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<document id="964396FAD49E0CA3EAF22EF27B67E924" ID-DOI="10.5281/zenodo.170066" ID-GBIF-Dataset="e3737643-9288-451a-b814-4f22256e41a4" ID-ISSN="1175­5326" ID-Zenodo-Dep="170066" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459749874227" checkinUser="plazi" docAuthor="Peters, J. G., Flowers, R. W., Hubbard, M. D., Domínguez, E. &amp; Savage, H. M." docDate="2005" docId="03F787BB0333FFCDF9733A81F15AC5C9" docLanguage="en" docName="zt01054p060.pdf" docOrigin="Zootaxa 1054" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Tikuna bilineata Needham &amp; Murphy, comb. n." docType="treatment" docVersion="9" lastPageNumber="52" masterDocId="FFCEFFC30332FFCCF87B3B65F310C04F" masterDocTitle="New records and combinations for Neotropical Leptophlebiidae (Ephemeroptera)" masterLastPageNumber="60" masterPageNumber="51" pageNumber="52" updateTime="1698211586579" updateUser="plazi">
<mods:mods id="CE8FC6701B4F9A36A01155E986C3D739" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C5F9EDA009A1989803399329ABB7D41F">
<mods:title id="80D9A1D2E0ABCBD7426DDAD807FD5FE9">New records and combinations for Neotropical Leptophlebiidae (Ephemeroptera)</mods:title>
</mods:titleInfo>
<mods:name id="B56A17DD0A1E317B14BF5625DB4BE872" type="personal">
<mods:role id="941B3C7CDD601A87A5F63C0016258DB1">
<mods:roleTerm id="3E676E26434E64610FE28C0DBF6C1E19">Author</mods:roleTerm>
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<mods:namePart id="57D204ED5EBBCC90AC8FCF914F97E30F">Peters, J. G.</mods:namePart>
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<mods:namePart id="79A0F1DDC4DAA04F19DEFFAFA81329C6">Flowers, R. W.</mods:namePart>
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<mods:name id="F397AA008A8C92CA9296B577D1E6FE13" type="personal">
<mods:role id="7C5C1A2D9FE11AD6A3D19D5DBD42444A">
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<mods:namePart id="B4845E982E860A9BFC378012D94946FE">Hubbard, M. D.</mods:namePart>
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<mods:role id="A44AFA78EB24E4EA2D2295DBCBB1855A">
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<mods:namePart id="B1A1A7AFC31F4913154FF019440CBE24">Domínguez, E.</mods:namePart>
</mods:name>
<mods:name id="D941F4BF1DA825B2A2B4492BD8EDC5C8" type="personal">
<mods:role id="4AF9A5E6887AA924A9E070810947F70A">
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<mods:namePart id="F4D25C1A2A56E0905A29F09CF337C068">Savage, H. M.</mods:namePart>
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<mods:typeOfResource id="F4870AF5028D2DED6DFB7DEF80320B8E">text</mods:typeOfResource>
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<mods:titleInfo id="D1C10BB6953A0FFB1482D09CBACF47EC">
<mods:title id="D1885A0C7D92A3491B92FBD7F9B332FD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8B0E51F41453ACE9AAF911FAB88B912C">
<mods:date id="B56205E020E816A211AD1BF66321DA2F">2005</mods:date>
<mods:detail id="270D08D5CF3A2D8E1ED8EC8FA100574F" type="volume">
<mods:number id="D4A4BCC836606FF2F39C1E629D85FE57">1054</mods:number>
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<mods:start id="30930EA3DFF1B95A637EFA6DFF6E40A2">51</mods:start>
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<mods:classification id="234C1B6B99D629E2F817111DCFD42677">journal article</mods:classification>
<mods:identifier id="B6D9C4F3F1201FF6744AF1580A913990" type="DOI">10.5281/zenodo.170066</mods:identifier>
<mods:identifier id="F7222898EFC3595F7B322A455B0B1237" type="GBIF-Dataset">e3737643-9288-451a-b814-4f22256e41a4</mods:identifier>
<mods:identifier id="E073BA0346888620E906CD5193E0A814" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="4ECC98C721056A31B209342A71F52C22" type="Zenodo-Dep">170066</mods:identifier>
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<treatment id="03F787BB0333FFCDF9733A81F15AC5C9" ID-DOI="http://doi.org/10.5281/zenodo.6267038" ID-GBIF-Taxon="119337556" ID-Zenodo-Dep="6267038" LSID="urn:lsid:plazi:treatment:03F787BB0333FFCDF9733A81F15AC5C9" httpUri="http://treatment.plazi.org/id/03F787BB0333FFCDF9733A81F15AC5C9" lastPageNumber="52" pageId="1" pageNumber="52">
<subSubSection id="C34465260333FFCDF9733A81F073C1B2" box="[264,867,483,510]" pageId="1" pageNumber="52" type="nomenclature">
<paragraph id="8BE136AD0333FFCDF9733A81F073C1B2" blockId="1.[264,867,483,510]" box="[264,867,483,510]" pageId="1" pageNumber="52">
<heading id="D0A981C10333FFCDF9733A81F073C1B2" bold="true" box="[264,867,483,510]" fontSize="11" level="1" pageId="1" pageNumber="52" reason="1">
<emphasis id="B92AEABF0333FFCDF9733A81F073C1B2" bold="true" box="[264,867,483,510]" pageId="1" pageNumber="52">
<taxonomicName id="4C5E4D2E0333FFCDF9733A81F1E1C1B2" ID-CoL="56W77" authority="Needham &amp; Murphy" authorityName="Needham &amp; Murphy" box="[264,753,483,510]" class="Insecta" family="Leptophlebiidae" genus="Tikuna" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="bilineata" status="comb. nov.">
<emphasis id="B92AEABF0333FFCDF9733A81F2DFC1B1" bold="true" box="[264,463,484,510]" italics="true" pageId="1" pageNumber="52">Tikuna bilineata</emphasis>
(Needham &amp; Murphy)
</taxonomicName>
<taxonomicNameLabel id="A21957C40333FFCDFA833A86F073C1B2" box="[760,867,483,509]" pageId="1" pageNumber="52">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34465260333FFCDF973394FF7CFC3B9" pageId="1" pageNumber="52" type="reference_group">
<paragraph id="8BE136AD0333FFCDF973394FF0ECC20F" blockId="1.[264,1020,553,577]" box="[264,1020,553,577]" pageId="1" pageNumber="52">
<treatmentCitationGroup id="AB4E11830333FFCDF973394FF0ECC20F" box="[264,1020,553,577]" pageId="1" pageNumber="52">
<taxonomicName id="4C5E4D2E0333FFCDF973394FF028C20F" authority="Needham &amp; Murphy 1924:48" authorityName="Needham &amp; Murphy" authorityPageNumber="48" authorityYear="1924" box="[264,824,553,577]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="bilineata">
<emphasis id="B92AEABF0333FFCDF973394FF2E1C20E" box="[264,497,554,577]" italics="true" pageId="1" pageNumber="52">Choroterpes bilineata</emphasis>
<treatmentCitation id="0AFF10BC0333FFCDF982394CF028C20F" author="Needham" box="[505,824,553,576]" page="48" pageId="1" pageNumber="52" year="1924">
<bibRefCitation id="EFCF4B5C0333FFCDF982394CF008C20F" author="Needham" box="[505,792,553,576]" pageId="1" pageNumber="59" refString="Needham, J. G. &amp; Murphy, H. E. (1924) Neotropical Mayflies. Bulletin of the Lloyd Library, 24, Entomological Series, 4, 3 - 79." type="journal article" year="1924">Needham &amp; Murphy 1924</bibRefCitation>
:48
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AFF10BC0333FFCDFB38394CF0E8C20F" author="Traver" box="[835,1016,553,576]" page="156" pageId="1" pageNumber="52" year="1947">
<bibRefCitation id="EFCF4B5C0333FFCDFB38394CF0DAC20F" author="Traver" box="[835,970,553,576]" pageId="1" pageNumber="60" refString="Traver, J. R. (1947) Notes on Neotropical mayflies. Part II. Family Baetidae, Subfamily Leptophlebiinae. Revista de Entomologia, Rio de Janeiro, 18, 149 - 160." type="journal article" year="1947">Traver 1947</bibRefCitation>
:156
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE136AD0333FFCDF9733911F7CFC3B9" blockId="1.[264,1323,628,1414]" pageId="1" pageNumber="52">
The genus
<taxonomicName id="4C5E4D2E0333FFCDF9F53911F2CCC2C2" ID-CoL="7WWK" box="[398,476,628,653]" class="Insecta" family="Leptophlebiidae" genus="Tikuna" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEABF0333FFCDF9F53911F2CCC2C2" box="[398,476,628,653]" italics="true" pageId="1" pageNumber="52">Tikuna</emphasis>
</taxonomicName>
was established by
<bibRefCitation id="EFCF4B5C0333FFCDFAA93911F0D3C2C1" box="[722,963,628,654]" pageId="1" pageNumber="59" refString="Savage, H. M., Flowers, R. W. &amp; Porras V., W. (2005) Rediscovery of Choroterpes atramentum in Costa Rica, type species of Tikuna new genus (Ephemeroptera: Leptophlebiidae: Atalophlebiinae), and its role in the &quot; Great American Interchange. &quot; Zootaxa, 932, 1 - 14." type="journal article">Savage et al. (2005)</bibRefCitation>
and
<taxonomicName id="4C5E4D2E0333FFCDFC793911F2B1C2F9" authority="Traver, 1947" authorityName="Traver" authorityYear="1947" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="atramentum">
<emphasis id="B92AEABF0333FFCDFC793911F63BC2C2" box="[1026,1323,628,653]" italics="true" pageId="1" pageNumber="52">Choroterpes atramentum</emphasis>
<bibRefCitation id="EFCF4B5C0333FFCDF97339F9F2B1C2F9" author="Traver" box="[264,417,668,694]" pageId="1" pageNumber="60" refString="Traver, J. R. (1947) Notes on Neotropical mayflies. Part II. Family Baetidae, Subfamily Leptophlebiinae. Revista de Entomologia, Rio de Janeiro, 18, 149 - 160." type="journal article" year="1947">Traver, 1947</bibRefCitation>
</taxonomicName>
, was designated as the
<typeStatus id="54E5880F0333FFCDFAC339F9F1FAC2F9" box="[696,746,668,694]" pageId="1" pageNumber="52">type</typeStatus>
species. Following this publication, two species from South and Central
<collectingCountry id="F349763D0333FFCDFA4239A1F18EC291" box="[569,670,708,734]" name="United States of America" pageId="1" pageNumber="52">America</collectingCountry>
remain incorrectly placed in
<taxonomicName id="4C5E4D2E0333FFCDFC6F39A1F7B5C292" ID-CoL="3P6W" box="[1044,1189,708,733]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEABF0333FFCDFC6F39A1F7B5C292" box="[1044,1189,708,733]" italics="true" pageId="1" pageNumber="52">Choroterpes</emphasis>
</taxonomicName>
. The first,
<taxonomicName id="4C5E4D2E0333FFCDF9733989F048C349" authority="Needham &amp; Murphy, 1924" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[264,856,748,774]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="bilineata">
<emphasis id="B92AEABF0333FFCDF9733989F11AC34A" box="[264,522,748,773]" italics="true" pageId="1" pageNumber="52">Choroterpes bilineata</emphasis>
<bibRefCitation id="EFCF4B5C0333FFCDFA6F3989F048C349" author="Needham" box="[532,856,748,774]" pageId="1" pageNumber="59" refString="Needham, J. G. &amp; Murphy, H. E. (1924) Neotropical Mayflies. Bulletin of the Lloyd Library, 24, Entomological Series, 4, 3 - 79." type="journal article" year="1924">Needham &amp; Murphy, 1924</bibRefCitation>
</taxonomicName>
, originally described from female imagos and male subimagos from La Chorrera, Putumayo Dist.,
<collectingCountry id="F349763D0333FFCDFBA43871F705C361" box="[991,1045,788,814]" name="Peru" pageId="1" pageNumber="52">Peru</collectingCountry>
(now a part of
<collectingCountry id="F349763D0333FFCDFCAA3871F220C319" name="Colombia" pageId="1" pageNumber="52">Colombia</collectingCountry>
), also belongs to the genus
<taxonomicName id="4C5E4D2E0333FFCDFAFF3859F1C4C31A" ID-CoL="7WWK" box="[644,724,828,853]" class="Insecta" family="Leptophlebiidae" genus="Tikuna" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEABF0333FFCDFAFF3859F1C4C31A" box="[644,724,828,853]" italics="true" pageId="1" pageNumber="52">Tikuna</emphasis>
</taxonomicName>
, requiring the new combination
<emphasis id="B92AEABF0333FFCDFC1A3859F63BC31A" box="[1121,1323,828,853]" italics="true" pageId="1" pageNumber="52">
<taxonomicName id="4C5E4D2E0333FFCDFC1A3859F637C31A" ID-CoL="56W77" box="[1121,1319,828,853]" class="Insecta" family="Leptophlebiidae" genus="Tikuna" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="bilineata">Tikuna bilineata</taxonomicName>
.
</emphasis>
The
<typeStatus id="54E5880F0333FFCDF9473801F2B2C331" box="[316,418,868,894]" pageId="1" pageNumber="52" type="holotype">holotype</typeStatus>
and
<typeStatus id="54E5880F0333FFCDF9A13801F15AC331" box="[474,586,868,894]" pageId="1" pageNumber="52" type="paratype">paratypes</typeStatus>
(Cornell University) were studied by one of us (H.M. Savage); additional specimens deposited at the
<collectingCountry id="F349763D0333FFCDFAB738E9F019C3E9" box="[716,777,908,934]" name="United States of America" pageId="1" pageNumber="52">U. S.</collectingCountry>
National Museum of Natural History, Florida A&amp;M University, and the Zoologische Sammlung des Bayerischen Staates were recorded from
<collectingCountry id="F349763D0333FFCDF93338B9F283C3B9" box="[328,403,988,1014]" name="Brazil" pageId="1" pageNumber="52">Brazil</collectingCountry>
,
<collectingCountry id="F349763D0333FFCDF9E438B9F113C3B9" box="[415,515,988,1014]" name="Ecuador" pageId="1" pageNumber="52">Ecuador</collectingCountry>
,
<collectingCountry id="F349763D0333FFCDFA7538B9F165C3B9" box="[526,629,988,1014]" name="Suriname" pageId="1" pageNumber="52">Surinam</collectingCountry>
, and
<collectingCountry id="F349763D0333FFCDFAC938B9F03FC3B9" box="[690,815,988,1014]" name="Venezuela" pageId="1" pageNumber="52">Venezuela</collectingCountry>
. New records for
<taxonomicName id="4C5E4D2E0333FFCDFB8538B9F793C3BA" ID-CoL="56W77" box="[1022,1155,988,1013]" class="Insecta" family="Leptophlebiidae" genus="Tikuna" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="bilineata">
<emphasis id="B92AEABF0333FFCDFB8538B9F793C3BA" box="[1022,1155,988,1013]" italics="true" pageId="1" pageNumber="52">T. bilineata</emphasis>
</taxonomicName>
follow:
</paragraph>
</subSubSection>
<subSubSection id="C34465260333FFCDF9433F61F15AC5C9" pageId="1" pageNumber="52" type="description">
<paragraph id="8BE136AD0333FFCDF9433F61F1F6C409" blockId="1.[264,1323,628,1414]" pageId="1" pageNumber="52">
<collectingCountry id="F349763D0333FFCDF9433F61F2B7C451" box="[312,423,1028,1054]" name="Brazil" pageId="1" pageNumber="52">BRAZIL</collectingCountry>
:
<specimenCount id="9D58FD240333FFCDF9CC3F61F133C451" box="[439,547,1028,1054]" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
imago, Pará State, Rio Cururu, about
<quantity id="4CA69B480333FFCDFC763F61F77CC451" box="[1037,1132,1028,1054]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="52" unit="km" value="100.0">100 km</quantity>
above Mission Cururu, at light,
<date id="FFE0106D0333FFCDF9B13F49F15DC409" box="[458,589,1068,1094]" pageId="1" pageNumber="52" value="1962-01-25">25–I–1962</date>
, E.J. Fittkau.
</paragraph>
<paragraph id="8BE136AD0333FFCDF9433F31F0D4C4F1" blockId="1.[264,1323,628,1414]" pageId="1" pageNumber="52">
<collectingCountry id="F349763D0333FFCDF9433F31F2DBC421" box="[312,459,1108,1134]" name="Ecuador" pageId="1" pageNumber="52">ECUADOR</collectingCountry>
: Pastaza Prov., coll. J. Cohen:
<specimenCount id="9D58FD240333FFCDFB303F31F0D4C421" box="[843,964,1108,1134]" pageId="1" pageNumber="52" type="female">37 female</specimenCount>
imagos, Tzapino,
<quantity id="4CA69B480333FFCDFCD13F31F7E9C421" box="[1194,1273,1108,1134]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="52" unit="km" value="32.0">32 km</quantity>
NE Tigueno,
<geoCoordinate id="EE6A506A0333FFCDF9033F19F2DEC4D9" box="[376,462,1148,1174]" direction="south" orientation="latitude" pageId="1" pageNumber="52" precision="925" value="-1.1833334">1°11' S</geoCoordinate>
,
<geoCoordinate id="EE6A506A0333FFCDF9A73F19F15CC4D9" box="[476,588,1148,1174]" direction="west" orientation="longitude" pageId="1" pageNumber="52" precision="925" value="-77.23333">77°14' W</geoCoordinate>
,
<quantity id="4CA69B480333FFCDFA233F19F1B6C4D9" box="[600,678,1148,1174]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="52" unit="m" value="400.0">400 m</quantity>
,
<date id="FFE0106D0333FFCDFAC83F19F051C4D9" box="[691,833,1148,1174]" pageId="1" pageNumber="52" value="1976-05-25">25–V–1976</date>
;
<specimenCount id="9D58FD240333FFCDFB353F19F0A4C4D9" box="[846,948,1148,1174]" pageId="1" pageNumber="52" type="female">2 female</specimenCount>
imagos, Limoncocha (
<quantity id="4CA69B480333FFCDFCBF3F19F63AC4D9" box="[1220,1322,1148,1174]" metricMagnitude="5" metricUnit="m" metricValue="1.1265408" pageId="1" pageNumber="52" unit="mi" value="70.0">70 miles</quantity>
SE),
<date id="FFE0106D0333FFCDF9383FC1F2C0C4F1" box="[323,464,1188,1214]" pageId="1" pageNumber="52" value="1976-05-30">30–V–1976</date>
;
<specimenCount id="9D58FD240333FFCDF9A73FC1F152C4F1" box="[476,578,1188,1214]" pageId="1" pageNumber="52" type="female">3 female</specimenCount>
imagos, Cononaco,
<date id="FFE0106D0333FFCDFB493FC1F0D0C4F1" box="[818,960,1188,1214]" pageId="1" pageNumber="52" value="1976-05-30">30–V–1976</date>
.
</paragraph>
<paragraph id="8BE136AD0333FFCDF9433FA9F60FC4A9" blockId="1.[264,1323,628,1414]" box="[312,1311,1228,1254]" pageId="1" pageNumber="52">
<collectingCountry id="F349763D0333FFCDF9433FA9F2D4C4A9" box="[312,452,1228,1254]" name="Suriname" pageId="1" pageNumber="52">SURINAM</collectingCountry>
:
<specimenCount id="9D58FD240333FFCDF9B53FA9F124C4A9" box="[462,564,1228,1254]" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
imago, Wijne Dist., Moengo, Boven, 1/
<date id="FFE0106D0333FFCDFC7D3FA9F787C4A9" box="[1030,1175,1228,1254]" pageId="1" pageNumber="52" value="1927-05-28">28–V–1927</date>
, P.P. Babiy.
</paragraph>
<paragraph id="8BE136AD0333FFCDF9433F91F15AC5C9" blockId="1.[264,1323,628,1414]" pageId="1" pageNumber="52">
<collectingCountry id="F349763D0333FFCDF9433F91F2FBC541" box="[312,491,1268,1294]" name="Venezuela" pageId="1" pageNumber="52">VENEZUELA</collectingCountry>
: Zulia State:
<specimenCount id="9D58FD240333FFCDFAF63F91F1F0C541" box="[653,736,1268,1294]" pageId="1" pageNumber="52" type="male">1 male</specimenCount>
subimago, Dist. Mara, Río Socuy, Campamento Corpozulia,
<quantity id="4CA69B480333FFCDF9E53E79F2FDC579" box="[414,493,1308,1334]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="52" unit="km" value="50.0">50 km</quantity>
W of Carrasquero, 6/
<date id="FFE0106D0333FFCDFB783E79F092C579" box="[771,898,1308,1334]" pageId="1" pageNumber="52" value="1979-10-07">7–X–1979</date>
, H.M. Savage &amp; R.A. Romero;
<specimenCount id="9D58FD240333FFCDFD603E79F248C511" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
imago, Perija El Tucuco, Mission El Tucuco, Río El Tucuco, 1/
<quantity id="4CA69B480333FFCDFC373E21F79BC511" box="[1100,1163,1348,1374]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="52" unit="km" value="2.0">2 km</quantity>
from church, 1/
<date id="FFE0106D0333FFCDF9603E09F28DC5C9" box="[283,413,1388,1414]" pageId="1" pageNumber="52" value="1979-10-05">5–X–1979</date>
, H.M. Savage.
</paragraph>
</subSubSection>
</treatment>
</document>
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