135 lines
12 KiB
XML
135 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.204.2740" ID-GBIF-Dataset="c9f2525c-d111-4a5d-b242-8ba4d5c55d2f" ID-PMC="PMC3391719" ID-Pensoft-Pub="1313-2970-204-1" ID-PubMed="22787417" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-204-1" ModsDocOrigin="ZooKeys 204" ModsDocTitle="Snakefly diversity in Early Cretaceous amber from Spain (Neuropterida, Raphidioptera)" checkinTime="1451248933081" checkinUser="pensoft" docAuthor="Fuente, Ricardo Perez-de la, Penalver, Enrique, Delclos, Xavier & Engel, Michael S." docDate="2012" docId="4CEF55152367960D400D22FC5D5C0077" docLanguage="en" docName="ZooKeys 204: 1-40" docOrigin="ZooKeys 204" docSource="http://dx.doi.org/10.3897/zookeys.204.2740" docTitle="Styporaphidia hispanica Perez-de la Fuente, Penalver, Delclos & Engel, sp. n." docType="treatment" docVersion="4" lastPageNumber="13" masterDocId="FFA6FFC19C69FFCD6A5CFF8AFFFA647B" masterDocTitle="Snakefly diversity in Early Cretaceous amber from Spain (Neuropterida, Raphidioptera)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="13" updateTime="1668154032987" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Snakefly diversity in Early Cretaceous amber from Spain (Neuropterida, Raphidioptera)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fuente, Ricardo Perez-de la</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Penalver, Enrique</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Delclos, Xavier</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Engel, Michael S.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>204</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>40</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.204.2740</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.204.2740</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-204-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152036179" LSID="urn:lsid:zoobank.org:act:0AAD19B5-EF08-4B10-B704-DBF6D56A0D6F" httpUri="http://treatment.plazi.org/id/4CEF55152367960D400D22FC5D5C0077" lastPageId="13" lastPageNumber="13" pageId="12" pageNumber="13">
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<subSubSection pageId="12" pageNumber="13" type="nomenclature">
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<paragraph pageId="12" pageNumber="13">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:0AAD19B5-EF08-4B10-B704-DBF6D56A0D6F" authority="Perez-de la Fuente, Penalver, Delclos & Engel" class="Insecta" family="Mesoraphidiidae" genus="Styporaphidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Styporaphidia hispanica" order="Raphidioptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="hispanica">
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Styporaphidia? hispanica
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<normalizedToken originalValue="Pérez-de">Perez-de</normalizedToken>
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la Fuente,
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<normalizedToken originalValue="Peñalver">Penalver</normalizedToken>
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,
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<normalizedToken originalValue="Delclòs">Delclos</normalizedToken>
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& Engel
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</taxonomicName>
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<taxonomicNameLabel pageId="12" pageNumber="13">sp. n.</taxonomicNameLabel>
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Fig. 6
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="13" type="holotype">
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<paragraph pageId="12" pageNumber="13">Holotype.</paragraph>
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<paragraph pageId="12" pageNumber="13">
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MCNA 9343, from
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<normalizedToken originalValue="Peñacerrada">Penacerrada</normalizedToken>
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I amber; hind wing fragment and abdominal apex, including the genitalia.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="13" type="diagnosis">
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<paragraph pageId="12" pageNumber="13">Diagnosis.</paragraph>
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<paragraph pageId="12" pageNumber="13">
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The new species is similar to
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<taxonomicName class="Insecta" family="Mesoraphidiidae" genus="Styporaphidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Styporaphidia magia" order="Raphidioptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="magia">Styporaphidia magia</taxonomicName>
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Engel and Ren, 2008 from the Late Jurassic of Inner Mongolia, China, in the presence of two pterostigmal crossveins.
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<taxonomicName class="Insecta" family="Mesoraphidiidae" genus="Styporaphidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Styporaphidia hispanica" order="Raphidioptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="hispanica">Styporaphidia? hispanica</taxonomicName>
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sp. n.differs in that the distance between 1ptc and 2ptc is three times the distance between 2ptc and the end of the pterostigma (two times in
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<taxonomicName class="Insecta" family="Mesoraphidiidae" genus="Styporaphidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Styporaphidia magia" order="Raphidioptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="magia">Styporaphidia magia</taxonomicName>
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), the forking of Rs at the apicalmost r-rs crossvein (rather than prior to it in
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<taxonomicName class="Insecta" family="Mesoraphidiidae" genus="Styporaphidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Styporaphidia magia" order="Raphidioptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="magia">Styporaphidia magia</taxonomicName>
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), and R meeting the apicalmost r-rs crossvein beyond the pterostigma (within in
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<taxonomicName class="Insecta" family="Mesoraphidiidae" genus="Styporaphidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Styporaphidia magia" order="Raphidioptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="magia">Styporaphidia magia</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="13" type="description">
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<paragraph pageId="12" pageNumber="13">Description.</paragraph>
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<paragraph pageId="12" pageNumber="13">Male. Hind wing. Length as preserved 3.5, maximum width as preserved 2.5; wing apex relatively rounded; C especially thick when compared to other veins; pterostigma almost with constant width along its entire length, infumate; pterostigma with two crossveins, distalmost crossvein oblique and slightly arcuate, proximal crossvein apparently straight, distance between 1ptc and 2ptc three times distance between 2ptc and end of pterostigma; Rs with two branches, forking at r-rs crossvein; R meeting apicalmost r-rs crossvein beyond pterostigma; rs-ma crossvein meeting MA after its distalmost fork; MA with two branches. Abdomen. Gonocoxites 9 with a few long setae;gonostyli 9 segment rather short, rounded (not acute), slightly upcurved;tergite 10 (+11?) with distal setae;paired, contiguous, acute genital structures located dorsad to gonostyli 9, interpreted as distalmost part of parameres (Fig. 6A).</paragraph>
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<caption pageId="12" pageNumber="13">
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<paragraph pageId="12" pageNumber="13">
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Figure 6.
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<taxonomicName class="Insecta" family="Mesoraphidiidae" genus="Styporaphidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Styporaphidia hispanica" order="Raphidioptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="hispanica">Styporaphidia? hispanica</taxonomicName>
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sp. n., holotype MCNA 9343. A hind wing apical fragment and genitalia; arrow 1 points to gonostyli 9, whereas arrow 2 points to the paired genital sclerites interpreted as the distalmost part of the parameres B drawing of preserved hind wing apical fragment. Scale bar = 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="13" type="etymology">
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<paragraph pageId="12" pageNumber="13">Etymology.</paragraph>
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<paragraph pageId="12" pageNumber="13">The specific epithet refers to the occurrence of this species in ancient Spain (Hispania in Latin).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="comments">
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<paragraph pageId="12" pageNumber="13">Comments.</paragraph>
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<paragraph lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
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Although the base of the pterostigma is not preserved and it is accordingly impossible to ascertain if it was diffuse (i.e., lacking a crossvein), this species is tentatively placed in the genus
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<taxonomicName class="Insecta" family="Mesoraphidiidae" genus="Styporaphidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Styporaphidia" order="Raphidioptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Styporaphidia</taxonomicName>
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owing to the presence of two pterostigmal crossveins. The presence of two pterostigmal crossveins is a rare feature among the
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<taxonomicName genus="Raphidioptera" lsidName="Raphidioptera" pageId="12" pageNumber="13" rank="genus">Raphidioptera</taxonomicName>
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and otherwise known in a few other raphidiopterans, i.e, namely the baissopterids
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<taxonomicName class="Insecta" family="Baissopteridae" genus="Baissoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baissoptera kolosnitsynae" order="Raphidioptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="kolosnitsynae">Baissoptera kolosnitsynae</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Baissopteridae" genus="Baissoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baissoptera pulchra" order="Raphidioptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pulchra">Baissoptera pulchra</taxonomicName>
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(
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<bibRefCitation author="Martynova, OM" journalOrPublisher="Paleontologicheskiy Zhurnal" pageId="30" pageNumber="31" pagination="73 - 83" title="Recent and fossil snakeflies (Insecta, Raphidioptera)." volume="1961" year="1961">Martynova 1961</bibRefCitation>
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: p. 81, fig. 7;
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<bibRefCitation pageId="12" pageNumber="13">Martins-Neto and Nel 1992</bibRefCitation>
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: p. 428, figs 2, 3), but the present fossil is clearly not a baissopterid. The fragment is considered as corresponding to a hind wing due to the more distal disposition of both crossveins as occurs in
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<taxonomicName class="Insecta" family="Mesoraphidiidae" genus="Styporaphidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Styporaphidia magia" order="Raphidioptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="magia">Styporaphidia magia</taxonomicName>
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(the distance between them being
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<pageBreakToken pageId="13" pageNumber="14" start="start">greater</pageBreakToken>
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than the distance between 2ptc and the end of the pterostigma), and by the relative position of the abdomen. The veins C and R appear to be especially thickened close to the wing apex as occurs also in
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<taxonomicName class="Insecta" family="Mesoraphidiidae" genus="Styporaphidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Styporaphidia magia" order="Raphidioptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="magia">Styporaphidia magia</taxonomicName>
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(
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<bibRefCitation author="Engel, MS" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="30" pageNumber="31" pagination="188 - 193" title="New snakeflies from the Jiulongshan Formation of Inner Mongolia, China (Raphidioptera)." volume="81" year="2008">Engel and Ren 2008</bibRefCitation>
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: fig. 2), though this effect could be enhanced by the darkening of the margins of the pterostigma. Although the species is represented by a wing fragment and the genitalia, it is distinctive enough that it should be easy to associate with more complete material that may become available in the future.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |