154 lines
18 KiB
XML
154 lines
18 KiB
XML
<document id="DB5B30A29B0FFCA7E2F66FBDFC00EB62" ID-DOI="10.5281/zenodo.170066" ID-GBIF-Dataset="e3737643-9288-451a-b814-4f22256e41a4" ID-ISSN="11755326" ID-Zenodo-Dep="170066" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459749874227" checkinUser="plazi" docAuthor="Peters, J. G., Flowers, R. W., Hubbard, M. D., Domínguez, E. & Savage, H. M." docDate="2005" docId="03F787BB0336FFC8F9733811F226C6EC" docLanguage="en" docName="zt01054p060.pdf" docOrigin="Zootaxa 1054" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Hermanella costalis Navas, comb. n." docType="treatment" docVersion="9" lastPageNumber="55" masterDocId="FFCEFFC30332FFCCF87B3B65F310C04F" masterDocTitle="New records and combinations for Neotropical Leptophlebiidae (Ephemeroptera)" masterLastPageNumber="60" masterPageNumber="51" pageNumber="55" updateTime="1698211586579" updateUser="plazi">
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<mods:titleInfo id="C5D32DF66549882C73290F2F31180A82">
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<mods:title id="A45A2C6BDCBC262C45BABBAABB06ED1A">New records and combinations for Neotropical Leptophlebiidae (Ephemeroptera)</mods:title>
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<mods:namePart id="459D24EBBFD64BA34021B9081BCFCBFD">Peters, J. G.</mods:namePart>
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<mods:namePart id="2F759D5DFF543497441C0F06F31CF562">Flowers, R. W.</mods:namePart>
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<mods:namePart id="7E226EDADF3AAF92D5B816016BB7B3EF">Hubbard, M. D.</mods:namePart>
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<mods:namePart id="0485A9C2372C7830B2ECACFF2335B9FB">Domínguez, E.</mods:namePart>
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<mods:namePart id="78B8EF4CC193A806EFD65A6101C67E93">Savage, H. M.</mods:namePart>
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<mods:typeOfResource id="A1E8D2AF90330F9714F30091619F572B">text</mods:typeOfResource>
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<mods:title id="F5A2FC1A47CE407FB7F0AD9BBEF74379">Zootaxa</mods:title>
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<mods:part id="D86D8703E0046684EA3A973EBD8C8DF4">
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<mods:date id="BC7ABA24A9A82FB32C016E3BA222B0EE">2005</mods:date>
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<mods:number id="519DE3CC2B574DC2345361DF93854428">1054</mods:number>
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<treatment id="03F787BB0336FFC8F9733811F226C6EC" ID-DOI="http://doi.org/10.5281/zenodo.6267046" ID-GBIF-Taxon="119337552" ID-Zenodo-Dep="6267046" LSID="urn:lsid:plazi:treatment:03F787BB0336FFC8F9733811F226C6EC" httpUri="http://treatment.plazi.org/id/03F787BB0336FFC8F9733811F226C6EC" lastPageNumber="55" pageId="4" pageNumber="55">
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<subSubSection id="C34465260336FFC8F9733811F2D4C3FE" pageId="4" pageNumber="55" type="nomenclature">
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<paragraph id="8BE136AD0336FFC8F9733811F1C3C3C1" blockId="4.[264,723,884,945]" box="[264,723,884,910]" pageId="4" pageNumber="55">
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<heading id="D0A981C10336FFC8F9733811F1C3C3C1" bold="true" box="[264,723,884,910]" fontSize="11" level="1" pageId="4" pageNumber="55" reason="1">
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<emphasis id="B92AEABF0336FFC8F9733811F1C3C3C1" bold="true" box="[264,723,884,910]" pageId="4" pageNumber="55">
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<taxonomicName id="4C5E4D2E0336FFC8F9733811F171C3C1" ID-CoL="6LS6T" authority="Navas" authorityName="Navas" box="[264,609,884,910]" class="Insecta" family="Leptophlebiidae" genus="Hermanella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="costalis" status="comb. nov.">
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<emphasis id="B92AEABF0336FFC8F9733811F2E9C3C1" bold="true" box="[264,505,884,910]" italics="true" pageId="4" pageNumber="55">Hermanella costalis</emphasis>
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(Navás)
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</taxonomicName>
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<taxonomicNameLabel id="A21957C40336FFC8FA133811F1C3C3C1" box="[616,723,884,910]" pageId="4" pageNumber="55">comb. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE136AD0336FFC8F97338F2F2D4C3FE" blockId="4.[264,723,884,945]" box="[264,452,919,945]" pageId="4" pageNumber="55">
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(
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<figureCitation id="13652A280336FFC8F96B38F2F2ABC3FE" box="[272,443,919,945]" captionStart="FIGURE 10 – 15" captionStartId="5.[264,368,1546,1570]" captionTargetBox="[264,1322,285,1532]" captionTargetId="figure@5.[249,1338,281,1537]" captionTargetPageId="5" captionText="FIGURE 10 – 15, holotype of Thraulus costalis: 10, fore wing; 11, hind wing enlarged (actual size about 1.5 mm); 12, lateral view of body (with left foreleg); 13, detail of foretarsal segments (outer surface); 14 – 15, male genitalia with detail. Photographs by C. Kutzscher. (Editing: Fig. 12, pin removed; Fig. 11, 14 – 15 details outlined.)" httpUri="https://zenodo.org/record/170068/files/figure.png" pageId="4" pageNumber="55">Figures 10–15</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34465260336FFC8F9733882F226C6EC" pageId="4" pageNumber="55" type="reference_group">
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<paragraph id="8BE136AD0336FFC8F9733882F03EC3B2" blockId="4.[264,814,998,1022]" box="[264,814,998,1022]" pageId="4" pageNumber="55">
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<treatmentCitationGroup id="AB4E11830336FFC8F9733882F03EC3B2" box="[264,814,998,1022]" pageId="4" pageNumber="55">
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<taxonomicName id="4C5E4D2E0336FFC8F9733882F161C3B2" authority="Navas, 1934:26" authorityName="Navas" authorityPageNumber="26" authorityYear="1934" box="[264,625,998,1022]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="costalis">
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<emphasis id="B92AEABF0336FFC8F9733882F2D0C3B1" box="[264,448,999,1022]" italics="true" pageId="4" pageNumber="55">Thraulus costalis</emphasis>
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<treatmentCitation id="0AFF10BC0336FFC8F9BC3883F161C3B2" author="Navas" box="[455,625,998,1021]" page="26" pageId="4" pageNumber="55" year="1934">
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<bibRefCitation id="EFCF4B5C0336FFC8F9BC3883F141C3B2" author="Navas" box="[455,593,998,1021]" pageId="4" pageNumber="59" refString="Navas, L. (1934) Insectos Suramericanos. Octava Serie. La Revista de la Academia de Ciencias de Madrid, 31, 9 - 28" type="journal article" year="1934">Navás, 1934</bibRefCitation>
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:26
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="0AFF10BC0336FFC8FA073883F03AC3B2" author="Traver" box="[636,810,998,1021]" page="73" pageId="4" pageNumber="55" year="1960">
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<bibRefCitation id="EFCF4B5C0336FFC8FA073883F019C3B2" author="Traver" box="[636,777,998,1021]" pageId="4" pageNumber="60" refString="Traver, J. R. (1960) Uruguayan mayflies-Family Leptophlebiidae: Part III. Revista de la Sociedad Uruguaya de Entomologia, 4, 73 - 85, 1 plate." type="book chapter" year="1960">Traver, 1960</bibRefCitation>
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:73
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BE136AD0336FFC8F9733F54F226C6EC" blockId="4.[264,1324,1073,1699]" pageId="4" pageNumber="55">
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<taxonomicName id="4C5E4D2E0336FFC8F9733F54F164C404" authority="Navas, 1934" authorityName="Navas" authorityYear="1934" box="[264,628,1073,1099]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="costalis">
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<emphasis id="B92AEABF0336FFC8F9733F54F2C4C405" box="[264,468,1073,1098]" italics="true" pageId="4" pageNumber="55">Thraulus costalis</emphasis>
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<bibRefCitation id="EFCF4B5C0336FFC8F9A53F54F164C404" author="Navas" box="[478,628,1073,1099]" pageId="4" pageNumber="59" refString="Navas, L. (1934) Insectos Suramericanos. Octava Serie. La Revista de la Academia de Ciencias de Madrid, 31, 9 - 28" type="journal article" year="1934">Navás, 1934</bibRefCitation>
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</taxonomicName>
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, was described from
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<collectingCountry id="F349763D0336FFC8FB0F3F54F0ACC404" box="[884,956,1073,1099]" name="Brazil" pageId="4" pageNumber="55">Brazil</collectingCountry>
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(Barão Homem de Mello, Rio de Janeiro) from a male imago in poor condition.
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<bibRefCitation id="EFCF4B5C0336FFC8FB0F3F3CF70AC43C" author="Traver" box="[884,1050,1113,1139]" pageId="4" pageNumber="60" refString="Traver, J. R. (1960) Uruguayan mayflies-Family Leptophlebiidae: Part III. Revista de la Sociedad Uruguaya de Entomologia, 4, 73 - 85, 1 plate." type="book chapter" year="1960">Traver (1960)</bibRefCitation>
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stated that it was not
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<taxonomicName id="4C5E4D2E0336FFC8F9733FE4F262C4D5" ID-CoL="648CZ" box="[264,370,1153,1178]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="B92AEABF0336FFC8F9733FE4F262C4D5" box="[264,370,1153,1178]" italics="true" pageId="4" pageNumber="55">Thraulus</emphasis>
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</taxonomicName>
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, but she did not reassign the species. Photographs of the
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<typeStatus id="54E5880F0336FFC8FC6E3FE4F76CC4D4" box="[1045,1148,1153,1179]" pageId="4" pageNumber="55" type="holotype">holotype</typeStatus>
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were prepared by C. Kutzscher of the Deutsches Entomologisches Institut im ZALF; P. Grant, who examined the
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<typeStatus id="54E5880F0336FFC8F9143FB4F2C6C4A4" box="[367,470,1233,1259]" pageId="4" pageNumber="55" type="holotype">holotype</typeStatus>
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in 1985, provided additional notes on its genitalia. The wings (
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<figureCitation id="13652A280336FFC8FCBC3FB4F234C55C" captionStart="FIGURE 10 – 15" captionStartId="5.[264,368,1546,1570]" captionTargetBox="[264,1322,285,1532]" captionTargetId="figure@5.[249,1338,281,1537]" captionTargetPageId="5" captionText="FIGURE 10 – 15, holotype of Thraulus costalis: 10, fore wing; 11, hind wing enlarged (actual size about 1.5 mm); 12, lateral view of body (with left foreleg); 13, detail of foretarsal segments (outer surface); 14 – 15, male genitalia with detail. Photographs by C. Kutzscher. (Editing: Fig. 12, pin removed; Fig. 11, 14 – 15 details outlined.)" httpUri="https://zenodo.org/record/170068/files/figure.png" pageId="4" pageNumber="55">Fig. 10– 11</figureCitation>
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) and genitalia (
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<figureCitation id="13652A280336FFC8F9A03F9CF14BC55C" box="[475,603,1273,1299]" captionStart="FIGURE 10 – 15" captionStartId="5.[264,368,1546,1570]" captionTargetBox="[264,1322,285,1532]" captionTargetId="figure@5.[249,1338,281,1537]" captionTargetPageId="5" captionText="FIGURE 10 – 15, holotype of Thraulus costalis: 10, fore wing; 11, hind wing enlarged (actual size about 1.5 mm); 12, lateral view of body (with left foreleg); 13, detail of foretarsal segments (outer surface); 14 – 15, male genitalia with detail. Photographs by C. Kutzscher. (Editing: Fig. 12, pin removed; Fig. 11, 14 – 15 details outlined.)" httpUri="https://zenodo.org/record/170068/files/figure.png" pageId="4" pageNumber="55">Fig. 14–15</figureCitation>
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) clearly place the species in the
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<taxonomicName id="4C5E4D2E0336FFC8FBAE3F9CF771C55D" ID-CoL="4VTV" box="[981,1121,1273,1298]" class="Insecta" family="Leptophlebiidae" genus="Hermanella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="B92AEABF0336FFC8FBAE3F9CF771C55D" box="[981,1121,1273,1298]" italics="true" pageId="4" pageNumber="55">Hermanella</emphasis>
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</taxonomicName>
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generic complex as defined by
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<bibRefCitation id="EFCF4B5C0336FFC8F9D43E44F008C574" author="Dominguez" box="[431,792,1313,1339]" pageId="4" pageNumber="59" refString="Dominguez, E. & Flowers, R. W. (1989) A revision of Hermanella and related genera (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) from Subtropical South America. Annals of the Entomological Society of America, 82, 555 - 573." type="journal article" year="1989">Domínguez & Flowers (1989)</bibRefCitation>
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, but the condition of the styliger plate (blunt median protuberance and no paired dorsal projections) does not fit the definition of any described genus of the complex. Also, the ventral projection on each penis lobe is broad in
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<taxonomicName id="4C5E4D2E0336FFC8F9733EFCF284C5FD" ID-CoL="4VTV" box="[264,404,1433,1458]" class="Insecta" family="Leptophlebiidae" genus="Hermanella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="B92AEABF0336FFC8F9733EFCF284C5FD" box="[264,404,1433,1458]" italics="true" pageId="4" pageNumber="55">Hermanella</emphasis>
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</taxonomicName>
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and slender in
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<taxonomicName id="4C5E4D2E0336FFC8FA323EFCF1ADC5FD" box="[585,701,1433,1458]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="costalis">
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<emphasis id="B92AEABF0336FFC8FA323EFCF1ADC5FD" box="[585,701,1433,1458]" italics="true" pageId="4" pageNumber="55">T. costalis</emphasis>
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</taxonomicName>
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. Although
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<taxonomicName id="4C5E4D2E0336FFC8FB3D3EFCF0ACC5FD" box="[838,956,1433,1458]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="costalis">
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<emphasis id="B92AEABF0336FFC8FB3D3EFCF0ACC5FD" box="[838,956,1433,1458]" italics="true" pageId="4" pageNumber="55">T. costalis</emphasis>
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</taxonomicName>
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is not a species of
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<taxonomicName id="4C5E4D2E0336FFC8FCE53EFCF63AC5FD" ID-CoL="4VTV" box="[1182,1322,1433,1458]" class="Insecta" family="Leptophlebiidae" genus="Hermanella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="B92AEABF0336FFC8FCE53EFCF63AC5FD" box="[1182,1322,1433,1458]" italics="true" pageId="4" pageNumber="55">Hermanella</emphasis>
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</taxonomicName>
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in the restricted sense, we prefer not to establish a new genus until the nymphs are known, and thus treat this species as
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<taxonomicName id="4C5E4D2E0336FFC8FA193E8CF0A5C64C" ID-CoL="6LS6T" authority="Navas" authorityName="Navas" box="[610,949,1513,1539]" class="Insecta" family="Leptophlebiidae" genus="Hermanella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="costalis" status="comb. nov.">
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<emphasis id="B92AEABF0336FFC8FA193E8CF05FC64D" box="[610,847,1513,1538]" italics="true" pageId="4" pageNumber="55">Hermanella costalis</emphasis>
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(Navás)
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</taxonomicName>
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<taxonomicNameLabel id="A21957C40336FFC8FBC63E8CF79DC64C" box="[957,1165,1513,1539]" pageId="4" pageNumber="55">new combination</taxonomicNameLabel>
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. A set of figures of the imago is included for purposes of future identification (
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<figureCitation id="13652A280336FFC8FC5D3D74F7B7C664" box="[1062,1191,1553,1579]" captionStart="FIGURE 10 – 15" captionStartId="5.[264,368,1546,1570]" captionTargetBox="[264,1322,285,1532]" captionTargetId="figure@5.[249,1338,281,1537]" captionTargetPageId="5" captionText="FIGURE 10 – 15, holotype of Thraulus costalis: 10, fore wing; 11, hind wing enlarged (actual size about 1.5 mm); 12, lateral view of body (with left foreleg); 13, detail of foretarsal segments (outer surface); 14 – 15, male genitalia with detail. Photographs by C. Kutzscher. (Editing: Fig. 12, pin removed; Fig. 11, 14 – 15 details outlined.)" httpUri="https://zenodo.org/record/170068/files/figure.png" pageId="4" pageNumber="55">Fig. 1015</figureCitation>
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). Note that the torn hind wing is hyaline and that the color in
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<figureCitation id="13652A280336FFC8FB023D5CF0DFC61C" box="[889,975,1593,1619]" captionStart="FIGURE 10 – 15" captionStartId="5.[264,368,1546,1570]" captionTargetBox="[264,1322,285,1532]" captionTargetId="figure@5.[249,1338,281,1537]" captionTargetPageId="5" captionText="FIGURE 10 – 15, holotype of Thraulus costalis: 10, fore wing; 11, hind wing enlarged (actual size about 1.5 mm); 12, lateral view of body (with left foreleg); 13, detail of foretarsal segments (outer surface); 14 – 15, male genitalia with detail. Photographs by C. Kutzscher. (Editing: Fig. 12, pin removed; Fig. 11, 14 – 15 details outlined.)" httpUri="https://zenodo.org/record/170068/files/figure.png" pageId="4" pageNumber="55">Fig. 11</figureCitation>
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is that of the body showing behind the wing. The only hind wing color is a brown basal strip in the costal area (
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<figureCitation id="13652A280336FFC8FC803D04F238C6EC" captionStart="FIGURE 10 – 15" captionStartId="5.[264,368,1546,1570]" captionTargetBox="[264,1322,285,1532]" captionTargetId="figure@5.[249,1338,281,1537]" captionTargetPageId="5" captionText="FIGURE 10 – 15, holotype of Thraulus costalis: 10, fore wing; 11, hind wing enlarged (actual size about 1.5 mm); 12, lateral view of body (with left foreleg); 13, detail of foretarsal segments (outer surface); 14 – 15, male genitalia with detail. Photographs by C. Kutzscher. (Editing: Fig. 12, pin removed; Fig. 11, 14 – 15 details outlined.)" httpUri="https://zenodo.org/record/170068/files/figure.png" pageId="4" pageNumber="55">Fig. 12</figureCitation>
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).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |