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<document id="C43E98BADC9FC071107609396110CAEB" ID-DOI="10.5281/zenodo.275090" ID-GBIF-Dataset="3b4dc4c4-2b0d-4328-a28d-0f778deb8622" ID-ISSN="1175-5326" ID-Zenodo-Dep="275090" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461100627665" checkinUser="plazi" docAuthor="Lamboj, Anton" docDate="2009" docId="03F787A23A72BA2FFF0EFFDCFAE78A53" docLanguage="en" docName="zt02173p048.pdf" docOrigin="Zootaxa 2173" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Enigmatochromis lucanusi Lamboj, 2009, new species" docType="treatment" docVersion="8" lastPageNumber="47" masterDocId="FFCEFFDA3A76BA29FF99FF4BFFC68B1F" masterDocTitle="A new dwarf cichlid genus and species (Teleostei, Cichlidae) from Guinea, West Africa" masterLastPageNumber="48" masterPageNumber="41" pageNumber="45" updateTime="1698591197457" updateUser="plazi">
<mods:mods id="7BBAA4524B28DB27022EA0F908E07865" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AC14436853C9DDA82028E1546CD01999">
<mods:title id="C0A911CD46A6429DB3581C6C37BC9928">A new dwarf cichlid genus and species (Teleostei, Cichlidae) from Guinea, West Africa</mods:title>
</mods:titleInfo>
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<mods:namePart id="6B53894F604572D9818D35EFF41C5863">Lamboj, Anton</mods:namePart>
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<mods:typeOfResource id="9F017075551127B2CDB6D1A65B8F350B">text</mods:typeOfResource>
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<mods:title id="9EDFFD463835C9C21AEB9B0348F68FBF">Zootaxa</mods:title>
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<mods:part id="9997C93F36E8230312B542D10C5EAD27">
<mods:date id="B811BFBD5457CDD4E5C1B466FD1BA159">2009</mods:date>
<mods:detail id="DF79844C2615D0FF1CD281322E075473" type="volume">
<mods:number id="5C0B9D450398532844256341660EEF85">2173</mods:number>
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<mods:classification id="8D9EB38EEB6692F39482460916F814B3">journal article</mods:classification>
<mods:identifier id="B4961427670B8208CA31ABC5E4D6E532" type="DOI">10.5281/zenodo.275090</mods:identifier>
<mods:identifier id="8F62BAB8A2D3BE4E897E0A9455C8DCBD" type="GBIF-Dataset">3b4dc4c4-2b0d-4328-a28d-0f778deb8622</mods:identifier>
<mods:identifier id="52D88DB5571BA23AC0378684EF47B0B1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9C56DBBCDB6341901E66594A47B05A93" type="Zenodo-Dep">275090</mods:identifier>
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</mods:mods>
<treatment id="03F787A23A72BA2FFF0EFFDCFAE78A53" ID-DOI="http://doi.org/10.5281/zenodo.6214563" ID-GBIF-Taxon="119621689" ID-Zenodo-Dep="6214563" LSID="urn:lsid:plazi:treatment:03F787A23A72BA2FFF0EFFDCFAE78A53" httpUri="http://treatment.plazi.org/id/03F787A23A72BA2FFF0EFFDCFAE78A53" lastPageId="6" lastPageNumber="47" pageId="4" pageNumber="45">
<subSubSection id="C344653F3A72BA2DFF0EFFDCFE538BCB" pageId="4" pageNumber="45" type="nomenclature">
<paragraph id="8BE136B43A72BA2DFF0EFFDCFDBB8BAE" blockId="4.[151,637,151,212]" box="[151,637,151,177]" pageId="4" pageNumber="45">
<heading id="D0A981D83A72BA2DFF0EFFDCFDBB8BAE" bold="true" box="[151,637,151,177]" fontSize="11" level="1" pageId="4" pageNumber="45" reason="1">
<emphasis id="B92AEAA63A72BA2DFF0EFFDCFDBB8BAE" bold="true" box="[151,637,151,177]" pageId="4" pageNumber="45">
<taxonomicName id="4C5E4D373A72BA2DFF0EFFDCFE198BAE" ID-CoL="39V5N" box="[151,479,151,177]" class="Actinopterygii" family="Cichlidae" genus="Enigmatochromis" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="45" phylum="Chordata" rank="species" species="lucanusi" status="sp. nov.">
<emphasis id="B92AEAA63A72BA2DFF0EFFDCFE198BAE" bold="true" box="[151,479,151,177]" italics="true" pageId="4" pageNumber="45">Enigmatochromis lucanusi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21957DD3A72BA2DFE77FFDCFDBB8BAE" box="[494,637,151,177]" pageId="4" pageNumber="45" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE136B43A72BA2DFF0EFFF1FE538BCB" blockId="4.[151,637,151,212]" box="[151,405,186,212]" pageId="4" pageNumber="45">
(
<figureCitation id="13652A313A72BA2DFF06FFF1FF378BCB" box="[159,241,186,212]" captionStart="FIGURE 1" captionStartId="1.[151,258,1339,1363]" captionTargetBox="[157,1429,784,1309]" captionTargetId="figure@1.[151,1436,773,1314]" captionTargetPageId="1" captionText="FIGURE 1. Enigmatochromis lucanusi, NMW 95063, holotype, male, 44.7 mm SL; Guinea: Small creek near the village of Fria, north of Conakry. Scale bar = 10 mm." httpUri="https://zenodo.org/record/275091/files/figure.png" pageId="4" pageNumber="45">Figs. 1</figureCitation>
&amp;
<figureCitation id="13652A313A72BA2DFE8FFFF1FEE18BCB" box="[278,295,186,212]" captionStart="FIGURE 2" captionStartId="2.[151,255,1792,1816]" captionTargetBox="[151,1434,705,1765]" captionTargetId="figure@2.[151,1436,705,1766]" captionTargetPageId="2" captionText="FIGURE 2. Photographs showing coloration of Enigmatochromis lucanusi, nov. sp.: A) male, wild collected specimen in aquaria, ~ 40 mm SL, not preserved, displaying submissive coloration; B) male, wild collected specimen in aquaria, ~ 42 mm SL, not preserved, displaying aggressive coloration; C) female, wild collected specimen in aquaria, ~ 38 mm SL, not preserved, displaying submissive coloration; D) female, wild collected specimen in aquaria, ~ 39 mm SL, not preserved, displaying aggressive coloration; E) female, wild collected specimen in aquaria, ~ 38 mm SL, with juveniles at the age of approx. one week after swimming free and size of less than 12 mm TL, not preserved; F) aquarium bred juveniles with the age of app. one week after swimming free, size less than 12 mm TL, not preserved." httpUri="https://zenodo.org/record/275092/files/figure.png" pageId="4" pageNumber="45">2</figureCitation>
; Table 1)
</paragraph>
</subSubSection>
<subSubSection id="C344653F3A72BA2DFF0EFE43FD458A00" box="[151,643,264,287]" pageId="4" pageNumber="45" type="description">
<paragraph id="8BE136B43A72BA2DFF0EFE43FD458A00" blockId="4.[151,643,264,287]" box="[151,643,264,287]" pageId="4" pageNumber="45">
<taxonomicName id="4C5E4D373A72BA2DFF0EFE43FEFA8A00" box="[151,316,264,287]" class="Actinopterygii" family="Cichlidae" genus="Pelvicachromis" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis id="B92AEAA63A72BA2DFF0EFE43FEFA8A00" box="[151,316,264,287]" italics="true" pageId="4" pageNumber="45">Pelvicachromis</emphasis>
</taxonomicName>
sp. “Blue Fin” (
<bibRefCitation id="EFCF4B453A72BA2DFE73FE43FDBA8A00" author="Lamboj" box="[490,636,264,287]" pageId="4" pageNumber="48" refString="Lamboj, A. (2005) Pelvicachromis species from Guinea. Cichlid News, 14 (4), 6 - 13." type="journal article" year="2005">Lamboj 2005</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C344653F3A72BA2DFF0EFE05FBCC8981" pageId="4" pageNumber="45" type="materials_examined">
<paragraph id="8BE136B43A72BA2DFF0EFE05FAF48A90" blockId="4.[151,1437,334,2029]" pageId="4" pageNumber="45">
<emphasis id="B92AEAA63A72BA2DFF0EFE05FED28A77" bold="true" box="[151,276,334,360]" pageId="4" pageNumber="45">
<typeStatus id="54E588163A72BA2DFF0EFE05FEC98A77" box="[151,271,334,360]" pageId="4" pageNumber="45" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED4FAE713A72BA2DFE85FE05FEAE8A77" box="[284,360,334,360]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="45">NMW</collectionCode>
95063, male,
<quantity id="4CA69B513A72BA2DFD8EFE05FD458A77" box="[535,643,334,360]" metricMagnitude="-2" metricUnit="m" metricValue="4.47" pageId="4" pageNumber="45" unit="mm" value="44.7">44.7 mm</quantity>
SL;
<collectingCountry id="F34976243A72BA2DFD59FE05FCDA8A77" box="[704,796,334,360]" name="Guinea-Bissau" pageId="4" pageNumber="45">Guinea</collectingCountry>
: Foto River near the town of Fria, north of Conakry, Konkoure River system.
<geoCoordinate id="EE6A50733A72BA2DFE25FE3EFDA08A90" box="[444,614,373,399]" direction="north" orientation="latitude" pageId="4" pageNumber="45" precision="1" value="10.334567">10° 20.074’ N</geoCoordinate>
,
<geoCoordinate id="EE6A50733A72BA2DFDEBFE3EFCE68A90" box="[626,800,373,399]" direction="west" orientation="longitude" pageId="4" pageNumber="45" precision="1" value="-13.72385">13° 43.431’ W</geoCoordinate>
, collected by W. M. Traore,
<date id="FFE010743A72BA2DFBF4FE3EFAE88A90" box="[1133,1326,373,399]" pageId="4" pageNumber="45" value="2007-11">November 2007</date>
.
</paragraph>
<paragraph id="8BE136B43A72BA2DFF5CFED0FBCC8981" blockId="4.[151,1437,334,2029]" pageId="4" pageNumber="45">
<emphasis id="B92AEAA63A72BA2DFF5CFED0FE8C8AAA" bold="true" box="[197,330,411,437]" pageId="4" pageNumber="45">
<typeStatus id="54E588163A72BA2DFF5CFED0FE838AAA" box="[197,325,411,437]" pageId="4" pageNumber="45" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED4FAE713A72BA2DFECBFED7FE5B8AA9" box="[338,413,412,438]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="45">NMW</collectionCode>
95064,
<specimenCount id="9D58FD3D3A72BA2DFE64FED7FD9D8AA9" box="[509,603,412,438]" pageId="4" pageNumber="45" type="male">2 males</specimenCount>
,
<specimenCount id="9D58FD3D3A72BA2DFDF1FED7FD178AA9" box="[616,721,412,438]" pageId="4" pageNumber="45" type="female">1 female</specimenCount>
,
<quantity id="4CA69B513A72BA2DFD44FED7FC4F8AA9" box="[733,905,412,438]" metricMagnitude="-2" metricUnit="m" metricValue="3.9099999999999993" metricValueMax="4.529999999999999" metricValueMin="3.29" pageId="4" pageNumber="45" unit="mm" value="39.099999999999994" valueMax="45.3" valueMin="32.9">32.9–45.3 mm</quantity>
SL, same collection data as
<typeStatus id="54E588163A72BA2DFB42FED7FA838AA9" box="[1243,1349,412,438]" pageId="4" pageNumber="45" type="holotype">holotype</typeStatus>
.
<collectionCode id="ED4FAE713A72BA2DFAC8FED7FA5A8AA9" box="[1361,1436,412,438]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="45">NMW</collectionCode>
95065,
<specimenCount id="9D58FD3D3A72BA2DFF68FE89FE978AC3" box="[241,337,450,476]" pageId="4" pageNumber="45" type="male">2 males</specimenCount>
,
<specimenCount id="9D58FD3D3A72BA2DFEC7FE89FE138AC3" box="[350,469,450,476]" pageId="4" pageNumber="45" type="female">2 females</specimenCount>
, 1 undet.,
<quantity id="4CA69B513A72BA2DFDCAFE89FCC78AC3" box="[595,769,450,476]" metricMagnitude="-2" metricUnit="m" metricValue="3.67" metricValueMax="4.06" metricValueMin="3.28" pageId="4" pageNumber="45" unit="mm" value="36.7" valueMax="40.6" valueMin="32.8">32.8–40.6 mm</quantity>
SL, same locality as
<typeStatus id="54E588163A72BA2DFB9CFE89FBB78AC3" box="[1029,1137,450,476]" pageId="4" pageNumber="45" type="holotype">holotype</typeStatus>
, but collected in
<date id="FFE010743A72BA2DFAA6FE89FF1E891C" pageId="4" pageNumber="45" value="2004-01">January 2004</date>
;
<specimenCount id="9D58FD3D3A72BA2DFF7FFEA2FE93891C" box="[230,341,489,515]" pageId="4" pageNumber="45" type="female">1 female</specimenCount>
, 1 undet. used for C&amp;S after taking counts and measurements.
<collectionCode id="ED4FAE713A72BA2DFBF0FEA2FB70891C" box="[1129,1206,489,515]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="45">NMW</collectionCode>
95066,
<specimenCount id="9D58FD3D3A72BA2DFA84FEA2FA46891C" box="[1309,1408,489,515]" pageId="4" pageNumber="45" type="male">5 males</specimenCount>
,
<specimenCount id="9D58FD3D3A72BA2DFA14FEA2FF338935" pageId="4" pageNumber="45" type="female">4 females</specimenCount>
,
<quantity id="4CA69B513A72BA2DFE98FD5BFE6B8935" box="[257,429,528,554]" metricMagnitude="-2" metricUnit="m" metricValue="3.8549999999999995" metricValueMax="4.43" metricValueMin="3.28" pageId="4" pageNumber="45" unit="mm" value="38.55" valueMax="44.3" valueMin="32.8">32.8–44.3 mm</quantity>
SL, same locality as
<typeStatus id="54E588163A72BA2DFD32FD5BFCD78935" box="[683,785,528,554]" pageId="4" pageNumber="45" type="holotype">holotype</typeStatus>
but collected in
<date id="FFE010743A72BA2DFC40FD5BFB658935" box="[985,1187,528,554]" pageId="4" pageNumber="45" value="2006-02-02">
February 2006,
<specimenCount id="9D58FD3D3A72BA2DFB0DFD5BFB658935" box="[1172,1187,528,554]" pageId="4" pageNumber="45" type="male">2</specimenCount>
</date>
males and
<specimenCount id="9D58FD3D3A72BA2DFAB3FD5BFA5A8935" box="[1322,1436,528,554]" pageId="4" pageNumber="45" type="female">2 females</specimenCount>
used for C&amp;S after taking counts and measurements.
<collectionCode id="ED4FAE713A72BA2DFC89FD7DFCAC894F" LSID="urn:lsid:biocol.org:col:34925" box="[784,874,566,592]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="45">AMNH</collectionCode>
238830,
<specimenCount id="9D58FD3D3A72BA2DFC43FD7DFBF1894F" box="[986,1079,566,592]" pageId="4" pageNumber="45" type="male">2 males</specimenCount>
,
<specimenCount id="9D58FD3D3A72BA2DFBDDFD7DFB70894F" box="[1092,1206,566,592]" pageId="4" pageNumber="45" type="female">2 females</specimenCount>
<quantity id="4CA69B513A72BA2DFB27FD7DFAAC894F" box="[1214,1386,566,592]" metricMagnitude="-2" metricUnit="m" metricValue="3.76" metricValueMax="4.4399999999999995" metricValueMin="3.08" pageId="4" pageNumber="45" unit="mm" value="37.6" valueMax="44.4" valueMin="30.8">30.8–44.4 mm</quantity>
SL, same locality as
<typeStatus id="54E588163A72BA2DFEF3FD16FE1D8968" box="[362,475,605,631]" pageId="4" pageNumber="45" type="holotype">holotype</typeStatus>
, but collected in
<date id="FFE010743A72BA2DFD2CFD16FCA78968" box="[693,865,605,631]" pageId="4" pageNumber="45" value="2004-01">January 2004</date>
.
<collectionCode id="ED4FAE713A72BA2DFCF7FD16FC0E8968" box="[878,968,605,631]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="45">MRAC</collectionCode>
2008-015-P-1-4,
<specimenCount id="9D58FD3D3A72BA2DFB30FD16FACB8968" box="[1193,1293,605,631]" pageId="4" pageNumber="45" type="male">2 males</specimenCount>
,
<specimenCount id="9D58FD3D3A72BA2DFA82FD16FA5E8968" box="[1307,1432,605,631]" pageId="4" pageNumber="45" type="female">2 females</specimenCount>
,
<quantity id="4CA69B513A72BA2DFF0EFDCFFE878981" box="[151,321,644,670]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.39" metricValueMin="3.31" pageId="4" pageNumber="45" unit="mm" value="38.5" valueMax="43.9" valueMin="33.1">33.1–43.9 mm</quantity>
SL, same locality as
<typeStatus id="54E588163A72BA2DFDA2FDCFFD678981" box="[571,673,644,670]" pageId="4" pageNumber="45" type="holotype">holotype</typeStatus>
but collected in
<date id="FFE010743A72BA2DFCFCFDCFFBC08981" box="[869,1030,644,670]" pageId="4" pageNumber="45" value="2008-01">January 2008</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C344653F3A72BA2DFF5CFDE1FA5489F4" pageId="4" pageNumber="45" type="description">
<paragraph id="8BE136B43A72BA2DFF5CFDE1FA5489F4" blockId="4.[151,1437,334,2029]" pageId="4" pageNumber="45">
<emphasis id="B92AEAA63A72BA2DFF5CFDE1FEA389DB" bold="true" box="[197,357,682,708]" pageId="4" pageNumber="45">None-types.</emphasis>
In addition, 8 pairs of wild collected fishes and their F1-descendents were used for behavioural observations, but not included in morphological examinations or
<typeStatus id="54E588163A72BA2DFBBDFD9AFB9189F4" box="[1060,1111,721,747]" pageId="4" pageNumber="45">type</typeStatus>
series or other collections.
</paragraph>
</subSubSection>
<subSubSection id="C344653F3A72BA2DFF5CFDBCFDEF880D" box="[197,553,759,786]" pageId="4" pageNumber="45" type="diagnosis">
<paragraph id="8BE136B43A72BA2DFF5CFDBCFDEF880D" blockId="4.[151,1437,334,2029]" box="[197,553,759,786]" pageId="4" pageNumber="45">
<emphasis id="B92AEAA63A72BA2DFF5CFDBCFE82880E" bold="true" box="[197,324,759,785]" pageId="4" pageNumber="45">Diagnosis.</emphasis>
Same as for genus.
</paragraph>
</subSubSection>
<subSubSection id="C344653F3A72BA2CFF5CFC55FCDD8D89" lastPageId="5" lastPageNumber="46" pageId="4" pageNumber="45" type="description">
<paragraph id="8BE136B43A72BA2DFF5CFC55FF338840" blockId="4.[151,1437,334,2029]" pageId="4" pageNumber="45">
<emphasis id="B92AEAA63A72BA2DFF5CFC55FE988827" bold="true" box="[197,350,798,824]" pageId="4" pageNumber="45">Description.</emphasis>
Measurements and meristic counts for
<typeStatus id="54E588163A72BA2DFCAAFC55FC5C8827" box="[819,922,798,824]" pageId="4" pageNumber="45" type="holotype">holotype</typeStatus>
and 25 syntopic
<typeStatus id="54E588163A72BA2DFBFEFC55FB1E8827" box="[1127,1240,798,824]" pageId="4" pageNumber="45" type="paratype">paratypes</typeStatus>
are presented in Table 1.
</paragraph>
<paragraph id="8BE136B43A72BA2DFF5FFC27FB1F8F3F" blockId="4.[151,1437,334,2029]" pageId="4" pageNumber="45">Small cichlid species with body moderately gracile, dorsal head profile smoothly rounded and short snout. Ventral body wall gently rounded to caudal peduncle. Caudal fin rounded. Caudal peduncle short, always deeper than long. Sexual dimorphism well-developed with males usually 10–15% larger than females, with soft dorsal and anal fin rays more elongated. First ray of pelvic fin always longest in males, in females of equal length or longer than second ray. In males, tips of pelvic fin reaching origin of anal fin.</paragraph>
<paragraph id="8BE136B43A72BA2DFF5CFB66FCDD8F88" blockId="4.[151,1437,334,2029]" pageId="4" pageNumber="45">
<emphasis id="B92AEAA63A72BA2DFF5CFB66FE318F58" bold="true" box="[197,503,1069,1095]" pageId="4" pageNumber="45">Osteology and dentition.</emphasis>
Infraorbital bones series with lachrymal and three additional tubular bones and a gap between 2nd and 3rd tubular infraorbital; lachrymal with four openings of laterosensory system. 23–25 total vertebrae (12–14 abdominal and 11 or 12 caudal).
</paragraph>
<paragraph id="8BE136B43A72BA2DFF5FFBEFFC0B8E46" blockId="4.[151,1437,334,2029]" pageId="4" pageNumber="45">Premaxilla with one or two, dentary with one to three rows of regularly set unicuspid teeth. Anteriorly in the lower jaw a few teeth oriented posteriorly, not buccally. Lower pharyngeal bone triangular, with unicuspid to weakly bicuspid teeth on lateral parts of this bone, and asymmetric bicuspid teeth in the central field. Gill rakers on first gill arch. Eight to twelve tuberculate gill rakers on ceratobranchials, 4–7 pointed gill rakers on epibranchials. Well-developed hanging pad on roof of the pharynx.</paragraph>
<paragraph id="8BE136B43A72BA2DFF5CFA2EFBE28D5B" blockId="4.[151,1437,334,2029]" pageId="4" pageNumber="45">
<emphasis id="B92AEAA63A72BA2DFF5CFA2EFEDE8E60" bold="true" box="[197,280,1381,1407]" pageId="4" pageNumber="45">Scales.</emphasis>
Cycloid, one or two rows of scales on cheek; four horizontal rows on opercle. Dark spot on outer edge of opercle unscaled. Chest-scales smaller than body scales, four or five scales between pectoral and pelvic fins. Upper lateral-line separated from dorsal-fin base anteriorly by three scales, at the 8th pored scale by one or one and a half scales, and at last pored scale by no or a half scale. End of upper lateral-line never overlapping lower lateral-line, and separated from beginning of lower lateral line by one to four rows of scales. About ¼ of caudal fin covered with scales, and all other fins unscaled.
</paragraph>
<paragraph id="8BE136B43A72BA2DFF5CF91BFDC08CBF" blockId="4.[151,1437,334,2029]" pageId="4" pageNumber="45">
<emphasis id="B92AEAA63A72BA2DFF5CF91BFE8A8D75" bold="true" box="[197,332,1616,1642]" pageId="4" pageNumber="45">Coloration</emphasis>
. Live specimens of both sexes (
<figureCitation id="13652A313A72BA2DFD5CF91BFCC88D75" box="[709,782,1616,1642]" captionStart="FIGURE 2" captionStartId="2.[151,255,1792,1816]" captionTargetBox="[151,1434,705,1765]" captionTargetId="figure@2.[151,1436,705,1766]" captionTargetPageId="2" captionText="FIGURE 2. Photographs showing coloration of Enigmatochromis lucanusi, nov. sp.: A) male, wild collected specimen in aquaria, ~ 40 mm SL, not preserved, displaying submissive coloration; B) male, wild collected specimen in aquaria, ~ 42 mm SL, not preserved, displaying aggressive coloration; C) female, wild collected specimen in aquaria, ~ 38 mm SL, not preserved, displaying submissive coloration; D) female, wild collected specimen in aquaria, ~ 39 mm SL, not preserved, displaying aggressive coloration; E) female, wild collected specimen in aquaria, ~ 38 mm SL, with juveniles at the age of approx. one week after swimming free and size of less than 12 mm TL, not preserved; F) aquarium bred juveniles with the age of app. one week after swimming free, size less than 12 mm TL, not preserved." httpUri="https://zenodo.org/record/275092/files/figure.png" pageId="4" pageNumber="45">Fig. 2</figureCitation>
): Head and body pale brown to greyish brown. Dorsum somewhat darker than ventral parts of body. Dark scaleless spot on the outer edge of opercle, with a light blue margin on posterior edge. Upper lip brownish to yellow-brown, lower lip greyish to brown. Throat and ventral parts of branchiostegal membranes yellow, more prominent in dominant specimens. Dark longitudinal stripe sometimes visible on sides (in stressed, submissive, breeding and guarding individuals), at about height of lower lateral-line, from posterior edge of eye to end of caudal peduncle, but not extending onto caudal fin. Dark stripe from anterior edge of the eye to the middle of upper lip. Upper edge of eye golden-yellow. Lachrymal bluish, with a red to reddish horizontal stripe from angle of the mouth to anterior edge of preopercular. Pectoral fin clear.
</paragraph>
<paragraph id="8BE136B43A72BA2CFF5CF8E7FC138A39" blockId="4.[151,1437,334,2029]" lastBlockId="5.[151,1437,152,2034]" lastPageId="5" lastPageNumber="46" pageId="4" pageNumber="45">
<emphasis id="B92AEAA63A72BA2DFF5CF8E7FE3D8CD9" bold="true" box="[197,507,1964,1990]" pageId="4" pageNumber="45">Male specific coloration.</emphasis>
Dorsal fin pale yellow, with red margin, followed by a thin white submargin, morer prominent in anterior portion of this fin. Upper half of caudal fin yellow, lower half reddish. Soft ray parts of dorsal fin and upper half of caudal fin with some to numerous black dots in individual variation. Base of anal fin yellowish, rest of this fin reddish to violet. Anterior edge of pelvic fins yellow, first soft ray dark grey to black, other parts of this fin pale reddish to pale violet. Body scales with pale greyish margins. Most ventral parts of opercle, suborpercle, interopercle and chest are yellow.
</paragraph>
<paragraph id="8BE136B43A73BA2CFF5CFE79FEC28944" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="46">
<emphasis id="B92AEAA63A73BA2CFF5CFE79FDCA8A53" bold="true" box="[197,524,306,332]" pageId="5" pageNumber="46">Female specific coloration</emphasis>
. Spiny portion of dorsal fin iridescent blue, sometimes extending to first one to two soft rays, with a red margin, followed by a thin dark to black submargin. Rest of fin clear. One or two black dots in most posterior part of iridescent blue coloration in many individuals. Caudal fin clear to very pale reddish, without patterns or marks. Anal-fin base and most anterior parts bluish, rest of the fin clear to pale reddish. Anterior edge of pelvic fins black, as first soft ray dark, other parts of this fin reddish with thin blue vertical lines in some specimens. Lower parts of opercle, subopercle and interopercle are blue, also most anterior parts of chest. Flanks and belly region, up to about upper lateral line or higher in some individuals, are wine red.
</paragraph>
<paragraph id="8BE136B43A73BA2CFF5CFD23FF1589D0" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="46">
Juveniles of both sexes (before getting adult coloration) exhibit a pattern of three or four rows of irregular dark spots on brown coloration, up to about
<quantity id="4CA69B513A73BA2CFD36FDC5FCE989B7" box="[687,815,654,680]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="5" pageNumber="46" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
SL. With increasing size, sex-specific coloration is seen.
</paragraph>
<paragraph id="8BE136B43A73BA2CFF5CFD90FE58888F" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="46">
<emphasis id="B92AEAA63A73BA2CFF5CFD90FD8489EA" bold="true" box="[197,578,731,757]" pageId="5" pageNumber="46">Preserved specimen coloration</emphasis>
General coloration of both sexes: Head and body brown, darker dorsally. Cheek, throat, pre-pelvic and pre-pectoral regions of flanks and chest pale light brown. Lips greyish. Dark spot on outer edge of opercle. A dark longitudinal stripe visible in some individuals, reaching from posterior edge of eye to end of caudal peduncle, but not extending into the caudal fin. Dark bar from anterior edge of eye to angle of mouth.
</paragraph>
<paragraph id="8BE136B43A73BA2CFF5FFCD6FEA98F4D" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="46">
<emphasis id="B92AEAA63A73BA2CFF5FFCD6FE3788A8" bold="true" box="[198,497,925,951]" pageId="5" pageNumber="46">Male specific coloration</emphasis>
. The general preserved coloration of adult males is shown in the
<typeStatus id="54E588163A73BA2CFB6CFCD6FA9D88A8" box="[1269,1371,925,951]" pageId="5" pageNumber="46" type="holotype">holotype</typeStatus>
(
<figureCitation id="13652A313A73BA2CFAF5FCD6FF6088C1" captionStart="FIGURE 1" captionStartId="1.[151,258,1339,1363]" captionTargetBox="[157,1429,784,1309]" captionTargetId="figure@1.[151,1436,773,1314]" captionTargetPageId="1" captionText="FIGURE 1. Enigmatochromis lucanusi, NMW 95063, holotype, male, 44.7 mm SL; Guinea: Small creek near the village of Fria, north of Conakry. Scale bar = 10 mm." httpUri="https://zenodo.org/record/275091/files/figure.png" pageId="5" pageNumber="46">Fig. 1</figureCitation>
). Anterior portion of dorsal fin greyish. Posterior portion of dorsal fin and upper parts of caudal fin yellowish to greyish, with numerous dark spots. Lower parts of caudal fin pale grey. Anal fin dusky grey to dark grey anteriorly, posterior portion pale greyish to pale reddish. Pelvic fins dark grey to reddish grey, and pectoral fins clear.
</paragraph>
<paragraph id="8BE136B43A73BA2CFF5FFB15FB918FF3" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="46">
<emphasis id="B92AEAA63A73BA2CFF5FFB15FDD28F67" bold="true" box="[198,532,1118,1144]" pageId="5" pageNumber="46">Female specific coloration.</emphasis>
Belly dark grey to reddish grey. Anterior portion of dorsal fin bluish grey to dark grey, posterior portion of this fin clear with no up to two black dots. Caudal fin clear without any markings. Anal fin base dark grey, sometimes with a pale reddish flush, outer parts of fin clear. Pelvic fins reddish to reddish brown, with dark to black anterior edge, and pectoral fins clear.
</paragraph>
<paragraph id="8BE136B43A73BA2CFF5CFBB3FCDD8D89" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="46">
<emphasis id="B92AEAA63A73BA2CFF5CFBB3FE0F8E0D" bold="true" box="[197,457,1272,1298]" pageId="5" pageNumber="46">Breeding behaviour.</emphasis>
In aquaria, this species is a monogamous, pair bonding, cave spawner. Eggs are guarded by both sexes, but more intensively by the female. Hatching occurs after three days post-spawn. Larvae are normally deposited on the bottom of the cave, rarely in other caves nearby the original cave. Juveniles are free swimming 8 or 9 days post-hatching, and are guarded by both parents for about 5 to 6 weeks. Breeding and guarding individuals of both sexes regularly exhibit more aggressive and intensive coloration. The dark, longitudinal stripe that is typical for breeding and guarding specimens of both sexes in many other cave breeders of the chromidotilapiine lineage (e.g.,
<taxonomicName id="4C5E4D373A73BA2CFC4AFAAAFB508EE5" box="[979,1174,1505,1530]" class="Actinopterygii" family="Cichlidae" genus="Pelvicachromis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B92AEAA63A73BA2CFC4AFAAAFB508EE5" box="[979,1174,1505,1530]" italics="true" pageId="5" pageNumber="46">Pelvicachromis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5E4D373A73BA2CFB3EFAAAFA988EE5" box="[1191,1374,1505,1530]" class="Actinopterygii" family="Cichlidae" genus="Congochromis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B92AEAA63A73BA2CFB3EFAAAFA988EE5" box="[1191,1374,1505,1530]" italics="true" pageId="5" pageNumber="46">Congochromis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C5E4D373A73BA2CFF0EF943FEAA8D3E" box="[151,364,1544,1569]" class="Actinopterygii" family="Cichlidae" genus="Parananochromis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B92AEAA63A73BA2CFF0EF943FEAA8D3E" box="[151,364,1544,1569]" italics="true" pageId="5" pageNumber="46">Parananochromis</emphasis>
</taxonomicName>
) is prominently visible in males, but is more rarely and weakly visible in females. In this character, females of
<taxonomicName id="4C5E4D373A73BA2CFE2AF964FD518D57" box="[435,663,1583,1608]" class="Actinopterygii" family="Cichlidae" genus="Enigmatochromis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B92AEAA63A73BA2CFE2AF964FD518D57" box="[435,663,1583,1608]" italics="true" pageId="5" pageNumber="46">Enigmatochromis</emphasis>
</taxonomicName>
differ from females of
<taxonomicName id="4C5E4D373A73BA2CFC57F964FB518D57" box="[974,1175,1583,1608]" class="Actinopterygii" family="Cichlidae" genus="Pelvicachromis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B92AEAA63A73BA2CFC57F964FB518D57" box="[974,1175,1583,1608]" italics="true" pageId="5" pageNumber="46">Pelvicachromis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5E4D373A73BA2CFB30F964FAA38D57" box="[1193,1381,1583,1608]" class="Actinopterygii" family="Cichlidae" genus="Congochromis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B92AEAA63A73BA2CFB30F964FAA38D57" box="[1193,1381,1583,1608]" italics="true" pageId="5" pageNumber="46">Congochromis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5E4D373A73BA2CFF0EF91EFEB78D71" box="[151,369,1621,1646]" class="Actinopterygii" family="Cichlidae" genus="Parananochromis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="B92AEAA63A73BA2CFF0EF91EFEB78D71" box="[151,369,1621,1646]" italics="true" pageId="5" pageNumber="46">Parananochromis</emphasis>
</taxonomicName>
, where a prominent dark, longitudinal stripe is typical for females (and only rarely for males) during the first 2 to 4 weeks when guarding fry.
</paragraph>
</subSubSection>
<subSubSection id="C344653F3A73BA2FFF5FF9E9FAE78A53" lastPageId="6" lastPageNumber="47" pageId="5" pageNumber="46" type="distribution">
<paragraph id="8BE136B43A73BA2CFF5FF9E9FEDF8C15" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="46">
<emphasis id="B92AEAA63A73BA2CFF5FF9E9FEA18DA3" bold="true" box="[198,359,1698,1724]" pageId="5" pageNumber="46">Distribution.</emphasis>
This species is thus far only known from
<typeStatus id="54E588163A73BA2CFCE1F9E9FC6D8DA3" box="[888,939,1698,1724]" pageId="5" pageNumber="46">type</typeStatus>
locality, the Foto River, what is a small savannah river near the bauxite mining town of Fria in coastal
<collectingCountry id="F34976243A73BA2CFCE7F982FC108DFC" box="[894,982,1737,1763]" name="Guinea-Bissau" pageId="5" pageNumber="46">Guinea</collectingCountry>
, north of Conakry,
<geoCoordinate id="EE6A50733A73BA2CFB22F982FAA08DFC" box="[1211,1382,1737,1763]" direction="north" orientation="latitude" pageId="5" pageNumber="46" precision="1" value="10.334567">10° 20.074’ N</geoCoordinate>
,
<geoCoordinate id="EE6A50733A73BA2CFAEAF982FED38C15" direction="west" orientation="longitude" pageId="5" pageNumber="46" precision="1" value="-13.72385">13° 43.431’ W</geoCoordinate>
.
</paragraph>
<paragraph id="8BE136B43A73BA2CFF5FF85DFEB38CBB" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="46">
<emphasis id="B92AEAA63A73BA2CFF5FF85DFEEB8C2F" bold="true" box="[198,301,1814,1840]" pageId="5" pageNumber="46">Habitat.</emphasis>
Detailed habit data for the
<typeStatus id="54E588163A73BA2CFDEBF85DFD628C2F" box="[626,676,1814,1840]" pageId="5" pageNumber="46">type</typeStatus>
locality was provided by O. Lucanus. The river was 3–6 meters wide, and during the dry season only
<quantity id="4CA69B513A73BA2CFDD5F876FD068C48" box="[588,704,1853,1879]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" metricValueMax="9.0" metricValueMin="3.0" pageId="5" pageNumber="46" unit="cm" value="60.0" valueMax="90.0" valueMin="30.0">30–90 cm</quantity>
deep with water visibility ~
<quantity id="4CA69B513A73BA2CFB9EF876FB8B8C48" box="[1031,1101,1853,1879]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="46" unit="m" value="1.2">1.2 m</quantity>
. Water parameters measured during the dry season in February, 2006 were: General hardness 0, Carbonate hardness 0, pH 5.8, and temperature 24° C.
</paragraph>
<paragraph id="8BE136B43A73BA2FFF5FF8FAFEA28BE0" blockId="5.[151,1437,152,2034]" lastBlockId="6.[151,1436,152,332]" lastPageId="6" lastPageNumber="47" pageId="5" pageNumber="46">
This site was ringed by dense forest and often fallen trees obstructing access to the water in many places. Margins of the river were densely overgrown by
<emphasis id="B92AEAA63A73BA2CFD7AF893FC178CEE" box="[739,977,2008,2033]" italics="true" pageId="5" pageNumber="46">
<taxonomicName id="4C5E4D373A73BA2CFD7AF893FC0A8CEE" box="[739,972,2008,2033]" class="Liliopsida" family="Araceae" genus="Anubias" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="46" phylum="Tracheophyta" rank="species" species="lanceolata">Anubias lanceolata</taxonomicName>
,
</emphasis>
and exposed boulders and fallen trees have large clusters of
<emphasis id="B92AEAA63A70BA2FFE03FFD3FDBC8BAE" box="[410,634,152,177]" italics="true" pageId="6" pageNumber="47">Bolibitis heudelotti</emphasis>
and other ferns (
<figureCitation id="13652A313A70BA2FFCDDFFD3FC4D8BAD" box="[836,907,152,178]" captionStart="FIGURE 3" captionStartId="6.[151,256,850,874]" captionTargetBox="[466,1121,390,826]" captionTargetId="figure@6.[466,1121,390,826]" captionTargetPageId="6" captionText="FIGURE 3. Type locality of Enigmatochromis lucanusi, Guinea: Foto River, near the town of Fria, north of Conakry. 10 ° 20.074 ’ N, 13 ° 43.431 ’ W. Photo by O. Lucanus." httpUri="https://zenodo.org/record/275093/files/figure.png" pageId="6" pageNumber="47">Fig. 3</figureCitation>
). The river substrate consisted of fine gravel with a few larger rocks and boulders. Dense growths of
<taxonomicName id="4C5E4D373A70BA2FFCDDFFF4FC018BC7" box="[836,967,191,216]" class="Liliopsida" family="Hydrocharitaceae" genus="Vallisneria" kingdom="Plantae" order="Alismatales" pageId="6" pageNumber="47" phylum="Tracheophyta" rank="genus">
<emphasis id="B92AEAA63A70BA2FFCDDFFF4FC018BC7" box="[836,967,191,216]" italics="true" pageId="6" pageNumber="47">Vallisneria</emphasis>
</taxonomicName>
were found where sunlight penetrated through the trees.
</paragraph>
<paragraph id="8BE136B43A70BA2FFF5FFE47FAE78A53" blockId="6.[151,1436,152,332]" pageId="6" pageNumber="47">
<taxonomicName id="4C5E4D373A70BA2FFF5FFE47FDC68A3A" box="[198,512,268,293]" class="Actinopterygii" family="Cichlidae" genus="Enigmatochromis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="lucanusi">
<emphasis id="B92AEAA63A70BA2FFF5FFE47FDC68A3A" box="[198,512,268,293]" italics="true" pageId="6" pageNumber="47">Enigmatochromis lucanusi</emphasis>
</taxonomicName>
occured syntopically at this site with
<taxonomicName id="4C5E4D373A70BA2FFC25FE47FB088A3A" box="[956,1230,268,293]" class="Actinopterygii" family="Cichlidae" genus="Pelvicachromis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="humilis">
<emphasis id="B92AEAA63A70BA2FFC25FE47FB088A3A" box="[956,1230,268,293]" italics="true" pageId="6" pageNumber="47">Pelvicachromis humilis</emphasis>
</taxonomicName>
.
<emphasis id="B92AEAA63A70BA2FFF5FFE79FE8D8A53" bold="true" box="[198,331,306,332]" pageId="6" pageNumber="47">Etymology</emphasis>
. The species name is dedicated to the collector, aquarist and friend Oliver Lucanus.
</paragraph>
</subSubSection>
</treatment>
</document>
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