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<document id="BCDEFAEE304AF083E25B6F50807F790B" ID-DOI="10.11646/phytotaxa.99.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="5072471" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625523958576" checkinUser="felipe" docAuthor="Sang, Julia, Kiew, Ruth &amp; Geri, Connie" docDate="2013" docId="A7507658EA7FFF84C5AB39093EDDF95F" docLanguage="en" docName="Phytotaxa.99.1.1-34.pdf" docOrigin="Phytotaxa 99 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.99.1.1" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Begonia rhodoneura S. Julia 2013, spec. nov." docType="treatment" docVersion="3" lastPageNumber="24" masterDocId="5B690E20EA69FF93C53C38513C3EFF8A" masterDocTitle="Revision of Begonia (Begoniaceae) from the Melinau Limestone in Gunung Mulu National Park and Gunung Buda National Park, Sarawak, Borneo, including thirteen new species" masterLastPageNumber="34" masterPageNumber="1" pageNumber="23" updateTime="1698991887890" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="B204144711C08A6CD4338DD13F312B20" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="218DA324A5FFE1F2507F6456183EABC9">
<mods:title id="EA6ADDB2116866CAC1473F4B602CE33D">Revision of Begonia (Begoniaceae) from the Melinau Limestone in Gunung Mulu National Park and Gunung Buda National Park, Sarawak, Borneo, including thirteen new species</mods:title>
</mods:titleInfo>
<mods:name id="CA9B8EC042BA2DAC1750240D8B545DD3" type="personal">
<mods:role id="08926AB2DE435044FC29FAB080423F2B">
<mods:roleTerm id="5DD00EC3D6D1B1AF72AFC18A5719F542">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DECECFC96E5E910D1605052B04B66DB7">Sang, Julia</mods:namePart>
<mods:affiliation id="D96AF8F5C1C2D378FAFB8370B081B9B7">Botanical Research Centre, SARAWAK FORESTRY, Kuching, Sarawak, Malaysia; e-mail: juliasang @ sarawakforestry. com; conniegeri @ sarawakforestry. com</mods:affiliation>
<mods:nameIdentifier id="C270E949541406F10B0752869931693F" type="email">juliasang@sarawakforestry.com</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="1586D689123C0184778C87F4012E295D">Kiew, Ruth</mods:namePart>
<mods:affiliation id="70F6530574BC6E7EFDA3906E316B873E">Forest Research Institute Malaysia, Kepong, Malaysia; e-mail: ruth @ frim. gov. my</mods:affiliation>
</mods:name>
<mods:name id="578882B50763B8211FAF970306BE0E89" type="personal">
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<mods:roleTerm id="66688550F9CAD5B0A3C396FE30E91E8F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B49B936815E18C436A3298CA29EBD9E6">Geri, Connie</mods:namePart>
<mods:affiliation id="7877D00438860A4920CE1D4A3179E7B5">Botanical Research Centre, SARAWAK FORESTRY, Kuching, Sarawak, Malaysia; e-mail: juliasang @ sarawakforestry. com; conniegeri @ sarawakforestry. com</mods:affiliation>
<mods:nameIdentifier id="AFBBA1671F635F72BBA881F4CB96497C" type="email">juliasang@sarawakforestry.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="3E1D77DDE1AFCB95256F96297A0761E0">text</mods:typeOfResource>
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<mods:title id="07D5BA1EC73792886B14BF528C99551E">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="ECEA2BD03F9B9E9A8B71009F23A24246">
<mods:date id="1EDB30B1CCFB3A98A51434312CC20BB5">2013</mods:date>
<mods:detail id="6330F532EBE48B26ABB17E430AD6F740" type="pubDate">
<mods:number id="0F5C1C286EC43A0E302319DA486D106D">2013-05-09</mods:number>
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<mods:detail id="5A1A0F2C45D30F392039B9534BBEE19B" type="volume">
<mods:number id="29EA37522010DD61FD472540670630E3">99</mods:number>
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<mods:number id="5C400078F328C4698CA41DD92F31D973">1</mods:number>
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<mods:start id="AD9EC31B749439216BB20D0E54A7CB93">1</mods:start>
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<mods:url id="19B2AB36A948068022FFC9E16DFED78B">http://dx.doi.org/10.11646/phytotaxa.99.1.1</mods:url>
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<mods:classification id="B5D3C24F89B9802632F331EDB078ED0C">journal article</mods:classification>
<mods:identifier id="490D5BF8B698C2F285A357D12C67BF12" type="DOI">10.11646/phytotaxa.99.1.1</mods:identifier>
<mods:identifier id="711EB9881F5BCCA99512E06D0BEDEC3A" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="F1F2CCF68CABB5E87B4828B18A1062C0" type="Zenodo-Dep">5072471</mods:identifier>
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<treatment id="A7507658EA7FFF84C5AB39093EDDF95F" LSID="urn:lsid:plazi:treatment:A7507658EA7FFF84C5AB39093EDDF95F" httpUri="http://treatment.plazi.org/id/A7507658EA7FFF84C5AB39093EDDF95F" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<subSubSection id="67E394C5EA7FFF85C5AB39093F1FFEF9" box="[151,801,344,371]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="2F46C74EEA7FFF85C5AB39093F1FFEF9" blockId="22.[151,801,344,371]" box="[151,801,344,371]" pageId="22" pageNumber="23">
<emphasis id="1D8D1B5CEA7FFF85C5AB39093C82FEF8" bold="true" box="[151,188,344,370]" pageId="22" pageNumber="23">10.</emphasis>
<taxonomicName id="E8F9BCCDEA7FFF85C5FF39083E29FEF9" ID-CoL="7SH7K" authority="S.Julia" authorityName="S. Julia" authorityYear="2013" box="[195,535,345,371]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="rhodoneura" status="sp. nov.">
<emphasis id="1D8D1B5CEA7FFF85C5FF39083D80FEF9" bold="true" box="[195,446,345,371]" italics="true" pageId="22" pageNumber="23">Begonia rhodoneura</emphasis>
S.Julia
</taxonomicName>
,
<emphasis id="1D8D1B5CEA7FFF85C71F39083EA8FEF8" box="[547,662,345,370]" italics="true" pageId="22" pageNumber="23">
<taxonomicNameLabel id="06BEA627EA7FFF85C71F39083EA8FEF8" box="[547,662,345,370]" pageId="22" pageNumber="23" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="B7C2DBCBEA7FFF85C79839083F26FEF9" box="[676,792,345,371]" captionStart="FIGURE 13" captionStartId="23.[151,245,1387,1408]" captionTargetBox="[159,1429,157,1369]" captionTargetId="figure-214@23.[151,1436,155,1385]" captionTargetPageId="23" captionText="FIGURE 13. Begonia rhodoneura. A. habit; B. juvenile plant; C. male flowers; D. female flower taken from specimen preserved in spirit; E. fruit taken from specimen preserved in spirit. Photos from SFC 2801 by Julia Sang." figureDoi="http://doi.org/10.5281/zenodo.5072497" httpUri="https://zenodo.org/record/5072497/files/figure.png" pageId="22" pageNumber="23">Figure 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="67E394C5EA7FFF85C5AB39F73DFDFD21" pageId="22" pageNumber="23" type="description">
<paragraph id="2F46C74EEA7FFF85C5AB39F73D51FE4A" blockId="22.[151,367,422,448]" box="[151,367,422,448]" pageId="22" pageNumber="23">
Sect.
<taxonomicName id="E8F9BCCDEA7FFF85C5E439F63D51FE4A" authorityName="A. de Candolle" authorityYear="1859" baseAuthorityName="Klotzsch" baseAuthorityYear="1854" box="[216,367,423,448]" class="Liliopsida" family="Petermanniaceae" genus="Petermannia" kingdom="Plantae" order="Liliales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="genus">
<emphasis id="1D8D1B5CEA7FFF85C5E439F63D51FE4A" box="[216,367,423,448]" italics="true" pageId="22" pageNumber="23">Petermannia</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="2F46C74EEA7FFF85C5AB39A53DFDFD21" blockId="22.[151,1436,500,683]" pageId="22" pageNumber="23">
In habit and leaf shape, this species resembles
<taxonomicName id="E8F9BCCDEA7FFF85C7B739A53F8BFD81" authority="K. G. Pearce" authorityName="K. G. Pearce" authorityYear="2003" box="[651,949,500,523]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="kachak">
<emphasis id="1D8D1B5CEA7FFF85C7B739A53F09FD81" box="[651,823,500,523]" italics="true" pageId="22" pageNumber="23">Begonia kachak</emphasis>
K.G.Pearce
</taxonomicName>
from Niah National Park but it is different in its more or less sessile inflorescence with
<specimenCount id="39FF0CC7EA7FFF85C7BF3A453EC6FDA1" box="[643,760,532,555]" pageId="22" pageNumber="23" type="female">one female</specimenCount>
flower in the leaf axil that is distally much branched with many small white male flowers. The male tepals measure 4.5–5 ×
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compared with the longer inflorescence of
<taxonomicName id="E8F9BCCDEA7FFF85C0B93A653D2CFDE1" authorityName="K. G. Pearce" authorityYear="2003" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="kachak">
<emphasis id="1D8D1B5CEA7FFF85C0B93A653D2CFDE1" italics="true" pageId="22" pageNumber="23">B. kachak</emphasis>
</taxonomicName>
that is up to
<quantity id="E8016AABEA7FFF85C4A33A053DDCFDE1" box="[415,482,596,619]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" pageId="22" pageNumber="23" unit="cm" value="29.0">29 cm</quantity>
long of which the peduncle is
<quantity id="E8016AABEA7FFF85C6103A053F51FDE1" box="[812,879,596,619]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="22" pageNumber="23" unit="cm" value="14.0">14 cm</quantity>
long. The female flowers are in pairs and the tepals measure 7 ×
<quantity id="E8016AABEA7FFF85C4733A253D9DFD01" box="[335,419,628,651]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="22" pageNumber="23" unit="mm" value="5.5">5.5 mm</quantity>
. In addition, its fruits are much narrower (19–)22(–30) × 10–13(–17) mm compared with ca. 19 ×
<quantity id="E8016AABEA7FFF85C5E03AC53D18FD21" box="[220,294,660,683]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="22" pageNumber="23" unit="mm" value="16.0">16 mm</quantity>
for
<taxonomicName id="E8F9BCCDEA7FFF85C46F3AC53D82FD21" authorityName="K. G. Pearce" authorityYear="2003" box="[339,444,660,683]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="kachak">
<emphasis id="1D8D1B5CEA7FFF85C46F3AC53D82FD21" box="[339,444,660,683]" italics="true" pageId="22" pageNumber="23">B. kachak</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="67E394C5EA7FFF85C5AA3A823D1DF92D" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="2F46C74EEA7FFF85C5AA3A82384CFC81" blockId="22.[150,1436,723,779]" pageId="22" pageNumber="23">
<emphasis id="1D8D1B5CEA7FFF85C5AA3A823CE9FD61" bold="true" box="[150,215,723,747]" pageId="22" pageNumber="23">Type:</emphasis>
—
<materialsCitation id="9F91CD13EA7FFF85C5CF3A853850FC81" collectingDate="2012-04-15" collectionCode="SAR, E, K, KEP, L, SAN, SING" country="Malaysia" county="Marudi District" location="Gunung Benarat" municipality="Gunung Mulu National Park" pageId="22" pageNumber="23" specimenCode="SFC 2801" specimenCount="1" stateProvince="Sarawak" typeStatus="holotype">
<collectingCountry id="57EE87DEEA7FFF85C5CF3A853DBEFD61" box="[243,384,724,747]" name="Malaysia" pageId="22" pageNumber="23">MALAYSIA</collectingCountry>
. Borneo.
<collectingRegion id="ED3D09ACEA7FFF85C4DA3A853E7BFD61" box="[486,581,724,747]" country="Malaysia" name="Sarawak" pageId="22" pageNumber="23">Sarawak</collectingRegion>
.
<collectingCounty id="C627BFC2EA7FFF85C7733A853EC7FD61" box="[591,761,724,747]" pageId="22" pageNumber="23">Marudi District</collectingCounty>
:
<collectingMunicipality id="CF225D34EA7FFF85C63F3A853809FD61" box="[771,1079,724,747]" pageId="22" pageNumber="23">Gunung Mulu National Park</collectingMunicipality>
,
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,
<date id="5B47E18EEA7FFF85C1C33A8539A6FD61" box="[1279,1432,724,747]" pageId="22" pageNumber="23" value="2012-04-15">
<collectingDate id="4B031866EA7FFF85C1C33A8539A6FD61" box="[1279,1432,724,747]" pageId="22" pageNumber="23" value="2012-04-15">15 April 2012</collectingDate>
</date>
,
<emphasis id="1D8D1B5CEA7FFF85C5FB3AA53D94FC81" box="[199,426,756,779]" italics="true" pageId="22" pageNumber="23">
<location id="2A269195EA7FFF85C5FB3AA53CC2FC81" LSID="urn:lsid:plazi:treatment:A7507658EA7FFF84C5AB39093EDDF95F:2A269195EA7FFF85C5FB3AA53CC2FC81" box="[199,252,756,779]" country="Malaysia" county="Marudi District" municipality="Gunung Mulu National Park" name="Julia" pageId="22" pageNumber="23" stateProvince="Sarawak">Julia</location>
et al.
<specimenCode id="7F5F6F35EA7FFF85C4023AA53D94FC81" box="[318,426,756,779]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="22" pageNumber="23">SFC 2801</specimenCode>
</emphasis>
(
<typeStatus id="F04279ECEA7FFF85C48B3AA53E28FC81" box="[439,534,756,779]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
<collectionCode id="49E85F8BEA7FFF85C7213AA53E67FC81" box="[541,601,756,779]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="22" pageNumber="23" type="Herbarium">SAR</collectionCode>
!; isotypes
<collectionCode id="49E85F8BEA7FFF85C7FA3AA53EE2FC81" box="[710,732,756,779]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="22" pageNumber="23" type="Herbarium">E</collectionCode>
!,
<collectionCode id="49E85F8BEA7FFF85C7D03AA53F38FC81" box="[748,774,756,779]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="22" pageNumber="23" type="Herbarium">K</collectionCode>
!,
<collectionCode id="49E85F8BEA7FFF85C62A3AA53F6EFC81" box="[790,848,756,779]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="22" pageNumber="23" type="Herbarium">KEP</collectionCode>
!,
<collectionCode id="49E85F8BEA7FFF85C6623AA53F4AFC81" box="[862,884,756,779]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="22" pageNumber="23" type="Herbarium">L</collectionCode>
!,
<collectionCode id="49E85F8BEA7FFF85C6B93AA53FFCFC81" box="[901,962,756,779]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="22" pageNumber="23" type="Herbarium">SAN</collectionCode>
!,
<collectionCode id="49E85F8BEA7FFF85C6ED3AA53832FC81" box="[977,1036,756,779]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="22" pageNumber="23" type="Herbarium">SAR</collectionCode>
!,
<collectionCode id="49E85F8BEA7FFF85C1263AA5385CFC81" box="[1050,1122,756,779]" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="22" pageNumber="23" type="Herbarium">SING</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="2F46C74EEA7FFF85C5AB3B6B3D1DF92D" blockId="22.[151,1436,826,1703]" pageId="22" pageNumber="23">
Scrambling horizontally to
<quantity id="E8016AABEA7FFF85C4D43B6B3E10FCDE" box="[488,558,826,852]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="23" unit="m" value="1.5">1.5 m</quantity>
tall, bushy and branching, sometimes creeping on rock surfaces or tree trunks. Stems reddish brown, slender, woody, minutely ferrugineous, slightly swollen at nodes, reddish, internodes (2–)
<quantity id="E8016AABEA7FFF85C4743BD93DF0FC28" box="[328,462,904,930]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.5" metricValueMin="2.5" pageId="22" pageNumber="23" unit="cm" value="4.0" valueMax="5.5" valueMin="2.5">2.5–5.5 cm</quantity>
long,
<quantity id="E8016AABEA7FFF85C7263BD93E45FC28" box="[538,635,904,930]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="22" pageNumber="23" unit="mm" value="7.0" valueMax="9.0" valueMin="5.0">5–9 mm</quantity>
diameter. Stipules brownish red, lanceolate, ca. 14 ×
<quantity id="E8016AABEA7FFF85C1C93BD93903FC28" box="[1269,1341,904,930]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="23" unit="mm" value="3.0">3 mm</quantity>
, margin entire, apex setose, caducous and leaving a circular scar. Leaves alternate, distant or arranged close together. Petioles minutely ferrugineous and dark red,
<quantity id="E8016AABEA7FFF85C78F3B843F1BFC65" box="[691,805,981,1007]" metricMagnitude="-2" metricUnit="m" metricValue="8.25" metricValueMax="9.5" metricValueMin="7.0" pageId="22" pageNumber="23" unit="cm" value="8.25" valueMax="9.5" valueMin="7.0">7–9.5 cm</quantity>
, slender. Laminas dark green, paler beneath, slightly velvety above, fleshy when fresh, papery when dried, very oblique, broadly ovate, 6–14.5 ×
<quantity id="E8016AABEA7FFF85C02F3BAD39A9FB9C" box="[1299,1431,1020,1046]" metricMagnitude="-2" metricUnit="m" metricValue="9.75" metricValueMax="13.5" metricValueMin="6.0" pageId="22" pageNumber="23" unit="cm" value="9.75" valueMax="13.5" valueMin="6.0">6–13.5 cm</quantity>
, asymmetric, broad side
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, base cordate overlapping, basal lobes rounded,
<quantity id="E8016AABEA7FFF85C1603C7338F3FBB6" box="[1116,1229,1058,1084]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="22" pageNumber="23" unit="cm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 cm</quantity>
, margin minutely toothed, apex acuminate. Venation palmate, main veins glabrous beneath, 2 pairs at the base, with 2 pairs along the midrib and 2 veins in basal lobe, branching twice before reaching the margin, impressed above, slightly prominent beneath, veins red below, dark red above. Inflorescences axillary, red, erect and shorter than petioles, protogynous, with
<specimenCount id="39FF0CC7EA7FFF85C7203CEC3EA0FB5D" box="[540,670,1213,1239]" pageId="22" pageNumber="23" type="female">one female</specimenCount>
flower directly in the leaf axil with the male flowers above on a panicle with a total length (3.5–)7(–11.5) cm long. Bracts elliptic, pale green, ca. 12 ×
<quantity id="E8016AABEA7FFF85C19E3CB538D7FB74" box="[1186,1257,1252,1278]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="mm" value="5.0">5 mm</quantity>
, margin entire, caducous. Uppermost bracts below the male flower greenish, ovate, ca. 10
<quantity id="E8016AABEA7FFF85C11E3D5B3854FAAE" box="[1058,1130,1290,1316]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="22" pageNumber="23" unit="mm" value="8.0">8 mm</quantity>
, margin entire, caducous. Male flower: pedicel red ca.
<quantity id="E8016AABEA7FFF85C4DB3D603E11FAC1" box="[487,559,1329,1355]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="22" pageNumber="23" unit="mm" value="7.0">7 mm</quantity>
; tepals 2, red at the base becoming white towards the tip, glabrous outside, rounded, 4.5–5 ×
<quantity id="E8016AABEA7FFF85C4583D093D95FAF8" box="[356,427,1368,1394]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="mm" value="5.0">5 mm</quantity>
, margin entire, apex rounded; stamens more than 40, cluster globose, stalk
<quantity id="E8016AABEA7FFF85C02A3D093967FAF8" box="[1302,1369,1368,1394]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="23" unit="mm" value="1.0">1 mm</quantity>
long; filament ca.
<quantity id="E8016AABEA7FFF85C40E3D2F3D46FA12" box="[306,376,1406,1432]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="23" unit="mm" value="1.0">1 mm</quantity>
long, anthers pale yellow, obovate, ca. 1 ×
<quantity id="E8016AABEA7FFF85C6AB3D2F3FC6FA12" box="[919,1016,1406,1432]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="mm" value="0.5">0.5 mm</quantity>
, apex emarginate. Female flower: pedicel
<quantity id="E8016AABEA7FFF85C5CE3DF43D79FA35" box="[242,327,1445,1471]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="23" unit="mm" value="10.0">10 mm</quantity>
, pink, with short hairs; ovary greenish, oblong, ca. 12 ×
<quantity id="E8016AABEA7FFF85C6EA3DF43823FA35" box="[982,1053,1445,1471]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="mm" value="5.0">5 mm</quantity>
, minutely ferrugineous, wings 3, equal, locules 3, placental branches 2 per locule; tepals 5, greenish, elliptic, ca. 10 ×
<quantity id="E8016AABEA7FFF85C1B03D9D38EAFA6C" box="[1164,1236,1484,1510]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="22" pageNumber="23" unit="mm" value="7.0">7 mm</quantity>
, margin toothed, apex acute; styles ca.
<quantity id="E8016AABEA7FFF85C4943DA33DD0F986" box="[424,494,1522,1548]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="23" unit="mm" value="2.0">2 mm</quantity>
long, free to base and bifurcating with greenish papillose stigma forming a continuous twisted band. Fruits pendent; pedicel
<quantity id="E8016AABEA7FFF85C7EA3E483F6AF9B9" box="[726,852,1561,1587]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="22" pageNumber="23" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
long, stiff and recurved; capsule oblong, 19–22(– 30) × 10–13(–17) mm, glabrous, locules 3, wing unequal, narrowed proximally,
<quantity id="E8016AABEA7FFF85C1743E113897F9D0" box="[1096,1193,1600,1626]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="22" pageNumber="23" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
wide, stiffly papery, dehiscing between the wings and locules. Seeds barrel-shaped, ca.
<quantity id="E8016AABEA7FFF85C69C3E373FC7F90A" box="[928,1017,1638,1664]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="mm" value="0.5">0.5 mm</quantity>
long, collar cells more than half the seed length.
</paragraph>
</subSubSection>
<subSubSection id="67E394C5EA7FFF85C5FA3E8B3D16F805" pageId="22" pageNumber="23" type="distribution">
<paragraph id="2F46C74EEA7FFF85C5FA3E8B3E43F8C8" blockId="22.[151,1436,1754,2012]" pageId="22" pageNumber="23">
<emphasis id="1D8D1B5CEA7FFF85C5FA3E8B3D57F97E" bold="true" box="[198,361,1754,1780]" pageId="22" pageNumber="23">Distribution:</emphasis>
—
<collectingCountry id="57EE87DEEA7FFF85C4BB3E8B3E1CF97E" box="[391,546,1754,1780]" name="Malaysia" pageId="22" pageNumber="23">MALAYSIA</collectingCountry>
. Borneo.
<collectingRegion id="ED3D09ACEA7FFF85C7983E8B3F33F97E" box="[676,781,1754,1780]" country="Malaysia" name="Sarawak" pageId="22" pageNumber="23">Sarawak</collectingRegion>
. Marudi District: Gunung Mulu National Park and Gunung Buda National Park. Endemic in the Melinau limestone. Widespread from Lobang Cina, Gunung Benarat, Bukit Agung and Gunung Buda.
</paragraph>
<paragraph id="2F46C74EEA7FFF85C5FA3F1F3D16F805" blockId="22.[151,1436,1754,2012]" pageId="22" pageNumber="23">
<emphasis id="1D8D1B5CEA7FFF85C5FA3F1F3D0EF8E2" bold="true" box="[198,304,1870,1896]" pageId="22" pageNumber="23">Habitat:</emphasis>
—Common. In lowland limestone forest on limestone boulders or mossy rock faces, at
<quantity id="E8016AABEA7FFF85C00A3F1F3C90F805" metricMagnitude="2" metricUnit="m" metricValue="1.685" metricValueMax="2.35" metricValueMin="1.02" pageId="22" pageNumber="23" unit="m" value="168.5" valueMax="235.0" valueMin="102.0">102–235 m</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="67E394C5EA7FFF84C5FA3FCA3EDDF95F" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="etymology">
<paragraph id="2F46C74EEA7FFF85C5FA3FCA3CD0F856" blockId="22.[151,1436,1754,2012]" pageId="22" pageNumber="23">
<emphasis id="1D8D1B5CEA7FFF85C5FA3FCA3D69F83F" bold="true" box="[198,343,1947,1973]" pageId="22" pageNumber="23">Etymology:</emphasis>
—It takes its name from its conspicuous fine dark red veins that contrast with the bright green lamina.
</paragraph>
<caption id="7B8697C6EA7EFF84C5AB3D3A382CFA2B" ID-DOI="http://doi.org/10.5281/zenodo.5072497" ID-Zenodo-Dep="5072497" httpUri="https://zenodo.org/record/5072497/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,245,1387,1408]" targetBox="[159,1429,157,1369]" targetPageId="23">
<paragraph id="2F46C74EEA7EFF84C5AB3D3A382CFA2B" blockId="23.[151,1436,1387,1441]" pageId="23" pageNumber="24">
<emphasis id="1D8D1B5CEA7EFF84C5AB3D3A3D27FA0A" bold="true" box="[151,281,1387,1408]" pageId="23" pageNumber="24">FIGURE 13.</emphasis>
<taxonomicName id="E8F9BCCDEA7EFF84C41C3D3D3DD6FA0B" authorityName="S. Julia" authorityYear="2013" box="[288,488,1388,1409]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="rhodoneura">
<emphasis id="1D8D1B5CEA7EFF84C41C3D3D3DD6FA0B" box="[288,488,1388,1409]" italics="true" pageId="23" pageNumber="24">Begonia rhodoneura</emphasis>
</taxonomicName>
. A. habit; B. juvenile plant; C. male flowers; D. female flower taken from specimen preserved in spirit; E. fruit taken from specimen preserved in spirit. Photos from SFC 2801 by Julia Sang.
</paragraph>
</caption>
<paragraph id="2F46C74EEA7EFF84C5FA3D823930F902" blockId="23.[151,1436,1491,1749]" pageId="23" pageNumber="24">
<materialsCitation id="9F91CD13EA7EFF84C5FA3D8238D5F9EB" accessionNumber="S 49472, S 49413" collectionCode="SAN, SAR, SING, K, E, KEP, S, L" collectorName="Lobang Cina &amp; Abang Mohtar &amp; Othman &amp; Anderson &amp; Keng" country="Malaysia" county="Marudi District" location="Julia" municipality="Gunung Mulu National Park - Bukit Agung" pageId="23" pageNumber="24" specimenCode="SFC 2813, SFC 2822, SFC 2833, S 4452" specimenCount="2" stateProvince="Sarawak" typeStatus="paratype">
<emphasis id="1D8D1B5CEA7EFF84C5FA3D823F2AFA67" bold="true" box="[198,788,1491,1517]" pageId="23" pageNumber="24">
Additional specimens examined (
<typeStatus id="F04279ECEA7EFF84C7B83D823F39FA67" box="[644,775,1491,1517]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
):
</emphasis>
—
<collectingCountry id="57EE87DEEA7EFF84C60E3D823FF3FA67" box="[818,973,1491,1517]" name="Malaysia" pageId="23" pageNumber="24">MALAYSIA</collectingCountry>
. Borneo.
<collectingRegion id="ED3D09ACEA7EFF84C1683D823883FA67" box="[1108,1213,1491,1517]" country="Malaysia" name="Sarawak" pageId="23" pageNumber="24">Sarawak</collectingRegion>
.
<collectingCounty id="C627BFC2EA7EFF84C1EF3D8239A6FA67" box="[1235,1432,1491,1517]" pageId="23" pageNumber="24">Marudi District</collectingCounty>
:
<collectingMunicipality id="CF225D34EA7EFF84C5AB3DAB3EA4F99E" box="[151,666,1530,1556]" pageId="23" pageNumber="24">Gunung Mulu National Park –Bukit Agung</collectingMunicipality>
,
<emphasis id="1D8D1B5CEA7EFF84C79A3DAB3F9DF999" box="[678,931,1530,1555]" italics="true" pageId="23" pageNumber="24">
<location id="2A269195EA7EFF84C79A3DAB3EDEF999" LSID="urn:lsid:plazi:treatment:A7507658EA7FFF84C5AB39093EDDF95F:2A269195EA7EFF84C79A3DAB3EDEF999" box="[678,736,1530,1555]" country="Malaysia" county="Marudi District" municipality="Gunung Mulu National Park - Bukit Agung" name="Julia" pageId="23" pageNumber="24" stateProvince="Sarawak">Julia</location>
et al.
<specimenCode id="7F5F6F35EA7EFF84C6103DAB3F9DF999" box="[812,931,1530,1555]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="23" pageNumber="24">SFC 2813</specimenCode>
</emphasis>
(
<collectionCode id="49E85F8BEA7EFF84C68F3DAB3FCCF99E" box="[947,1010,1530,1556]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="23" pageNumber="24" type="Herbarium">SAN</collectionCode>
,
<collectionCode id="49E85F8BEA7EFF84C6C23DAB3804F99E" box="[1022,1082,1530,1556]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="23" pageNumber="24" type="Herbarium">SAR</collectionCode>
,
<collectionCode id="49E85F8BEA7EFF84C17B3DAB38AEF99E" box="[1095,1168,1530,1556]" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="23" pageNumber="24" type="Herbarium">SING</collectionCode>
),
<emphasis id="1D8D1B5CEA7EFF84C1983DAB3924F999" box="[1188,1306,1530,1555]" italics="true" pageId="23" pageNumber="24">
<specimenCode id="7F5F6F35EA7EFF84C1983DAB3924F999" box="[1188,1306,1530,1555]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="23" pageNumber="24">SFC 2822</specimenCode>
</emphasis>
(
<collectionCode id="49E85F8BEA7EFF84C0103DAB397AF99E" box="[1324,1348,1530,1556]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="23" pageNumber="24" type="Herbarium">K</collectionCode>
,
<collectionCode id="49E85F8BEA7EFF84C06D3DAB39B1F99E" box="[1361,1423,1530,1556]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="23" pageNumber="24" type="Herbarium">SAR</collectionCode>
);
<location id="2A269195EA7EFF84C5AB3E703D61F9B1" LSID="urn:lsid:plazi:treatment:A7507658EA7FFF84C5AB39093EDDF95F:2A269195EA7EFF84C5AB3E703D61F9B1" box="[151,351,1569,1595]" country="Malaysia" county="Marudi District" municipality="Gunung Mulu National Park - Bukit Agung" name="Gunung Benarat" pageId="23" pageNumber="24" stateProvince="Sarawak">Gunung Benarat</location>
,
<emphasis id="1D8D1B5CEA7EFF84C4503E703E56F9B0" box="[364,616,1569,1594]" italics="true" pageId="23" pageNumber="24">
<location id="2A269195EA7EFF84C4503E703D9DF9B0" LSID="urn:lsid:plazi:treatment:A7507658EA7FFF84C5AB39093EDDF95F:2A269195EA7EFF84C4503E703D9DF9B0" box="[364,419,1569,1594]" country="Malaysia" county="Marudi District" municipality="Gunung Mulu National Park - Bukit Agung" name="Beai" pageId="23" pageNumber="24" stateProvince="Sarawak">Beai</location>
et al.
<specimenCode id="7F5F6F35EA7EFF84C4CC3E703E56F9B0" box="[496,616,1569,1594]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="23" pageNumber="24">SFC 2833</specimenCode>
</emphasis>
(
<collectionCode id="49E85F8BEA7EFF84C7463E703EB1F9B1" box="[634,655,1569,1595]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="23" pageNumber="24" type="Herbarium">E</collectionCode>
,
<collectionCode id="49E85F8BEA7EFF84C7A13E703E8BF9B1" box="[669,693,1569,1595]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="23" pageNumber="24" type="Herbarium">K</collectionCode>
,
<collectionCode id="49E85F8BEA7EFF84C7FE3E703EC4F9B1" box="[706,762,1569,1595]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="23" pageNumber="24" type="Herbarium">KEP</collectionCode>
,
<collectionCode id="49E85F8BEA7EFF84C63B3E703F7BF9B1" box="[775,837,1569,1595]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="23" pageNumber="24" type="Herbarium">SAR</collectionCode>
),
<emphasis id="1D8D1B5CEA7EFF84C6673E70388FF9B0" box="[859,1201,1569,1594]" italics="true" pageId="23" pageNumber="24">
<location id="2A269195EA7EFF84C6673E703834F9B0" LSID="urn:lsid:plazi:treatment:A7507658EA7FFF84C5AB39093EDDF95F:2A269195EA7EFF84C6673E703834F9B0" box="[859,1034,1569,1594]" country="Malaysia" county="Marudi District" municipality="Gunung Mulu National Park - Bukit Agung" name="Abang Mohtar" pageId="23" pageNumber="24" stateProvince="Sarawak">Abang Mohtar</location>
et al
<accessionNumber id="30AA5AADEA7EFF84C16C3E70388FF9B0" box="[1104,1201,1569,1594]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S49472" pageId="23" pageNumber="24" type="EnaNcbi">
<collectionCode id="49E85F8BEA7EFF84C16C3E703860F9B0" box="[1104,1118,1569,1594]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="23" pageNumber="24" type="Herbarium">S</collectionCode>
49472
</accessionNumber>
</emphasis>
(
<collectionCode id="49E85F8BEA7EFF84C1FF3E7038E2F9B1" box="[1219,1244,1569,1595]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="23" pageNumber="24" type="Herbarium">K</collectionCode>
,
<collectionCode id="49E85F8BEA7EFF84C1D53E70391FF9B1" box="[1257,1313,1569,1595]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="23" pageNumber="24" type="Herbarium">KEP</collectionCode>
,
<collectionCode id="49E85F8BEA7EFF84C0123E70397DF9B1" box="[1326,1347,1569,1595]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="23" pageNumber="24" type="Herbarium">L</collectionCode>
,
<collectionCode id="49E85F8BEA7EFF84C06C3E7039B1F9B1" box="[1360,1423,1569,1595]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="23" pageNumber="24" type="Herbarium">SAR</collectionCode>
);
<collectorName id="820CA298EA7EFF84C5AB3E163D0DF9EB" box="[151,307,1607,1633]" pageId="23" pageNumber="24">Lobang Cina</collectorName>
,
<emphasis id="1D8D1B5CEA7EFF84C47C3E163EE4F9EA" box="[320,730,1607,1632]" italics="true" pageId="23" pageNumber="24">
<collectorName id="820CA298EA7EFF84C47C3E163DD2F9EA" box="[320,492,1607,1632]" pageId="23" pageNumber="24">Abang Mohtar</collectorName>
&amp;
<collectorName id="820CA298EA7EFF84C7283E163E4DF9EA" box="[532,627,1607,1632]" pageId="23" pageNumber="24">Othman</collectorName>
<accessionNumber id="30AA5AADEA7EFF84C7463E163EE4F9EA" box="[634,730,1607,1632]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S49413" pageId="23" pageNumber="24" type="EnaNcbi">
<collectionCode id="49E85F8BEA7EFF84C7463E163EB7F9EA" box="[634,649,1607,1632]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="23" pageNumber="24" type="Herbarium">S</collectionCode>
49413
</accessionNumber>
</emphasis>
(
<collectionCode id="49E85F8BEA7EFF84C7D73E163F17F9EB" box="[747,809,1607,1633]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="23" pageNumber="24" type="Herbarium">SAR</collectionCode>
),
<emphasis id="1D8D1B5CEA7EFF84C6033E163850F9EA" box="[831,1134,1607,1632]" italics="true" pageId="23" pageNumber="24">
<collectorName id="820CA298EA7EFF84C6033E163F91F9EA" box="[831,943,1607,1632]" pageId="23" pageNumber="24">Anderson</collectorName>
&amp;
<collectorName id="820CA298EA7EFF84C6EA3E16382AF9EA" box="[982,1044,1607,1632]" pageId="23" pageNumber="24">Keng</collectorName>
<specimenCode id="7F5F6F35EA7EFF84C1213E163850F9EA" box="[1053,1134,1607,1632]" collectionCode="S" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="23" pageNumber="24" type="Herbarium">S 4452</specimenCode>
</emphasis>
(
<collectionCode id="49E85F8BEA7EFF84C1433E1638A9F9EB" box="[1151,1175,1607,1633]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="23" pageNumber="24" type="Herbarium">K</collectionCode>
,
<collectionCode id="49E85F8BEA7EFF84C1993E1638DDF9EB" box="[1189,1251,1607,1633]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="23" pageNumber="24" type="Herbarium">SAR</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="9F91CD13EA7EFF84C1CB3E163937F902" collectionCode="KEP, SAR, SING, E" country="Malaysia" location="Julia" municipality="Gunung Buda National Park - Gunung Buda" pageId="23" pageNumber="24" specimenCode="SFC 2824, B 8342" specimenCount="1">
<collectingMunicipality id="CF225D34EA7EFF84C1CB3E163DC3F902" pageId="23" pageNumber="24">Gunung Buda National Park – Gunung Buda</collectingMunicipality>
,
<emphasis id="1D8D1B5CEA7EFF84C7343E3F3F3CF90D" box="[520,770,1646,1671]" italics="true" pageId="23" pageNumber="24">
<location id="2A269195EA7EFF84C7343E3F3E7CF90D" LSID="urn:lsid:plazi:treatment:A7507658EA7FFF84C5AB39093EDDF95F:2A269195EA7EFF84C7343E3F3E7CF90D" box="[520,578,1646,1671]" country="Malaysia" municipality="Gunung Buda National Park - Gunung Buda" name="Julia" pageId="23" pageNumber="24">Julia</location>
et al.
<specimenCode id="7F5F6F35EA7EFF84C7B03E3F3F3CF90D" box="[652,770,1646,1671]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="23" pageNumber="24">SFC 2824</specimenCode>
</emphasis>
(
<collectionCode id="49E85F8BEA7EFF84C62D3E3F3F74F902" box="[785,842,1646,1672]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="23" pageNumber="24" type="Herbarium">KEP</collectionCode>
,
<collectionCode id="49E85F8BEA7EFF84C6693E3F3FAFF902" box="[853,913,1646,1672]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="23" pageNumber="24" type="Herbarium">SAR</collectionCode>
,
<collectionCode id="49E85F8BEA7EFF84C6A13E3F3FD8F902" box="[925,998,1646,1672]" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="23" pageNumber="24" type="Herbarium">SING</collectionCode>
),
<emphasis id="1D8D1B5CEA7EFF84C6C63E3F38AFF90D" box="[1018,1169,1646,1671]" italics="true" pageId="23" pageNumber="24">
Burtt
<specimenCode id="7F5F6F35EA7EFF84C1023E3F38AFF90D" box="[1086,1169,1646,1671]" collectionCode="B" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="23" pageNumber="24" type="Herbarium">B 8342</specimenCode>
</emphasis>
(
<collectionCode id="49E85F8BEA7EFF84C19E3E3F3889F902" box="[1186,1207,1646,1672]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="23" pageNumber="24" type="Herbarium">E</collectionCode>
,
<collectionCode id="49E85F8BEA7EFF84C1FF3E3F393FF902" box="[1219,1281,1646,1672]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="23" pageNumber="24" type="Herbarium">SAR</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="2F46C74EEA7EFF84C5FA3EC53EDDF95F" blockId="23.[151,1436,1491,1749]" pageId="23" pageNumber="24">
<emphasis id="1D8D1B5CEA7EFF84C5FA3EC53D2BF924" bold="true" box="[198,277,1684,1710]" pageId="23" pageNumber="24">Notes:</emphasis>
—The most common begonia on limestone outcrops and isolated boulders at Lobang
<collectingCountry id="57EE87DEEA7EFF84C0613EC539A9F924" box="[1373,1431,1684,1710]" name="China" pageId="23" pageNumber="24">Cina</collectingCountry>
, Gunung Benarat, Gunung Buda and Bukit Agung.
</paragraph>
</subSubSection>
</treatment>
</document>
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