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<document id="5F318B38FFCB2C80C715673D8DF06DC8" ID-DOI="10.5281/zenodo.5161403" ID-ISSN="1942-1354" ID-Zenodo-Dep="5161403" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628118715709" checkinUser="felipe" docAuthor="Gilbert, Arthur J." docDate="2011" docId="03F787A4FF82FFD4F9D40B0FFCD7F94E" docLanguage="en" docName="InsectaMundi.0200.1-57.pdf" docOrigin="Insecta Mundi 2011 (200)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Phrynocepha pseudocapitata Gilbert 2011, sp. nov." docType="treatment" docVersion="3" lastPageNumber="49" masterDocId="FFCEFFDCFFB3FFE6F9140E29FFDAFFD4" masterDocTitle="A review and clarification of the alticine genera Hemiphrynus Horn 1889 and Phrynocepha Baly 1861 (Coleoptera: Chrysomelidae: Galerucinae: Alticini)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="48" updateTime="1699035131958" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="3D55FDF93105EEB74CE2FCC37E6FDBC6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="406F6CCE2F0CE22B99165D6EB75D6B06">
<mods:title id="FAD0BB3D561158FD319DCC478EEF34D0">A review and clarification of the alticine genera Hemiphrynus Horn 1889 and Phrynocepha Baly 1861 (Coleoptera: Chrysomelidae: Galerucinae: Alticini)</mods:title>
</mods:titleInfo>
<mods:name id="55F47AF87035B7B0C5BF089B48DEA03D" type="personal">
<mods:role id="A0A61A38D05330F76D5B2AB430DC5954">
<mods:roleTerm id="2985F8BE070F9D792228610FA6AF5816">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DE027BBDD252F698219AA87963BA9DAE">Gilbert, Arthur J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="9D173F2D10830B845168C3AEBEEE2239">text</mods:typeOfResource>
<mods:relatedItem id="CC3B114D9214E04153D1E95A995389D3" type="host">
<mods:titleInfo id="44747B519D4EC660B408D9A8EAAF9BE4">
<mods:title id="DC713884D95BA78176814EB49AF823D2">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="AC1223571F0829F863EF058B729BD692">
<mods:date id="A4DBCE2B5EB7501D993BEF9C70C614D3">2011</mods:date>
<mods:detail id="421B33CF647DF31E41C176B3C42CE59A" type="pubDate">
<mods:number id="E2D2A9F1C4690262E4981FE9F81051A0">2011-11-02</mods:number>
</mods:detail>
<mods:detail id="D6AB8616A260C0D658B54426A4765BD4" type="volume">
<mods:number id="4680B3C05A1E51AE4729071D15CF2FAD">2011</mods:number>
</mods:detail>
<mods:detail id="E846D0E14546C66C1DD82FD4ED86C24D" type="issue">
<mods:number id="DD5643C0FE7B884A78550835B22F0FA1">200</mods:number>
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<mods:start id="3025BDF7BD7BD182B62FFB5EEA2D0671">1</mods:start>
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<mods:classification id="69F49A8C08AA5AFFE3427A8724462DED">journal article</mods:classification>
<mods:identifier id="8378EB0D23EE0C6D6E2291FAC1ABDAE1" type="DOI">10.5281/zenodo.5161403</mods:identifier>
<mods:identifier id="1C8C4BB7A628C668E94CC1E1F53F85DE" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="9D4C64EBBB9B3F5A0F963F1C4E763EC9" type="Zenodo-Dep">5161403</mods:identifier>
</mods:mods>
<treatment id="03F787A4FF82FFD4F9D40B0FFCD7F94E" LSID="urn:lsid:plazi:treatment:03F787A4FF82FFD4F9D40B0FFCD7F94E" httpUri="http://treatment.plazi.org/id/03F787A4FF82FFD4F9D40B0FFCD7F94E" lastPageId="50" lastPageNumber="49" pageId="49" pageNumber="48">
<subSubSection id="C3446539FF82FFD7F9D40B0FFCF6FAEB" box="[192,812,1318,1343]" pageId="49" pageNumber="48" type="nomenclature">
<paragraph id="8BE136B2FF82FFD7F9D40B0FFCF6FAEB" blockId="49.[192,812,1318,1375]" box="[192,812,1318,1343]" pageId="49" pageNumber="48">
<heading id="D0A981DEFF82FFD7F9D40B0FFCF6FAEB" bold="true" box="[192,812,1318,1343]" fontSize="10" level="2" pageId="49" pageNumber="48" reason="3">
<emphasis id="B92AEAA0FF82FFD7F9D40B0FFCF6FAEB" bold="true" box="[192,812,1318,1343]" pageId="49" pageNumber="48">
<taxonomicName id="4C5E4D31FF82FFD7F9D40B0FFD61FAEB" authority="Gilbert" authorityName="Gilbert" authorityYear="2011" box="[192,699,1318,1343]" class="Insecta" family="Chrysomelidae" genus="Phrynocepha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="48" phylum="Arthropoda" rank="species" species="pseudocapitata" status="sp. nov.">
<emphasis id="B92AEAA0FF82FFD7F9D40B0FFD8AFAEA" bold="true" box="[192,592,1318,1342]" italics="true" pageId="49" pageNumber="48">Phrynocepha pseudocapitata</emphasis>
Gilbert
</taxonomicName>
<taxonomicNameLabel id="A21957DBFF82FFD7FBD70B0EFCF6FAEB" box="[707,812,1319,1343]" pageId="49" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3446539FF82FFD7F9D40B6EFE1EFA8B" box="[192,452,1351,1375]" pageId="49" pageNumber="48" type="description">
<paragraph id="8BE136B2FF82FFD7F9D40B6EFE1EFA8B" blockId="49.[192,812,1318,1375]" box="[192,452,1351,1375]" pageId="49" pageNumber="48">
(
<figureCitation id="13652A37FF82FFD7F9DD0B6EFEE2FA8B" box="[201,312,1351,1375]" captionStart="Figures 5-10" captionStartId="9.[278,374,1721,1743]" captionTargetBox="[279,1354,195,1708]" captionTargetId="figure-17@9.[278,1354,195,1709]" captionTargetPageId="9" captionText="Figures 5-10. Male heads (fgs = frontogenal suture, jl = jowl, lc = lateral carina). 5) Hemiphrynus intermedius, lateral view. 6) H. intermedius, frontal view. 7) Phrynocepha deyrollei, lateral view. 8) P. deyrollei, frontal view. 9) P. pseudocapitata, lateral view. 10) P. pseudocapitata, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5161409" httpUri="https://zenodo.org/record/5161409/files/figure.png" pageId="49" pageNumber="48">Fig. 9-10</figureCitation>
,
<figureCitation id="13652A37FF82FFD7F8500B6EFEBEFA8B" box="[324,356,1351,1375]" captionStart="Figures 51-54" captionStartId="34.[192,288,1453,1475]" captionTargetBox="[192,1440,192,1442]" captionTargetId="figure-235@34.[192,1440,192,1442]" captionTargetPageId="34" captionText="Figures 51-54. Pronota of Phrynocepha. 51) P. marciae. 52) P. pseudocapitata. 53) P. pueblae. 54) P. punctulata." figureDoi="http://doi.org/10.5281/zenodo.5161439" httpUri="https://zenodo.org/record/5161439/files/figure.png" pageId="49" pageNumber="48">52</figureCitation>
,
<figureCitation id="13652A37FF82FFD7F8650B6EFE4BFA8B" box="[369,401,1351,1375]" captionStart="Figures 60-63" captionStartId="37.[192,288,1450,1472]" captionTargetBox="[192,1440,192,1441]" captionTargetId="figure-192@37.[192,1440,192,1441]" captionTargetPageId="37" captionText="Figures 60-63. Aedeagi, lateral view. 60) Phrynocepha pseudocapitata. 61) P. pueblae. 62) P. pulchella. 63) P. punctulata." figureDoi="http://doi.org/10.5281/zenodo.5161449" httpUri="https://zenodo.org/record/5161449/files/figure.png" pageId="49" pageNumber="48">60</figureCitation>
,
<figureCitation id="13652A37FF82FFD7F8890B6EFE61FA8B" box="[413,443,1351,1375]" captionStart="Figures 78-79" captionStartId="43.[192,288,1855,1877]" captionTargetBox="[192,1440,195,1843]" captionTargetId="figure-24@43.[192,1440,195,1843]" captionTargetPageId="43" captionText="Figures 78-79. Distribution of Phrynocepha in Mexico." figureDoi="http://doi.org/10.5281/zenodo.5161459" httpUri="https://zenodo.org/record/5161459/files/figure.png" pageId="49" pageNumber="48">78</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3446539FF82FFD7F9D40BAEFE8AF9EB" pageId="49" pageNumber="48" type="materials_examined">
<paragraph id="8BE136B2FF82FFD7F9D40BAEFDA7FA0B" blockId="49.[192,1441,1415,1599]" pageId="49" pageNumber="48">
<emphasis id="B92AEAA0FF82FFD7F9D40BAEFE7DFA4B" bold="true" box="[192,423,1415,1439]" pageId="49" pageNumber="48">Type Specimens.</emphasis>
<materialsCitation id="3B363CEFFF82FFD7F8B90BAEFC61FA6B" collectingDate="1987-05-24" collectorName="D. A. Rider &amp; E. G. and T. J. Riley" country="Mexico" location="Hwy" pageId="49" pageNumber="48" specimenCount="2" stateProvince="Chiapas" typeStatus="holotype">
<emphasis id="B92AEAA0FF82FFD7F8B90BAEFDF1FA4B" bold="true" box="[429,555,1415,1439]" pageId="49" pageNumber="48">
<typeStatus id="54E58810FF82FFD7F8B90BAEFDF1FA4B" box="[429,555,1415,1439]" pageId="49" pageNumber="48" type="holotype">Holotype</typeStatus>
</emphasis>
(male) and
<emphasis id="B92AEAA0FF82FFD7FBAF0BAEFCF1FA4B" bold="true" box="[699,811,1415,1439]" pageId="49" pageNumber="48">
<typeStatus id="54E58810FF82FFD7FBAF0BAEFCF1FA4B" box="[699,811,1415,1439]" pageId="49" pageNumber="48" type="allotype">allotype</typeStatus>
</emphasis>
(female):
<emphasis id="B92AEAA0FF82FFD7FAB70BAEFB6DFA4B" bold="true" box="[931,1207,1415,1439]" pageId="49" pageNumber="48">
<collectingCountry id="F3497622FF82FFD7FAB70BAEFBC4FA4B" box="[931,1054,1415,1439]" name="Mexico" pageId="49" pageNumber="48">MEXICO</collectingCountry>
.
<collectingRegion id="499AF850FF82FFD7FD3E0BAEFB68FA4B" box="[1066,1202,1415,1439]" country="Mexico" name="Chiapas" pageId="49" pageNumber="48">CHIAPAS</collectingRegion>
:
</emphasis>
<location id="8E816069FF82FFD7FDA90BAEFB2CFA4B" LSID="urn:lsid:plazi:treatment:03F787A4FF82FFD4F9D40B0FFCD7F94E:8E816069FF82FFD7FDA90BAEFB2CFA4B" box="[1213,1270,1415,1439]" country="Mexico" name="Hwy" pageId="49" pageNumber="48" stateProvince="Chiapas">Hwy</location>
195
<locationDeviation id="9A532F90FF82FFD7FC240BAEFEC9FA6A" location="Ixtapa" pageId="49" pageNumber="48">
<quantity id="4CA69B57FF82FFD7FC240BAEFA59FA4B" box="[1328,1411,1415,1439]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="49" pageNumber="48" unit="km" value="4.5">4.5 km</quantity>
N Ixtapa
</locationDeviation>
, 3,000,
<date id="FFE01072FF82FFD7F8630B8EFDCDFA6B" box="[375,535,1447,1471]" pageId="49" pageNumber="48" value="1987-05-24">
<collectingDate id="EFA4E99AFF82FFD7F8630B8EFDCDFA6B" box="[375,535,1447,1471]" pageId="49" pageNumber="48" value="1987-05-24">24 May 1987</collectingDate>
</date>
,
<collectorName id="26AB5364FF82FFD7FB300B8EFD6BFA6B" box="[548,689,1447,1471]" pageId="49" pageNumber="48">D. A. Rider</collectorName>
,
<collectorName id="26AB5364FF82FFD7FBA90B8EFC61FA6B" box="[701,955,1447,1471]" pageId="49" pageNumber="48">E. G. and T. J. Riley</collectorName>
</materialsCitation>
.
<typeStatus id="54E58810FF82FFD7FAD10B8EFBEEFA6B" box="[965,1076,1447,1471]" pageId="49" pageNumber="48" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54E58810FF82FFD7FD650B8EFB09FA6B" box="[1137,1235,1447,1471]" pageId="49" pageNumber="48" type="allotype">allotype</typeStatus>
deposited in the collection of Texas A&amp;M University.
</paragraph>
<paragraph id="8BE136B2FF82FFD7F9E40BCEFE8AF9EB" blockId="49.[192,1441,1415,1599]" pageId="49" pageNumber="48">
<materialsCitation id="3B363CEFFF82FFD7F9E40BCEFCC5F9CB" collectingDate="1987-05-24" collectionCode="AJGC, AMNH, BMNH, CASC, EGRC, MCZC, USNM" collectorName="D. A. Rider &amp; E. G. and T. J. Riley" country="Mexico" location="Hwy" pageId="49" pageNumber="48" specimenCount="93" specimenCount-female="49" specimenCount-male="44" stateProvince="Chiapas" typeStatus="paratype">
<emphasis id="B92AEAA0FF82FFD7F9E40BCEFEA6FA2B" bold="true" box="[240,380,1511,1535]" pageId="49" pageNumber="48">
<typeStatus id="54E58810FF82FFD7F9E40BCEFEA6FA2B" box="[240,380,1511,1535]" pageId="49" pageNumber="48" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D58FD3BFF82FFD7F89D0BCEFE2FFA2B" box="[393,501,1511,1535]" pageId="49" pageNumber="48" type="male">44 males</specimenCount>
and
<specimenCount id="9D58FD3BFF82FFD7FB3A0BCEFD6AFA2B" box="[558,688,1511,1535]" pageId="49" pageNumber="48" type="female">49 females</specimenCount>
) same data as holotype (6) [
<collectionCode id="ED4FAE77FF82FFD7FD190BCEFB87FA2B" box="[1037,1117,1511,1535]" pageId="49" pageNumber="48">AJGC</collectionCode>
], (2) [
<collectionCode id="ED4FAE77FF82FFD7FD8E0BCEFB23FA2B" box="[1178,1273,1511,1535]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="49" pageNumber="48">AMNH</collectionCode>
], (4) [
<collectionCode id="ED4FAE77FF82FFD7FC220BCEFA4FFA2A" box="[1334,1429,1511,1534]" pageId="49" pageNumber="48">BMNH</collectionCode>
], (4) [
<collectionCode id="ED4FAE77FF82FFD7F9FA082EFE9AF9CB" box="[238,320,1543,1567]" pageId="49" pageNumber="48">CASC</collectionCode>
], (72) [
<collectionCode id="ED4FAE77FF82FFD7F89B082EFE3EF9CB" box="[399,484,1543,1567]" pageId="49" pageNumber="48">EGRC</collectionCode>
], (2) [
<collectionCode id="ED4FAE77FF82FFD7FB30082EFDA6F9CB" box="[548,636,1543,1567]" pageId="49" pageNumber="48">MCZC</collectionCode>
], (2) [
<collectionCode id="ED4FAE77FF82FFD7FBA8082EFCC2F9CB" box="[700,792,1543,1567]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="49" pageNumber="48" type="Museum">USNM</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B363CEFFF82FFD7FA3E082EFE96F9EB" collectingDate="1990-06-26" collectionCode="RHTC" collectorName="El Chorreadero &amp; R. Turnbow" country="Mexico" pageId="49" pageNumber="48" specimenCount="1" stateProvince="Chiapas" typeStatus="paratype">
<collectorName id="26AB5364FF82FFD7FA3E082EFC30F9CB" box="[810,1002,1543,1567]" pageId="49" pageNumber="48">El Chorreadero</collectorName>
, mv + bl,
<date id="FFE01072FF82FFD7FD4A082EFADEF9CB" box="[1118,1284,1543,1567]" pageId="49" pageNumber="48" value="1990-06-26">
<collectingDate id="EFA4E99AFF82FFD7FD4A082EFADEF9CB" box="[1118,1284,1543,1567]" pageId="49" pageNumber="48" value="1990-06-26">26 June 1990</collectingDate>
</date>
,
<collectorName id="26AB5364FF82FFD7FC1B082EFA7AF9CB" box="[1295,1440,1543,1567]" pageId="49" pageNumber="48">R. Turnbow</collectorName>
(1) [
<collectionCode id="ED4FAE77FF82FFD7F9FB080EFE9FF9EB" box="[239,325,1575,1599]" pageId="49" pageNumber="48">RHTC</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3446539FF82FFD4F9D4084EFB9EFB2E" lastPageId="50" lastPageNumber="49" pageId="49" pageNumber="48" type="description">
<paragraph id="8BE136B2FF82FFD4F9D4084EFDFBFCEE" blockId="49.[192,1440,1639,1919]" lastBlockId="50.[192,1441,194,826]" lastPageId="50" lastPageNumber="49" pageId="49" pageNumber="48">
<emphasis id="B92AEAA0FF82FFD7F9D4084EFEB1F9AB" bold="true" box="[192,363,1639,1663]" pageId="49" pageNumber="48">Description.</emphasis>
Male (
<typeStatus id="54E58810FF82FFD7F8AA084EFDF6F9AB" box="[446,556,1639,1663]" pageId="49" pageNumber="48" type="holotype">holotype</typeStatus>
). Body length
<quantity id="4CA69B57FF82FFD7FBCD084EFC9FF9AA" box="[729,837,1639,1663]" metricMagnitude="-3" metricUnit="m" metricValue="6.47" pageId="49" pageNumber="48" unit="mm" value="6.47">6.47 mm</quantity>
; width at elytral humeri
<quantity id="4CA69B57FF82FFD7FD6C084EFB3EF9AA" box="[1144,1252,1639,1663]" metricMagnitude="-3" metricUnit="m" metricValue="2.76" pageId="49" pageNumber="48" unit="mm" value="2.76">2.76 mm</quantity>
; form elongate. Elytra bluish-black; head, pronotum, mesosternum, legs yellow-orange.
<emphasis id="B92AEAA0FF82FFD7FD5608AEFB51F94B" bold="true" box="[1090,1163,1671,1695]" pageId="49" pageNumber="48">Head</emphasis>
elongate, with curved carina ventrally extending from behind eye to front edge of postclypeus (
<figureCitation id="13652A37FF82FFD7FD2D088EFB5BF96B" box="[1081,1153,1703,1727]" captionStart="Figures 5-10" captionStartId="9.[278,374,1721,1743]" captionTargetBox="[279,1354,195,1708]" captionTargetId="figure-17@9.[278,1354,195,1709]" captionTargetPageId="9" captionText="Figures 5-10. Male heads (fgs = frontogenal suture, jl = jowl, lc = lateral carina). 5) Hemiphrynus intermedius, lateral view. 6) H. intermedius, frontal view. 7) Phrynocepha deyrollei, lateral view. 8) P. deyrollei, frontal view. 9) P. pseudocapitata, lateral view. 10) P. pseudocapitata, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5161409" httpUri="https://zenodo.org/record/5161409/files/figure.png" pageId="49" pageNumber="48">Fig. 9</figureCitation>
), appearing “jowl-like;” genal length 1.76x longer than maximum length of eye; vertex shining, impunctate; postclypeus with prefrontal ridge shallowly, broadly emarginate; an acute, elevated frontal carina extends between antennal fossae, curving upward before descending to front edge of postclypeus; anteclypeus entire, same color as labrum; labrum orange, rounded, with six long, evenly spaced setae; antennal fossae separated by distance slightly less than width of antennal fossa (fossal width measured from inside edges); frontal tubercles flat, distinct, separated by coronal suture; area in front of antennal fossa with sunken appearance, bordered by acute lateral carina extending from eye to front edge of postclypeus; frontogenal suture indistinct; interocular distance 1.80x wider than width of both eyes combined; eyes nearly round, very small relative to head; antennae stout, 0.75x length of body; antennomeres 1-3 orange; 4-11 dark brown; antennomere 3 longest; antennomere 2 shortest; antennomere 4 longer than 5-11; antennomeres 5-11 subequal in length.
<emphasis id="B92AEAA0FF81FFD4FA220F6BFC18FE8E" bold="true" box="[822,962,322,346]" pageId="50" pageNumber="49">Pronotum</emphasis>
dull, smooth, reticulate, not obviously punctate; lateral margins tapering to apex, widest behind middle; basolateral projections small, obtuse; anterolateral projections rectangular, with corners rounded, not strongly projected; basal depression poorly defined.
<emphasis id="B92AEAA0FF81FFD4F8630F88FDD8FE6D" bold="true" box="[375,514,417,441]" pageId="50" pageNumber="49">Scutellum</emphasis>
dark brown, triangular, impunctate.
<emphasis id="B92AEAA0FF81FFD4FAD00F8BFBC1FE6E" bold="true" box="[964,1051,418,442]" pageId="50" pageNumber="49">Elytra</emphasis>
distinctly reticulate, irregularly and inconspicuously punctate; humeral costa not obvious; humeral depression indistinct.
<emphasis id="B92AEAA0FF81FFD4FC180FEBFE61FE2E" bold="true" pageId="50" pageNumber="49">Prosternal intercoxal process</emphasis>
elevated ventrally over plane of prosternum and even with plane of tip of procoxae, narrowly, distinctly separating coxae, rising to a small wedge-shaped knob midway between procoxae.
<emphasis id="B92AEAA0FF81FFD4F9D40C0BFE9BFDEE" bold="true" box="[192,321,546,570]" pageId="50" pageNumber="49">Procoxae</emphasis>
globose.
<emphasis id="B92AEAA0FF81FFD4F8BA0C0BFD40FDED" bold="true" box="[430,666,545,570]" pageId="50" pageNumber="49">Procoxal cavities</emphasis>
open.
<emphasis id="B92AEAA0FF81FFD4FBF30C0BFC78FDEE" bold="true" box="[743,930,546,570]" pageId="50" pageNumber="49">Mesosternum</emphasis>
orange.
<emphasis id="B92AEAA0FF81FFD4FD1D0C0BFB93FDEE" bold="true" box="[1033,1097,546,570]" pageId="50" pageNumber="49">Legs</emphasis>
orange; tarsi orange-brown, darker than femora or tibiae; pro- and mesofemora enlarged; tarsomere 1 of protarsi enlarged, heartshaped, wider than tarsomere 3; tarsomere 1 of mesotarsi enlarged, more elongate than tarsomere 1 of protarsi, but still wider than tarsomere 3 of mesotarsi; tarsomere 1 of metatarsi enlarged, more elongate than tarsomere 1 of pro- or mesotarsi, wider than tarsomere 3 of metatarsi; all tibiae stout, sulcate, expanded apically, with distinct central longitudinal carina.
<emphasis id="B92AEAA0FF81FFD4FA8F0CE8FBC4FD0D" bold="true" box="[923,1054,705,729]" pageId="50" pageNumber="49">Abdomen</emphasis>
dark brown, shiny, impunctate, sparsely pubescent; fifth ventrite broadly emarginate with small, central, orange convexity.
<emphasis id="B92AEAA0FF81FFD4FC0F0CC8FA7AFD2D" bold="true" box="[1307,1440,737,761]" pageId="50" pageNumber="49">Aedeagus</emphasis>
slightly, evenly arched; basal piece about 20% of total length (
<figureCitation id="13652A37FF81FFD4FAD20D2BFBC4FCCD" box="[966,1054,769,794]" captionStart="Figures 60-63" captionStartId="37.[192,288,1450,1472]" captionTargetBox="[192,1440,192,1441]" captionTargetId="figure-192@37.[192,1440,192,1441]" captionTargetPageId="37" captionText="Figures 60-63. Aedeagi, lateral view. 60) Phrynocepha pseudocapitata. 61) P. pueblae. 62) P. pulchella. 63) P. punctulata." figureDoi="http://doi.org/10.5281/zenodo.5161449" httpUri="https://zenodo.org/record/5161449/files/figure.png" pageId="50" pageNumber="49">Fig. 60</figureCitation>
); ventral end slightly widened, with no distinct tip (
<figureCitation id="13652A37FF81FFD4F8AF0D0BFDC9FCEE" box="[443,531,802,826]" captionStart="Figures 64-71" captionStartId="39.[192,288,1259,1281]" captionTargetBox="[192,1440,195,1245]" captionTargetId="figure-247@39.[192,1440,195,1245]" captionTargetPageId="39" captionText="Figures 64-71. Tips of aedeagi, ventral view. 64) Phrynocepha capitata. 65) P. deyrollei. 66) P. kendrae. 67) P. marciae. 68) P. pueblae. 69) P. pulchella. 70) P. punctulata. 71) P. pseudocapitata." figureDoi="http://doi.org/10.5281/zenodo.5161451" httpUri="https://zenodo.org/record/5161451/files/figure.png" pageId="50" pageNumber="49">Fig. 71</figureCitation>
).
</paragraph>
<paragraph id="8BE136B2FF81FFD4F9D40D4BFC3FFBCE" blockId="50.[192,1440,866,1050]" pageId="50" pageNumber="49">
<emphasis id="B92AEAA0FF81FFD4F9D40D4BFEFCFCAE" bold="true" box="[192,294,866,890]" pageId="50" pageNumber="49">Female</emphasis>
(
<typeStatus id="54E58810FF81FFD4F82E0D4BFE78FCAE" box="[314,418,866,890]" pageId="50" pageNumber="49" type="allotype">allotype</typeStatus>
). Similar to
<typeStatus id="54E58810FF81FFD4FB500D4BFD6BFCAE" box="[580,689,866,890]" pageId="50" pageNumber="49" type="holotype">holotype</typeStatus>
, differing in the following characters: size slightly smaller (length
<quantity id="4CA69B57FF81FFD4F80B0DA8FE51FC4D" box="[287,395,897,921]" metricMagnitude="-3" metricUnit="m" metricValue="6.29" pageId="50" pageNumber="49" unit="mm" value="6.29">6.29 mm</quantity>
; width
<quantity id="4CA69B57FF81FFD4F8F00DABFD8BFC4E" box="[484,593,898,922]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="50" pageNumber="49" unit="mm" value="2.7">2.70 mm</quantity>
); tarsomere 1 not broadly enlarged on pro- or mesotarsi; fifth ventrite not modified; each elytron with short, distinct humeral costa and 3-4 additional vague costae (light and angles must be right to observe the costae); “jowl” less pronounced laterally (in frontal view the margin of the “jowl” in the male is visible which gives the male a broader head than the female below the eyes); all tibiae similar, slender, gradually expanded apically.
</paragraph>
<paragraph id="8BE136B2FF81FFD4F9D40A68FB9EFB2E" blockId="50.[192,1440,1089,1274]" pageId="50" pageNumber="49">
<emphasis id="B92AEAA0FF81FFD4F9D40A68FE96FB8D" bold="true" box="[192,332,1089,1113]" pageId="50" pageNumber="49">Variation.</emphasis>
Male: length
<quantity id="4CA69B57FF81FFD4F8E10A68FD6EFB8D" box="[501,692,1089,1114]" metricMagnitude="-3" metricUnit="m" metricValue="6.56" metricValueMax="7.47" metricValueMin="5.65" pageId="50" pageNumber="49" unit="mm" value="6.56" valueMax="7.47" valueMin="5.65">5.65 7.47 mm</quantity>
; width at humeri
<quantity id="4CA69B57FF81FFD4FA9B0A6BFB95FB8D" box="[911,1103,1090,1114]" metricMagnitude="-3" metricUnit="m" metricValue="2.635" metricValueMax="3.18" metricValueMin="2.09" pageId="50" pageNumber="49" unit="mm" value="2.635" valueMax="3.18" valueMin="2.09">2.09 3.18 mm</quantity>
. Female: length
<quantity id="4CA69B57FF81FFD4FC0E0A68FF28FBAD" metricMagnitude="-3" metricUnit="m" metricValue="6.324999999999999" metricValueMax="7.18" metricValueMin="5.47" pageId="50" pageNumber="49" unit="mm" value="6.324999999999999" valueMax="7.18" valueMin="5.47">5.47 7.18 mm</quantity>
; width
<quantity id="4CA69B57FF81FFD4F8440A4BFDCFFBAD" box="[336,533,1122,1146]" metricMagnitude="-3" metricUnit="m" metricValue="2.71" metricValueMax="3.13" metricValueMin="2.29" pageId="50" pageNumber="49" unit="mm" value="2.71" valueMax="3.13" valueMin="2.29">2.29 3.13 mm</quantity>
. The stout or slender appearance of the antennae, as represented by the fourth antennomere (length to width ratio), varies. In the specimens selected, the ratio of the fourth antennomere of the male is stout, approximately 2.5x longer than wide, while in the female the ratio of the fourth antennomere is even more stout, approximately 2.2x longer than wide. In the female, the number and degree of additional elytral costae varies among individuals.
</paragraph>
</subSubSection>
<subSubSection id="C3446539FF81FFD4F9D40B0BFDD3FAAE" pageId="50" pageNumber="49" type="discussion">
<paragraph id="8BE136B2FF81FFD4F9D40B0BFDD3FAAE" blockId="50.[192,1440,1314,1402]" pageId="50" pageNumber="49">
<emphasis id="B92AEAA0FF81FFD4F9D40B0BFE9EFAEE" bold="true" box="[192,324,1314,1338]" pageId="50" pageNumber="49">Remarks.</emphasis>
The only obvious external difference between this species and
<taxonomicName id="4C5E4D31FF81FFD4FD4A0B0BFB3CFAED" authorityName="Jacoby" authorityYear="1884" box="[1118,1254,1314,1337]" class="Insecta" family="Chrysomelidae" genus="Phrynocepha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="49" phylum="Arthropoda" rank="species" species="capitata">
<emphasis id="B92AEAA0FF81FFD4FD4A0B0BFBAAFAED" box="[1118,1136,1314,1337]" italics="true" pageId="50" pageNumber="49">P</emphasis>
.
<emphasis id="B92AEAA0FF81FFD4FD950B0BFB3CFAED" box="[1153,1254,1314,1337]" italics="true" pageId="50" pageNumber="49">capitata</emphasis>
</taxonomicName>
is the smooth, lightly punctured pronotum (
<figureCitation id="13652A37FF81FFD4FB3D0B6BFDA5FA8D" box="[553,639,1345,1370]" captionStart="Figures 51-54" captionStartId="34.[192,288,1453,1475]" captionTargetBox="[192,1440,192,1442]" captionTargetId="figure-235@34.[192,1440,192,1442]" captionTargetPageId="34" captionText="Figures 51-54. Pronota of Phrynocepha. 51) P. marciae. 52) P. pseudocapitata. 53) P. pueblae. 54) P. punctulata." figureDoi="http://doi.org/10.5281/zenodo.5161439" httpUri="https://zenodo.org/record/5161439/files/figure.png" pageId="50" pageNumber="49">Fig. 52</figureCitation>
) and the distinct carina on the metatibiae. See “Remarks” for
<taxonomicName id="4C5E4D31FF81FFD4FC930B6BFEFFFAAD" authorityName="Jacoby" authorityYear="1884" class="Insecta" family="Chrysomelidae" genus="Phrynocepha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="49" phylum="Arthropoda" rank="species" species="capitata">
<emphasis id="B92AEAA0FF81FFD4FC930B6BFA43FA8D" box="[1415,1433,1346,1369]" italics="true" pageId="50" pageNumber="49">P</emphasis>
.
<emphasis id="B92AEAA0FF81FFD4F9D40B4BFEFFFAAD" box="[192,293,1378,1401]" italics="true" pageId="50" pageNumber="49">capitata</emphasis>
</taxonomicName>
and other species.
</paragraph>
</subSubSection>
<subSubSection id="C3446539FF81FFD4F9D40B88FD57FA0E" pageId="50" pageNumber="49" type="distribution">
<paragraph id="8BE136B2FF81FFD4F9D40B88FD57FA0E" blockId="50.[192,1440,1441,1498]" pageId="50" pageNumber="49">
<emphasis id="B92AEAA0FF81FFD4F9D40B88FEAEFA6D" bold="true" box="[192,372,1441,1465]" pageId="50" pageNumber="49">Distribution.</emphasis>
Known only from the
<typeStatus id="54E58810FF81FFD4FB900B8BFD63FA6E" box="[644,697,1442,1466]" pageId="50" pageNumber="49">type</typeStatus>
locality in the state of
<collectingRegion id="499AF850FF81FFD4FAC50B8BFBE3FA6E" box="[977,1081,1442,1466]" country="Mexico" name="Chiapas" pageId="50" pageNumber="49">Chiapas</collectingRegion>
,
<collectingCountry id="F3497622FF81FFD4FD510B8BFB45FA6E" box="[1093,1183,1442,1466]" name="Mexico" pageId="50" pageNumber="49">Mexico</collectingCountry>
. All of the specimens were collected at
<quantity id="4CA69B57FF81FFD4F8810BEBFDD7FA0E" box="[405,525,1474,1498]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="50" pageNumber="49" unit="ft" value="3000.0">3,000 feet</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="C3446539FF81FFD4F9D4082BFE55F9CE" box="[192,399,1538,1562]" pageId="50" pageNumber="49" type="biology_ecology">
<paragraph id="8BE136B2FF81FFD4F9D4082BFE55F9CE" blockId="50.[192,399,1538,1562]" box="[192,399,1538,1562]" pageId="50" pageNumber="49">
<emphasis id="B92AEAA0FF81FFD4F9D4082BFED2F9CE" bold="true" box="[192,264,1538,1562]" pageId="50" pageNumber="49">Host.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3446539FF81FFD4F9D4086BFCD7F94E" pageId="50" pageNumber="49" type="etymology">
<paragraph id="8BE136B2FF81FFD4F9D4086BFC70F98D" blockId="50.[192,938,1602,1626]" box="[192,938,1602,1626]" pageId="50" pageNumber="49">
<emphasis id="B92AEAA0FF81FFD4F9D4086BFE86F98E" bold="true" box="[192,348,1602,1626]" pageId="50" pageNumber="49">Etymology.</emphasis>
Named for its very close likeness to
<taxonomicName id="4C5E4D31FF81FFD4FA0A086BFC79F98D" authorityName="Jacoby" authorityYear="1884" box="[798,931,1602,1625]" class="Insecta" family="Chrysomelidae" genus="Phrynocepha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="49" phylum="Arthropoda" rank="species" species="capitata">
<emphasis id="B92AEAA0FF81FFD4FA0A086BFCEAF98D" box="[798,816,1602,1625]" italics="true" pageId="50" pageNumber="49">P</emphasis>
.
<emphasis id="B92AEAA0FF81FFD4FA2A086BFC79F98D" box="[830,931,1602,1625]" italics="true" pageId="50" pageNumber="49">capitata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE136B2FF81FFD4F9D408A8FCD7F94E" blockId="50.[192,781,1665,1690]" box="[192,781,1665,1690]" pageId="50" pageNumber="49">
<emphasis id="B92AEAA0FF81FFD4F9D408A8FE35F94D" bold="true" box="[192,495,1665,1689]" pageId="50" pageNumber="49">Specimens Examined.</emphasis>
See “
<typeStatus id="54E58810FF81FFD4FB2008ABFDABF94E" box="[564,625,1666,1690]" pageId="50" pageNumber="49">Type</typeStatus>
Specimens.”
</paragraph>
</subSubSection>
</treatment>
</document>