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<document id="3C7663AA1A0561F98855FDEB268AB98F" ID-DOI="10.37828/em.2023.61.9" ID-ISSN="2336-9744" ID-Zenodo-Dep="13246732" ID-ZooBank="urn:lsid:zoobank.org:pub:4EB022DC-076A-4C1C-BFBD-01F9B4342F0B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722972049171" checkinUser="felipe" docAuthor="Dubatolov, Vladimir V., Korb, Stanislav K. &amp; Yakovlev, Roman V." docDate="2023" docId="601E954AFFA9FFE412C2272B8735DAAB" docLanguage="en" docName="EcolMontenegrina.61.88-123.pdf" docOrigin="Ecologica Montenegrina 61" docSource="http://dx.doi.org/10.37828/em.2023.61.9" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Triphysa issykkulica Dubatolov, Korb &amp; Yakovlev 2023, sp.n." docType="treatment" docUuid="EADEB292-5523-45B1-8F90-470BBB147E25" docUuidSource="ZooBank" docVersion="2" lastPageNumber="102" masterDocId="9C27ED32FFA4FFEA1302231E833CD90B" masterDocTitle="A review of the genus Triphysa Zeller, 1858 (Lepidoptera, Satyridae)" masterLastPageNumber="123" masterPageNumber="88" pageNumber="101" updateTime="1722979451847" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9444B9A6236C8C3554296B6589CAA90C">A review of the genus Triphysa Zeller, 1858 (Lepidoptera, Satyridae)</mods:title>
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<mods:namePart id="C3A43DFBE04CE349430C4AF1639C1EE5">Dubatolov, Vladimir V.</mods:namePart>
<mods:nameIdentifier id="B88C787CDF3B458D199FC9DE43BD760D" type="ORCID">0000-0001-7687-2102</mods:nameIdentifier>
<mods:affiliation id="C07B8E73BC1861D2CB9C080113236FF3">Institute of Systematics and Ecology of Animals, Siberian Branch of Russian Academy of Sciences, Frunze str. 11, Novosibirsk 630091 Russia. https: // orcid. org / 0000 - 0001 - 7687 - 2102</mods:affiliation>
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<mods:namePart id="186428EE8B0F3BE220A6DBE995B9AE7E">Korb, Stanislav K.</mods:namePart>
<mods:affiliation id="A90AF052F7D8B317A4D5B5BAA79B8A84">Russian Entomological Society, Nizhny Novgorod Division P. O. Box 97, Nizhny Novgorod 603009 Russia.</mods:affiliation>
<mods:nameIdentifier id="BD72A558486E5713953CFBAF71DC7FA3" type="email">stanislavkorb@list.ru</mods:nameIdentifier>
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<mods:namePart id="615D215697A5A3B44DAAAD36D9CB75F6">Yakovlev, Roman V.</mods:namePart>
<mods:nameIdentifier id="1E44D8100EC120E981A9CDE112A0F4FD" type="ORCID">0000-0001-9512-8709</mods:nameIdentifier>
<mods:affiliation id="F6B2F917C3B894BEE798804718BDC034">Altai State University, pr. Lenina 61, Barnaul, 656049, Russia. https: // orcid. org / 0000 - 0001 - 9512 - 8709 * Corresponding author. E-mail: stanislavkorb @ list. ru; https: // orcid. org / 0000 - 0002 - 1120 - 424 X</mods:affiliation>
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<mods:title id="F80B8C3331AB09D2A4FE9F6D6E4B0DE6">Ecologica Montenegrina</mods:title>
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<mods:part id="85BA041A9C23C427172B40C93F40F756">
<mods:date id="52BF7C81F265588025C1E88ED507D7D5">2023</mods:date>
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<mods:number id="868079C6C49386C50EE0FB32CD3D0E46">2023-03-19</mods:number>
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<mods:number id="A672D1917333D2139D24A63CA25CE914">61</mods:number>
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<subSubSection id="A0AD77D7FFA9FFE712C2272B8634DD66" pageId="13" pageNumber="101" type="nomenclature">
<paragraph id="E808245CFFA9FFE712C2272B874FDD44" blockId="13.[300,1288,1077,1167]" box="[448,1139,1077,1103]" pageId="13" pageNumber="101">
<heading id="B3409330FFA9FFE712C2272B874FDD44" box="[448,1139,1077,1103]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="101" reason="3">
<emphasis id="DAC3F84EFFA9FFE712C2272B874FDD44" bold="true" box="[448,1139,1077,1103]" pageId="13" pageNumber="101">
<taxonomicName id="2FB75FDFFFA9FFE712C2272B870DDD44" authority="Dubatolov, Korb &amp; Yakovlev" authorityName="Dubatolov, Korb &amp; Yakovlev" authorityYear="2023" box="[448,1073,1077,1103]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="101" phylum="Arthropoda" rank="species" species="issykkulica" status="sp. nov.">
<emphasis id="DAC3F84EFFA9FFE712C2272B8186DD44" bold="true" box="[448,698,1077,1103]" italics="true" pageId="13" pageNumber="101">Triphysa issykkulica</emphasis>
Dubatolov, Korb &amp; Yakovlev
</taxonomicName>
<taxonomicNameLabel id="C1F04535FFA9FFE7173A272B874FDD44" box="[1080,1139,1077,1103]" pageId="13" pageNumber="101" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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https://zoobank.org/
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urn:lsid:zoobank.org:act:
<uuid id="9C111E89FFA9FFE71004274B8634DD66" box="[774,1288,1109,1133]" pageId="13" pageNumber="101">EADEB292-5523-45B1-8F90-470BBB147E25</uuid>
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<subSubSection id="A0AD77D7FFA9FFE7119D276A82B0DFE4" pageId="13" pageNumber="101" type="description">
<paragraph id="E808245CFFA9FFE7119D276A80A9DD84" blockId="13.[300,1288,1077,1167]" box="[671,917,1140,1167]" pageId="13" pageNumber="101">
<figureCitation id="708C38D9FFA9FFE7119D276A8020DD84" box="[671,796,1140,1167]" captionStart="Figure 2" captionStartId="6.[151,226,1670,1694]" captionTargetBox="[259,1324,199,1630]" captionTargetId="figure-8@6.[254,1333,189,1651]" captionTargetPageId="6" captionText="Figure 2. Triphysa sp., imago: a – T. striatula urumtshiensis, holotype (ZISP), upperside, b – underside; c – T. issykkulica sp. n., holotype (ZISP), upperside, d – underside; e – T. issykkulica sp. n., paratype, male, Kirgiziya, Issyk- Kul’ reg., Sarydzhaz, S slope of Inylchek Mts., Tashkoro, 3100 m, 4.07.1986, I. Pljustch (ISEAN), upperside, f – underside; g – T. issykkulica sp. n., paratype, female, 03.05.2009, Kazakhstan, W of Almaty, 800 m., leg. P.Egorov (SKNN), upperside, h – underside; i – T. issykkulica pljushchi ssp. n., holotype (ZMKU), upperside, j – underside; k – T. issykkulica pljushchi ssp. n., paratype (ZMKU), upperside, l – underside; m – T. nervosa glacialis, lectotype (MHUB), upperside, n – underside; o – T. nervosa glacialis, paralectotype, female (MHUB), upperside, p – underside; q – T. nervosa glacialis, male, SE Altai, Ukok plateau, Mai-pak, H – 2300 m, 29.06-3.07.1997, R. Yakovlev (RYB), upperside, r – underside; s – T. nervosa glacialis, female, SE Altai, Ukok plateau, Mai-pak, H – 2300 m, 29.06- 3.07.1997, R. Yakovlev (RYB), upperside, t – underside; u – T. nervosa glacialis, male, W. Buryariya, Mondy, 21.06.2002 (RYB), upperside, v – underside; w – T. nervosa glacialis, female, W. Buryariya, Mondy, 21.06.2002 (RYB), upperside, x – underside." figureDoi="http://doi.org/10.5281/zenodo.13246738" httpUri="https://zenodo.org/record/13246738/files/figure.png" pageId="13" pageNumber="101">Figs. 2c–h</figureCitation>
,
<figureCitation id="708C38D9FFA9FFE7102B276A807ADD84" box="[809,838,1140,1167]" captionStart="Figure 6" captionStartId="12.[151,226,1968,1992]" captionTargetBox="[211,1376,217,1925]" captionTargetId="figure-8@12.[210,1376,189,1249]" captionTargetPageId="12" captionText="Figure 6. Distribution map of Triphysa species: a (above) – general map; b (below) – northern-eastern part of area of T. nervosa nervosa." figureDoi="http://doi.org/10.5281/zenodo.13246750" httpUri="https://zenodo.org/record/13246750/files/figure.png" pageId="13" pageNumber="101">6a</figureCitation>
,
<figureCitation id="708C38D9FFA9FFE71051276A8058DD84" box="[851,868,1140,1167]" captionStart="Figures 8–9" captionStartId="19.[151,237,1958,1982]" captionTargetBox="[162,1425,193,1926]" captionTargetId="figure-22@19.[160,1427,189,1927]" captionTargetPageId="19" captionText="Figures 8–9. Habitats of Triphysa: 8 – T. phryne phryne, Russia, Volgograd reg. (photo by A. Samus’); 9 – T. issykkulica, SE Kazakhstan, Alma-Ata reg. (photo by S. Korb)." figureDoi="http://doi.org/10.5281/zenodo.13246756" httpUri="https://zenodo.org/record/13246756/files/figure.png" pageId="13" pageNumber="101">9</figureCitation>
,
<figureCitation id="708C38D9FFA9FFE71072276A80ADDD84" box="[880,913,1140,1167]" captionStart="Figures 20–24" captionStartId="26.[151,237,1964,1988]" captionTargetBox="[251,1354,202,1932]" captionTargetId="figure-8@26.[222,1366,189,1946]" captionTargetPageId="26" captionText="Figures 20–24. Male genitalia (lateral projection) of Triphysa: 20 – T. phryne, Volgograd reg.; 21 – T. phryne kasikoporana, holotype; 22 – T. striatula, cotype; 23 – T. striatula urumtshiensis, holotype; 24 – T. issykkulica, holotype." figureDoi="http://doi.org/10.5281/zenodo.13246772" httpUri="https://zenodo.org/record/13246772/files/figure.png" pageId="13" pageNumber="101">24</figureCitation>
.
</paragraph>
<paragraph id="E808245CFFA9FFE7139527A680A6DF2F" blockId="13.[151,1437,1208,2044]" pageId="13" pageNumber="101">
<emphasis id="DAC3F84EFFA9FFE7139527A6821FDDD9" bold="true" box="[151,291,1208,1234]" italics="true" pageId="13" pageNumber="101">Description</emphasis>
.
<emphasis id="DAC3F84EFFA9FFE7123627A6824CDDD8" box="[308,368,1208,1235]" italics="true" pageId="13" pageNumber="101">Male</emphasis>
. Forewing length in
<typeStatus id="370C9AFEFFA9FFE7116927A681EEDDD8" box="[619,722,1208,1235]" pageId="13" pageNumber="101" type="holotype">holotype</typeStatus>
<quantity id="2F4F89B9FFA9FFE711D927A6800DDDD8" box="[731,817,1208,1235]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="101" unit="mm" value="16.0">16 mm</quantity>
, in
<typeStatus id="370C9AFEFFA9FFE7105C27A780F3DDD8" box="[862,975,1209,1235]" pageId="13" pageNumber="101" type="paratype">paratypes</typeStatus>
<quantity id="2F4F89B9FFA9FFE710D527A68767DDD8" box="[983,1115,1208,1235]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="13" pageNumber="101" unit="mm" value="15.5" valueMax="17.0" valueMin="14.0">14–17 mm</quantity>
. Upperside brown. Fringes gray or brownish. Underside brown, with whitegreyish suffusion along the veins; forewing under surface with incomplete submarginal row of black round spots with white centers (no spot in anal region of this wing is present); hindwing underside with full submarginal row of black spots with white centers; each spot in submarginal series is located in yellowish oval elongated stroke. Discal cell of hindwing underside with yellowish short stroke.
<emphasis id="DAC3F84EFFA9FFE712B9267F812FDC77" box="[443,531,1377,1404]" italics="true" pageId="13" pageNumber="101">Female</emphasis>
. Forewing length
<quantity id="2F4F89B9FFA9FFE711FE267F80BFDC77" box="[764,899,1377,1404]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="13" pageNumber="101" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
. Wing upperside, fringes and abdomen are white, submarginal pattern of underside is visible. Underside grayish-brown, veins bright, white, and on the under surface of forewing is located an incomplete submarginal row of black rounded spots with white centers (no spot in anal region of this wing is present); on the hindwing underside the full submarginal row of black spots with white centers is present. Each spot in submarginal row is located inside of yellowish oval stroke, in some specimens these strokes form an almost full band.
</paragraph>
<paragraph id="E808245CFFA9FFE713F4253582B0DFE4" blockId="13.[151,1437,1208,2044]" pageId="13" pageNumber="101">
<emphasis id="DAC3F84EFFA9FFE713F42535829ADF4E" bold="true" box="[246,422,1579,1605]" italics="true" pageId="13" pageNumber="101">Male genitalia</emphasis>
. Tegumen helmet-like with long transverse fold from apex to base. Uncus long, with wide, rounded obtuse apex and broad base of trapezoidal shape. Gnathos short, massive, semi-circular. Valva long, with wide base and narrow pointed apex. Ventral part of valva is wrapped. Saccus elongated, narrow, cylindrical, with smoothly rounded apex. Aedeagus massive, long (longer than valva), tapering to apex. Middle section of aedeagus is covered by small teeth, apex has large strongly sclerotized teeth.
<specimenCount id="FEB1EFD5FFA9FFE7166525AC822DDFE4" count="5" pageId="13" pageNumber="101" type="generic">Five specimens</specimenCount>
dissected.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFA9FFE713F425E88053DE9C" pageId="13" pageNumber="101" type="diagnosis">
<paragraph id="E808245CFFA9FFE713F425E88053DE9C" blockId="13.[151,1437,1208,2044]" pageId="13" pageNumber="101">
<emphasis id="DAC3F84EFFA9FFE713F425E88253DE1B" bold="true" box="[246,367,1782,1808]" italics="true" pageId="13" pageNumber="101">Diagnosis</emphasis>
: From all other species of the genus, this one is well distinguished by the strongly sclerotized teeth on the apex of aedeagus (in other species it is not present) and by the shape of valva (especially its caudal part) and tegumen. The good external difference between the new species and others is in the antenna apex color: in the new species it is always light, yellow, or with very small dark top, whereas in other species it can be completely dark or at least half dark.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFA9FFE413F4248181B7D848" lastPageId="14" lastPageNumber="102" pageId="13" pageNumber="101" type="materials_examined">
<paragraph id="E808245CFFA9FFE413F4248181B7D848" blockId="13.[151,1437,1208,2044]" lastBlockId="14.[151,1437,195,425]" lastPageId="14" lastPageNumber="102" pageId="13" pageNumber="101">
<emphasis id="DAC3F84EFFA9FFE713F4248182E1DEB2" bold="true" box="[246,477,1951,1977]" italics="true" pageId="13" pageNumber="101">Material examined</emphasis>
.
<materialsCitation id="58DF2E01FFA9FFE712EC24808236DED0" collectingDate="2009-05-03" collectionCode="ZMMU" collectorName="P. Egorov" country="Kazakhstan" elevation="800" location="Almaty" pageId="13" pageNumber="101" specimenCount="1" stateProvince="Almaty Oblysy" typeStatus="holotype">
<typeStatus id="370C9AFEFFA9FFE712EC24808162DEB2" box="[494,606,1950,1977]" pageId="13" pageNumber="101" type="holotype">Holotype</typeStatus>
, male,
<collectingCountry id="90A064CCFFA9FFE711B42480807EDEB2" box="[694,834,1950,1977]" name="Kazakhstan" pageId="13" pageNumber="101">Kazakhstan</collectingCountry>
, W of
<collectingRegion id="2A73EABEFFA9FFE71095248080CEDEB2" box="[919,1010,1950,1977]" country="Kazakhstan" name="Almaty Oblysy" pageId="13" pageNumber="101">Almaty</collectingRegion>
,
<quantity id="2F4F89B9FFA9FFE7170224808773DEB2" box="[1024,1103,1950,1977]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="101" unit="m" value="800.0">
<elevation id="639AC36FFFA9FFE7170224808773DEB2" box="[1024,1103,1950,1977]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="101" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="9C09029CFFA9FFE7175F248087DEDEB2" box="[1117,1250,1950,1977]" pageId="13" pageNumber="101" value="2009-05-03">
<collectingDate id="8C4DFB74FFA9FFE7175F248087DEDEB2" box="[1117,1250,1950,1977]" pageId="13" pageNumber="101" value="2009-05-03">03-V-2009</collectingDate>
</date>
, leg.
<collectorName id="4542418AFFA9FFE71627248186A0DEB2" box="[1317,1436,1951,1977]" pageId="13" pageNumber="101">P. Egorov</collectorName>
(
<collectionCode id="8EA6BC99FFA9FFE7139D24DE823DDED0" box="[159,257,1984,2011]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="13" pageNumber="101" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="58DF2E01FFA9FFE7121A24DF806EDEF7" collectingDate="1986-07-04" collectionCode="ISEAN, IPK" collectorName="Sarydzhaz, S &amp; I. Pljustch" country="Kazakhstan" county="Kirgiziya" elevation="3100" location="Inylchek Mts." municipality="Issyk-Kul'" pageId="13" pageNumber="101" specimenCount="9" specimenCount-female="6" specimenCount-male="3" stateProvince="Almaty Oblysy" typeStatus="paratype">
<typeStatus id="370C9AFEFFA9FFE7121A24DF82B2DED0" box="[280,398,1985,2011]" pageId="13" pageNumber="101" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="FEB1EFD5FFA9FFE7129F24DE82C2DED0" box="[413,510,1984,2011]" count="3" pageId="13" pageNumber="101" type="male">3 males</specimenCount>
,
<specimenCount id="FEB1EFD5FFA9FFE7110F24DE81B8DED0" box="[525,644,1984,2011]" count="6" pageId="13" pageNumber="101" type="female">6 females</specimenCount>
,
<collectingCounty id="01695CD0FFA9FFE7119124DE8039DED0" box="[659,773,1984,2011]" pageId="13" pageNumber="101">Kirgiziya</collectingCounty>
,
<collectingMunicipality id="086CBE26FFA9FFE7101624DE80AEDED0" box="[788,914,1984,2011]" pageId="13" pageNumber="101">Issyk-Kul’</collectingMunicipality>
reg.,
<collectorName id="4542418AFFA9FFE710DE24DE8744DED0" box="[988,1144,1984,2011]" pageId="13" pageNumber="101">Sarydzhaz, S</collectorName>
slope of
<location id="ED687287FFA9FFE717ED24DE86ABDED0" LSID="urn:lsid:plazi:treatment:601E954AFFA9FFE412C2272B8735DAAB:ED687287FFA9FFE717ED24DE86ABDED0" box="[1263,1431,1984,2011]" country="Kazakhstan" county="Kirgiziya" municipality="Issyk-Kul'" name="Inylchek Mts." pageId="13" pageNumber="101" stateProvince="Almaty Oblysy">Inylchek Mts.</location>
,
<location id="ED687287FFA9FFE7139524FF8235DEF7" LSID="urn:lsid:plazi:treatment:601E954AFFA9FFE412C2272B8735DAAB:ED687287FFA9FFE7139524FF8235DEF7" box="[151,265,2017,2044]" country="Kazakhstan" county="Kirgiziya" municipality="Issyk-Kul'" name="Tashkoro" pageId="13" pageNumber="101" stateProvince="Almaty Oblysy">Tashkoro</location>
,
<quantity id="2F4F89B9FFA9FFE7121A24FF824BDEF7" box="[280,375,2017,2044]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="13" pageNumber="101" unit="m" value="3100.0">
<elevation id="639AC36FFFA9FFE7121A24FF824BDEF7" box="[280,375,2017,2044]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="13" pageNumber="101" unit="m" value="3100.0">3100 m</elevation>
</quantity>
,
<date id="9C09029CFFA9FFE7128424FF8133DEF7" box="[390,527,2017,2044]" pageId="13" pageNumber="101" value="1986-07-04">
<collectingDate id="8C4DFB74FFA9FFE7128424FF8133DEF7" box="[390,527,2017,2044]" pageId="13" pageNumber="101" value="1986-07-04">4-VII-1986</collectingDate>
</date>
,
<collectorName id="4542418AFFA9FFE7111C24FC81A5DEF7" box="[542,665,2017,2044]" pageId="13" pageNumber="101">I. Pljustch</collectorName>
(
<collectionCode id="8EA6BC99FFA9FFE711A924FF803ADEF7" box="[683,774,2017,2044]" pageId="13" pageNumber="101">ISEAN</collectionCode>
;
<collectionCode id="8EA6BC99FFA9FFE7101424FF8076DEF7" box="[790,842,2017,2044]" country="Germany" httpUri="http://grbio.org/cool/2wtr-g6nj" name="Institut fuer Pflanzenschutzforschung" pageId="13" pageNumber="101">IPK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFA9FFE4106024FF81AAD9D5" collectingDate="2009-05-03" collectionCode="SKNN" collectorName="P. Egorov" country="Kazakhstan" elevation="800" lastPageId="14" lastPageNumber="102" location="Almaty" pageId="13" pageNumber="101" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Almaty Oblysy" typeStatus="paratype">
<specimenCount id="FEB1EFD5FFA9FFE7106024FF80FEDEF7" box="[866,962,2017,2044]" count="2" pageId="13" pageNumber="101" type="male">2 males</specimenCount>
,
<specimenCount id="FEB1EFD5FFA9FFE710D024FF8775DEF7" box="[978,1097,2017,2044]" count="2" pageId="13" pageNumber="101" type="female">2 females</specimenCount>
,
<collectingCountry id="90A064CCFFA9FFE7175A24FF87D8DEF7" box="[1112,1252,2017,2044]" name="Kazakhstan" pageId="13" pageNumber="101">Kazakhstan</collectingCountry>
, W of
<collectingRegion id="2A73EABEFFA9FFE7163F24FF86A4DEF7" box="[1341,1432,2017,2044]" country="Kazakhstan" name="Almaty Oblysy" pageId="13" pageNumber="101">Almaty</collectingRegion>
,
<quantity id="2F4F89B9FFAAFFE4139523DD83D8D9D5" box="[151,228,195,222]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="102" unit="m" value="800.0">
<elevation id="639AC36FFFAAFFE4139523DD83D8D9D5" box="[151,228,195,222]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="102" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="9C09029CFFAAFFE413F223DD8248D9D5" box="[240,372,195,222]" pageId="14" pageNumber="102" value="2009-05-03">
<collectingDate id="8C4DFB74FFAAFFE413F223DD8248D9D5" box="[240,372,195,222]" pageId="14" pageNumber="102" value="2009-05-03">03-V-2009</collectingDate>
</date>
, leg.
<collectorName id="4542418AFFAAFFE412B023DA8114D9D5" box="[434,552,196,222]" pageId="14" pageNumber="102">P. Egorov</collectorName>
(
<collectionCode id="8EA6BC99FFAAFFE4113523DD81B2D9D5" box="[567,654,195,222]" pageId="14" pageNumber="102">SKNN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFAAFFE411A023DD8115D80B" collectingDate="2009-06-29" collectionCode="SKNN" collectorName="S. K. Korb" country="Kazakhstan" county="Kirghizstan" elevation="1600" location="Krasnyi Kanyon" municipality="Kirghiz Mts." pageId="14" pageNumber="102" specimenCount="1" specimenCount-male="1" stateProvince="Almaty Oblysy" typeStatus="paratype">
<specimenCount id="FEB1EFD5FFAAFFE411A023DD81C9D9D5" box="[674,757,195,222]" count="1" pageId="14" pageNumber="102" type="male">1 male</specimenCount>
,
<collectingCounty id="01695CD0FFAAFFE4100223DD80B0D9D5" box="[768,908,195,222]" pageId="14" pageNumber="102">Kirghizstan</collectingCounty>
,
<collectingMunicipality id="086CBE26FFAAFFE4109A23DD870DD9D5" box="[920,1073,195,222]" pageId="14" pageNumber="102">Kirghiz Mts.</collectingMunicipality>
,
<location id="ED687287FFAAFFE4173F23DD863DD9D5" LSID="urn:lsid:plazi:treatment:601E954AFFA9FFE412C2272B8735DAAB:ED687287FFAAFFE4173F23DD863DD9D5" box="[1085,1281,195,222]" country="Kazakhstan" county="Kirghizstan" municipality="Kirghiz Mts." name="Krasnyi Kanyon" pageId="14" pageNumber="102" stateProvince="Almaty Oblysy">Krasnyi Kanyon</location>
,
<quantity id="2F4F89B9FFAAFFE4160F23DD8654D9D5" box="[1293,1384,195,222]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="102" unit="m" value="1600.0">
<elevation id="639AC36FFFAAFFE4160F23DD8654D9D5" box="[1293,1384,195,222]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="102" unit="m" value="1600.0">1600 m</elevation>
</quantity>
,
<date id="9C09029CFFAAFFE4167623DD83C1D80B" pageId="14" pageNumber="102" value="2009-06-29">
<collectingDate id="8C4DFB74FFAAFFE4167623DD83C1D80B" pageId="14" pageNumber="102" value="2009-06-29">29- VI-2009</collectingDate>
</date>
, leg.
<collectorName id="4542418AFFAAFFE4123D23FB8285D80B" box="[319,441,229,256]" pageId="14" pageNumber="102">S.K. Korb</collectorName>
(
<collectionCode id="8EA6BC99FFAAFFE412C823FB811CD80B" box="[458,544,229,256]" pageId="14" pageNumber="102">SKNN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFAAFFE4113423FB8721D829" collectingDate="2009-06-30" collectingDateMax="2009-07-01" collectingDateMin="2009-06-30" collectionCode="SKNN" collectorName="A. Shaposhnikov" country="Kazakhstan" county="Kirghizstan" elevation="1700" location="Kok-Moynok - Kochkor" municipality="Terskey Ala-Too Mts." pageId="14" pageNumber="102" specimenCount="1" specimenCount-male="1" stateProvince="Almaty Oblysy" typeStatus="paratype">
<specimenCount id="FEB1EFD5FFAAFFE4113423FB81B6D80B" box="[566,650,229,256]" count="1" pageId="14" pageNumber="102" type="male">1 male</specimenCount>
,
<collectingCounty id="01695CD0FFAAFFE4119523FB801FD80B" box="[663,803,229,256]" pageId="14" pageNumber="102">Kirghizstan</collectingCounty>
,
<collectingMunicipality id="086CBE26FFAAFFE4103223FB8701D80B" box="[816,1085,229,256]" pageId="14" pageNumber="102">Terskey Ala-Too Mts.</collectingMunicipality>
, 4th km road
<location id="ED687287FFAAFFE417E623FB823DD829" LSID="urn:lsid:plazi:treatment:601E954AFFA9FFE412C2272B8735DAAB:ED687287FFAAFFE417E623FB823DD829" country="Kazakhstan" county="Kirghizstan" municipality="Terskey Ala-Too Mts." name="Kok-Moynok - Kochkor" pageId="14" pageNumber="102" stateProvince="Almaty Oblysy">Kok-Moynok – Kochkor</location>
,
<quantity id="2F4F89B9FFAAFFE4121122198248D829" box="[275,372,263,290]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="102" unit="m" value="1700.0">
<elevation id="639AC36FFFAAFFE4121122198248D829" box="[275,372,263,290]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="102" unit="m" value="1700.0">1700 m</elevation>
</quantity>
,
<date id="9C09029CFFAAFFE41284221981B2D829" box="[390,654,263,290]" pageId="14" pageNumber="102" value="2009-06-30" valueMax="2009-07-01" valueMin="2009-06-30">
<collectingDate id="8C4DFB74FFAAFFE41284221981B2D829" box="[390,654,263,290]" pageId="14" pageNumber="102" value="2009-06-30" valueMax="2009-07-01" valueMin="2009-06-30">30-VI – 01-VII-2009</collectingDate>
</date>
, leg.
<collectorName id="4542418AFFAAFFE411DA22198094D829" box="[728,936,263,290]" pageId="14" pageNumber="102">A. Shaposhnikov</collectorName>
(
<collectionCode id="8EA6BC99FFAAFFE410BC22198729D829" box="[958,1045,263,290]" pageId="14" pageNumber="102">SKNN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFAAFFE4172D221981BBD848" collectionCode="SCR" collectorName="A. Zhdanko" country="Kazakhstan" elevation="1500" location="Sjugoty Mts." municipality="Tian-Shan" pageId="14" pageNumber="102" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Almaty Oblysy" typeStatus="paratype">
<specimenCount id="FEB1EFD5FFAAFFE4172D221987B4D829" box="[1071,1160,263,290]" count="1" pageId="14" pageNumber="102" type="male">1 male</specimenCount>
,
<specimenCount id="FEB1EFD5FFAAFFE4179822198635D829" box="[1178,1289,263,290]" count="1" pageId="14" pageNumber="102" type="female">1 female</specimenCount>
,
<collectingMunicipality id="086CBE26FFAAFFE41618221986A4D829" box="[1306,1432,263,290]" pageId="14" pageNumber="102">Tian-Shan</collectingMunicipality>
,
<location id="ED687287FFAAFFE413952236820ED848" LSID="urn:lsid:plazi:treatment:601E954AFFA9FFE412C2272B8735DAAB:ED687287FFAAFFE413952236820ED848" box="[151,306,296,323]" country="Kazakhstan" municipality="Tian-Shan" name="Sjugoty Mts." pageId="14" pageNumber="102" stateProvince="Almaty Oblysy">Sjugoty Mts.</location>
,
<quantity id="2F4F89B9FFAAFFE4123F223682A5D848" box="[317,409,296,323]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="102" unit="m" value="1500.0">
<elevation id="639AC36FFFAAFFE4123F223682A5D848" box="[317,409,296,323]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="102" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<collectorName id="4542418AFFAAFFE412A72236810DD848" box="[421,561,296,323]" pageId="14" pageNumber="102">A. Zhdanko</collectorName>
(
<collectionCode id="8EA6BC99FFAAFFE4114322368143D848" box="[577,639,296,323]" pageId="14" pageNumber="102">SCR</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFAAFFE413F422558056DB45" pageId="14" pageNumber="102" type="distribution">
<paragraph id="E808245CFFAAFFE413F422558750D8A2" blockId="14.[151,1437,195,425]" pageId="14" pageNumber="102">
<emphasis id="DAC3F84EFFAAFFE413F4225582B5D86E" bold="true" box="[246,393,331,357]" italics="true" pageId="14" pageNumber="102">Distribution</emphasis>
.
<collectingCountry id="90A064CCFFAAFFE4129F22558118D86E" box="[413,548,331,357]" name="Kyrgyzstan" pageId="14" pageNumber="102">Kyrgyzstan</collectingCountry>
(Issyk-Kul’ reg., Central Tian- Shan: Enilchek Mts., Khan-Tengri peak area; North Tian-Shan: Kirgizskii Mts.) (
<bibRefCitation id="8C2659ADFFAAFFE411C52272808BD88C" author="Tshikolovets, V. V." box="[711,951,364,391]" pageId="14" pageNumber="102" refId="ref23956" refString="Tshikolovets, V. V. (2005) The Butterflies of Kyrgyzstan. Kyiv-Brno, Tshikolovets Publications, 511 p." type="book" year="2005">Tshikolovets 2005</bibRefCitation>
),
<collectingCountry id="90A064CCFFAAFFE410D922728758D88C" box="[987,1124,364,391]" name="Kazakhstan" pageId="14" pageNumber="102">Kazakhstan</collectingCountry>
(Transili Alatau Mts.) (nominotypical subspecies) and West Tian-Shan (Talasskij Alatau) (
<taxonomicName id="2FB75FDFFFAAFFE410B922908767D8A2" authorityName="Dubatolov, Korb et Yakovlev" authorityYear="2023" box="[955,1115,398,425]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="102" phylum="Arthropoda" rank="subSpecies" species="issykkulica" subSpecies="pljushchi">
ssp.
<emphasis id="DAC3F84EFFAAFFE410F222908767D8A2" box="[1008,1115,398,425]" italics="true" pageId="14" pageNumber="102">pljushchi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="E808245CFFAAFFE4127A22EA862DDB27" blockId="14.[291,1297,500,590]" pageId="14" pageNumber="102">
<emphasis id="DAC3F84EFFAAFFE4127A22EA8787DB05" bold="true" box="[376,1211,500,526]" pageId="14" pageNumber="102">
<taxonomicName id="2FB75FDFFFAAFFE4127A22EA875CDB05" authority="Dubatolov, Korb et Yakovlev" authorityName="Dubatolov, Korb et Yakovlev" authorityYear="2023" box="[376,1120,500,526]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="102" phylum="Arthropoda" rank="subSpecies" species="issykkulica" status="subsp. nov." subSpecies="pljushchi">
<emphasis id="DAC3F84EFFAAFFE4127A22EA81D5DB05" bold="true" box="[376,745,500,526]" italics="true" pageId="14" pageNumber="102">Triphysa issykkulica pljushchi</emphasis>
Dubatolov, Korb et Yakovlev
</taxonomicName>
,
<taxonomicNameLabel id="C1F04535FFAAFFE4176F22EA8787DB05" box="[1133,1211,500,526]" pageId="14" pageNumber="102" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
https://zoobank.org/
<uri id="9C26285EFFAAFFE412FB210A862DDB27" box="[505,1297,532,556]" pageId="14" pageNumber="102">
urn:lsid:zoobank.org:act:
<uuid id="9C111E89FFAAFFE411FE210A862DDB27" box="[764,1297,532,556]" pageId="14" pageNumber="102">3777473A-AA6B-4D81-A6D8-02D25BBE662C</uuid>
</uri>
</paragraph>
<paragraph id="E808245CFFAAFFE411CB212D8056DB45" blockId="14.[291,1297,500,590]" box="[713,874,563,590]" pageId="14" pageNumber="102">
<figureCitation id="708C38D9FFAAFFE411CB212D8006DB45" box="[713,826,563,590]" captionStart="Figure 2" captionStartId="6.[151,226,1670,1694]" captionTargetBox="[259,1324,199,1630]" captionTargetId="figure-8@6.[254,1333,189,1651]" captionTargetPageId="6" captionText="Figure 2. Triphysa sp., imago: a – T. striatula urumtshiensis, holotype (ZISP), upperside, b – underside; c – T. issykkulica sp. n., holotype (ZISP), upperside, d – underside; e – T. issykkulica sp. n., paratype, male, Kirgiziya, Issyk- Kul’ reg., Sarydzhaz, S slope of Inylchek Mts., Tashkoro, 3100 m, 4.07.1986, I. Pljustch (ISEAN), upperside, f – underside; g – T. issykkulica sp. n., paratype, female, 03.05.2009, Kazakhstan, W of Almaty, 800 m., leg. P.Egorov (SKNN), upperside, h – underside; i – T. issykkulica pljushchi ssp. n., holotype (ZMKU), upperside, j – underside; k – T. issykkulica pljushchi ssp. n., paratype (ZMKU), upperside, l – underside; m – T. nervosa glacialis, lectotype (MHUB), upperside, n – underside; o – T. nervosa glacialis, paralectotype, female (MHUB), upperside, p – underside; q – T. nervosa glacialis, male, SE Altai, Ukok plateau, Mai-pak, H – 2300 m, 29.06-3.07.1997, R. Yakovlev (RYB), upperside, r – underside; s – T. nervosa glacialis, female, SE Altai, Ukok plateau, Mai-pak, H – 2300 m, 29.06- 3.07.1997, R. Yakovlev (RYB), upperside, t – underside; u – T. nervosa glacialis, male, W. Buryariya, Mondy, 21.06.2002 (RYB), upperside, v – underside; w – T. nervosa glacialis, female, W. Buryariya, Mondy, 21.06.2002 (RYB), upperside, x – underside." figureDoi="http://doi.org/10.5281/zenodo.13246738" httpUri="https://zenodo.org/record/13246738/files/figure.png" pageId="14" pageNumber="102">Figs. 2i–l</figureCitation>
,
<figureCitation id="708C38D9FFAAFFE41045212D805ADB45" box="[839,870,563,590]" captionStart="Figure 6" captionStartId="12.[151,226,1968,1992]" captionTargetBox="[211,1376,217,1925]" captionTargetId="figure-8@12.[210,1376,189,1249]" captionTargetPageId="12" captionText="Figure 6. Distribution map of Triphysa species: a (above) – general map; b (below) – northern-eastern part of area of T. nervosa nervosa." figureDoi="http://doi.org/10.5281/zenodo.13246750" httpUri="https://zenodo.org/record/13246750/files/figure.png" pageId="14" pageNumber="102">6a</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFAAFFE4139521698794DBFC" pageId="14" pageNumber="102" type="description">
<paragraph id="E808245CFFAAFFE4139521698794DBFC" blockId="14.[151,1437,631,928]" pageId="14" pageNumber="102">
<emphasis id="DAC3F84EFFAAFFE413952169821FDB9A" bold="true" box="[151,291,631,657]" italics="true" pageId="14" pageNumber="102">Description</emphasis>
.
<emphasis id="DAC3F84EFFAAFFE412362169824CDB99" box="[308,368,631,658]" italics="true" pageId="14" pageNumber="102">Male</emphasis>
. Forewing length
<quantity id="2F4F89B9FFAAFFE41149216981F3DB99" box="[587,719,631,658]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="14" pageNumber="102" unit="mm" value="18.5" valueMax="19.0" valueMin="18.0">18–19 mm</quantity>
. Postdiscal eyespots on the forewing underside with a ledge between the cubital and other spots; cubital spots are are slightly larger. Upperside with no brightening, underside with small eyespots.
<emphasis id="DAC3F84EFFAAFFE4112321A48145DBDE" box="[545,633,698,725]" italics="true" pageId="14" pageNumber="102">Female</emphasis>
. Forewing length
<quantity id="2F4F89B9FFAAFFE4106621A480FDDBDE" box="[868,961,698,725]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="102" unit="mm" value="19.0">19 mm</quantity>
. Upperside light yellow. Underside is characteristic for this species; eyespots are very small with slightly visible white centers.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFAAFFE413F421E08137DA30" pageId="14" pageNumber="102" type="diagnosis">
<paragraph id="E808245CFFAAFFE413F421E08137DA30" blockId="14.[151,1437,631,928]" pageId="14" pageNumber="102">
<emphasis id="DAC3F84EFFAAFFE413F421E08253DA13" bold="true" box="[246,367,766,792]" italics="true" pageId="14" pageNumber="102">Diagnosis</emphasis>
. Bordering on veins is narrow, without any extension (in nominotypical subspecies it has extension, so looks quite wide).
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFAAFFE413F4205C8087DA75" pageId="14" pageNumber="102" type="materials_examined">
<paragraph id="E808245CFFAAFFE413F4205C8087DA75" blockId="14.[151,1437,631,928]" pageId="14" pageNumber="102">
<emphasis id="DAC3F84EFFAAFFE413F4205C829BDA57" bold="true" box="[246,423,834,860]" italics="true" pageId="14" pageNumber="102">Type material</emphasis>
.
<materialsCitation id="58DF2E01FFAAFFE412C2205F8239DA75" collectingDate="1996-06-09" collectionCode="ZMKU" collectorName="W. Kirgiziya &amp; I. Pljustch" location="Talasskii Mts." municipality="Manas" pageId="14" pageNumber="102" specimenCount="1" typeStatus="holotype">
<typeStatus id="370C9AFEFFAAFFE412C2205F810CDA57" box="[448,560,833,860]" pageId="14" pageNumber="102" type="holotype">Holotype</typeStatus>
, male,
<collectorName id="4542418AFFAAFFE4119A205C8003DA57" box="[664,831,833,860]" pageId="14" pageNumber="102">W. Kirgiziya</collectorName>
,
<location id="ED687287FFAAFFE41057205F8734DA57" LSID="urn:lsid:plazi:treatment:601E954AFFA9FFE412C2272B8735DAAB:ED687287FFAAFFE41057205F8734DA57" box="[853,1032,833,860]" municipality="Manas" name="Talasskii Mts." pageId="14" pageNumber="102">Talasskii Mts.</location>
,
<collectingMunicipality id="086CBE26FFAAFFE4171C205C8753DA57" box="[1054,1135,834,860]" pageId="14" pageNumber="102">Manas</collectingMunicipality>
,
<date id="9C09029CFFAAFFE41784205F8639DA57" box="[1158,1285,833,860]" pageId="14" pageNumber="102" value="1996-06-09">
<collectingDate id="8C4DFB74FFAAFFE41784205F8639DA57" box="[1158,1285,833,860]" pageId="14" pageNumber="102" value="1996-06-09">9-VI-1996</collectingDate>
</date>
,
<collectorName id="4542418AFFAAFFE41619205C86A1DA57" box="[1307,1437,833,860]" pageId="14" pageNumber="102">I. Pljustch</collectorName>
(
<collectionCode id="8EA6BC99FFAAFFE4139D207D83C0DA75" box="[159,252,867,894]" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" pageId="14" pageNumber="102">ZMKU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="58DF2E01FFAAFFE41212207A808BDA75" box="[272,951,867,894]" collectingDate="1996-06-09" collectionCode="IPK" collectorName="W. Kirgiziya &amp; I. Pljustch" location="Talasskii Mts." municipality="Manas" pageId="14" pageNumber="102" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="370C9AFEFFAAFFE41212207A82BBDA75" box="[272,391,868,894]" pageId="14" pageNumber="102" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="FEB1EFD5FFAAFFE41293207D82DFDA75" box="[401,483,867,894]" count="1" pageId="14" pageNumber="102" type="male">1 male</specimenCount>
,
<specimenCount id="FEB1EFD5FFAAFFE412ED207D8164DA75" box="[495,600,867,894]" count="1" pageId="14" pageNumber="102" type="female">1 female</specimenCount>
, same locality and data (
<collectionCode id="8EA6BC99FFAAFFE4107B207D8092DA75" box="[889,942,867,894]" country="Germany" httpUri="http://grbio.org/cool/2wtr-g6nj" name="Institut fuer Pflanzenschutzforschung" pageId="14" pageNumber="102">IPK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFAAFFE413F4209B8735DAAB" box="[246,1033,901,928]" pageId="14" pageNumber="102" type="distribution">
<paragraph id="E808245CFFAAFFE413F4209B8735DAAB" blockId="14.[151,1437,631,928]" box="[246,1033,901,928]" pageId="14" pageNumber="102">
<emphasis id="DAC3F84EFFAAFFE413F4209B82B5DA94" bold="true" box="[246,393,901,927]" italics="true" pageId="14" pageNumber="102">Distribution</emphasis>
.
<collectingCountry id="90A064CCFFAAFFE412952098811FDAAB" box="[407,547,902,928]" name="Kyrgyzstan" pageId="14" pageNumber="102">Kyrgyzstan</collectingCountry>
: West Tian-Shan: Talassky Ala-Tau Mts.
</paragraph>
</subSubSection>
</treatment>
</document>
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