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<document id="0FFCDFD2E954B7410D53B1639E605311" ID-DOI="10.1111/zoj.12362" ID-ISSN="0024-4082" ID-Zenodo-Dep="10109299" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630235092894" checkinUser="marcus" docAuthor="Dellapé, Pablo M., Melo, María C. &amp; Henry, Thomas J." docDate="2016" docId="69250127E907506EFC113489FE15ED58" docLanguage="en" docName="zoj12362.pdf" docOrigin="Zoological Journal of the Linnean Society 177 (1)" docSource="http://dx.doi.org/10.1111/zoj.12362" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Heraeus chamamecinus Dellapé &amp; Melo &amp; Henry 2016, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="67" masterDocId="951C795FE9235048FF9E3541FFCCEE5A" masterDocTitle="A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species" masterLastPageNumber="134" masterPageNumber="29" pageNumber="65" updateTime="1700019240479" updateUser="ExternalLinkService">
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<mods:title id="CCD55F24CF124EB0210771009B7B47C6">A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species</mods:title>
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<mods:namePart id="FCCE649656A4044C636DEE3233DBB5A4">Dellapé, Pablo M.</mods:namePart>
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<subSubSection id="A996E3BAE907506CFC113489FAE8EF87" box="[911,1316,456,477]" pageId="36" pageNumber="65" type="nomenclature">
<paragraph id="E133B031E907506CFC113489FAE8EF87" blockId="36.[911,1316,456,477]" box="[911,1316,456,477]" pageId="36" pageNumber="65">
<heading id="BA7B075DE907506CFC113489FAE8EF87" box="[911,1316,456,477]" centered="true" fontSize="9" level="2" pageId="36" pageNumber="65" reason="4">
<taxonomicName id="268CCBB2E907506CFC113489FB0EEF86" ID-CoL="3KXLW" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[911,1218,456,477]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="66" phylum="Arthropoda" rank="species" species="chamamecinus" status="sp. nov.">HERAEUS CHAMAMECINUS</taxonomicName>
<emphasis id="D3F86C23E907506CFB57348AFAE8EF87" bold="true" box="[1225,1316,456,477]" pageId="36" pageNumber="65">
<taxonomicNameLabel id="C8CBD158E907506CFB57348AFAE8EF87" box="[1225,1316,456,477]" pageId="36" pageNumber="65" rank="species">SP. NOV.</taxonomicNameLabel>
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<subSubSection id="A996E3BAE907506EFC0B34AEFE15ED58" lastPageId="38" lastPageNumber="67" pageId="36" pageNumber="65" type="description">
<paragraph id="E133B031E907506CFC0B34AEFAD2EC5F" blockId="36.[917,1310,494,517]" box="[917,1310,494,517]" pageId="36" pageNumber="65">
(
<figureCitation id="79B7ACB4E907506CFC0334AEFBD1EC5E" box="[925,1053,495,516]" captionStart="Figure 8" captionStartId="17.[164,243,1376,1395]" captionTargetBox="[164,1444,194,1347]" captionTargetId="figure-152@17.[164,1444,194,1347]" captionTargetPageId="17" captionText="Figure 8. Aedeagus. A, Heraeus coquilletti Barber, 1914; B, Heraeus costalis sp. nov.; C, Heraeus setosus sp. nov.; D, Heraeus caliginosus Slater &amp; Baranowski, 1994; E, Heraeus guttatus (Dallas, 1852); F, Heraeus chamamecinus sp. nov., anterior view; G, Heraeus chamamecinus sp. nov., lateral view; H, Heraeus illitus Distant, 1882; I, Heraeus tiputini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128591" httpUri="https://zenodo.org/record/10128591/files/figure.png" pageId="36" pageNumber="65">FIGS 8FG</figureCitation>
,
<figureCitation id="79B7ACB4E907506CFBB734AFFB97EC5E" box="[1065,1115,494,516]" captionStart="Figure 15" captionStartId="32.[144,228,1610,1629]" captionTargetBox="[144,1424,192,1579]" captionTargetId="figure-66@32.[144,1424,192,1580]" captionTargetPageId="32" captionText="Figure 15. Habitus dorsal: A, Heraeus annulatus sp. nov.; B, Heraeus antennalis sp. nov.; C, Heraeus chamamecinus sp. nov.; D, Heraeus illitus Distant, 1882; E, Heraeus inca sp. nov.; F, Heraeus panamaensis sp. nov.; G, Heraeus similis sp. nov.; H, Heraeus spinosus sp. nov. I, Heraeus splendens sp. nov.; J, Heraeus tiputini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128605" httpUri="https://zenodo.org/record/10128605/files/figure.png" pageId="36" pageNumber="65">15C</figureCitation>
,
<figureCitation id="79B7ACB4E907506CFBF934AFFB54EC5E" box="[1127,1176,494,516]" captionStart="Figure 16" captionStartId="33.[164,241,1062,1081]" captionTargetBox="[325,1281,195,1030]" captionTargetId="figure-363@33.[323,1283,193,1032]" captionTargetPageId="33" captionText="Figure 16. Legs: A, Heraeus annulatus sp. nov.; B, Heraeus antennalis sp. nov.; C, Heraeus chamamecinus sp. nov.; D, Heraeus illitus Distant, 1882; E, Heraeus inca sp. nov.; F, Heraeus panamaensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128607" httpUri="https://zenodo.org/record/10128607/files/figure.png" pageId="36" pageNumber="65">16C</figureCitation>
,
<figureCitation id="79B7ACB4E907506CFB3B34AEFB21EC5E" box="[1189,1261,495,516]" captionStart="Figure 17" captionStartId="34.[144,223,1714,1733]" captionTargetBox="[241,1332,196,1675]" captionTargetId="figure-0@34.[229,1338,192,1684]" captionTargetPageId="34" captionText="Figure 17. Male genitalia. Heraeus annulatus sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus antennalis sp. nov.: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus chamamecinus sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus illitus Distant, 1882: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128609" httpUri="https://zenodo.org/record/10128609/files/figure.png" pageId="36" pageNumber="65">17IL</figureCitation>
,
<figureCitation id="79B7ACB4E907506CFB6434AEFADAEC5F" box="[1274,1302,495,517]" captionStart="Figure 19" captionStartId="37.[164,243,1166,1185]" captionTargetBox="[168,1440,199,1131]" captionTargetId="figure-228@37.[164,1444,195,1136]" captionTargetPageId="37" captionText="Figure 19. Distributional map of Heraeus chamamecinus sp. nov. and Heraeus illitus Distant, 1882." figureDoi="http://doi.org/10.5281/zenodo.10128613" httpUri="https://zenodo.org/record/10128613/files/figure.png" pageId="36" pageNumber="65">19</figureCitation>
)
</paragraph>
<paragraph id="E133B031E907506CFCB83757FC5AEC71" blockId="36.[806,1421,534,983]" box="[806,918,534,555]" pageId="36" pageNumber="65">
<emphasis id="D3F86C23E907506CFCB83757FC5AEC71" box="[806,918,534,555]" italics="true" pageId="36" pageNumber="65">Diagnosis</emphasis>
</paragraph>
<paragraph id="E133B031E907506CFCB83775FC17ED45" blockId="36.[806,1421,534,983]" pageId="36" pageNumber="65">Strongly setose. Ocelli located posteriorly. Distiflagellomere without or with only a weakly developed pale band. Profemur irregularly pigmented. Metafemur with a conspicuous dark apical band. Posterior margin of pygophore in dorsal view weakly bilobed; aedeagus with spines laterally to the ejaculatory reservoir and on the sclerotized vesical lobes forming an arc.</paragraph>
<paragraph id="E133B031E907506CFCA13668FA9BED8D" blockId="36.[806,1421,534,983]" pageId="36" pageNumber="65">
<taxonomicName id="268CCBB2E907506CFCA13668FB89ED67" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[831,1093,808,830]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="65" phylum="Arthropoda" rank="species" species="chamamecinus" status="sp. nov.">
<emphasis id="D3F86C23E907506CFCA13668FB89ED67" box="[831,1093,808,830]" italics="true" pageId="36" pageNumber="65">Heraeus chamamecinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E907506CFBD53668FB50ED64" box="[1099,1180,809,830]" pageId="36" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
shares with
<taxonomicName id="268CCBB2E907506CFAB23668FA41ED67" authorityName="Distant" authorityYear="1882" box="[1324,1421,808,830]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="65" phylum="Arthropoda" rank="species" species="illitus">
<emphasis id="D3F86C23E907506CFAB23668FA41ED67" box="[1324,1421,808,830]" italics="true" pageId="36" pageNumber="65">H. illitus</emphasis>
</taxonomicName>
and
<taxonomicName id="268CCBB2E907506CFCC23609FC07ED06" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[860,971,839,861]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="65" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="D3F86C23E907506CFCC23609FC07ED06" box="[860,971,839,861]" italics="true" pageId="36" pageNumber="65">H. similis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E907506CFC4C3606FBEAED06" box="[978,1062,839,860]" pageId="36" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
a highly setose body, and the presence of a slightly bilobed posterior margin of pygophore, but it can be distinguished by the absence of a well-defined pale band on distiflagellomere, and the short labium not surpassing the mesocoxae.
</paragraph>
<paragraph id="E133B031E907506CFCB83141FBEDEA4E" blockId="36.[806,1422,1023,1902]" box="[806,1057,1023,1045]" pageId="36" pageNumber="65">
<emphasis id="D3F86C23E907506CFCB83141FBEDEA4E" box="[806,1057,1023,1045]" italics="true" pageId="36" pageNumber="65">
Description (
<figureCitation id="79B7ACB4E907506CFC2736BEFBD5EA4E" box="[953,1049,1023,1044]" captionStart="Figure 15" captionStartId="32.[144,228,1610,1629]" captionTargetBox="[144,1424,192,1579]" captionTargetId="figure-66@32.[144,1424,192,1580]" captionTargetPageId="32" captionText="Figure 15. Habitus dorsal: A, Heraeus annulatus sp. nov.; B, Heraeus antennalis sp. nov.; C, Heraeus chamamecinus sp. nov.; D, Heraeus illitus Distant, 1882; E, Heraeus inca sp. nov.; F, Heraeus panamaensis sp. nov.; G, Heraeus similis sp. nov.; H, Heraeus spinosus sp. nov. I, Heraeus splendens sp. nov.; J, Heraeus tiputini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128605" httpUri="https://zenodo.org/record/10128605/files/figure.png" pageId="36" pageNumber="65">Fig. 15C</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="E133B031E907506CFCB8315FFB79EA69" blockId="36.[806,1422,1023,1902]" box="[806,1205,1053,1076]" pageId="36" pageNumber="65">Total length 6.18. Strongly setose.</paragraph>
<paragraph id="E133B031E907506CFCA1317DFBE9E861" blockId="36.[806,1422,1023,1902]" pageId="36" pageNumber="65">
<emphasis id="D3F86C23E907506CFCA1317DFC4DEA0B" box="[831,897,1084,1105]" italics="true" pageId="36" pageNumber="65">Head:</emphasis>
Convex dorsally, dark brown, shiny, coriaceus, with short recumbent and long erect setae. Head length 1.25, width 1.01, postocular length 0.46. Eyes protruding, not surpassing dorsal margin of head in lateral view. Ocelli placed posterior to an imaginary line passing posterior border of eyes. Interocular width 0.51, interocellar width 0.34. Labium pale brown, apex of article IV darker; with erect setae, extending to mesocoxae. Labial segment lengths: I 0.80, II 0.88, III 0.78, and IV 0.32. Antennal tubercles parallel. Antennae pale brown, except distal third of basiflagellomere and distiflagellomere darker; distiflagellomere without pale band; all segments with short recumbent and long erect setae, pedicel with abundant erect setae, equal to or longer than diameter of segment. Antennal lengths: scape 0.48, pedicel 1.18, basiflagellomere 0.96, distiflagellomere 1.02.
</paragraph>
<paragraph id="E133B031E907506DFCA13305FE9BE823" blockId="36.[806,1422,1023,1902]" lastBlockId="37.[164,779,1268,1902]" lastPageId="37" lastPageNumber="66" pageId="36" pageNumber="65">
<emphasis id="D3F86C23E907506CFCA13305FC5BE803" box="[831,919,1604,1625]" italics="true" pageId="36" pageNumber="65">Thorax:</emphasis>
Pronotum pruinose; dark brown, posterior lobe with irregular pale areas, and humeral angles dark brown with a small pale spot. Surface punctate, more conspicuous on posterior lobe. Anterior lobe with long erect and short recumbent setae, posterior lobe with erect setae shorter than setae of anterior lobe. Collar length 0.08, anterior lobe length 0.62, posteri- or lobe length 0.54; anterior lobe width 0.99, posteri- or lobe width 1.57. Pleurae brown. Evaporative area extended. Scutellum brown, pruinose, punctate, setae similar to setae on anterior lobe. Hemelytra pruinose, with erect and semi-erect setae. Colour pale brown, costal margin pale on proximal three-quarters, with a subapical corial spot, dark pattern as in
<figureCitation id="79B7ACB4E906506DFDE43011FD2FEB3F" box="[634,739,1359,1382]" captionStart="Figure 15" captionStartId="32.[144,228,1610,1629]" captionTargetBox="[144,1424,192,1579]" captionTargetId="figure-66@32.[144,1424,192,1580]" captionTargetPageId="32" captionText="Figure 15. Habitus dorsal: A, Heraeus annulatus sp. nov.; B, Heraeus antennalis sp. nov.; C, Heraeus chamamecinus sp. nov.; D, Heraeus illitus Distant, 1882; E, Heraeus inca sp. nov.; F, Heraeus panamaensis sp. nov.; G, Heraeus similis sp. nov.; H, Heraeus spinosus sp. nov. I, Heraeus splendens sp. nov.; J, Heraeus tiputini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128605" httpUri="https://zenodo.org/record/10128605/files/figure.png" pageId="37" pageNumber="66">Figure 15</figureCitation>
(C); membrane with a pale, ovoid, apical spot, veins pale. Legs: Pale brown, coxae, profemur, and pretarsus darker; profemur irregularly pigmented; metafemur with a conspicuous dark band apically, less evident on mesofemur. Meso- and metatibia with a subproximal dark band, apex of tibiae darker (
<figureCitation id="79B7ACB4E906506DFDBB3346FD4BE847" box="[549,647,1543,1565]" captionStart="Figure 16" captionStartId="33.[164,241,1062,1081]" captionTargetBox="[325,1281,195,1030]" captionTargetId="figure-363@33.[323,1283,193,1032]" captionTargetPageId="33" captionText="Figure 16. Legs: A, Heraeus annulatus sp. nov.; B, Heraeus antennalis sp. nov.; C, Heraeus chamamecinus sp. nov.; D, Heraeus illitus Distant, 1882; E, Heraeus inca sp. nov.; F, Heraeus panamaensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128607" httpUri="https://zenodo.org/record/10128607/files/figure.png" pageId="37" pageNumber="66">Fig. 16C</figureCitation>
). Legs with abundant erect setae, longest on profemur. Profemur with spines restricted to apical two-thirds. Protibia slightly curved.
</paragraph>
<caption id="B5F3E0B9E906506DFF3A31CFFACCEAFB" ID-DOI="http://doi.org/10.5281/zenodo.10128613" ID-Zenodo-Dep="10128613" box="[164,1280,1166,1185]" httpUri="https://zenodo.org/record/10128613/files/figure.png" pageId="37" pageNumber="66" startId="37.[164,243,1166,1185]" targetBox="[168,1440,199,1131]" targetPageId="37">
<paragraph id="E133B031E906506DFF3A31CFFACCEAFB" blockId="37.[164,1280,1166,1185]" box="[164,1280,1166,1185]" pageId="37" pageNumber="66">
<emphasis id="D3F86C23E906506DFF3A31CFFED5EAFB" bold="true" box="[164,281,1166,1185]" pageId="37" pageNumber="66">Figure 19.</emphasis>
Distributional map of
<emphasis id="D3F86C23E906506DFD8A31CFFC45EAFB" bold="true" box="[532,905,1166,1185]" pageId="37" pageNumber="66">
<taxonomicName id="268CCBB2E906506DFD8A31CFFCE1EAFB" authority="Dellapé &amp; Melo &amp; Henry, 2016" authorityName="Dellapé &amp; Melo &amp; Henry" authorityYear="2016" box="[532,813,1166,1185]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="66" phylum="Arthropoda" rank="species" species="chamamecinus" status="sp. nov.">
<emphasis id="D3F86C23E906506DFD8A31CFFCE1EAFB" bold="true" box="[532,813,1166,1185]" italics="true" pageId="37" pageNumber="66">Heraeus chamamecinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8CBD158E906506DFCAD31CFFC45EAFB" box="[819,905,1166,1185]" pageId="37" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="268CCBB2E906506DFC5C31CEFB30EAFB" authority="Distant, 1882" authorityName="Distant" authorityYear="1882" box="[962,1276,1166,1185]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="66" phylum="Arthropoda" rank="species" species="illitus">
<emphasis id="D3F86C23E906506DFC5C31CEFB93EAFB" box="[962,1119,1166,1185]" italics="true" pageId="37" pageNumber="66">Heraeus illitus</emphasis>
<bibRefCitation id="851DCDC0E906506DFBF631CFFB30EAFB" author="Distant WL" box="[1128,1276,1166,1185]" pageId="37" pageNumber="66" pagination="173 - 220" refId="ref76470" refString="Distant WL. 1882. Lygaeidae. In: Godman FD, Salvin O, eds. Biologia Centrali Americana, Insecta. Rhynchota. Hemiptera- Heteroptera, Vol. I. London: Published by the editors by R. H. Porter, 173 - 220." type="book chapter" year="1882">Distant, 1882</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="E133B031E906506DFF2233C3FC58EB1D" blockId="37.[164,779,1268,1902]" lastBlockId="37.[826,1441,1268,1351]" pageId="37" pageNumber="66">
<emphasis id="D3F86C23E906506DFF2233C3FEE2E8CD" box="[188,302,1666,1687]" italics="true" pageId="37" pageNumber="66">Abdomen:</emphasis>
Brown with abundant short recumbent setae. Male genitalia: Pygophore (
<figureCitation id="79B7ACB4E906506DFDE833E1FCCDE8EC" box="[630,769,1696,1718]" captionStart="Figure 17" captionStartId="34.[144,223,1714,1733]" captionTargetBox="[241,1332,196,1675]" captionTargetId="figure-0@34.[229,1338,192,1684]" captionTargetPageId="34" captionText="Figure 17. Male genitalia. Heraeus annulatus sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus antennalis sp. nov.: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus chamamecinus sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus illitus Distant, 1882: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128609" httpUri="https://zenodo.org/record/10128609/files/figure.png" pageId="37" pageNumber="66">Fig. 17I, J</figureCitation>
) subquadrangular, with posterior margin slightly bilobed and depressed towards aperture, anterior margin of dorsal aperture rounded, inner projections subquadrangular. Parameres:
<figureCitation id="79B7ACB4E906506DFE70325BFD9AE96A" box="[494,598,1818,1840]" captionStart="Figure 17" captionStartId="34.[144,223,1714,1733]" captionTargetBox="[241,1332,196,1675]" captionTargetId="figure-0@34.[229,1338,192,1684]" captionTargetPageId="34" captionText="Figure 17. Male genitalia. Heraeus annulatus sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus antennalis sp. nov.: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus chamamecinus sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus illitus Distant, 1882: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128609" httpUri="https://zenodo.org/record/10128609/files/figure.png" pageId="37" pageNumber="66">Figure 17</figureCitation>
(K, L). Aedeagus (
<figureCitation id="79B7ACB4E906506DFF333278FEFFE915" box="[173,307,1849,1871]" captionStart="Figure 8" captionStartId="17.[164,243,1376,1395]" captionTargetBox="[164,1444,194,1347]" captionTargetId="figure-152@17.[164,1444,194,1347]" captionTargetPageId="17" captionText="Figure 8. Aedeagus. A, Heraeus coquilletti Barber, 1914; B, Heraeus costalis sp. nov.; C, Heraeus setosus sp. nov.; D, Heraeus caliginosus Slater &amp; Baranowski, 1994; E, Heraeus guttatus (Dallas, 1852); F, Heraeus chamamecinus sp. nov., anterior view; G, Heraeus chamamecinus sp. nov., lateral view; H, Heraeus illitus Distant, 1882; I, Heraeus tiputini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10128591" httpUri="https://zenodo.org/record/10128591/files/figure.png" pageId="37" pageNumber="66">Fig. 8F, G</figureCitation>
): conjunctiva with an anterior lobe basal to ejaculatory reservoir; vesica with spines lateral to ejaculatory reservoir, forming an arc on the sclerotized vesical lobes;
<emphasis id="D3F86C23E906506DFBC23052FAFFEB72" box="[1116,1331,1299,1320]" italics="true" pageId="37" pageNumber="66">processus gonopori</emphasis>
long and slender.
</paragraph>
<paragraph id="E133B031E906506DFCA43037FB7EEBD1" blockId="37.[826,1441,1398,1481]" box="[826,1202,1398,1419]" pageId="37" pageNumber="66">
<emphasis id="D3F86C23E906506DFCA43037FB7EEBD1" box="[826,1202,1398,1419]" italics="true" pageId="37" pageNumber="66">
Variability observed in
<typeStatus id="3E370E93E906506DFBDA3037FB7EEBD1" box="[1092,1202,1398,1419]" pageId="37" pageNumber="66" type="paratype">paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="E133B031E906506DFCA430D5FBE9EB93" blockId="37.[826,1441,1398,1481]" pageId="37" pageNumber="66">Some specimens show a weakly developed pale band on distiflagellomere.</paragraph>
<paragraph id="E133B031E906506DFCA430B9FC09E857" blockId="37.[826,1185,1528,1581]" box="[826,965,1528,1549]" pageId="37" pageNumber="66">
<emphasis id="D3F86C23E906506DFCA430B9FC09E857" box="[826,965,1528,1549]" italics="true" pageId="37" pageNumber="66">Distribution</emphasis>
</paragraph>
<paragraph id="E133B031E906506DFCA43357FB6DE877" blockId="37.[826,1185,1528,1581]" box="[826,1185,1558,1581]" pageId="37" pageNumber="66">
<collectingCountry id="999BF0A1E906506DFCA43357FC61E876" box="[826,941,1558,1580]" name="Argentina" pageId="37" pageNumber="66">Argentina</collectingCountry>
and
<collectingCountry id="999BF0A1E906506DFC723357FBFEE876" box="[1004,1074,1558,1580]" name="Brazil" pageId="37" pageNumber="66">Brazil</collectingCountry>
(
<figureCitation id="79B7ACB4E906506DFBDA3357FB59E877" box="[1092,1173,1558,1581]" captionStart="Figure 19" captionStartId="37.[164,243,1166,1185]" captionTargetBox="[168,1440,199,1131]" captionTargetId="figure-228@37.[164,1444,195,1136]" captionTargetPageId="37" captionText="Figure 19. Distributional map of Heraeus chamamecinus sp. nov. and Heraeus illitus Distant, 1882." figureDoi="http://doi.org/10.5281/zenodo.10128613" httpUri="https://zenodo.org/record/10128613/files/figure.png" pageId="37" pageNumber="66">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="E133B031E906506DFCA4331AFC7DE82A" blockId="37.[826,1441,1627,1741]" box="[826,945,1627,1648]" pageId="37" pageNumber="66">
<emphasis id="D3F86C23E906506DFCA4331AFC7DE82A" box="[826,945,1627,1648]" italics="true" pageId="37" pageNumber="66">Etymology</emphasis>
</paragraph>
<paragraph id="E133B031E906506DFCA4333BFB5DE897" blockId="37.[826,1441,1627,1741]" pageId="37" pageNumber="66">
The specific epithet is a noun in aposition and refers to the typical music style (Chamamé) from the Argentinian province of
<collectingRegion id="23487ED3E906506DFB8C33F6FB42E897" box="[1042,1166,1719,1741]" country="Argentina" name="Corrientes" pageId="37" pageNumber="66">Corrientes</collectingRegion>
.
</paragraph>
<paragraph id="E133B031E906506DFCA433BCFC1BE94B" blockId="37.[826,1440,1788,1902]" box="[826,983,1788,1810]" pageId="37" pageNumber="66">
<emphasis id="D3F86C23E906506DFCA433BCFC1BE94B" box="[826,983,1788,1810]" italics="true" pageId="37" pageNumber="66">Type material</emphasis>
</paragraph>
<paragraph id="E133B031E906506DFCA4325BFC46E934" blockId="37.[826,1440,1788,1902]" pageId="37" pageNumber="66">
<materialsCitation id="51E4BA6CE906506DFCA4325BFC46E934" collectingDate="2003-10-30" collectionCode="MLP" collectorName="M. C. Melo" country="Argentina" location="Reserva Santa Maria" municipality="Ituzaingo" pageId="37" pageNumber="66" specimenCount="1" specimenCount-male="1" stateProvince="Corrientes" typeStatus="holotype">
<emphasis id="D3F86C23E906506DFCA4325BFC64E975" box="[826,936,1818,1839]" italics="true" pageId="37" pageNumber="66">
<typeStatus id="3E370E93E906506DFCA4325BFC6DE975" box="[826,929,1818,1839]" pageId="37" pageNumber="66" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="F78A7BB8E906506DFC2A325BFC01E96A" box="[948,973,1818,1840]" pageId="37" pageNumber="66" type="male"></specimenCount>
,
<emphasis id="D3F86C23E906506DFC7E325BFB46E96A" bold="true" box="[992,1162,1818,1840]" pageId="37" pageNumber="66">
<collectingCountry id="999BF0A1E906506DFC7E325BFB46E96A" box="[992,1162,1818,1840]" name="Argentina" pageId="37" pageNumber="66">ARGENTINA</collectingCountry>
</emphasis>
,
<emphasis id="D3F86C23E906506DFB03325AFAD9E96A" box="[1181,1301,1819,1840]" italics="true" pageId="37" pageNumber="66">
<collectingRegion id="23487ED3E906506DFB03325AFAD9E96A" box="[1181,1301,1819,1840]" country="Argentina" name="Corrientes" pageId="37" pageNumber="66">Corrientes</collectingRegion>
</emphasis>
:
<collectingMunicipality id="01572A4BE906506DFAB7325BFA50E96A" box="[1321,1436,1818,1840]" pageId="37" pageNumber="66">Ituzaingo</collectingMunicipality>
,
<location id="E453E6EAE906506DFCA43278FBF1E915" LSID="urn:lsid:plazi:treatment:69250127E907506EFC113489FE15ED58:E453E6EAE906506DFCA43278FBF1E915" box="[826,1085,1849,1871]" country="Argentina" municipality="Ituzaingo" name="Reserva Santa Maria" pageId="37" pageNumber="66" stateProvince="Corrientes">Reserva Santa María</location>
,
<date id="953296F1E906506DFBD33278FB0BE914" box="[1101,1223,1849,1870]" pageId="37" pageNumber="66" value="2003-10-30">
<collectingDate id="85766F19E906506DFBD33278FB0BE914" box="[1101,1223,1849,1870]" pageId="37" pageNumber="66" value="2003-10-30">30-X-2003</collectingDate>
</date>
, T. luz,
<collectorName id="4C79D5E7E906506DFAB43278FA6CE915" box="[1322,1440,1849,1871]" pageId="37" pageNumber="66">M.C. Melo</collectorName>
(
<collectionCode id="879D28F4E906506DFCDC3219FCB3E934" box="[834,895,1880,1902]" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="37" pageNumber="66">MLP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="E133B031E905506EFF0E3587FE15ED58" blockId="38.[144,759,196,770]" pageId="38" pageNumber="67">
<materialsCitation id="51E4BA6CE905506EFF0E3587FD3EEE81" box="[144,754,196,219]" collectingDate="2003-10-30" collectionCode="MLP" collectorName="M. C. Melo" country="Argentina" location="Reserva Santa Maria" municipality="Ituzaingo" pageId="38" pageNumber="67" specimenCount="36" specimenCount-female="14" specimenCount-male="22" stateProvince="Corrientes" typeStatus="paratype">
<emphasis id="D3F86C23E905506EFF0E3587FECFEE81" box="[144,259,198,219]" italics="true" pageId="38" pageNumber="67">
<typeStatus id="3E370E93E905506EFF0E3587FF31EE81" box="[144,253,198,219]" pageId="38" pageNumber="67" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="F78A7BB8E905506EFE943584FEF0EE80" box="[266,316,196,218]" pageId="38" pageNumber="67" type="male">22♂</specimenCount>
,
<specimenCount id="F78A7BB8E905506EFED43584FEB4EE80" box="[330,376,196,218]" pageId="38" pageNumber="67" type="female">14♀</specimenCount>
, same data as for holotype (
<collectionCode id="879D28F4E905506EFD2F3584FD27EE81" box="[689,747,197,219]" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="38" pageNumber="67">MLP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFF3635A2FD48EEA0" box="[168,644,227,250]" collectingDate="2003-10-30" collectionCode="USNM" collectorName="M. C. Melo" country="Argentina" location="Reserva Santa Maria" municipality="Ituzaingo" pageId="38" pageNumber="67" specimenCount="14" specimenCount-female="8" specimenCount-male="6" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFF3635A2FF01EEA3" box="[168,205,227,249]" pageId="38" pageNumber="67" type="male">6♂</specimenCount>
,
<specimenCount id="F78A7BB8E905506EFF4535A5FF30EEA3" box="[219,252,227,249]" pageId="38" pageNumber="67" type="female">8♀</specimenCount>
, same data as for holotype (
<collectionCode id="879D28F4E905506EFDB035A5FDB1EEA0" box="[558,637,228,250]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="38" pageNumber="67" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFD1135A5FD45EF42" collectingDate="2003-10-30" collectionCode="MLP" collectorName="P. M. Dellape" country="Argentina" location="Reserva Santa Maria" municipality="Ituzaingo" pageId="38" pageNumber="67" specimenCount="1" specimenCount-female="1" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFD1135A5FD7CEEA3" box="[655,688,227,249]" pageId="38" pageNumber="67" type="female">1♀</specimenCount>
, same data as for holotype,
<collectorName id="4C79D5E7E905506EFE3D3442FDFFEF42" box="[419,563,258,280]" pageId="38" pageNumber="67">P.M. Dellapé</collectorName>
(
<collectionCode id="879D28F4E905506EFDDB3443FD4DEF42" box="[581,641,258,280]" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="38" pageNumber="67">MLP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFD083443FD76EF6D" collectingDate="2003-10-30" collectionCode="MLP" collectorName="M. Chayle" country="Argentina" location="Reserva Santa Maria" municipality="Ituzaingo" pageId="38" pageNumber="67" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFD083443FD71EF42" box="[662,701,258,280]" pageId="38" pageNumber="67" type="male">2♂</specimenCount>
,
<specimenCount id="F78A7BB8E905506EFD533443FD3CEF42" box="[717,752,258,280]" pageId="38" pageNumber="67" type="female">4♀</specimenCount>
, same data as for holotype,
<collectorName id="4C79D5E7E905506EFE713460FDA8EF6D" box="[495,612,289,311]" pageId="38" pageNumber="67">M. Chayle</collectorName>
(
<collectionCode id="879D28F4E905506EFDE83460FD7EEF6D" box="[630,690,289,311]" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="38" pageNumber="67">MLP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFD573460FD95EF0F" collectingDate="2003-10-31" collectionCode="MLP" collectorName="M. C. Coscaron" country="Argentina" pageId="38" pageNumber="67" specimenCount="1" specimenCount-male="1" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFD573460FD3CEF6C" box="[713,752,288,310]" pageId="38" pageNumber="67" type="male">1♂</specimenCount>
,
<date id="953296F1E905506EFF36347EFED3EF0E" box="[168,287,319,340]" pageId="38" pageNumber="67" value="2003-10-31">
<collectingDate id="85766F19E905506EFF36347EFED3EF0E" box="[168,287,319,340]" pageId="38" pageNumber="67" value="2003-10-31">31-X-2003</collectingDate>
</date>
,
<collectorName id="4C79D5E7E905506EFEB3347EFE1FEF0F" box="[301,467,319,341]" pageId="38" pageNumber="67">M.C. Coscarón</collectorName>
col. (
<collectionCode id="879D28F4E905506EFD8B347EFD9DEF0F" box="[533,593,319,341]" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="38" pageNumber="67">MLP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFDF6347EFDF0EF2E" collectingDate="2003-10-29" collectionCode="AMNH" collectorName="P. M. Dellape" country="Argentina" pageId="38" pageNumber="67" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFDF6347EFD42EF0F" box="[616,654,319,341]" pageId="38" pageNumber="67" type="male">2♂</specimenCount>
,
<specimenCount id="F78A7BB8E905506EFD01347EFD0EEF0F" box="[671,706,319,341]" pageId="38" pageNumber="67" type="female">3♀</specimenCount>
,
<date id="953296F1E905506EFD4D347EFF30EF29" pageId="38" pageNumber="67" value="2003-10-29">
<collectingDate id="85766F19E905506EFD4D347EFF30EF29" pageId="38" pageNumber="67" value="2003-10-29">29- X-2003</collectingDate>
</date>
,
<collectorName id="4C79D5E7E905506EFE95341FFE57EF2E" box="[267,411,350,372]" pageId="38" pageNumber="67">P.M. Dellapé</collectorName>
col. (
<collectionCode id="879D28F4E905506EFE41341FFDF9EF2E" box="[479,565,350,372]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="38" pageNumber="67">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFDD2341FFDCDEFC9" collectingDate="2003-10-29" collectionCode="USNM" collectorName="P. Dellape" country="Argentina" pageId="38" pageNumber="67" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFDD2341FFDBFEF29" box="[588,627,349,371]" pageId="38" pageNumber="67" type="male">2♂</specimenCount>
,
<specimenCount id="F78A7BB8E905506EFD1A341FFD6BEF29" box="[644,679,349,371]" pageId="38" pageNumber="67" type="female">1♀</specimenCount>
,
<date id="953296F1E905506EFD27341FFF2DEFC8" pageId="38" pageNumber="67" value="2003-10-29">
<collectingDate id="85766F19E905506EFD27341FFF2DEFC8" pageId="38" pageNumber="67" value="2003-10-29">29-X- 2003</collectingDate>
</date>
,
<collectorName id="4C79D5E7E905506EFF6F343CFEAFEFC8" box="[241,355,380,402]" pageId="38" pageNumber="67">P. Dellapé</collectorName>
col. (
<collectionCode id="879D28F4E905506EFE38343CFE35EFC9" box="[422,505,381,403]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="38" pageNumber="67" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFD91343DFE11EFEB" collectingDate="2003-10-29" collectionCode="MLP" collectorName="P. M. Dellape" country="Argentina" pageId="38" pageNumber="67" specimenCount="13" specimenCount-female="7" specimenCount-male="6" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFD91343DFDFAEFC8" box="[527,566,380,402]" pageId="38" pageNumber="67" type="male">6♂</specimenCount>
,
<specimenCount id="F78A7BB8E905506EFDD6343CFDA6EFC8" box="[584,618,380,402]" pageId="38" pageNumber="67" type="female">7♀</specimenCount>
,
<date id="953296F1E905506EFDE2343CFD3EEFC8" box="[636,754,381,402]" pageId="38" pageNumber="67" value="2003-10-29">
<collectingDate id="85766F19E905506EFDE2343CFD3EEFC8" box="[636,754,381,402]" pageId="38" pageNumber="67" value="2003-10-29">29-X-2003</collectingDate>
</date>
,
<collectorName id="4C79D5E7E905506EFF3634DDFE8AEFEB" box="[168,326,411,433]" pageId="38" pageNumber="67">P.M. Dellapé</collectorName>
col. (
<collectionCode id="879D28F4E905506EFE0A34DAFE19EFEB" box="[404,469,411,433]" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="38" pageNumber="67">MLP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFE7134DAFE53EF8A" collectingDate="2003-04-27" collectionCode="MLP" collectorName="M. C. Melo" country="Argentina" pageId="38" pageNumber="67" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFE7134DAFDDBEFEB" box="[495,535,411,433]" pageId="38" pageNumber="67" type="male">1♂</specimenCount>
,
<specimenCount id="F78A7BB8E905506EFDB334DAFD9DEFEB" box="[557,593,411,433]" pageId="38" pageNumber="67" type="female">1♀</specimenCount>
,
<date id="953296F1E905506EFDF934DAFD3EEFEA" box="[615,754,411,432]" pageId="38" pageNumber="67" value="2003-04-27">
<collectingDate id="85766F19E905506EFDF934DAFD3EEFEA" box="[615,754,411,432]" pageId="38" pageNumber="67" value="2003-04-27">27-IV-2003</collectingDate>
</date>
,
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col. (
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)
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;
<materialsCitation id="51E4BA6CE905506EFE3534FBFD3FEF8A" box="[427,755,441,464]" collectionCode="MLP" collectorName="P. M. Dellape" country="Argentina" pageId="38" pageNumber="67" specimenCount="3" specimenCount-male="3" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFE3534FBFE1EEF95" box="[427,466,441,463]" pageId="38" pageNumber="67" type="male">3♂</specimenCount>
,
<collectorName id="4C79D5E7E905506EFE7F34FBFDBDEF8A" box="[481,625,442,464]" pageId="38" pageNumber="67">P.M. Dellapé</collectorName>
col. (
<collectionCode id="879D28F4E905506EFD3134FBFD27EF8A" box="[687,747,442,464]" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="38" pageNumber="67">MLP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFF363498FEC3EFB4" box="[168,271,473,494]" country="Argentina" location="Tucuman" pageId="38" pageNumber="67" specimenCount="1" stateProvince="Corrientes" typeStatus="paratype">
<emphasis id="D3F86C23E905506EFF363498FEC3EFB4" box="[168,271,473,494]" italics="true" pageId="38" pageNumber="67">
<location id="E453E6EAE905506EFF363498FEC3EFB4" LSID="urn:lsid:plazi:treatment:69250127E907506EFC113489FE15ED58:E453E6EAE905506EFF363498FEC3EFB4" box="[168,271,473,494]" country="Argentina" name="Tucuman" pageId="38" pageNumber="67" stateProvince="Corrientes">Tucumán</location>
</emphasis>
</materialsCitation>
:
<materialsCitation id="51E4BA6CE905506EFE833499FD9AEC57" collectionCode="USNM" collectorName="Ciudad Universitaria &amp; J. F. G. Clarke" country="Argentina" elevation="800" pageId="38" pageNumber="67" specimenCount="1" specimenCount-male="1" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFE833499FE8FEFB4" box="[285,323,472,494]" pageId="38" pageNumber="67" type="male">1♂</specimenCount>
,
<collectorName id="4C79D5E7E905506EFECF3499FDF2EFB4" box="[337,574,472,494]" pageId="38" pageNumber="67">Ciudad Universitaria</collectorName>
,
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<quantity id="26741DD4E905506EFDD43499FD42EFB4" box="[586,654,472,494]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="38" pageNumber="67" unit="m" value="800.0">800 m</quantity>
a.s.l.
</elevation>
, 18- II-[19]59,
<collectorName id="4C79D5E7E905506EFEAC34B6FE15EC57" box="[306,473,503,525]" pageId="38" pageNumber="67">J.F.G. Clarke</collectorName>
(
<collectionCode id="879D28F4E905506EFE6B34B6FD83EC57" box="[501,591,503,525]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="38" pageNumber="67" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFDF034B6FD3EEC76" collectingDate="1982-02-12" collectionCode="IFML" collectorName="R. Golbach" country="Argentina" location="Rio Salas" municipality="Depto Burruyacu" pageId="38" pageNumber="67" specimenCount="1" specimenCount-female="1" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFDF034B6FD5EEC56" box="[622,658,502,524]" pageId="38" pageNumber="67" type="female">1♀</specimenCount>
,
<collectingMunicipality id="01572A4BE905506EFD3334B6FEEDEC71" pageId="38" pageNumber="67">Depto Burruyacú</collectingMunicipality>
,
<location id="E453E6EAE905506EFEB23754FE5AEC71" LSID="urn:lsid:plazi:treatment:69250127E907506EFC113489FE15ED58:E453E6EAE905506EFEB23754FE5AEC71" box="[300,406,533,555]" country="Argentina" municipality="Depto Burruyacu" name="Rio Salas" pageId="38" pageNumber="67" stateProvince="Corrientes">Rio Salas</location>
,
<date id="953296F1E905506EFE3C3757FDDAEC71" box="[418,534,534,555]" pageId="38" pageNumber="67" value="1982-02-12">
<collectingDate id="85766F19E905506EFE3C3757FDDAEC71" box="[418,534,534,555]" pageId="38" pageNumber="67" value="1982-02-12">12-II-1982</collectingDate>
</date>
,
<collectorName id="4C79D5E7E905506EFDBF3757FD55EC71" box="[545,665,533,555]" pageId="38" pageNumber="67">R. Golbach</collectorName>
(
<collectionCode id="879D28F4E905506EFD393757FD27EC76" box="[679,747,534,556]" pageId="38" pageNumber="67">IFML</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFF363775FD3EEC10" box="[168,754,564,586]" collectingDate="1966-09-18" collectionCode="IFML" collectorName="L. Stange" country="Argentina" location="Horco Molle" pageId="38" pageNumber="67" specimenCount="1" specimenCount-female="1" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFF363775FF07EC10" box="[168,203,564,586]" pageId="38" pageNumber="67" type="female">1♀</specimenCount>
,
<location id="E453E6EAE905506EFF433775FEA2EC10" LSID="urn:lsid:plazi:treatment:69250127E907506EFC113489FE15ED58:E453E6EAE905506EFF433775FEA2EC10" box="[221,366,564,586]" country="Argentina" name="Horco Molle" pageId="38" pageNumber="67" stateProvince="Corrientes">Horco Molle</location>
, 1/
<date id="953296F1E905506EFE0E3775FDD8EC10" box="[400,532,564,586]" pageId="38" pageNumber="67" value="1966-09-18">
<collectingDate id="85766F19E905506EFE0E3775FDD8EC10" box="[400,532,564,586]" pageId="38" pageNumber="67" value="1966-09-18">18-IX-1966</collectingDate>
</date>
,
<collectorName id="4C79D5E7E905506EFDBD3775FD5DEC13" box="[547,657,564,585]" pageId="38" pageNumber="67">L. Stange</collectorName>
(
<collectionCode id="879D28F4E905506EFD3A3775FD26EC10" box="[676,746,564,586]" pageId="38" pageNumber="67">IFML</collectionCode>
)
</materialsCitation>
;
<emphasis id="D3F86C23E905506EFF363712FEE7EC32" box="[168,299,595,616]" italics="true" pageId="38" pageNumber="67">Entre Rios</emphasis>
:
<materialsCitation id="51E4BA6CE905506EFEA13713FD8DEC33" box="[319,577,594,617]" collectingDate="1994-10" collectionCode="USNM" country="Argentina" pageId="38" pageNumber="67" specimenCount="5" specimenCount-male="5" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFEA13713FEAAEC32" box="[319,358,594,616]" pageId="38" pageNumber="67" type="male">5♂</specimenCount>
,
<date id="953296F1E905506EFEE43712FE02EC32" box="[378,462,595,616]" pageId="38" pageNumber="67" value="1994-10">
<collectingDate id="85766F19E905506EFEE43712FE02EC32" box="[378,462,595,616]" pageId="38" pageNumber="67" value="1994-10">X-1994</collectingDate>
</date>
(
<collectionCode id="879D28F4E905506EFE7D3712FDF5EC33" box="[483,569,595,617]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="38" pageNumber="67" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFDCC3712FE6CECDD" collectionCode="USNM" collectorName="Foerster" country="Argentina" pageId="38" pageNumber="67" specimenCount="1" specimenCount-female="1" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFDCC3712FDB9EC32" box="[594,629,594,616]" pageId="38" pageNumber="67" type="female">1♀</specimenCount>
, X-[19]93, E.
<collectorName id="4C79D5E7E905506EFF573733FEE1ECDD" box="[201,301,626,647]" pageId="38" pageNumber="67">Foerster</collectorName>
(
<collectionCode id="879D28F4E905506EFEDC3730FE54ECDD" box="[322,408,625,647]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="38" pageNumber="67" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFE2E3730FE91ECFC" collectionCode="USNM" collectorName="Pena &amp; Ugarte" country="Argentina" pageId="38" pageNumber="67" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Corrientes" typeStatus="paratype">
<specimenCount id="F78A7BB8E905506EFE2E3730FE1BECDD" box="[432,471,625,647]" pageId="38" pageNumber="67" type="male">1♂</specimenCount>
,
<specimenCount id="F78A7BB8E905506EFE753730FDC2ECDD" box="[491,526,625,647]" pageId="38" pageNumber="67" type="female">1♀</specimenCount>
, X-[19]93,
<collectorName id="4C79D5E7E905506EFD043733FD19ECDD" box="[666,725,626,647]" pageId="38" pageNumber="67">Peña</collectorName>
&amp;
<collectorName id="4C79D5E7E905506EFF3637D1FF3AECFF" box="[168,246,656,677]" pageId="38" pageNumber="67">Ugarte</collectorName>
(
<collectionCode id="879D28F4E905506EFE9B37D1FE9BECFC" box="[261,343,656,678]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="38" pageNumber="67" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="51E4BA6CE905506EFEF637D1FD9BEC9F" collectionCode="CNC" collectorName="Banhado Purane &amp; E. G. &amp; E. A. Monroe" country="Brazil" location="BRAZIL" pageId="38" pageNumber="67" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="D3F86C23E905506EFEF637D1FE18ECFF" bold="true" box="[360,468,656,677]" pageId="38" pageNumber="67">
<collectingCountry id="999BF0A1E905506EFEF637D1FE03ECFF" box="[360,463,656,677]" name="Brazil" pageId="38" pageNumber="67">BRAZIL</collectingCountry>
:
</emphasis>
<specimenCount id="F78A7BB8E905506EFE4537D1FE30ECFF" box="[475,508,655,677]" pageId="38" pageNumber="67" type="female">1♀</specimenCount>
,
<collectorName id="4C79D5E7E905506EFD9437D1FD04ECFF" box="[522,712,656,678]" pageId="38" pageNumber="67">Banhado Purane</collectorName>
, 14- II-[19]02,
<collectorName id="4C79D5E7E905506EFE8237EEFDCFEC9E" box="[284,515,687,708]" pageId="38" pageNumber="67">E.G. &amp; E.A. Monroe</collectorName>
(
<collectionCode id="879D28F4E905506EFD8A37EEFD9CEC9F" box="[532,592,687,709]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="38" pageNumber="67">CNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="51E4BA6CE905506EFDFA37EEFE19ED58" collectingDate="1985-11-24" collectionCode="USNM" collectorName="dos Pinhais &amp; T. J. Henry" country="Brazil" location="Sao Jose" pageId="38" pageNumber="67" specimenCount="1" specimenCount-male="1" stateProvince="Parana" typeStatus="paratype">
<emphasis id="D3F86C23E905506EFDFA37EEFD75EC9E" box="[612,697,687,708]" italics="true" pageId="38" pageNumber="67">
<collectingRegion id="23487ED3E905506EFDFA37EEFD75EC9E" box="[612,697,687,708]" country="Brazil" name="Parana" pageId="38" pageNumber="67">Paraná</collectingRegion>
</emphasis>
:
<specimenCount id="F78A7BB8E905506EFD5737EEFD3CEC9E" box="[713,752,686,708]" pageId="38" pageNumber="67" type="male">1♂</specimenCount>
,
<locationDeviation id="F081A913E905506EFF36378CFE4DECB9" box="[168,385,717,739]" location="Curitiba" pageId="38" pageNumber="67">
<quantity id="26741DD4E905506EFF36378CFF3DECB9" box="[168,241,717,739]" metricMagnitude="4" metricUnit="m" metricValue="5.8" pageId="38" pageNumber="67" unit="km" value="58.0">58 km</quantity>
S Curitiba
</locationDeviation>
,
<location id="E453E6EAE905506EFE0C378CFE31ECB8" LSID="urn:lsid:plazi:treatment:69250127E907506EFC113489FE15ED58:E453E6EAE905506EFE0C378CFE31ECB8" box="[402,509,717,738]" country="Brazil" name="Sao Jose" pageId="38" pageNumber="67" stateProvince="Parana">Sao Jose</location>
<collectorName id="4C79D5E7E905506EFD94378CFD57ECB9" box="[522,667,717,739]" pageId="38" pageNumber="67">dos Pinhais</collectorName>
,
<date id="953296F1E905506EFD32378CFF2DED5B" pageId="38" pageNumber="67" value="1985-11-24">
<collectingDate id="85766F19E905506EFD32378CFF2DED5B" pageId="38" pageNumber="67" value="1985-11-24">24-XI- 1985</collectingDate>
</date>
,
<collectorName id="4C79D5E7E905506EFF6E37ADFEA4ED5B" box="[240,360,748,769]" pageId="38" pageNumber="67">T.J. Henry</collectorName>
(
<collectionCode id="879D28F4E905506EFEE737ADFE01ED58" box="[377,461,748,770]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="38" pageNumber="67" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>