Agency" pageId="10" pageNumber="688" refId="ref19011" refString="National Geospatial Intelligence Agency. 2017. NGA GEOnet Names Server (GNS). geonames. nga. mil / gns / html / index. html. (accessed 22 May 2017)." type="book" year="2017">National Geospatial Intelligence Agency 2017</bibRefCitation>
|
||
) —
|
||
<geoCoordinate id="EE6A506EFF8AFF904BCEFD4C7D5FFD2D" box="[685,793,645,666]" degrees="5" direction="south" minutes="11.62" orientation="latitude" pageId="10" pageNumber="688" precision="9" value="-5.1936665">5° 11.62´S</geoCoordinate>
|
||
<geoCoordinate id="EE6A506EFF8AFF904A7CFD4C7DD9FD2D" box="[799,927,645,666]" degrees="141" direction="east" minutes="20.43" orientation="longitude" pageId="10" pageNumber="688" precision="9" value="141.3405">141° 20.43´E</geoCoordinate>
|
||
— are associated with an elevation of
|
||
<quantity id="4CA69B4CFF8AFF904A4CFD697DC4FD02" box="[815,898,672,693]" metricMagnitude="3" metricUnit="m" metricValue="2.506" pageId="10" pageNumber="688" unit="m" value="2506.0">2,506 m</quantity>
|
||
(
|
||
<bibRefCitation id="EFCF4B58FF8AFF904AF6FD697A1CFD02" author="Google Earth" box="[917,1114,672,693]" pageId="10" pageNumber="688" refId="ref18014" refString="Google Earth. 2017. Earth version 7.1. 8.3036. Google Inc., Mountain View, CA." type="book" year="2017">Google Earth 2017</bibRefCitation>
|
||
). The highest point in the vicinity of these coordinates is at
|
||
<quantity id="4CA69B4CFF8AFF904BCEFD1C7CB9FD5D" box="[685,767,725,746]" metricMagnitude="3" metricUnit="m" metricValue="2.782" pageId="10" pageNumber="688" unit="m" value="2782.0">2,782 m</quantity>
|
||
elevation, suggesting a tentative summit elevation for this massif. Regardless of elevation, the area surrounding Mt. Fugilil is forested, supporting this species’ occupation of montane forest habitat.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |