176 lines
18 KiB
XML
176 lines
18 KiB
XML
<document id="ACF4451C1C2E169378794CB7B86FDA7E" ID-DOI="10.1649/0010-065X(2001)055[0433:NSOHKC]2.0.CO;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900383" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622828115451" checkinUser="carolina" docAuthor="Ratcliffe, Brett C." docDate="2001" docId="4C4A87F74452E030FE309066FD4BFD38" docLanguage="en" docName="TheColeopBull.55.4.433-443.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (4)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Hemiphileurus warneri Ratcliffe 2001, new species" docType="treatment" docVersion="3" lastPageNumber="441" masterDocId="B073FF8F4455E038FF98931DFFFEFFA1" masterDocTitle="New Species Of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae) From Mexico, Guatemala, Colombia, And Brazil" masterLastPageNumber="443" masterPageNumber="433" pageNumber="440" updateTime="1698955217201" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="0C27C7A8E67067423DBA8F691DE44E41" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="699DB24EE0CEA653957B05EF47ADA42B">
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<mods:title id="5CD94E146F56AEAB9E3EF229D79CDB6A">New Species Of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae) From Mexico, Guatemala, Colombia, And Brazil</mods:title>
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</mods:titleInfo>
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<mods:name id="E081FB5E2313612531690273634DE682" type="personal">
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<mods:role id="289273E37AF553DC4E54EDFEBDB06AC9">
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<mods:roleTerm id="65B0A4C005A2561AB56D7D08FD3F655C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0E57AAB8731777B4CEE2E50CA1693F86">Ratcliffe, Brett C.</mods:namePart>
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<mods:affiliation id="D606F8D9662E8AEB3B75835D4EEC68D2">Systematics Research Collections W 436 Nebraska Hall University of Nebraska Lincoln, NE 68588 0514, U. S. A.</mods:affiliation>
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<mods:nameIdentifier id="B19FB809FF5F2821D96F2ABF18086126" type="email">bratcliffe1@unl.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="2790FC4A2B6112F79ACC6AE72ADD39B6">text</mods:typeOfResource>
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<mods:titleInfo id="24AC48DE1E95278E16AF72A14373EF92">
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<mods:title id="ED03771F939C3095580EA6C259CD2593">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="66D5C01BEE6A1A7313D092ACD3B6F928">
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<mods:date id="C9A294F92E7560338A4D3EF26948E942">2001</mods:date>
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<mods:detail id="E62D467A37263BAC79CEBAD957360878" type="pubDate">
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<mods:number id="711B74289B412D6CFF1A86D93F53CDE1">2001-12-01</mods:number>
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</mods:detail>
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<mods:detail id="B2AEF83EADF10B92A5A601123E024B49" type="volume">
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<mods:number id="3B51D550475F397539B266D4660F75AE">55</mods:number>
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</mods:detail>
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<mods:detail id="AC1C88F4CBE15225121380AAC9B01938" type="issue">
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<mods:number id="B97D8CED0E21001BD28D9875949C4624">4</mods:number>
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</mods:detail>
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<mods:extent id="4DA5BE9895D8F4A51AE2EB90807D8048" unit="page">
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<mods:start id="1780DAA4ADF5F1E4320B650B1A2BBF2F">433</mods:start>
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<mods:end id="E23339535E40A26F803B0E68E37C58F8">443</mods:end>
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<mods:classification id="C87B7DBD1DE989E995994F865A914666">journal article</mods:classification>
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<mods:identifier id="B673F8ED9D8AB21D8EE1C88354E0CADE" type="DOI">10.1649/0010-065X(2001)055[0433:NSOHKC]2.0.CO;2</mods:identifier>
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<mods:identifier id="B38617BF807AD878E4484ACCE97D6F40" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="1D9750D83E94C7E9126138BD5B516AC3" type="Zenodo-Dep">4900383</mods:identifier>
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</mods:mods>
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<treatment id="4C4A87F74452E030FE309066FD4BFD38" LSID="urn:lsid:plazi:treatment:4C4A87F74452E030FE309066FD4BFD38" httpUri="http://treatment.plazi.org/id/4C4A87F74452E030FE309066FD4BFD38" lastPageId="8" lastPageNumber="441" pageId="7" pageNumber="440">
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<subSubSection id="8CF9656A4452E03FFE309066FD20FC0A" pageId="7" pageNumber="440" type="nomenclature">
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<paragraph id="C45C36E14452E03FFE309066FC6BFC31" blockId="7.[424,917,891,939]" box="[424,917,891,912]" pageId="7" pageNumber="440">
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<heading id="9F14818D4452E03FFE309066FC6BFC31" box="[424,917,891,912]" centered="true" fontSize="9" level="2" pageId="7" pageNumber="440" reason="2">
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<taxonomicName id="03E34D624452E03FFE309066FCF8FC31" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2001" box="[424,774,891,912]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="440" phylum="Arthropoda" rank="species" species="warneri" status="sp. nov.">
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<emphasis id="F697EAF34452E03FFE309066FD61FC31" box="[424,671,891,912]" italics="true" pageId="7" pageNumber="440">Hemiphileurus warneri</emphasis>
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Ratcliffe
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</taxonomicName>
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,
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||
<emphasis id="F697EAF34452E03FFC8A9066FC6BFC31" bold="true" box="[786,917,891,912]" pageId="7" pageNumber="440">
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<taxonomicNameLabel id="EDA457884452E03FFC8A9066FC6BFC31" box="[786,917,891,912]" pageId="7" pageNumber="440" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="C45C36E14452E03FFDD6908BFD20FC0A" blockId="7.[424,917,891,939]" box="[590,734,918,939]" pageId="7" pageNumber="440">(Figs. 16–18)</paragraph>
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</subSubSection>
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<subSubSection id="8CF9656A4452E03FFE8890D8FCFFF9E1" pageId="7" pageNumber="440" type="materials_examined">
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<paragraph id="C45C36E14452E03FFE8890D8FC84FBB0" blockId="7.[246,1078,965,1041]" pageId="7" pageNumber="440">
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<emphasis id="F697EAF34452E03FFE8890D8FE44FC7B" bold="true" box="[272,442,965,986]" pageId="7" pageNumber="440">Type Material.</emphasis>
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<materialsCitation id="748B3CBC4452E03FFE5B90D8FC9CFC57" collectingDate="1991-06-07" collectionCode="V" collectorName="E Los Piedras & W. B. Warner." country="Mexico" location="Hwy" pageId="7" pageNumber="440" specimenCount="1" stateProvince="Nayarit" typeStatus="holotype">
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<typeStatus id="1B5888434452E03FFE5B90D8FDDAFC7B" box="[451,548,965,986]" pageId="7" pageNumber="440" type="holotype">Holotype</typeStatus>
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labeled ‘‘
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<collectingCountry id="BCF476714452E03FFD1390D8FD02FC7B" box="[651,764,965,986]" name="Mexico" pageId="7" pageNumber="440">MEXICO</collectingCountry>
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:
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<collectingRegion id="0627F8034452E03FFC9090D8FCA7FC7B" box="[776,857,965,986]" country="Mexico" name="Nayarit" pageId="7" pageNumber="440">Nayarit</collectingRegion>
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,
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<location id="C13C603A4452E03FFCFE90D8FC67FC7B" LSID="urn:lsid:plazi:treatment:4C4A87F74452E030FE309066FD4BFD38:C13C603A4452E03FFCFE90D8FC67FC7B" box="[870,921,965,986]" country="Mexico" name="Hwy" pageId="7" pageNumber="440" stateProvince="Nayarit">Hwy</location>
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200, 20 km
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<collectorName id="691653374452E03FFBBE90D8FE8BFC57" pageId="7" pageNumber="440">E Los Piedras</collectorName>
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,
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<date id="B05D10214452E03FFE1890FCFE0BFC57" box="[384,501,993,1014]" pageId="7" pageNumber="440" value="1991-06-07">
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||
<collectingDate id="A019E9C94452E03FFE1890FCFE0BFC57" box="[384,501,993,1014]" pageId="7" pageNumber="440" value="1991-06-07">
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VI
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<quantity id="031B9B044452E03FFE3E90FCFE0BFC57" box="[422,501,993,1014]" metricMagnitude="5" metricUnit="m" metricValue="9.958685" metricValueMax="19.9173" metricValueMin="7.0E-5" pageId="7" pageNumber="440" unit="m" value="995868.5" valueMax="1991730.0" valueMin="7.0">7–1991</quantity>
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</collectingDate>
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</date>
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, 730 m, U.
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<collectionCode id="A2F2AE244452E03FFDF290FCFD80FC57" box="[618,638,993,1014]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="440" type="Museum">V</collectionCode>
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. light,
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<collectorName id="691653374452E03FFD5F90FCFCA4FC57" box="[711,858,993,1014]" pageId="7" pageNumber="440">W.B. Warner.</collectorName>
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’’
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</materialsCitation>
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<materialsCitation id="748B3CBC4452E03FFCF290FCFC88FBB0" collectorName="Lincoln, NE" country="Mexico" location="University of Nebraska State Museum" pageId="7" pageNumber="440" specimenCount="1" stateProvince="Nayarit" typeStatus="holotype">
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<typeStatus id="1B5888434452E03FFCF290FCFC35FC57" box="[874,971,993,1014]" pageId="7" pageNumber="440" type="holotype">Holotype</typeStatus>
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||
deposited at the
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<location id="C13C603A4452E03FFEA390E1FD2AFBB0" LSID="urn:lsid:plazi:treatment:4C4A87F74452E030FE309066FD4BFD38:C13C603A4452E03FFEA390E1FD2AFBB0" box="[315,724,1020,1041]" country="Mexico" name="University of Nebraska State Museum" pageId="7" pageNumber="440" stateProvince="Nayarit">University of Nebraska State Museum</location>
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(
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<collectorName id="691653374452E03FFD7D90E1FC90FBB0" box="[741,878,1020,1041]" pageId="7" pageNumber="440">Lincoln, NE</collectorName>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="C45C36E14452E03FFE959737FCFFF9E1" blockId="7.[246,1080,1066,1601]" pageId="7" pageNumber="440">
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<emphasis id="F697EAF34452E03FFE959737FE99FB9C" bold="true" box="[269,359,1066,1085]" pageId="7" pageNumber="440">
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<typeStatus id="1B5888434452E03FFE959737FE99FB9C" box="[269,359,1066,1085]" pageId="7" pageNumber="440" type="holotype">Holotype</typeStatus>
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</emphasis>
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. Male. Length 21.0 mm; width
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<quantity id="031B9B044452E03FFD0C9737FD20FB9C" box="[660,734,1066,1085]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="7" pageNumber="440" unit="mm" value="9.7">9.7 mm</quantity>
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. Color black.
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<emphasis id="F697EAF34452E03FFCF99737FC6DFB9C" box="[865,915,1066,1085]" italics="true" pageId="7" pageNumber="440">Head</emphasis>
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: Surface of frons completely, coarsely rugose, with median, deep fovea; a low, rounded tubercle present mesad of anterior border of each eye. Clypeus triangular, apex acute, reflexed; surface with sparse, moderatelysized punctures and with strong carina extending from clypeal apex to base of each tubercle. Interocular width equals 3.1 transverse eye diameters. Antenna 10segmented, club slightly longer than segments 2–7. Mandibles with external edges arcuate, apices acute.
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<emphasis id="F697EAF34452E03FFD9D97A3FD9CFB70" box="[517,610,1214,1233]" italics="true" pageId="7" pageNumber="440">Pronotum</emphasis>
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: Surface with moderately large punctures; punctures moderately dense, becoming denser in anterior fourth and in longitudinal furrow. Median, longitudinal furrow deep, narrow, extending from near base to near apex. Apex just behind marginal bead with small tubercle either side of midline. Base with complete marginal bead.
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<emphasis id="F697EAF34452E03FFE15963DFE39FA92" box="[397,455,1312,1331]" italics="true" pageId="7" pageNumber="440">Elytra</emphasis>
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: Surface with 5 distinct, punctate striae between suture and humeral umbone; punctures moderately large, ocellate, subcontiguous or separated from one another in each row by less than half a puncture length. Intervals convex, with sparse micropunctures.
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<taxonomicName id="03E34D624452E03FFE4D9674FD7DFADD" authority=": Surface" authorityName="Surface" box="[469,643,1385,1404]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="440" phylum="Chordata" rank="genus">
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<emphasis id="F697EAF34452E03FFE4D9674FDD0FADD" box="[469,558,1385,1404]" italics="true" pageId="7" pageNumber="440">Pygidium</emphasis>
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: Surface
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</taxonomicName>
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with punctures moderately dense, moderately large, weakly ocellate, setigerous; setae minute, tawny. In lateral view, surface regularly convex.
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<emphasis id="F697EAF34452E03FFEDF9687FE8DFA0C" box="[327,371,1434,1453]" italics="true" pageId="7" pageNumber="440">Legs</emphasis>
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: Foretibia with 4 subequally separated teeth. Median transverse carina on meso and metatibiae ending with acute spine on upper surface. Apex of posterior tibia with strong, acute spine on upper angle and with 7 (left) and 4 (right) short, broad spinules below spine. Apex of first tarsomere of posterior tarsus with strong, acute spine dorsally.
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<emphasis id="F697EAF34452E03FFED696E1FE72F9AE" box="[334,396,1532,1551]" italics="true" pageId="7" pageNumber="440">Venter</emphasis>
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: Prosternal process (Fig. 18) broad, moderate in length, apex flattened into transversely subrectangular, slightly concave plate, setae lacking. Last sternite with sparse, scattered punctures.
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<emphasis id="F697EAF34452E03FFE679533FD9BF9E0" box="[511,613,1582,1601]" italics="true" pageId="7" pageNumber="440">Parameres</emphasis>
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: Figures 16–17.
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</paragraph>
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</subSubSection>
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<subSubSection id="8CF9656A4452E030FE889543FED7FE9F" lastPageId="8" lastPageNumber="441" pageId="7" pageNumber="440" type="etymology">
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<paragraph id="C45C36E14452E03FFE889543FBC8F9D5" blockId="7.[272,1078,1630,1652]" box="[272,1078,1630,1652]" pageId="7" pageNumber="440">
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<emphasis id="F697EAF34452E03FFE889543FE71F9D2" bold="true" box="[272,399,1630,1651]" pageId="7" pageNumber="440">Etymology.</emphasis>
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I take great pleasure in naming this species after Bill Warner
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</paragraph>
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<paragraph id="C45C36E1445DE030FF619212FED7FE9F" blockId="8.[249,1082,271,665]" pageId="8" pageNumber="441">(Chandler, AZ) who collected the specimen and provided it to me for description.</paragraph>
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</subSubSection>
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<subSubSection id="8CF9656A445DE030FE8B9259FD4BFD38" pageId="8" pageNumber="441" type="discussion">
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<paragraph id="C45C36E1445DE030FE8B9259FED1FDC5" blockId="8.[249,1082,271,665]" pageId="8" pageNumber="441">
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<emphasis id="F697EAF3445DE030FE8B9259FE7EFEF8" bold="true" box="[275,384,324,345]" pageId="8" pageNumber="441">Remarks.</emphasis>
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<taxonomicName id="03E34D62445DE030FE149259FD78FEF8" authorityName="Ratcliffe" authorityYear="2001" box="[396,646,324,345]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="441" phylum="Arthropoda" rank="species" species="warneri">
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<emphasis id="F697EAF3445DE030FE149259FD78FEF8" box="[396,646,324,345]" italics="true" pageId="8" pageNumber="441">Hemiphileurus warneri</emphasis>
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</taxonomicName>
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is similar externally to
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<taxonomicName id="03E34D62445DE030FC019259FBFEFEF8" baseAuthorityName="LeConte" baseAuthorityYear="1854" box="[921,1024,324,345]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="441" phylum="Arthropoda" rank="species" species="illatus">
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<emphasis id="F697EAF3445DE030FC019259FBFEFEF8" box="[921,1024,324,345]" italics="true" pageId="8" pageNumber="441">H. illatus</emphasis>
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</taxonomicName>
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(Le Conte) and will key to this species in Endrödi (1985). The parameres of
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<taxonomicName id="03E34D62445DE030FBB89242FEB2FE2E" authorityName="Ratcliffe" authorityYear="2001" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="441" phylum="Arthropoda" rank="species" species="warneri">
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<emphasis id="F697EAF3445DE030FBB89242FEB2FE2E" italics="true" pageId="8" pageNumber="441">H. warneri</emphasis>
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</taxonomicName>
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(Figs. 16–17) are different than those of
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<taxonomicName id="03E34D62445DE030FC909267FC94FE2E" baseAuthorityName="LeConte" baseAuthorityYear="1854" box="[776,874,378,399]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="441" phylum="Arthropoda" rank="species" species="illatus">
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<emphasis id="F697EAF3445DE030FC909267FC94FE2E" box="[776,874,378,399]" italics="true" pageId="8" pageNumber="441">H. illatus</emphasis>
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</taxonomicName>
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(Figs. 19–20). The most striking difference between the parameres, aside from the obvious length proportions, is the almost entirely concave surface of each paramere that abruptly becomes convex at the apices in
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<taxonomicName id="03E34D62445DE030FD4892D7FCB9FE7E" authorityName="Ratcliffe" authorityYear="2001" box="[720,839,458,479]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="441" phylum="Arthropoda" rank="species" species="warneri">
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<emphasis id="F697EAF3445DE030FD4892D7FCB9FE7E" box="[720,839,458,479]" italics="true" pageId="8" pageNumber="441">H. warneri</emphasis>
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</taxonomicName>
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whereas in
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<taxonomicName id="03E34D62445DE030FC4B92D7FBC7FE7E" baseAuthorityName="LeConte" baseAuthorityYear="1854" box="[979,1081,458,479]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="441" phylum="Arthropoda" rank="species" species="illatus">
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<emphasis id="F697EAF3445DE030FC4B92D7FBC7FE7E" box="[979,1081,458,479]" italics="true" pageId="8" pageNumber="441">H. illatus</emphasis>
|
||
</taxonomicName>
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there is only an oblong depression on the center of the paramere. In addition, the base of the last sternite in
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<taxonomicName id="03E34D62445DE030FDC892E2FD48FDB5" baseAuthorityName="LeConte" baseAuthorityYear="1854" box="[592,694,511,532]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="441" phylum="Arthropoda" rank="species" species="illatus">
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<emphasis id="F697EAF3445DE030FDC892E2FD48FDB5" box="[592,694,511,532]" italics="true" pageId="8" pageNumber="441">H. illatus</emphasis>
|
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</taxonomicName>
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is densely punctate to transversely rugulose in a narrow band while in
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<taxonomicName id="03E34D62445DE030FD1D9107FD02FD8E" authorityName="Ratcliffe" authorityYear="2001" box="[645,764,538,559]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="441" phylum="Arthropoda" rank="species" species="warneri">
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<emphasis id="F697EAF3445DE030FD1D9107FD02FD8E" box="[645,764,538,559]" italics="true" pageId="8" pageNumber="441">H. warneri</emphasis>
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</taxonomicName>
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the entire surface of the last sternite is sparsely punctate and lacks any distinctive sculpturing across the base.
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</paragraph>
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<paragraph id="C45C36E1445DE030FE8B9174FD4BFD38" blockId="8.[249,1082,271,665]" pageId="8" pageNumber="441">
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<materialsCitation id="748B3CBC445DE030FE8B9174FD4FFD38" collectionCode="V" pageId="8" pageNumber="441" specimenCount="1" typeStatus="holotype">
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The
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||
<typeStatus id="1B588843445DE030FEDD9174FE5EFDDF" box="[325,416,617,638]" pageId="8" pageNumber="441" type="holotype">holotype</typeStatus>
|
||
was taken at a U.
|
||
<collectionCode id="A2F2AE24445DE030FDFE9174FD84FDDF" box="[614,634,617,638]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="441" type="Museum">V</collectionCode>
|
||
.
|
||
<collectingMethod id="1DA24EF6445DE030FD1F9174FD17FDDF" box="[647,745,617,638]" pageId="8" pageNumber="441">light trap</collectingMethod>
|
||
in lowland deciduous forest in a mesic canyon at the end of dry season
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |