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<document id="6B97958BBD2D2C615136F6EA7B816468" ID-DOI="10.11646/phytotaxa.99.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="5072471" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625523958576" checkinUser="felipe" docAuthor="Sang, Julia, Kiew, Ruth &amp; Geri, Connie" docDate="2013" docId="A7507658EA6CFF94C5AB3E733D1AFC1A" docLanguage="en" docName="Phytotaxa.99.1.1-34.pdf" docOrigin="Phytotaxa 99 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.99.1.1" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Begonia benaratensis S. Julia 2013, spec. nov." docType="treatment" docVersion="3" lastPageNumber="8" masterDocId="5B690E20EA69FF93C53C38513C3EFF8A" masterDocTitle="Revision of Begonia (Begoniaceae) from the Melinau Limestone in Gunung Mulu National Park and Gunung Buda National Park, Sarawak, Borneo, including thirteen new species" masterLastPageNumber="34" masterPageNumber="1" pageNumber="6" updateTime="1698991887890" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="4A8E98158CAB16C963E302412CB57FC7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9A98698E7D690B3922A88810E0A3A372">
<mods:title id="7F5A61550C2390327B251C08ADE76D45">Revision of Begonia (Begoniaceae) from the Melinau Limestone in Gunung Mulu National Park and Gunung Buda National Park, Sarawak, Borneo, including thirteen new species</mods:title>
</mods:titleInfo>
<mods:name id="6B9D254B3FB2D1056E56987A03D8EB41" type="personal">
<mods:role id="AA4DDC8286D16D46118FA02BAD186851">
<mods:roleTerm id="8828CEA0A4FD00CED2C0D7BDB5A81E9A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D02B3A3F1882ED4288C7C304E62E977B">Sang, Julia</mods:namePart>
<mods:affiliation id="C41372C8CD263E8783999CEE437B95B7">Botanical Research Centre, SARAWAK FORESTRY, Kuching, Sarawak, Malaysia; e-mail: juliasang @ sarawakforestry. com; conniegeri @ sarawakforestry. com</mods:affiliation>
<mods:nameIdentifier id="D96D24823FEF9BABA38CF2EB2678C272" type="email">juliasang@sarawakforestry.com</mods:nameIdentifier>
</mods:name>
<mods:name id="4B04F3ED71F516197A1146DC0DF9F9FA" type="personal">
<mods:role id="FDB78B90294E5125999B7D0A413D018E">
<mods:roleTerm id="4F0C91D50872EC43859065B0B6FCA393">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="22F1EEA0F5B15288267E440A7A58ABE6">Kiew, Ruth</mods:namePart>
<mods:affiliation id="ED8C3EB52592122D41E6CEF7F417023F">Forest Research Institute Malaysia, Kepong, Malaysia; e-mail: ruth @ frim. gov. my</mods:affiliation>
</mods:name>
<mods:name id="851E5120DC3CA741385EC9AE13E82CE1" type="personal">
<mods:role id="3F39112BA33734CA37EAFD0652B3D9F5">
<mods:roleTerm id="3BE44DA1422DFFEF23B60C4FEC1AAA53">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C865645B427EA0FAAD985ACB7AC15CA8">Geri, Connie</mods:namePart>
<mods:affiliation id="72CAF76526BAC647810EF93757FEA41B">Botanical Research Centre, SARAWAK FORESTRY, Kuching, Sarawak, Malaysia; e-mail: juliasang @ sarawakforestry. com; conniegeri @ sarawakforestry. com</mods:affiliation>
<mods:nameIdentifier id="BE73772DC86B156FEA975CFABCFA6A04" type="email">juliasang@sarawakforestry.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="69267B8E7FBE2F41199A74E7A878F640">text</mods:typeOfResource>
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<mods:titleInfo id="8EC2B2F86443687F87D63A3D0641E69B">
<mods:title id="994647A4DBF21EDD13F8CF3A2D6618A4">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="523415B021D73A037112118191FC26FF">
<mods:date id="5C2997283D81C02F8346CBCF2F77A3D6">2013</mods:date>
<mods:detail id="75ECC11499BCB51F7CF52051F562F4E7" type="pubDate">
<mods:number id="777E12880F3916A64803E5F971022726">2013-05-09</mods:number>
</mods:detail>
<mods:detail id="00F3819682E6C9AF6FF15375D27DBFA6" type="volume">
<mods:number id="4C09C82D3ED7DE63003C57E1C7E74BA8">99</mods:number>
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<mods:detail id="205E111C6DCFDA990DAB2398BDFDFE1E" type="issue">
<mods:number id="21A90BBE4ACCC9D80C738DD493887CF8">1</mods:number>
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<mods:start id="0D3A6B52632C70088F57792FC81F2F1B">1</mods:start>
<mods:end id="54A0D718564339F182219974238EAE6F">34</mods:end>
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<mods:location id="53AAABDAECF6908E09EEB8B0AD9B925A">
<mods:url id="FCD6CB47747F6F63653CA2D1302B4470">http://dx.doi.org/10.11646/phytotaxa.99.1.1</mods:url>
</mods:location>
<mods:classification id="FF49DD9DA4B1A6669934B72B5F1C3E29">journal article</mods:classification>
<mods:identifier id="87FF87397A56A30C1853A62082719E84" type="DOI">10.11646/phytotaxa.99.1.1</mods:identifier>
<mods:identifier id="9C3ED9E4DD58027F1A3021A158009C0E" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="BDE22897DF5D668A939C115AE8CB1407" type="Zenodo-Dep">5072471</mods:identifier>
</mods:mods>
<treatment id="A7507658EA6CFF94C5AB3E733D1AFC1A" LSID="urn:lsid:plazi:treatment:A7507658EA6CFF94C5AB3E733D1AFC1A" httpUri="http://treatment.plazi.org/id/A7507658EA6CFF94C5AB3E733D1AFC1A" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<subSubSection id="67E394C5EA6CFF96C5AB3E733F3DF9B7" box="[151,771,1570,1597]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="2F46C74EEA6CFF96C5AB3E733F3DF9B7" blockId="5.[151,771,1570,1597]" box="[151,771,1570,1597]" pageId="5" pageNumber="6">
<emphasis id="1D8D1B5CEA6CFF96C5AB3E733C93F9B6" bold="true" box="[151,173,1570,1596]" pageId="5" pageNumber="6">2.</emphasis>
<taxonomicName id="E8F9BCCDEA6CFF96C58F3E723E36F9B7" ID-CoL="7SDVQ" authority="S.Julia" authorityName="S. Julia" authorityYear="2013" box="[179,520,1571,1597]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="benaratensis" status="sp. nov.">
<emphasis id="1D8D1B5CEA6CFF96C58F3E723D8AF9B7" bold="true" box="[179,436,1571,1597]" italics="true" pageId="5" pageNumber="6">Begonia benaratensis</emphasis>
S.Julia
</taxonomicName>
,
<taxonomicNameLabel id="06BEA627EA6CFF96C72A3E723EB6F9B7" box="[534,648,1571,1597]" pageId="5" pageNumber="6" rank="species">
<emphasis id="1D8D1B5CEA6CFF96C72A3E723EBFF9B6" box="[534,641,1571,1596]" italics="true" pageId="5" pageNumber="6">spec. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="B7C2DBCBEA6CFF96C7AB3E723EC5F9B7" box="[663,763,1571,1597]" captionStart="FIGURE 4" captionStartId="6.[151,255,1552,1576]" captionTargetBox="[162,1427,153,1527]" captionTargetId="figure-224@6.[151,1436,143,1540]" captionTargetPageId="6" captionText="FIGURE 4. Begonia benaratensis. A. habit; B. protogynous inflorescence with a single open female flower and many young male flowers; C. male flowers; D. fruit. All photos from SFC 2804 by Julia Sang." figureDoi="http://doi.org/10.5281/zenodo.5072479" httpUri="https://zenodo.org/record/5072479/files/figure.png" pageId="5" pageNumber="6">Figure 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="67E394C5EA6CFF96C5AB3E213923F8E4" pageId="5" pageNumber="6" type="description">
<paragraph id="2F46C74EEA6CFF96C5AB3E213D51F900" blockId="5.[151,367,1648,1674]" box="[151,367,1648,1674]" pageId="5" pageNumber="6">
Sect.
<taxonomicName id="E8F9BCCDEA6CFF96C5E43E203D51F900" authorityName="A. de Candolle" authorityYear="1859" baseAuthorityName="Klotzsch" baseAuthorityYear="1854" box="[216,367,1649,1674]" class="Liliopsida" family="Petermanniaceae" genus="Petermannia" kingdom="Plantae" order="Liliales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="genus">
<emphasis id="1D8D1B5CEA6CFF96C5E43E203D51F900" box="[216,367,1649,1674]" italics="true" pageId="5" pageNumber="6">Petermannia</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="2F46C74EEA6CFF96C5AB3EE63923F8E4" blockId="5.[151,1436,1719,1902]" pageId="5" pageNumber="6">
It is most similar to
<taxonomicName id="E8F9BCCDEA6CFF96C4503EE63EF3F944" authority="Kiew &amp; S. Julia" authorityName="Kiew &amp; S. Julia" authorityYear="2013" box="[364,717,1719,1742]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="apiensis">
<emphasis id="1D8D1B5CEA6CFF96C4503EE63E1DF944" box="[364,547,1719,1742]" italics="true" pageId="5" pageNumber="6">Begonia apiensis</emphasis>
Kiew &amp; S.Julia
</taxonomicName>
in its small, glabrous leaves with the lamina as broad as long, in its two main veins with a further two in the basal lobe, in its petiole more-or-less the length of lamina, and the entire margin but
<taxonomicName id="E8F9BCCDEA6CFF96C47E3EA63DD6F884" authorityName="S. Julia" authorityYear="2013" box="[322,488,1783,1806]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="benaratensis">
<emphasis id="1D8D1B5CEA6CFF96C47E3EA63DD6F884" box="[322,488,1783,1806]" italics="true" pageId="5" pageNumber="6">B. benaratensis</emphasis>
</taxonomicName>
is different from
<taxonomicName id="E8F9BCCDEA6CFF96C79A3EA63F25F884" authorityName="Kiew &amp; S. Julia" authorityYear="2013" box="[678,795,1783,1806]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="apiensis">
<emphasis id="1D8D1B5CEA6CFF96C79A3EA63E88F884" box="[678,694,1783,1806]" italics="true" pageId="5" pageNumber="6">B</emphasis>
.
<emphasis id="1D8D1B5CEA6CFF96C7F83EA63F25F884" box="[708,795,1783,1806]" italics="true" pageId="5" pageNumber="6">apiensis</emphasis>
</taxonomicName>
in its creeping habit, larger stipules ca. 5 ×
<quantity id="E8016AABEA6CFF96C1CE3EA63911F884" box="[1266,1327,1783,1806]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="mm" value="3.0">3 mm</quantity>
(not ca. 3 ×
<quantity id="E8016AABEA6CFF96C5E23F463D0BF8A4" box="[222,309,1815,1838]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="6" unit="mm" value="2.5">2.5 mm</quantity>
), ovate or rounded lamina (not reniform), bigger bracts ca. 5 ×
<quantity id="E8016AABEA6CFF96C6CD3F46380EF8A4" box="[1009,1072,1815,1838]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="mm" value="3.0">3 mm</quantity>
(not ca. 4 ×
<quantity id="E8016AABEA6CFF96C1FE3F46393BF8A4" box="[1218,1285,1815,1838]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="6" unit="mm" value="2.0">2 mm</quantity>
), much larger tepals in female flower 8–10 ×
<quantity id="E8016AABEA6CFF96C72A3F663E51F8C4" box="[534,623,1847,1870]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="5" pageNumber="6" unit="mm" value="5.5" valueMax="8.0" valueMin="3.0">3–8 mm</quantity>
(not 2–2.5 ×
<quantity id="E8016AABEA6CFF96C63D3F663F4FF8C4" box="[769,881,1847,1870]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="5" pageNumber="6" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
), larger male flowers ca. 8 ×
<quantity id="E8016AABEA6CFF96C1973F6638D6F8C4" box="[1195,1256,1847,1870]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="6" unit="mm" value="8.0">8 mm</quantity>
(not 4.5–5.5 ×
<quantity id="E8016AABEA6CFF96C0B23F663CCAF8E4" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="6" unit="mm" value="4.0">4 mm</quantity>
), number of stamens 25–30 (not 15–20) and narrower fruits ca. 12 ×
<quantity id="E8016AABEA6CFF96C6E83F06382FF8E4" box="[980,1041,1879,1902]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="5.0">5 mm</quantity>
(not 10–18 ×
<quantity id="E8016AABEA6CFF96C1943F06392CF8E4" box="[1192,1298,1879,1902]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.9" metricValueMin="0.6" pageId="5" pageNumber="6" unit="mm" value="12.5" valueMax="19.0" valueMin="6.0">6–19 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="67E394C5EA6CFF94C5AB3FC73EB4FE4A" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="2F46C74EEA6CFF96C5AB3FC73818F844" blockId="5.[151,1436,1942,1998]" pageId="5" pageNumber="6">
<emphasis id="1D8D1B5CEA6CFF96C5AB3FC73CE9F824" bold="true" box="[151,215,1942,1966]" pageId="5" pageNumber="6">Type:</emphasis>
—
<materialsCitation id="9F91CD13EA6CFF96C5CF3FC6381CF844" collectingDate="2012-04-15" collectionCode="SAR, E, K, KEP, L, SING" country="Malaysia" county="Marudi District" location="Gunung Benarat" municipality="Gunung Mulu National Park" pageId="5" pageNumber="6" specimenCode="SFC 2804" specimenCount="1" stateProvince="Sarawak" typeStatus="holotype">
<collectingCountry id="57EE87DEEA6CFF96C5CF3FC63DBEF824" box="[243,384,1943,1966]" name="Malaysia" pageId="5" pageNumber="6">MALAYSIA</collectingCountry>
. Borneo.
<collectingRegion id="ED3D09ACEA6CFF96C4DA3FC63E7BF824" box="[486,581,1943,1966]" country="Malaysia" name="Sarawak" pageId="5" pageNumber="6">Sarawak</collectingRegion>
.
<collectingCounty id="C627BFC2EA6CFF96C7733FC63EC7F824" box="[591,761,1943,1966]" pageId="5" pageNumber="6">Marudi District</collectingCounty>
:
<collectingMunicipality id="CF225D34EA6CFF96C63F3FC63809F824" box="[771,1079,1943,1966]" pageId="5" pageNumber="6">Gunung Mulu National Park</collectingMunicipality>
,
<location id="2A269195EA6CFF96C17E3FC638CBF824" LSID="urn:lsid:plazi:treatment:A7507658EA6CFF94C5AB3E733D1AFC1A:2A269195EA6CFF96C17E3FC638CBF824" box="[1090,1269,1943,1966]" country="Malaysia" county="Marudi District" municipality="Gunung Mulu National Park" name="Gunung Benarat" pageId="5" pageNumber="6" stateProvince="Sarawak">Gunung Benarat</location>
,
<date id="5B47E18EEA6CFF96C1C33FC639A6F824" box="[1279,1432,1943,1966]" pageId="5" pageNumber="6" value="2012-04-15">
<collectingDate id="4B031866EA6CFF96C1C33FC639A6F824" box="[1279,1432,1943,1966]" pageId="5" pageNumber="6" value="2012-04-15">15 April 2012</collectingDate>
</date>
,
<emphasis id="1D8D1B5CEA6CFF96C5FB3FE63D94F844" box="[199,426,1975,1998]" italics="true" pageId="5" pageNumber="6">
<location id="2A269195EA6CFF96C5FB3FE63CC2F844" LSID="urn:lsid:plazi:treatment:A7507658EA6CFF94C5AB3E733D1AFC1A:2A269195EA6CFF96C5FB3FE63CC2F844" box="[199,252,1975,1998]" country="Malaysia" county="Marudi District" municipality="Gunung Mulu National Park" name="Julia" pageId="5" pageNumber="6" stateProvince="Sarawak">Julia</location>
et al.
<specimenCode id="7F5F6F35EA6CFF96C4023FE63D94F844" box="[318,426,1975,1998]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="5" pageNumber="6">SFC 2804</specimenCode>
</emphasis>
(
<typeStatus id="F04279ECEA6CFF96C48B3FE63E28F844" box="[439,534,1975,1998]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
<collectionCode id="49E85F8BEA6CFF96C7213FE63E67F844" box="[541,601,1975,1998]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="5" pageNumber="6" type="Herbarium">SAR</collectionCode>
!; isotypes
<collectionCode id="49E85F8BEA6CFF96C7FA3FE63EE2F844" box="[710,732,1975,1998]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="5" pageNumber="6" type="Herbarium">E</collectionCode>
!,
<collectionCode id="49E85F8BEA6CFF96C7D03FE63F38F844" box="[748,774,1975,1998]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="5" pageNumber="6" type="Herbarium">K</collectionCode>
!,
<collectionCode id="49E85F8BEA6CFF96C62A3FE63F6EF844" box="[790,848,1975,1998]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="5" pageNumber="6" type="Herbarium">KEP</collectionCode>
!,
<collectionCode id="49E85F8BEA6CFF96C6623FE63F4AF844" box="[862,884,1975,1998]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="5" pageNumber="6" type="Herbarium">L</collectionCode>
!,
<collectionCode id="49E85F8BEA6CFF96C6B93FE63FF5F844" box="[901,971,1975,1998]" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="5" pageNumber="6" type="Herbarium">SING</collectionCode>
!,
<collectionCode id="49E85F8BEA6CFF96C6E53FE63828F844" box="[985,1046,1975,1998]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="5" pageNumber="6" type="Herbarium">SAR</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<caption id="7B8697C6EA6FFF95C5AB3E413879F9C1" ID-DOI="http://doi.org/10.5281/zenodo.5072479" ID-Zenodo-Dep="5072479" httpUri="https://zenodo.org/record/5072479/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,255,1552,1576]" targetBox="[162,1427,153,1527]" targetPageId="6">
<paragraph id="2F46C74EEA6FFF95C5AB3E413879F9C1" blockId="6.[151,1435,1552,1611]" pageId="6" pageNumber="7">
<emphasis id="1D8D1B5CEA6FFF95C5AB3E413D24F9A2" bold="true" box="[151,282,1552,1576]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
<taxonomicName id="E8F9BCCDEA6FFF95C41E3E403E32F9A2" authorityName="S. Julia" authorityYear="2013" box="[290,524,1553,1576]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="benaratensis">
<emphasis id="1D8D1B5CEA6FFF95C41E3E403E32F9A2" box="[290,524,1553,1576]" italics="true" pageId="6" pageNumber="7">Begonia benaratensis</emphasis>
</taxonomicName>
. A. habit; B. protogynous inflorescence with a single open female flower and many young male flowers; C. male flowers; D. fruit. All photos from SFC 2804 by Julia Sang.
</paragraph>
</caption>
<paragraph id="2F46C74EEA6FFF94C5AB3E2C3EB4FE4A" blockId="6.[151,1436,1661,2036]" lastBlockId="7.[151,1436,150,448]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Creeping herb, rooting at nodes, sometimes branched. Whole plant glabrous. Stems red, ca.
<quantity id="E8016AABEA6FFF95C1E33E2C391AF91D" box="[1247,1316,1661,1687]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="mm" value="2.0">2 mm</quantity>
diameter; internodes 2.5–5(–6.5) cm long. Stipules triangular, ca. 5 ×
<quantity id="E8016AABEA6FFF95C6513EF53F8BF934" box="[877,949,1700,1726]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="7" unit="mm" value="3.0">3 mm</quantity>
, margin entire, apex pointed, persistent. Leaves alternate, distant, straight or sometime pendent. Petioles:
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, terete. Lamina matt plain green above, paler beneath, in life stiff and fleshy, oblique, reniform, (2–)3.5–4 × (3–)
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, asymmetric, broad side (1.7–)
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, base cordate but not overlapping, basal lobes
<quantity id="E8016AABEA6FFF95C6843F493814F8B8" box="[952,1066,1816,1842]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="6" pageNumber="7" unit="cm" value="0.85" valueMax="1.0" valueMin="0.7">0.7–1 cm</quantity>
, margin slightly red, with very distant teeth at vein endings, apex acute; main veins palmate, 2 pairs with an additional 2 veins in basal lobe, branching about halfway to margin, slightly prominent on both surfaces. Inflorescences axillary from the upper leaf axils, red, erect, racemose, protogynous,
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long with a single female flower below and many male flowers above. Bracts ovate, ca. 8 ×
<quantity id="E8016AABEA6FFF95C7A83FE23EE3F847" box="[660,733,1971,1997]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="mm" value="5.0">5 mm</quantity>
, margin entire, apex pointed, caducous. Uppermost bracts ovate, ca. 5 ×
<quantity id="E8016AABEA6FFF95C47F3F8B3DB2F87E" box="[323,396,2010,2036]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="7" unit="mm" value="3.0">3 mm</quantity>
, margin entire, apex pointed, caducous. Male flower: pedicel red,
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; tepals 2, white, glistening, rounded, ca. 8 ×
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, margin entire, apex rounded; stamens 25–30, cluster globose, stalked to
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long; filament to
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long; anthers yellow, oblong or obovate, 0.8–1 ×
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, apex emarginate. Female flower: pedicel ca.
<quantity id="E8016AABEA6EFF94C7BA38B43EF2FF75" box="[646,716,229,255]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="mm" value="5.0">5 mm</quantity>
long, red; ovary ca. 9 ×
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, wings 3, equal, locules 3, placental branches 2 per locule; tepals 5, pinkish, elliptic, 8–10 ×
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, margin entire, apex rounded; styles 3, ca.
<quantity id="E8016AABEA6EFF94C5E339633D1DFEC6" box="[223,291,306,332]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="8" unit="mm" value="2.0">2 mm</quantity>
length, bifurcating; stigma yellow, papillose forming a continuous twisted band. Fruits pendent; pedicel ca.
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long, recurved; capsule reddish, elliptic, 9–14 ×
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, glabrous, locules 3, wings equal, slightly pointed distally,
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wide, dehiscing between the wings and locules. Seeds: barrel-shaped, ca. 0.5 ×
<quantity id="E8016AABEA6EFF94C59239F73D34FE4A" box="[174,266,422,448]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="8" unit="mm" value="0.2">0.2 mm</quantity>
, collar cells half the seed length.
</paragraph>
</subSubSection>
<subSubSection id="67E394C5EA6EFF94C5FA39A23D16FD08" pageId="7" pageNumber="8" type="distribution">
<paragraph id="2F46C74EEA6EFF94C5FA39A23E4AFDBE" blockId="7.[151,1436,499,912]" pageId="7" pageNumber="8">
<emphasis id="1D8D1B5CEA6EFF94C5FA39A23D57FD87" bold="true" box="[198,361,499,525]" pageId="7" pageNumber="8">Distribution:</emphasis>
—
<collectingCountry id="57EE87DEEA6EFF94C4BB39A53E1CFD84" box="[391,546,500,526]" name="Malaysia" pageId="7" pageNumber="8">MALAYSIA</collectingCountry>
. Borneo.
<collectingRegion id="ED3D09ACEA6EFF94C7AA39A53EC1FD84" box="[662,767,500,526]" country="Malaysia" name="Sarawak" pageId="7" pageNumber="8">Sarawak</collectingRegion>
. Marudi District: Gunung Mulu National Park, Melinau limestone. Endemic on Gunung Benarat.
</paragraph>
<paragraph id="2F46C74EEA6EFF94C5FA3A113D16FD08" blockId="7.[151,1436,499,912]" pageId="7" pageNumber="8">
<emphasis id="1D8D1B5CEA6EFF94C5FA3A113D0EFDD0" bold="true" box="[198,304,576,602]" pageId="7" pageNumber="8">Habitat:</emphasis>
—Slightly damp and lightly shaded vertical cliff faces, sometime on mossy surfaces at
<quantity id="E8016AABEA6EFF94C0093A103C90FD08" metricMagnitude="2" metricUnit="m" metricValue="2.1399999999999997" metricValueMax="3.0" metricValueMin="1.28" pageId="7" pageNumber="8" unit="m" value="214.0" valueMax="300.0" valueMin="128.0">128–300 m</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="67E394C5EA6EFF94C5FA3ADF3D1AFC1A" pageId="7" pageNumber="8" type="etymology">
<paragraph id="2F46C74EEA6EFF94C5FA3ADF3905FD22" blockId="7.[151,1436,499,912]" box="[198,1339,654,680]" pageId="7" pageNumber="8">
<emphasis id="1D8D1B5CEA6EFF94C5FA3ADF3D69FD22" bold="true" box="[198,343,654,680]" pageId="7" pageNumber="8">Etymology:</emphasis>
—The species named after Gunung Benarat, the only known locality for this species.
</paragraph>
<paragraph id="2F46C74EEA6EFF94C5FA3AE43814FD7C" blockId="7.[151,1436,499,912]" pageId="7" pageNumber="8">
<emphasis id="1D8D1B5CEA6EFF94C5FA3AE43EE5FD45" bold="true" box="[198,731,693,719]" pageId="7" pageNumber="8">
Additional specimen examined (
<typeStatus id="F04279ECEA6EFF94C76A3AE43EF0FD45" box="[598,718,693,719]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
):
</emphasis>
—
<materialsCitation id="9F91CD13EA6EFF94C7C63AE4381BFD7C" collectionCode="K, SAN, SAR" country="Malaysia" county="Marudi District" location="Baei" municipality="Gunung Mulu National Park - Gunung Benarat" pageId="7" pageNumber="8" specimenCode="SFC 2835" specimenCount="1" stateProvince="Sarawak">
<collectingCountry id="57EE87DEEA6EFF94C7C63AE43FABFD45" box="[762,917,693,719]" name="Malaysia" pageId="7" pageNumber="8">MALAYSIA</collectingCountry>
. Borneo.
<collectingRegion id="ED3D09ACEA6EFF94C1393AE43850FD45" box="[1029,1134,693,719]" country="Malaysia" name="Sarawak" pageId="7" pageNumber="8">Sarawak</collectingRegion>
.
<collectingCounty id="C627BFC2EA6EFF94C1453AE4390CFD45" box="[1145,1330,693,719]" pageId="7" pageNumber="8">Marudi District</collectingCounty>
:
<collectingMunicipality id="CF225D34EA6EFF94C0023AE43E5FFD7C" pageId="7" pageNumber="8">Gunung Mulu National Park – Gunung Benarat</collectingMunicipality>
,
<emphasis id="1D8D1B5CEA6EFF94C7503A8D3F5FFD7F" box="[620,865,732,757]" italics="true" pageId="7" pageNumber="8">
<location id="2A269195EA6EFF94C7503A8D3E9CFD7F" LSID="urn:lsid:plazi:treatment:A7507658EA6CFF94C5AB3E733D1AFC1A:2A269195EA6EFF94C7503A8D3E9CFD7F" box="[620,674,732,757]" country="Malaysia" county="Marudi District" municipality="Gunung Mulu National Park - Gunung Benarat" name="Baei" pageId="7" pageNumber="8" stateProvince="Sarawak">Baei</location>
et al.
<specimenCode id="7F5F6F35EA6EFF94C7D73A8D3F5FFD7F" box="[747,865,732,757]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="7" pageNumber="8">SFC 2835</specimenCode>
</emphasis>
(
<collectionCode id="49E85F8BEA6EFF94C64E3A8D3FB4FD7C" box="[882,906,732,758]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="7" pageNumber="8" type="Herbarium">K</collectionCode>
,
<collectionCode id="49E85F8BEA6EFF94C6AA3A8D3FEDFD7C" box="[918,979,732,758]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="7" pageNumber="8" type="Herbarium">SAN</collectionCode>
,
<collectionCode id="49E85F8BEA6EFF94C6E33A8D3823FD7C" box="[991,1053,732,758]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="7" pageNumber="8" type="Herbarium">SAR</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="2F46C74EEA6EFF94C5FA3B533D1AFC1A" blockId="7.[151,1436,499,912]" pageId="7" pageNumber="8">
<emphasis id="1D8D1B5CEA6EFF94C5FA3B533D2BFC96" bold="true" box="[198,277,770,796]" pageId="7" pageNumber="8">Notes:</emphasis>
—The species is most similar to
<taxonomicName id="E8F9BCCDEA6EFF94C7953B533F12FC91" authorityName="Kiew &amp; S. Julia" authorityYear="2013" box="[681,812,770,795]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="apiensis">
<emphasis id="1D8D1B5CEA6EFF94C7953B533F12FC91" box="[681,812,770,795]" italics="true" pageId="7" pageNumber="8">B. apiensis</emphasis>
</taxonomicName>
but the habitat is completely different. It grows on lightly shaded mossy rock faces below
<quantity id="E8016AABEA6EFF94C74B3B783EFDFCC9" box="[631,707,809,835]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="8" unit="m" value="300.0">300 m</quantity>
compared with
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<emphasis id="1D8D1B5CEA6EFF94C6B53B783831FCC8" box="[905,1039,809,834]" italics="true" pageId="7" pageNumber="8">B. apiensis</emphasis>
</taxonomicName>
that is only found above
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<quantity id="E8016AABEA6EFF94C0743B7839A6FCC9" box="[1352,1432,809,835]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="8" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
Found in localized patches,
<taxonomicName id="E8F9BCCDEA6EFF94C4C83B013E8BFCE3" authorityName="S. Julia" authorityYear="2013" box="[500,693,848,873]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="benaratensis">
<emphasis id="1D8D1B5CEA6EFF94C4C83B013E8BFCE3" box="[500,693,848,873]" italics="true" pageId="7" pageNumber="8">B. benaratensis</emphasis>
</taxonomicName>
sometimes grows together with
<taxonomicName id="E8F9BCCDEA6EFF94C1703B0138C1FCE3" box="[1100,1279,848,873]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="rhodoneura">
<emphasis id="1D8D1B5CEA6EFF94C1703B0138C1FCE3" box="[1100,1279,848,873]" italics="true" pageId="7" pageNumber="8">B. rhodoneura</emphasis>
</taxonomicName>
that is more widespread.
</paragraph>
</subSubSection>
</treatment>
</document>
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