treatments-xml/data/44/38/87/443887FA6614FFB4FF6FFE47FE774C43.xml

<document id="1D19564809A1AA9722400EEF0ED11C3C" ID-DOI="10.11646/zootaxa.3741.4.2" ID-GBIF-Dataset="26b2bc4d-8855-43a3-b7ef-1541ed82c8ba" ID-ISSN="1175-5326" ID-Zenodo-Dep="247812" ID-ZooBank="6D837E20-C644-432D-9AAC-8F30D3B9854A" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460704005188" checkinUser="plazi" docAuthor="Behan-Pelletier, Valerie M. &amp; Walter, David E." docDate="2013" docId="443887FA6614FFB4FF6FFE47FE774C43" docLanguage="en" docName="zt03741p489.pdf" docOrigin="Zootaxa 3741 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tectoribates borealis Behan-Pelletier &amp; Walter, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="478" masterDocId="B801FF82661EFFA7FFF8FFFFFFD54B72" masterDocTitle="Phylogenetic relationships of Tectoribates: nymphal characters of new North American species place the genus in Tegoribatidae (Acari, Oribatida)" masterLastPageNumber="489" masterPageNumber="459" pageNumber="469" updateTime="1698356775934" updateUser="plazi">
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<mods:title id="D3069C949D741FE3CA720186E225D2C7">Phylogenetic relationships of Tectoribates: nymphal characters of new North American species place the genus in Tegoribatidae (Acari, Oribatida)</mods:title>
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<mods:namePart id="5A3BCD6017638C801FC072DF47B61083">Behan-Pelletier, Valerie M.</mods:namePart>
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<mods:namePart id="7A1882913395DA3EC387041385119504">Walter, David E.</mods:namePart>
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<mods:date id="43C0100D8A60BB0E46015AE5EEB05CAF">2013</mods:date>
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<treatment id="443887FA6614FFB4FF6FFE47FE774C43" ID-DOI="http://doi.org/10.5281/zenodo.5671483" ID-GBIF-Taxon="119583774" ID-Zenodo-Dep="5671483" LSID="urn:lsid:plazi:treatment:443887FA6614FFB4FF6FFE47FE774C43" httpUri="http://treatment.plazi.org/id/443887FA6614FFB4FF6FFE47FE774C43" lastPageId="19" lastPageNumber="478" pageId="10" pageNumber="469">
<subSubSection id="848B65676614FFADFF6FFE47FD084A81" pageId="10" pageNumber="469" type="nomenclature">
<paragraph id="CC2E36EC6614FFADFF6FFE47FE214AA0" blockId="10.[151,733,440,499]" box="[151,500,440,466]" pageId="10" pageNumber="469">
<heading id="976681806614FFADFF6FFE47FE214AA0" bold="true" box="[151,500,440,466]" fontSize="11" level="1" pageId="10" pageNumber="469" reason="1">
<emphasis id="FEE5EAFE6614FFADFF6FFE47FE214AA0" bold="true" box="[151,500,440,466]" pageId="10" pageNumber="469">
<taxonomicName id="0B914D6F6614FFADFF6FFE47FE5A4AA0" box="[151,399,440,466]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="469" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
<emphasis id="FEE5EAFE6614FFADFF6FFE47FE5A4AA0" bold="true" box="[151,399,440,466]" italics="true" pageId="10" pageNumber="469">Tectoribates borealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D657856614FFADFE6FFE47FE214AA0" box="[407,500,440,466]" pageId="10" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="CC2E36EC6614FFADFF6FFE25FD084A81" blockId="10.[151,733,440,499]" box="[151,733,474,499]" pageId="10" pageNumber="469">
(
<figureCitation id="54AA2A696614FFADFF67FE25FF3B4A81" box="[159,238,474,499]" captionStart="FIGURE 4" captionStartId="11.[151,250,1950,1972]" captionTargetBox="[286,1291,266,1939]" captionTargetId="figure@11.[285,1303,265,1940]" captionTargetPageId="11" captionText="FIGURE 4. Tectoribates borealis sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247816/files/figure.png" pageId="10" pageNumber="469">Figs. 4</figureCitation>
, 
<figureCitation id="54AA2A696614FFADFF02FE25FEDF4A81" box="[250,266,474,499]" captionStart="FIGURE 5" captionStartId="12.[151,250,1613,1635]" captionTargetBox="[217,1373,193,1596]" captionTargetId="figure@12.[211,1376,193,1600]" captionTargetPageId="12" captionText="FIGURE 5. Tectoribates borealis sp. nov., adult female, notogaster. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247817/files/figure.png" pageId="10" pageNumber="469">5</figureCitation>
, 
<figureCitation id="54AA2A696614FFADFEEEFE25FEF34A81" box="[278,294,474,499]" captionStart="FIGURE 6" captionStartId="13.[151,250,1919,1941]" captionTargetBox="[267,1320,196,1897]" captionTargetId="figure@13.[260,1326,193,1898]" captionTargetPageId="13" captionText="FIGURE 6. Tectoribates borealis sp. nov., adult female, ventral aspect, legs not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247818/files/figure.png" pageId="10" pageNumber="469">6</figureCitation>
, 7A–E, 8A–D, 9A–D, 10A–D, 11C–E)
</paragraph>
</subSubSection>
<subSubSection id="848B65676614FFADFF6FFDDDFAF44F41" pageId="10" pageNumber="469" type="materials_examined">
<paragraph id="CC2E36EC6614FFADFF6FFDDDFD314899" blockId="10.[151,1437,546,2014]" pageId="10" pageNumber="469">
<emphasis id="FEE5EAFE6614FFADFF6FFDDDFE504949" bold="true" box="[151,389,546,571]" pageId="10" pageNumber="469">Material examined.</emphasis>
<typeStatus id="132A884E6614FFADFE69FDDDFE284949" box="[401,509,546,571]" pageId="10" pageNumber="469" type="holotype">Holotype</typeStatus>
: adult female, 
<collectingCountry id="B486767C6614FFADFD48FDDDFCD24949" box="[688,775,546,571]" name="Canada" pageId="10" pageNumber="469">Canada</collectingCountry>
, Alberta, Cypress Hills Provincial Park, Firetower Area plateau, 4.8 km S. Elkwater, 
<date id="B82F102C6614FFADFE26FDB9FD5D492D" box="[478,648,582,607]" pageId="10" pageNumber="469" value="1978-07-22" valueMax="1978-07-29" valueMin="1978-07-22">22–29.vii.1978</date>
(E. E. Lindquist) from soil below elk and cow dung, deposited in the CNC, 
<typeStatus id="132A884E6614FFADFF27FD94FEDA49F1" box="[223,271,619,643]" pageId="10" pageNumber="469">type</typeStatus>
number 24149. 
<typeStatus id="132A884E6614FFADFE30FD93FDEF49F1" box="[456,570,620,643]" pageId="10" pageNumber="469" type="paratype">
<emphasis id="FEE5EAFE6614FFADFE30FD93FDEF49F1" box="[456,570,620,643]" italics="true" pageId="10" pageNumber="469">Paratypes</emphasis>
</typeStatus>
<specimenCount id="DA97FD656614FFADFDB9FD94FD4C49F1" box="[577,665,618,643]" pageId="10" pageNumber="469" type="adult">3 adults</specimenCount>
with same data as 
<typeStatus id="132A884E6614FFADFC88FD95FC0349F1" box="[880,982,618,643]" pageId="10" pageNumber="469" type="holotype">holotype</typeStatus>
; same location, 1 from moderately fresh buffalo dung, 1 from prairie herbage and soil below; Alberta: ABMI site OG-1650 (49.192046, -113.537695) 
<quantity id="0B699B096614FFADFA78FD71FF6E49B9" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="469" unit="km" value="15.0">15 km</quantity>
west of Cardson, ex grassland sod, 
<date id="B82F102C6614FFADFDB5FD4CFD0D49B9" box="[589,728,691,715]" pageId="10" pageNumber="469" value="2012-06-01">1 June 2012</date>
, 
<specimenCount id="DA97FD656614FFADFD1BFD4CFC8649B9" box="[739,851,690,715]" pageId="10" pageNumber="469" type="female">8 females</specimenCount>
; ABMI site 1642 NE (49.17302, -112.10323) northeast of Milk River, ex litter in agricultural field, 
<date id="B82F102C6614FFADFD4CFD28FC99499D" box="[692,844,727,751]" pageId="10" pageNumber="469" value="2012-05-31">31 May 2012</date>
, 
<specimenCount id="DA97FD656614FFADFCAFFD28FC13499D" box="[855,966,726,751]" pageId="10" pageNumber="469" type="female">2 females</specimenCount>
, 
<specimenCount id="DA97FD656614FFADFC29FD28FBF5499D" box="[977,1056,726,751]" pageId="10" pageNumber="469" type="male">1 male</specimenCount>
; ABMI site 1635 NW, (49.41763, -113.99814), 
<quantity id="0B699B096614FFADFECAFD05FEBA4861" box="[306,367,762,787]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="469" unit="km" value="5.0">5 km</quantity>
south-west Pincher Creek ex grass, sedge sod, 
<date id="B82F102C6614FFADFC5EFD05FBF54861" box="[934,1056,762,787]" pageId="10" pageNumber="469" value="2008-06-12">12.vi.2008</date>
, 
<specimenCount id="DA97FD656614FFADFBC8FD04FB4C4861" box="[1072,1177,762,787]" pageId="10" pageNumber="469" type="female">1 female</specimenCount>
; ABMI site 1411 SW (51.41224, -112.58736) 
<quantity id="0B699B096614FFADFE53FCE0FE334845" box="[427,486,798,823]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="469" unit="km" value="1.0">1 km</quantity>
NE 
<collectingCountry id="B486767C6614FFADFDE6FCE1FD544845" box="[542,641,798,823]" name="United Kingdom" pageId="10" pageNumber="469">Cambria</collectingCountry>
, ex litter in badlands, 
<specimenCount id="DA97FD656614FFADFC7AFCE0FC084845" box="[898,989,798,823]" pageId="10" pageNumber="469" type="male">2 males</specimenCount>
, 
<specimenCount id="DA97FD656614FFADFC13FCE0FB884845" box="[1003,1117,798,823]" pageId="10" pageNumber="469" type="female">2 females</specimenCount>
; ABMI site OG-1054-1NW (53.57580948, -112.461479) ex litter in open, disturbed site with herbs and ferns, 
<date id="B82F102C6614FFADFBD4FCBDFB164829" box="[1068,1219,834,859]" pageId="10" pageNumber="469" value="2011-05-25">25 May 2011</date>
, 
<specimenCount id="DA97FD656614FFADFB36FCBCFAC84829" box="[1230,1309,834,859]" pageId="10" pageNumber="469" type="male">1 male</specimenCount>
; 
<specimenCount id="DA97FD656614FFADFAD0FCBCFA4D4829" box="[1320,1432,834,859]" pageId="10" pageNumber="469" type="female">2 females</specimenCount>
; ABMI site 1362 SE, 
<date id="B82F102C6614FFADFE79FC98FDCD480D" box="[385,536,871,895]" pageId="10" pageNumber="469" value="2012-05-24">24 May 2012</date>
, ABMI site 1362 NE (51.75812, -112.46145) 
<quantity id="0B699B096614FFADFBE1FC98FB8A480D" box="[1049,1119,870,895]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="469" unit="km" value="40.0">40 km</quantity>
north-east of Drumheller ex litter in field, 
<date id="B82F102C6614FFADFECDFC74FE0548D1" box="[309,464,907,931]" pageId="10" pageNumber="469" value="2012-05-30">30 May 2012</date>
, 
<specimenCount id="DA97FD656614FFADFE25FC74FD8F48D1" box="[477,602,906,931]" pageId="10" pageNumber="469" type="female">11 females</specimenCount>
, 2 tritonymphs; ABMI site 2 SW (60.00878, -112.14697) ex bog with an understory of labrador tea, sphagnum, lichen in Wood Buffalo National Park, 
<date id="B82F102C6614FFADFBEBFC50FB7A48B5" box="[1043,1199,943,967]" pageId="10" pageNumber="469" value="2009-06-10">10 June 2009</date>
, 
<specimenCount id="DA97FD656614FFADFB44FC50FAF648B5" box="[1212,1315,942,967]" pageId="10" pageNumber="469" type="female">1 female</specimenCount>
. 
<typeStatus id="132A884E6614FFADFAD7FC50FA4948B5" box="[1327,1436,943,967]" pageId="10" pageNumber="469" type="paratype">Paratypes</typeStatus>
deposited in the PMAE.IZ, CNC, USNM, and RNC.
</paragraph>
<paragraph id="CC2E36EC6614FFADFF3FFC09FAF44F41" blockId="10.[151,1437,546,2014]" pageId="10" pageNumber="469">
<emphasis id="FEE5EAFE6614FFADFF3FFC09FE2E4F7D" bold="true" box="[199,507,1014,1039]" pageId="10" pageNumber="469">Other material examined.</emphasis>
Additional individuals from ABMI site 1411 SW (see above) and 1438 (51.10179, -111.48718) ex grass and sedge sod, 
<date id="B82F102C6614FFADFDCAFBE4FD694F41" box="[562,700,1051,1075]" pageId="10" pageNumber="469" value="2011-06-09">9 June 2011</date>
, were studied under the scanning electron microscope.
</paragraph>
</subSubSection>
<subSubSection id="848B65676614FFADFF3FFBC1FF6F4EB0" pageId="10" pageNumber="469" type="diagnosis">
<paragraph id="CC2E36EC6614FFADFF3FFBC1FF6F4EB0" blockId="10.[151,1437,546,2014]" pageId="10" pageNumber="469">
<emphasis id="FEE5EAFE6614FFADFF3FFBC1FEEF4F25" bold="true" box="[199,314,1086,1111]" pageId="10" pageNumber="469">Diagnosis</emphasis>
. Adult. Total length 325–340. Integument with foveae between genital and anal plates and posterolateral to genital plates. Rostral margin smooth to dentate, with medial V to U-shaped indentation. Notogaster anteriorly with distinct hexagonal pattern, visible in transmitted light. Notogastral porose areas 
<quantity id="0B699B096614FFADFABFFB79FA494FED" box="[1351,1436,1158,1183]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.54" metricValueMin="1.524" pageId="10" pageNumber="469" unit="in" value="8.0" valueMax="10.0" valueMin="6.0">6–10 in</quantity>
diameter. Notogastral setae barbed, 16–24, with seta 
<emphasis id="FEE5EAFE6614FFADFD12FB53FD234FB1" box="[746,758,1196,1219]" italics="true" pageId="10" pageNumber="469">c</emphasis>
longest; 
<emphasis id="FEE5EAFE6614FFADFCA5FB55FCAC4FB1" box="[861,889,1194,1219]" italics="true" pageId="10" pageNumber="469">lm</emphasis>
medial to Aa and 
<emphasis id="FEE5EAFE6614FFADFBB1FB55FB8A4FB1" box="[1097,1119,1194,1219]" italics="true" pageId="10" pageNumber="469">lp</emphasis>
anteromedial to A1; mutual distance of 
<emphasis id="FEE5EAFE6614FFADFEE3FB31FE814F99" box="[283,340,1230,1259]" italics="true" pageId="10" pageNumber="469">h1–h1</emphasis>
about 18–28, of 
<emphasis id="FEE5EAFE6614FFADFDE1FB30FD864F99" box="[537,595,1231,1259]" italics="true" pageId="10" pageNumber="469">p1–p1</emphasis>
37–40. Lamella about 71 long, of which cusp about 44; about 19 wide at level of insertion of seta 
<emphasis id="FEE5EAFE6614FFADFE51FB0AFE684E7C" box="[425,445,1269,1294]" italics="true" pageId="10" pageNumber="469">le</emphasis>
. Lamellar cusps effacing on prodorsum about 14 from base of prodorsum. Lamellar cleft extending posteriorly about 10 from region of fusion of lamella to prodorsum; distance between lamellae in this region (width of cleft) about 3. Lamellar cusp with two short tapered dentes, medial dens about 14 long, lateral dens shorter, about 12 long. Lamellar seta arising anteriorly on cusp, 38–44 long. Bothridial seta club-shaped, 51– 55. Seta 
<emphasis id="FEE5EAFE6614FFADFF01FA79FEC44EEF" box="[249,273,1414,1437]" italics="true" pageId="10" pageNumber="469">ex</emphasis>
about 15. Tutorium 71–81 long, of which cusp 28–32. Custodium about 15 long. Epimeral seta 
<emphasis id="FEE5EAFE6614FFADFAA8FA79FABF4EEC" box="[1360,1386,1414,1438]" italics="true" pageId="10" pageNumber="469">1c</emphasis>
30– 33.
</paragraph>
</subSubSection>
<subSubSection id="848B65676614FFB4FF3FFA32FE434F2F" lastPageId="19" lastPageNumber="478" pageId="10" pageNumber="469" type="description">
<paragraph id="CC2E36EC6614FFADFF3FFA32FEAC4D78" blockId="10.[151,1437,546,2014]" pageId="10" pageNumber="469">
<emphasis id="FEE5EAFE6614FFADFF3FFA32FE8C4E94" bold="true" box="[199,345,1485,1510]" pageId="10" pageNumber="469">Description.</emphasis>
Adults. 
<emphasis id="FEE5EAFE6614FFADFE4FFA31FD914E97" box="[439,580,1486,1509]" italics="true" pageId="10" pageNumber="469">Dimensions:</emphasis>
Total length: females (
<emphasis id="FEE5EAFE6614FFADFCBEFA30FC814E94" box="[838,852,1487,1510]" italics="true" pageId="10" pageNumber="469">n</emphasis>
= 10) 324 (307–340). Notogastral width: females (
<emphasis id="FEE5EAFE6614FFADFA76FA30FA494E94" box="[1422,1436,1487,1510]" italics="true" pageId="10" pageNumber="469">n</emphasis>
= 5) 205 (200–210).
</paragraph>
<paragraph id="CC2E36EC6614FFADFF3FF9E8FAD64DE8" blockId="10.[151,1437,546,2014]" pageId="10" pageNumber="469">
<emphasis id="FEE5EAFE6614FFADFF3FF9E8FE9B4D5C" box="[199,334,1559,1582]" italics="true" pageId="10" pageNumber="469">Integument:</emphasis>
Prodorsum, notogaster, pedotecta, anal plates, genital plates, coxisternum and mentum micropunctate (
<figureCitation id="54AA2A696614FFADFEB2F9C6FE734D20" box="[330,422,1593,1618]" captionStart="FIGURES 9 A – D" captionStartId="16.[151,264,1636,1658]" captionTargetBox="[151,1436,193,1614]" captionTargetId="figure@16.[151,1436,193,1615]" captionTargetPageId="16" captionText="FIGURES 9 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada; A, ventral region showing discidium, custodium and foveae on lateral of ventral plate; B, detail of foveae on notogaster, showing foveae mostly plugged with cerotegument; C, prodorsum, anterolateral aspect, with arrow to small teeth lateral to solenidia on tibia I; D, rostrum, laterofrontal aspect, with arrow to proximal solenidion on tarsus II. Scale bar: A, B, C = 10 µm, D = 1 µm." httpUri="https://zenodo.org/record/247820/files/figure.png" pageId="10" pageNumber="469">Fig. 9B</figureCitation>
); foveate between genital and anal plates and posterolateral of genital plates (
<figureCitation id="54AA2A696614FFADFAC4F9C6FA424D20" box="[1340,1431,1593,1618]" captionStart="FIGURES 8 A – D" captionStartId="15.[151,264,1836,1858]" captionTargetBox="[151,1436,193,1814]" captionTargetId="figure@15.[151,1436,193,1815]" captionTargetPageId="15" captionText="FIGURES 8 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, prodorsum, with arrow to small teeth lateral to solenidia on tarsus I, and arrow to exobothridial seta; C, habitus ventral, with arrow to foveae between genital and anal plates; D, gnathosoma, with arrow to band of tubercles on anterior of coxisternal region. Scale bar: A, B, C = 20 µm, D = 10 µm." httpUri="https://zenodo.org/record/247819/files/figure.png" pageId="10" pageNumber="469">Fig. 8C</figureCitation>
, arrow). Lamellae, pedotectum, mentum, coxisternal region with fine irregular striae (
<figureCitation id="54AA2A696614FFADFB93F9A2FB234D04" box="[1131,1270,1629,1654]" captionStart="FIGURES 9 A – D" captionStartId="16.[151,264,1636,1658]" captionTargetBox="[151,1436,193,1614]" captionTargetId="figure@16.[151,1436,193,1615]" captionTargetPageId="16" captionText="FIGURES 9 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada; A, ventral region showing discidium, custodium and foveae on lateral of ventral plate; B, detail of foveae on notogaster, showing foveae mostly plugged with cerotegument; C, prodorsum, anterolateral aspect, with arrow to small teeth lateral to solenidia on tibia I; D, rostrum, laterofrontal aspect, with arrow to proximal solenidion on tarsus II. Scale bar: A, B, C = 10 µm, D = 1 µm." httpUri="https://zenodo.org/record/247820/files/figure.png" pageId="10" pageNumber="469">Figs. 9A, C</figureCitation>
); tutorial cusp with distinct striae (
<figureCitation id="54AA2A696614FFADFE8DF97EFE184DE8" box="[373,461,1665,1690]" captionStart="FIGURES 9 A – D" captionStartId="16.[151,264,1636,1658]" captionTargetBox="[151,1436,193,1614]" captionTargetId="figure@16.[151,1436,193,1615]" captionTargetPageId="16" captionText="FIGURES 9 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada; A, ventral region showing discidium, custodium and foveae on lateral of ventral plate; B, detail of foveae on notogaster, showing foveae mostly plugged with cerotegument; C, prodorsum, anterolateral aspect, with arrow to small teeth lateral to solenidia on tibia I; D, rostrum, laterofrontal aspect, with arrow to proximal solenidion on tarsus II. Scale bar: A, B, C = 10 µm, D = 1 µm." httpUri="https://zenodo.org/record/247820/files/figure.png" pageId="10" pageNumber="469">Fig. 9C</figureCitation>
). Rostrum with short distinct striae close to medial indentation (
<figureCitation id="54AA2A696614FFADFB64F97EFB234DE8" box="[1180,1270,1665,1690]" captionStart="FIGURES 9 A – D" captionStartId="16.[151,264,1636,1658]" captionTargetBox="[151,1436,193,1614]" captionTargetId="figure@16.[151,1436,193,1615]" captionTargetPageId="16" captionText="FIGURES 9 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada; A, ventral region showing discidium, custodium and foveae on lateral of ventral plate; B, detail of foveae on notogaster, showing foveae mostly plugged with cerotegument; C, prodorsum, anterolateral aspect, with arrow to small teeth lateral to solenidia on tibia I; D, rostrum, laterofrontal aspect, with arrow to proximal solenidion on tarsus II. Scale bar: A, B, C = 10 µm, D = 1 µm." httpUri="https://zenodo.org/record/247820/files/figure.png" pageId="10" pageNumber="469">Fig. 9D</figureCitation>
).
</paragraph>
<paragraph id="CC2E36EC6614FFACFF3FF95AFC434BA6" blockId="10.[151,1437,546,2014]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="470" pageId="10" pageNumber="469">
<emphasis id="FEE5EAFE6614FFADFF3FF95AFE984DCC" box="[199,333,1701,1726]" italics="true" pageId="10" pageNumber="469">Prodorsum:</emphasis>
Rostrum with distinct, convex medial crest proximally, effacing distally, rostral margin smooth to weakly dentate, with medial V to U-shaped indentation (
<figureCitation id="54AA2A696614FFADFCD5F936FC6C4D90" box="[813,953,1737,1762]" captionStart="FIGURES 9 A – D" captionStartId="16.[151,264,1636,1658]" captionTargetBox="[151,1436,193,1614]" captionTargetId="figure@16.[151,1436,193,1615]" captionTargetPageId="16" captionText="FIGURES 9 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada; A, ventral region showing discidium, custodium and foveae on lateral of ventral plate; B, detail of foveae on notogaster, showing foveae mostly plugged with cerotegument; C, prodorsum, anterolateral aspect, with arrow to small teeth lateral to solenidia on tibia I; D, rostrum, laterofrontal aspect, with arrow to proximal solenidion on tarsus II. Scale bar: A, B, C = 10 µm, D = 1 µm." httpUri="https://zenodo.org/record/247820/files/figure.png" pageId="10" pageNumber="469">Figs. 9C, D</figureCitation>
). Rostrum with deep, cerotegument-filled groove, ventral to seta 
<emphasis id="FEE5EAFE6614FFADFE59F910FE6D4C74" box="[417,440,1775,1798]" italics="true" pageId="10" pageNumber="469">ro</emphasis>
, (
<figureCitation id="54AA2A696614FFADFE29F912FDF94C74" box="[465,556,1773,1798]" captionStart="FIGURES 9 A – D" captionStartId="16.[151,264,1636,1658]" captionTargetBox="[151,1436,193,1614]" captionTargetId="figure@16.[151,1436,193,1615]" captionTargetPageId="16" captionText="FIGURES 9 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada; A, ventral region showing discidium, custodium and foveae on lateral of ventral plate; B, detail of foveae on notogaster, showing foveae mostly plugged with cerotegument; C, prodorsum, anterolateral aspect, with arrow to small teeth lateral to solenidia on tibia I; D, rostrum, laterofrontal aspect, with arrow to proximal solenidion on tarsus II. Scale bar: A, B, C = 10 µm, D = 1 µm." httpUri="https://zenodo.org/record/247820/files/figure.png" pageId="10" pageNumber="469">Fig. 9C</figureCitation>
). Lamella about 71 long, of which cusp about 44; about 19 wide at level of insertion of seta 
<emphasis id="FEE5EAFE6614FFADFE98F8EEFEA14C58" box="[352,372,1809,1834]" italics="true" pageId="10" pageNumber="469">le</emphasis>
. Lamellar cusps effacing on prodorsum about 14 from base of prodorsum. Lamellar cleft extending posteriorly about 10 from region of fusion of lamella to prodorsum; distance between lamellae in this region (width of cleft) about 3. Lamellar cusp with two short, tapered dentes, medial dens about 14 long, lateral dens shorter, about 12 long (
<figureCitation id="54AA2A696614FFADFE22F882FDFF4CE4" box="[474,554,1917,1942]" captionStart="FIGURE 4" captionStartId="11.[151,250,1950,1972]" captionTargetBox="[286,1291,266,1939]" captionTargetId="figure@11.[285,1303,265,1940]" captionTargetPageId="11" captionText="FIGURE 4. Tectoribates borealis sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247816/files/figure.png" pageId="10" pageNumber="469">Figs. 4</figureCitation>
, 
<figureCitation id="54AA2A696614FFADFDCFF881FDAB4CE4" box="[567,638,1918,1942]" captionStart="FIGURES 8 A – D" captionStartId="15.[151,264,1836,1858]" captionTargetBox="[151,1436,193,1814]" captionTargetId="figure@15.[151,1436,193,1815]" captionTargetPageId="15" captionText="FIGURES 8 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, prodorsum, with arrow to small teeth lateral to solenidia on tarsus I, and arrow to exobothridial seta; C, habitus ventral, with arrow to foveae between genital and anal plates; D, gnathosoma, with arrow to band of tubercles on anterior of coxisternal region. Scale bar: A, B, C = 20 µm, D = 10 µm." httpUri="https://zenodo.org/record/247819/files/figure.png" pageId="10" pageNumber="469">8A, B</figureCitation>
). Seta 
<emphasis id="FEE5EAFE6614FFADFD34F880FD364CE4" box="[716,739,1919,1942]" italics="true" pageId="10" pageNumber="469">ro</emphasis>
51–58 long, strongly barbed along length, acuminate, curved anteromedially. Lamellar seta arising anteriorly on cusp, 38–44 long. heavily barbed, acuminate distally. Seta 
<emphasis id="FEE5EAFE6614FFADFA7EF85DFA494CCB" box="[1414,1436,1954,1977]" italics="true" pageId="10" pageNumber="469">in</emphasis>
67–76 long, barbed (subequally to 
<emphasis id="FEE5EAFE6614FFADFDDCF83AFDED4CAC" box="[548,568,1989,2014]" italics="true" pageId="10" pageNumber="469">le</emphasis>
), acuminate; mutual distance about 28 (
<figureCitation id="54AA2A696614FFADFBFBF83AFB9F4CAC" box="[1027,1098,1989,2014]" captionStart="FIGURE 4" captionStartId="11.[151,250,1950,1972]" captionTargetBox="[286,1291,266,1939]" captionTargetId="figure@11.[285,1303,265,1940]" captionTargetPageId="11" captionText="FIGURE 4. Tectoribates borealis sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247816/files/figure.png" pageId="10" pageNumber="469">Fig. 4</figureCitation>
). Bothridial seta club-shaped, 51–55, long, with barbed, rounded head; head directed anteromedially (
<figureCitation id="54AA2A696615FFACFC2DFF68FBF44BC2" box="[981,1057,151,176]" captionStart="FIGURE 4" captionStartId="11.[151,250,1950,1972]" captionTargetBox="[286,1291,266,1939]" captionTargetId="figure@11.[285,1303,265,1940]" captionTargetPageId="11" captionText="FIGURE 4. Tectoribates borealis sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247816/files/figure.png" pageId="11" pageNumber="470">Figs 4</figureCitation>
, 
<figureCitation id="54AA2A696615FFACFBC9FF67FB804BC2" box="[1073,1109,152,176]" captionStart="FIGURES 8 A – D" captionStartId="15.[151,264,1836,1858]" captionTargetBox="[151,1436,193,1814]" captionTargetId="figure@15.[151,1436,193,1815]" captionTargetPageId="15" captionText="FIGURES 8 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, prodorsum, with arrow to small teeth lateral to solenidia on tarsus I, and arrow to exobothridial seta; C, habitus ventral, with arrow to foveae between genital and anal plates; D, gnathosoma, with arrow to band of tubercles on anterior of coxisternal region. Scale bar: A, B, C = 20 µm, D = 10 µm." httpUri="https://zenodo.org/record/247819/files/figure.png" pageId="11" pageNumber="470">8A</figureCitation>
, 
<figureCitation id="54AA2A696615FFACFB9CFF67FB424BC2" box="[1124,1175,152,176]" captionStart="FIGURES 11 A – F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A – F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C – E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="11" pageNumber="470">11D</figureCitation>
). Seta 
<emphasis id="FEE5EAFE6615FFACFB11FF66FAD74BC2" box="[1257,1282,153,176]" italics="true" pageId="11" pageNumber="470">ex</emphasis>
thin, barbed, about 15 (
<figureCitation id="54AA2A696615FFACFEFFFF43FE8A4BA6" box="[263,351,188,213]" captionStart="FIGURES 8 A – D" captionStartId="15.[151,264,1836,1858]" captionTargetBox="[151,1436,193,1814]" captionTargetId="figure@15.[151,1436,193,1815]" captionTargetPageId="15" captionText="FIGURES 8 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, prodorsum, with arrow to small teeth lateral to solenidia on tarsus I, and arrow to exobothridial seta; C, habitus ventral, with arrow to foveae between genital and anal plates; D, gnathosoma, with arrow to band of tubercles on anterior of coxisternal region. Scale bar: A, B, C = 20 µm, D = 10 µm." httpUri="https://zenodo.org/record/247819/files/figure.png" pageId="11" pageNumber="470">Fig. 8B</figureCitation>
, arrow). Dorsophragmata separate at base (
<figureCitation id="54AA2A696615FFACFCBBFF43FC5C4BA6" box="[835,905,188,213]" captionStart="FIGURE 4" captionStartId="11.[151,250,1950,1972]" captionTargetBox="[286,1291,266,1939]" captionTargetId="figure@11.[285,1303,265,1940]" captionTargetPageId="11" captionText="FIGURE 4. Tectoribates borealis sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247816/files/figure.png" pageId="11" pageNumber="470">Fig. 4</figureCitation>
).
</paragraph>
<caption id="98EE66646615FFACFF6FF861FEB84CA1" httpUri="https://zenodo.org/record/247816/files/figure.png" pageId="11" pageNumber="470" targetBox="[286,1291,266,1939]" targetPageId="11">
<paragraph id="CC2E36EC6615FFACFF6FF861FEB84CA1" blockId="11.[151,1436,1950,2003]" pageId="11" pageNumber="470">
<emphasis id="FEE5EAFE6615FFACFF6FF861FEC74CC6" bold="true" box="[151,274,1950,1972]" pageId="11" pageNumber="470">FIGURE 4.</emphasis>
<taxonomicName id="0B914D6F6615FFACFEE0F861FE384CC6" box="[280,493,1950,1972]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="470" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
<emphasis id="FEE5EAFE6615FFACFEE0F861FE384CC6" box="[280,493,1950,1972]" italics="true" pageId="11" pageNumber="470">Tectoribates borealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D657856615FFACFE0BF861FD974CC6" box="[499,578,1950,1972]" pageId="11" pageNumber="470" rank="species">
<emphasis id="FEE5EAFE6615FFACFE0BF861FD974CC6" bold="true" box="[499,578,1950,1972]" pageId="11" pageNumber="470">sp. nov.</emphasis>
</taxonomicNameLabel>
, adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="98EE66646612FFABFF6FF9B2FBC74D11" box="[151,1042,1613,1635]" httpUri="https://zenodo.org/record/247817/files/figure.png" pageId="12" pageNumber="471" targetBox="[217,1373,193,1596]" targetPageId="12">
<paragraph id="CC2E36EC6612FFABFF6FF9B2FBC74D11" blockId="12.[151,1042,1613,1635]" box="[151,1042,1613,1635]" pageId="12" pageNumber="471">
<emphasis id="FEE5EAFE6612FFABFF6FF9B2FEC74D11" bold="true" box="[151,274,1613,1635]" pageId="12" pageNumber="471">FIGURE 5.</emphasis>
<taxonomicName id="0B914D6F6612FFABFEE1F9B2FE384D11" box="[281,493,1613,1635]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="471" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
<emphasis id="FEE5EAFE6612FFABFEE1F9B2FE384D11" box="[281,493,1613,1635]" italics="true" pageId="12" pageNumber="471">Tectoribates borealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D657856612FFABFE0CF9B2FD914D11" box="[500,580,1613,1635]" pageId="12" pageNumber="471" rank="species">
<emphasis id="FEE5EAFE6612FFABFE0CF9B2FD914D11" bold="true" box="[500,580,1613,1635]" pageId="12" pageNumber="471">sp. nov.</emphasis>
</taxonomicNameLabel>
, adult female, notogaster. Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="CC2E36EC6612FFABFF3FF975FAA44DB5" blockId="12.[151,1436,1674,2029]" pageId="12" pageNumber="471">
<emphasis id="FEE5EAFE6612FFABFF3FF975FDDF4DD1" box="[199,522,1674,1699]" italics="true" pageId="12" pageNumber="471">Lateral region of podosoma:</emphasis>
Tutorium 71–81 long, of which cusp 28–32. Custodium about 15 long (
<figureCitation id="54AA2A696612FFABFACDF975FA424DD1" box="[1333,1431,1674,1699]" captionStart="FIGURES 8 A – D" captionStartId="15.[151,264,1836,1858]" captionTargetBox="[151,1436,193,1814]" captionTargetId="figure@15.[151,1436,193,1815]" captionTargetPageId="15" captionText="FIGURES 8 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, prodorsum, with arrow to small teeth lateral to solenidia on tarsus I, and arrow to exobothridial seta; C, habitus ventral, with arrow to foveae between genital and anal plates; D, gnathosoma, with arrow to band of tubercles on anterior of coxisternal region. Scale bar: A, B, C = 20 µm, D = 10 µm." httpUri="https://zenodo.org/record/247819/files/figure.png" pageId="12" pageNumber="471">Figs. 8C</figureCitation>
, 
<figureCitation id="54AA2A696612FFABFF6FF950FF694DB5" box="[151,188,1711,1735]" captionStart="FIGURES 9 A – D" captionStartId="16.[151,264,1636,1658]" captionTargetBox="[151,1436,193,1614]" captionTargetId="figure@16.[151,1436,193,1615]" captionTargetPageId="16" captionText="FIGURES 9 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada; A, ventral region showing discidium, custodium and foveae on lateral of ventral plate; B, detail of foveae on notogaster, showing foveae mostly plugged with cerotegument; C, prodorsum, anterolateral aspect, with arrow to small teeth lateral to solenidia on tibia I; D, rostrum, laterofrontal aspect, with arrow to proximal solenidion on tarsus II. Scale bar: A, B, C = 10 µm, D = 1 µm." httpUri="https://zenodo.org/record/247820/files/figure.png" pageId="12" pageNumber="471">9A</figureCitation>
). Porose area Al about 
<quantity id="0B699B096612FFABFE39F950FE394DB5" box="[449,492,1710,1735]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="12" pageNumber="471" unit="in" value="8.0">8 in</quantity>
diameter (difficult to see because of proximal position on lateral of prodorsum).
</paragraph>
<paragraph id="CC2E36EC6612FFABFF3FF92BFE484C28" blockId="12.[151,1436,1674,2029]" pageId="12" pageNumber="471">
<emphasis id="FEE5EAFE6612FFABFF3FF92BFE994D99" box="[199,332,1748,1771]" italics="true" pageId="12" pageNumber="471">Notogaster:</emphasis>
Longer than wide: 1.2:1. Anteriorly with distinct hexagonal pattern (
<figureCitation id="54AA2A696612FFABFB92F92DFB6E4D99" box="[1130,1211,1746,1771]" captionStart="FIGURE 4" captionStartId="11.[151,250,1950,1972]" captionTargetBox="[286,1291,266,1939]" captionTargetId="figure@11.[285,1303,265,1940]" captionTargetPageId="11" captionText="FIGURE 4. Tectoribates borealis sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247816/files/figure.png" pageId="12" pageNumber="471">Figs. 4</figureCitation>
, 
<figureCitation id="54AA2A696612FFABFB31F92DFB0C4D99" box="[1225,1241,1746,1771]" captionStart="FIGURE 5" captionStartId="12.[151,250,1613,1635]" captionTargetBox="[217,1373,193,1596]" captionTargetId="figure@12.[211,1376,193,1600]" captionTargetPageId="12" captionText="FIGURE 5. Tectoribates borealis sp. nov., adult female, notogaster. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247817/files/figure.png" pageId="12" pageNumber="471">5</figureCitation>
, 
<figureCitation id="54AA2A696612FFABFB10F92CFACD4D99" box="[1256,1304,1747,1771]" captionStart="FIGURES 11 A – F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A – F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C – E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="12" pageNumber="471">11C</figureCitation>
). Octotaxic system expressed as porose areas 
<quantity id="0B699B096612FFABFDC4F909FD4F4C7D" box="[572,666,1782,1807]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.54" metricValueMin="1.524" pageId="12" pageNumber="471" unit="in" value="8.0" valueMax="10.0" valueMin="6.0">6–10 in</quantity>
diameter. Notogastral setae barbed, 16–24, with 
<emphasis id="FEE5EAFE6612FFABFB04F907FADD4C7D" box="[1276,1288,1784,1807]" italics="true" pageId="12" pageNumber="471">c</emphasis>
longest; 
<emphasis id="FEE5EAFE6612FFABFA78F909FA494C7D" box="[1408,1436,1782,1807]" italics="true" pageId="12" pageNumber="471">lm</emphasis>
posteromedial to Aa and 
<emphasis id="FEE5EAFE6612FFABFE48F8E5FE134C41" box="[432,454,1818,1843]" italics="true" pageId="12" pageNumber="471">lp</emphasis>
anterior to anteromedial of A1 (
<figureCitation id="54AA2A696612FFABFCC8F8E5FCAC4C41" box="[816,889,1818,1843]" captionStart="FIGURE 4" captionStartId="11.[151,250,1950,1972]" captionTargetBox="[286,1291,266,1939]" captionTargetId="figure@11.[285,1303,265,1940]" captionTargetPageId="11" captionText="FIGURE 4. Tectoribates borealis sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247816/files/figure.png" pageId="12" pageNumber="471">Figs 4</figureCitation>
, 
<figureCitation id="54AA2A696612FFABFC7DF8E5FC404C41" box="[901,917,1818,1843]" captionStart="FIGURE 5" captionStartId="12.[151,250,1613,1635]" captionTargetBox="[217,1373,193,1596]" captionTargetId="figure@12.[211,1376,193,1600]" captionTargetPageId="12" captionText="FIGURE 5. Tectoribates borealis sp. nov., adult female, notogaster. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247817/files/figure.png" pageId="12" pageNumber="471">5</figureCitation>
); mutual distance of 
<emphasis id="FEE5EAFE6612FFABFB78F8E5FB6F4C45" box="[1152,1210,1818,1847]" italics="true" pageId="12" pageNumber="471">h1–h1</emphasis>
about 24–28, of 
<emphasis id="FEE5EAFE6612FFABFA80F8E3FF784C2C" italics="true" pageId="12" pageNumber="471">p1– p1</emphasis>
about 37 (
<figureCitation id="54AA2A696612FFABFEDCF8BEFEA64C28" box="[292,371,1857,1882]" captionStart="FIGURE 4" captionStartId="11.[151,250,1950,1972]" captionTargetBox="[286,1291,266,1939]" captionTargetId="figure@11.[285,1303,265,1940]" captionTargetPageId="11" captionText="FIGURE 4. Tectoribates borealis sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247816/files/figure.png" pageId="12" pageNumber="471">Figs. 4</figureCitation>
, 
<figureCitation id="54AA2A696612FFABFE87F8BEFE5A4C28" box="[383,399,1857,1882]" captionStart="FIGURE 5" captionStartId="12.[151,250,1613,1635]" captionTargetBox="[217,1373,193,1596]" captionTargetId="figure@12.[211,1376,193,1600]" captionTargetPageId="12" captionText="FIGURE 5. Tectoribates borealis sp. nov., adult female, notogaster. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247817/files/figure.png" pageId="12" pageNumber="471">5</figureCitation>
).
</paragraph>
<paragraph id="CC2E36EC6612FFABFF3FF897FECF4C9F" blockId="12.[151,1436,1674,2029]" pageId="12" pageNumber="471">
<emphasis id="FEE5EAFE6612FFABFF3FF897FEAF4CF2" box="[199,378,1896,1921]" italics="true" pageId="12" pageNumber="471">Ventral Region:</emphasis>
Epimeral seta 
<emphasis id="FEE5EAFE6612FFABFDD3F896FD934CF3" box="[555,582,1897,1921]" italics="true" pageId="12" pageNumber="471">1c</emphasis>
more heavily barbed, and thicker than other epimeral setae, 30–33; other epimeral setae 10–16. Genital, aggenital, anal and adanal setae smooth; genital setae about 10, other setae about 7. Genital setae 3+3 or 2+4 (
<figureCitation id="54AA2A696612FFABFE3AF84FFDD24CBB" box="[450,519,1968,1993]" captionStart="FIGURE 6" captionStartId="13.[151,250,1919,1941]" captionTargetBox="[267,1320,196,1897]" captionTargetId="figure@13.[260,1326,193,1898]" captionTargetPageId="13" captionText="FIGURE 6. Tectoribates borealis sp. nov., adult female, ventral aspect, legs not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247818/files/figure.png" pageId="12" pageNumber="471">Fig. 6</figureCitation>
). Lyrifissure 
<emphasis id="FEE5EAFE6612FFABFD58F84FFD114CBB" box="[672,708,1968,1993]" italics="true" pageId="12" pageNumber="471">iad</emphasis>
anterolateral of anal plate. Postanal porose area (not illustrated) about 
<date id="B82F102C6612FFABFF24F82AFF2A4C9E" box="[220,255,2005,2029]" pageId="12" pageNumber="471">9 x</date>
5.
</paragraph>
<caption id="98EE66646613FFAAFF6FF880FB284CE7" box="[151,1277,1919,1941]" httpUri="https://zenodo.org/record/247818/files/figure.png" pageId="13" pageNumber="472" targetBox="[267,1320,196,1897]" targetPageId="13">
<paragraph id="CC2E36EC6613FFAAFF6FF880FB284CE7" blockId="13.[151,1277,1919,1941]" box="[151,1277,1919,1941]" pageId="13" pageNumber="472">
<emphasis id="FEE5EAFE6613FFAAFF6FF880FEC74CE7" bold="true" box="[151,274,1919,1941]" pageId="13" pageNumber="472">FIGURE 6.</emphasis>
<taxonomicName id="0B914D6F6613FFAAFEE1F880FE384CE7" box="[281,493,1919,1941]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="472" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
<emphasis id="FEE5EAFE6613FFAAFEE1F880FE384CE7" box="[281,493,1919,1941]" italics="true" pageId="13" pageNumber="472">Tectoribates borealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D657856613FFAAFE0CF880FD914CE7" box="[500,580,1919,1941]" pageId="13" pageNumber="472" rank="species">
<emphasis id="FEE5EAFE6613FFAAFE0CF880FD914CE7" bold="true" box="[500,580,1919,1941]" pageId="13" pageNumber="472">sp. nov.</emphasis>
</taxonomicNameLabel>
, adult female, ventral aspect, legs not illustrated. Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="CC2E36EC6610FFA9FF6FF880FD884CA7" blockId="14.[151,1436,1919,2005]" pageId="14" pageNumber="473">
<emphasis id="FEE5EAFE6610FFA9FF6FF880FEDD4CE7" bold="true" box="[151,264,1919,1941]" pageId="14" pageNumber="473">FIGURES</emphasis>
7A–E. 
<taxonomicName id="0B914D6F6610FFA9FEA1F880FDFA4CE7" box="[345,559,1919,1941]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="473" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
<emphasis id="FEE5EAFE6610FFA9FEA1F880FDFA4CE7" box="[345,559,1919,1941]" italics="true" pageId="14" pageNumber="473">Tectoribates borealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D657856610FFA9FDC0F880FD5F4CE7" box="[568,650,1919,1941]" pageId="14" pageNumber="473" rank="species">
<emphasis id="FEE5EAFE6610FFA9FDC0F880FD5F4CE7" bold="true" box="[568,650,1919,1941]" pageId="14" pageNumber="473">sp. nov.</emphasis>
</taxonomicNameLabel>
, adult female, legs I–IV and palp, all antiaxial aspect. Leg I, trochanter not illustrated. Leg II, trochanter not illustrated, with arrow to ventral tooth on genu and to proximal solenidion 
<emphasis id="FEE5EAFE6610FFA9FAD9F860FAEF4CC5" box="[1313,1338,1951,1975]" italics="true" pageId="14" pageNumber="473">ω1</emphasis>
. Leg III, trochanter not illustrated. Scale bar = 20 µm.
</paragraph>
<caption id="98EE66646611FFA8FF6FF8D3FCB44CEC" httpUri="https://zenodo.org/record/247819/files/figure.png" pageId="15" pageNumber="474" targetBox="[151,1436,193,1814]" targetPageId="15">
<paragraph id="CC2E36EC6611FFA8FF6FF8D3FCB44CEC" blockId="15.[151,1436,1836,1950]" pageId="15" pageNumber="474">
<emphasis id="FEE5EAFE6611FFA8FF6FF8D3FE864C30" bold="true" box="[151,339,1836,1858]" pageId="15" pageNumber="474">FIGURES 8A–D.</emphasis>
<taxonomicName id="0B914D6F6611FFA8FEA1F8D3FDFB4C30" box="[345,558,1836,1858]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="474" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
<emphasis id="FEE5EAFE6611FFA8FEA1F8D3FDFB4C30" box="[345,558,1836,1858]" italics="true" pageId="15" pageNumber="474">Tectoribates borealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D657856611FFA8FDCDF8D3FD534C30" box="[565,646,1836,1858]" pageId="15" pageNumber="474" rank="species">
<emphasis id="FEE5EAFE6611FFA8FDCDF8D3FD534C30" bold="true" box="[565,646,1836,1858]" pageId="15" pageNumber="474">sp. nov.</emphasis>
</taxonomicNameLabel>
, scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, prodorsum, with arrow to small teeth lateral to solenidia on tarsus I, and arrow to exobothridial seta; C, habitus ventral, with arrow to foveae between genital and anal plates; D, 
<taxonomicName id="0B914D6F6611FFA8FC59F896FBCB4C0D" box="[929,1054,1897,1919]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="15" pageNumber="479" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
, with arrow to band of tubercles on anterior of coxisternal region. Scale bar: A, B, C = 20 µm, D = 10 µm.
</paragraph>
</caption>
<caption id="98EE6664660EFFB7FF6FF99BFED54D87" httpUri="https://zenodo.org/record/247820/files/figure.png" pageId="16" pageNumber="475" targetBox="[151,1436,193,1614]" targetPageId="16">
<paragraph id="CC2E36EC660EFFB7FF6FF99BFED54D87" blockId="16.[151,1436,1636,1781]" pageId="16" pageNumber="475">
<emphasis id="FEE5EAFE660EFFB7FF6FF99BFE874D08" bold="true" box="[151,338,1636,1658]" pageId="16" pageNumber="475">FIGURES 9A–D.</emphasis>
<taxonomicName id="0B914D6F660EFFB7FEA1F99BFDFB4D08" box="[345,558,1636,1658]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="16" pageNumber="475" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
<emphasis id="FEE5EAFE660EFFB7FEA1F99BFDFB4D08" box="[345,558,1636,1658]" italics="true" pageId="16" pageNumber="475">Tectoribates borealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D65785660EFFB7FDCDF99BFD534D08" box="[565,646,1636,1658]" pageId="16" pageNumber="475" rank="species">
<emphasis id="FEE5EAFE660EFFB7FDCDF99BFD534D08" bold="true" box="[565,646,1636,1658]" pageId="16" pageNumber="475">sp. nov.</emphasis>
</taxonomicNameLabel>
, scanning electron microscope images of adult female from Alberta, Canada; A, ventral region showing discidium, custodium and foveae on lateral of ventral plate; B, detail of foveae on notogaster, showing foveae mostly plugged with cerotegument; C, prodorsum, anterolateral aspect, with arrow to small teeth lateral to solenidia on tibia I; D, rostrum, laterofrontal aspect, with arrow to proximal solenidion on tarsus II. Scale bar: A, B, C = 10 µm, D = 1 µm.
</paragraph>
</caption>
<caption id="98EE6664660FFFB6FF6FF89EFB314CA1" httpUri="https://zenodo.org/record/247821/files/figure.png" pageId="17" pageNumber="476" targetBox="[220,1350,193,1842]" targetPageId="17">
<paragraph id="CC2E36EC660FFFB6FF6FF89EFB314CA1" blockId="17.[151,1436,1889,2003]" pageId="17" pageNumber="476">
<emphasis id="FEE5EAFE660FFFB6FF6FF89EFE8A4C05" bold="true" box="[151,351,1889,1911]" pageId="17" pageNumber="476">FIGURES 10A–D.</emphasis>
<taxonomicName id="0B914D6F660FFFB6FE9EF89EFDEE4C05" box="[358,571,1889,1911]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="476" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
<emphasis id="FEE5EAFE660FFFB6FE9EF89EFDEE4C05" box="[358,571,1889,1911]" italics="true" pageId="17" pageNumber="476">Tectoribates borealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D65785660FFFB6FDBAF89EFD464C05" box="[578,659,1889,1911]" pageId="17" pageNumber="476" rank="species">
<emphasis id="FEE5EAFE660FFFB6FDBAF89EFD464C05" bold="true" box="[578,659,1889,1911]" pageId="17" pageNumber="476">sp. nov.</emphasis>
</taxonomicNameLabel>
, scanning electron microscope images of tritonymph from Alberta, Canada; A, habitus, dorsolateral aspect, with insert of cerotegumental platelets bottom right; B, habitus, ventrolateral aspect, insert of opisthonotal gland opening upper left; C, palptarsus, antiaxial aspect; D, detail of legs I and II, antiaxial aspect, with arrow to phylliform seta 
<emphasis id="FEE5EAFE660FFFB6FECFF842FE914CA1" box="[311,324,1981,2003]" italics="true" pageId="17" pageNumber="476">d</emphasis>
on femur III and coupled seta 
<emphasis id="FEE5EAFE660FFFB6FD83F842FD5D4CA1" box="[635,648,1981,2003]" italics="true" pageId="17" pageNumber="476">d</emphasis>
on tibia I. Scale bar: A, B = 50 µm, C, = 1 µm, D = 10 µm.
</paragraph>
</caption>
<caption id="98EE6664660CFFB5FF6FF8CCFB774C90" httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="18" pageNumber="477" targetBox="[163,1406,193,1798]" targetPageId="18">
<paragraph id="CC2E36EC660CFFB5FF6FF8CCFB774C90" blockId="18.[151,1436,1843,2018]" pageId="18" pageNumber="477">
<emphasis id="FEE5EAFE660CFFB5FF6FF8CCFE824C3B" bold="true" box="[151,343,1843,1865]" pageId="18" pageNumber="477">FIGURES 11A–F.</emphasis>
Differential interference contrast microscope images of adults: A, B: 
<taxonomicName id="0B914D6F660CFFB5FBE2F8CCFAEA4C3B" box="[1050,1343,1843,1865]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="477" phylum="Arthropoda" rank="species" species="alcescampestris" status="sp. nov.">
<emphasis id="FEE5EAFE660CFFB5FBE2F8CCFAEA4C3B" box="[1050,1343,1843,1865]" italics="true" pageId="18" pageNumber="477">Tectoribates alcescampestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D65785660CFFB5FABDF8CCFA404C3B" box="[1349,1429,1843,1865]" pageId="18" pageNumber="477" rank="species">
<emphasis id="FEE5EAFE660CFFB5FABDF8CCFA404C3B" bold="true" box="[1349,1429,1843,1865]" pageId="18" pageNumber="477">sp. nov.</emphasis>
</taxonomicNameLabel>
, adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, 
<taxonomicName id="0B914D6F660CFFB5FF6FF88FFEC44CF4" box="[151,273,1904,1926]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="18" pageNumber="479" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
with arrow to axillary saccule; C–E: 
<taxonomicName id="0B914D6F660CFFB5FD58F88FFCA24CF4" box="[672,887,1904,1926]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="477" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
<emphasis id="FEE5EAFE660CFFB5FD58F88FFCA24CF4" box="[672,887,1904,1926]" italics="true" pageId="18" pageNumber="477">Tectoribates borealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D65785660CFFB5FC79F88FFC014CF4" box="[897,980,1904,1926]" pageId="18" pageNumber="477" rank="species">
<emphasis id="FEE5EAFE660CFFB5FC79F88FFC014CF4" bold="true" box="[897,980,1904,1926]" pageId="18" pageNumber="477">sp. nov.</emphasis>
</taxonomicNameLabel>
, C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, 
<taxonomicName id="0B914D6F660CFFB5FDC0F852FCFE4CB1" box="[568,811,1965,1987]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="477" phylum="Arthropoda" rank="species" species="campestris" status="sp. nov.">
<emphasis id="FEE5EAFE660CFFB5FDC0F852FCFE4CB1" box="[568,811,1965,1987]" italics="true" pageId="18" pageNumber="477">Tectoribates campestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D65785660CFFB5FCCDF852FC5C4CB1" box="[821,905,1965,1987]" pageId="18" pageNumber="477" rank="species">
<emphasis id="FEE5EAFE660CFFB5FCCDF852FC5C4CB1" bold="true" box="[821,905,1965,1987]" pageId="18" pageNumber="477">sp. nov.</emphasis>
</taxonomicNameLabel>
, notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm.
</paragraph>
</caption>
<paragraph id="CC2E36EC660DFFB4FF3FFF68FC3D4BDD" blockId="19.[151,1437,151,1841]" box="[199,1000,151,176]" pageId="19" pageNumber="478">
<taxonomicName id="0B914D6F660DFFB4FF3FFF68FE164BC2" authority="Axillary" authorityName="Axillary" box="[199,451,151,176]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="19" pageNumber="478" phylum="Nematoda" rank="genus">
<emphasis id="FEE5EAFE660DFFB4FF3FFF68FE8A4BC2" box="[199,351,151,176]" italics="true" pageId="19" pageNumber="478">Gnathosoma:</emphasis>
Axillary
</taxonomicName>
saccule 
<date id="B82F102C660DFFB4FDDCFF67FD924BDD" box="[548,583,152,175]" pageId="19" pageNumber="478">7 x</date>
2. Relative lengths of setae: 
<emphasis id="FEE5EAFE660DFFB4FC73FF68FC344BC2" box="[907,993,151,176]" italics="true" pageId="19" pageNumber="478">m&gt;a&gt;h</emphasis>
.
</paragraph>
<paragraph id="CC2E36EC660DFFB4FF3FFF42FADF4B8A" blockId="19.[151,1437,151,1841]" pageId="19" pageNumber="478">
<emphasis id="FEE5EAFE660DFFB4FF3FFF42FED34BA6" box="[199,262,189,212]" italics="true" pageId="19" pageNumber="478">Legs:</emphasis>
(Figs. 7A–D). Setae 
<emphasis id="FEE5EAFE660DFFB4FE0DFF43FDD84BA7" box="[501,525,188,213]" italics="true" pageId="19" pageNumber="478">l”</emphasis>
of genu II short, spinous, more barbed than other setae on segment, about 13; 
<emphasis id="FEE5EAFE660DFFB4FA7CFF43FA494BA7" box="[1412,1436,188,213]" italics="true" pageId="19" pageNumber="478">l”</emphasis>
genu I tapered, about 22. Anterodorsal spines present on tibia I lateral to solenidia 
<emphasis id="FEE5EAFE660DFFB4FBCCFF1EFB964B8A" box="[1076,1091,225,248]" italics="true" pageId="19" pageNumber="478">φ</emphasis>
1 and 
<emphasis id="FEE5EAFE660DFFB4FB86FF1EFB584B8A" box="[1150,1165,225,248]" italics="true" pageId="19" pageNumber="478">φ</emphasis>
2 (
<figureCitation id="54AA2A69660DFFB4FB5DFF20FB294B8A" box="[1189,1276,223,248]" captionStart="FIGURES 8 A – D" captionStartId="15.[151,264,1836,1858]" captionTargetBox="[151,1436,193,1814]" captionTargetId="figure@15.[151,1436,193,1815]" captionTargetPageId="15" captionText="FIGURES 8 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, prodorsum, with arrow to small teeth lateral to solenidia on tarsus I, and arrow to exobothridial seta; C, habitus ventral, with arrow to foveae between genital and anal plates; D, gnathosoma, with arrow to band of tubercles on anterior of coxisternal region. Scale bar: A, B, C = 20 µm, D = 10 µm." httpUri="https://zenodo.org/record/247819/files/figure.png" pageId="19" pageNumber="478">Fig. 8B</figureCitation>
).
</paragraph>
<paragraph id="CC2E36EC660DFFB4FF3FFEF9FC4D4A6D" blockId="19.[151,1437,151,1841]" box="[199,920,262,287]" pageId="19" pageNumber="478">
<emphasis id="FEE5EAFE660DFFB4FF3FFEF9FE874A6D" bold="true" box="[199,338,262,287]" pageId="19" pageNumber="478">Description</emphasis>
<emphasis id="FEE5EAFE660DFFB4FEA0FEF9FDA24A6D" box="[344,631,262,287]" italics="true" pageId="19" pageNumber="478">Tritonymph: Dimensions:</emphasis>
Total length: (
<emphasis id="FEE5EAFE660DFFB4FCE4FEF7FCFF4A6D" box="[796,810,264,287]" italics="true" pageId="19" pageNumber="478">n</emphasis>
= 1) 307.
</paragraph>
<paragraph id="CC2E36EC660DFFB4FF3FFED3FC414A15" blockId="19.[151,1437,151,1841]" pageId="19" pageNumber="478">
<emphasis id="FEE5EAFE660DFFB4FF3FFED3FE904A31" box="[199,325,300,323]" italics="true" pageId="19" pageNumber="478">Integument</emphasis>
: Sclerotized with light brown coloration. Cerotegument of tightly packed platelets, about 
<quantity id="0B699B09660DFFB4FAABFED4FA494A31" box="[1363,1436,298,323]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="19" pageNumber="478" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">2–4 in</quantity>
width, covering all of body and leg segments (
<figureCitation id="54AA2A69660DFFB4FD66FEB1FCC54A15" box="[670,784,334,359]" captionStart="FIGURES 10 A – D" captionStartId="17.[151,264,1889,1911]" captionTargetBox="[220,1350,193,1842]" captionTargetId="figure@17.[220,1351,193,1843]" captionTargetPageId="17" captionText="FIGURES 10 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of tritonymph from Alberta, Canada; A, habitus, dorsolateral aspect, with insert of cerotegumental platelets bottom right; B, habitus, ventrolateral aspect, insert of opisthonotal gland opening upper left; C, palptarsus, antiaxial aspect; D, detail of legs I and II, antiaxial aspect, with arrow to phylliform seta d on femur III and coupled seta d on tibia I. Scale bar: A, B = 50 µm, C, = 1 µm, D = 10 µm." httpUri="https://zenodo.org/record/247821/files/figure.png" pageId="19" pageNumber="478">Figs. 10A</figureCitation>
, insert, B)).
</paragraph>
<paragraph id="CC2E36EC660DFFB4FF3FFE8DFAA94969" blockId="19.[151,1437,151,1841]" pageId="19" pageNumber="478">
<emphasis id="FEE5EAFE660DFFB4FF3FFE8DFE984AF9" box="[199,333,370,395]" italics="true" pageId="19" pageNumber="478">Prodorsum:</emphasis>
Aspis with transverse porose area posterior to setae 
<emphasis id="FEE5EAFE660DFFB4FC67FE8BFC604AF9" box="[927,949,372,395]" italics="true" pageId="19" pageNumber="478">in</emphasis>
and bothridia. Lateral porose areas present anterior of bothridia. Seta 
<emphasis id="FEE5EAFE660DFFB4FE2CFE67FE394ADD" box="[468,492,408,431]" italics="true" pageId="19" pageNumber="478">ro</emphasis>
finely barbed along length, tapered, about 21 long (
<figureCitation id="54AA2A69660DFFB4FB9DFE69FB044ADD" box="[1125,1233,406,431]" captionStart="FIGURES 10 A – D" captionStartId="17.[151,264,1889,1911]" captionTargetBox="[220,1350,193,1842]" captionTargetId="figure@17.[220,1351,193,1843]" captionTargetPageId="17" captionText="FIGURES 10 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of tritonymph from Alberta, Canada; A, habitus, dorsolateral aspect, with insert of cerotegumental platelets bottom right; B, habitus, ventrolateral aspect, insert of opisthonotal gland opening upper left; C, palptarsus, antiaxial aspect; D, detail of legs I and II, antiaxial aspect, with arrow to phylliform seta d on femur III and coupled seta d on tibia I. Scale bar: A, B = 50 µm, C, = 1 µm, D = 10 µm." httpUri="https://zenodo.org/record/247821/files/figure.png" pageId="19" pageNumber="478">Fig. 10B</figureCitation>
). Seta 
<emphasis id="FEE5EAFE660DFFB4FAD0FE69FAE84ADD" box="[1320,1341,406,431]" italics="true" pageId="19" pageNumber="478">le</emphasis>
barbed, phylliform, about 20 long, 10 at maximum width. Seta 
<emphasis id="FEE5EAFE660DFFB4FCECFE43FCFF4AA1" box="[788,810,444,467]" italics="true" pageId="19" pageNumber="478">in</emphasis>
barbed, phylliform, about 27 long. Mutual distance of setae 
<emphasis id="FEE5EAFE660DFFB4FF2DFE1FFE9B4A85" box="[213,334,478,503]" italics="true" pageId="19" pageNumber="478">ro-ro, le-le</emphasis>
and 
<emphasis id="FEE5EAFE660DFFB4FE7BFE1FFE6D4A85" box="[387,440,480,503]" italics="true" pageId="19" pageNumber="478">in-in</emphasis>
about 13, 18 and 50, respectively. Seta 
<emphasis id="FEE5EAFE660DFFB4FC89FE1FFC5F4A85" box="[881,906,480,503]" italics="true" pageId="19" pageNumber="478">ex</emphasis>
about 8 long, thick, heavily barbed, penicillate. Bothridial seta 56–59 long, barbed, clavate, head subequal in length to stalk, broader in shape than that of adult.
</paragraph>
<paragraph id="CC2E36EC660DFFB4FF3FFDD8FBA249F8" blockId="19.[151,1437,151,1841]" pageId="19" pageNumber="478">
<emphasis id="FEE5EAFE660DFFB4FF3FFDD8FE70494D" box="[199,421,551,575]" italics="true" pageId="19" pageNumber="478">Gastronotic Region</emphasis>
: Integument without sculpturing; orientation of plicae more transverse anteriorly, more longitudinal posteriorly (
<figureCitation id="54AA2A69660DFFB4FE4EFDB5FDF74911" box="[438,546,586,611]" captionStart="FIGURES 10 A – D" captionStartId="17.[151,264,1889,1911]" captionTargetBox="[220,1350,193,1842]" captionTargetId="figure@17.[220,1351,193,1843]" captionTargetPageId="17" captionText="FIGURES 10 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of tritonymph from Alberta, Canada; A, habitus, dorsolateral aspect, with insert of cerotegumental platelets bottom right; B, habitus, ventrolateral aspect, insert of opisthonotal gland opening upper left; C, palptarsus, antiaxial aspect; D, detail of legs I and II, antiaxial aspect, with arrow to phylliform seta d on femur III and coupled seta d on tibia I. Scale bar: A, B = 50 µm, C, = 1 µm, D = 10 µm." httpUri="https://zenodo.org/record/247821/files/figure.png" pageId="19" pageNumber="478">Fig. 10A</figureCitation>
). Gastronotal setae of 
<emphasis id="FEE5EAFE660DFFB4FCD5FDB3FC464911" box="[813,915,586,611]" italics="true" pageId="19" pageNumber="478">c, d, l, h</emphasis>
and 
<emphasis id="FEE5EAFE660DFFB4FC2AFDB3FC354911" box="[978,992,588,611]" italics="true" pageId="19" pageNumber="478">p</emphasis>
series about 14–20 long, with seta 
<emphasis id="FEE5EAFE660DFFB4FA70FDB3FA494915" box="[1416,1436,588,615]" italics="true" pageId="19" pageNumber="478">c3</emphasis>
shortest seta. Mutual distance setae 
<emphasis id="FEE5EAFE660DFFB4FDD0FD8EFD1049F8" box="[552,709,625,650]" italics="true" pageId="19" pageNumber="478">da-da, dm-dm</emphasis>
and 
<emphasis id="FEE5EAFE660DFFB4FD04FD8EFCE849F8" box="[764,829,625,650]" italics="true" pageId="19" pageNumber="478">dp-dp</emphasis>
41, 36 and 15, respectively.
</paragraph>
<paragraph id="CC2E36EC660DFFB4FF3FFD6AFAEF4868" blockId="19.[151,1437,151,1841]" pageId="19" pageNumber="478">
<emphasis id="FEE5EAFE660DFFB4FF3FFD6AFEA649DF" box="[199,371,661,686]" italics="true" pageId="19" pageNumber="478">Ventral Region</emphasis>
: Medial margins of epimeral plates weakly defined. Epimere I with narrow mentotectum developed laterally. Epimeral, genital, aggenital, anal and adanal setae smooth, acuminate, about 10 long. Porose integument surrounding opening of opisthonotal gland not evident (possibly because area covered by cerotegument). Epimeral setation 3-1-2-3. Genital, aggenital, anal and adanal setae 5, 1, 2, 3, respectively.
</paragraph>
<paragraph id="CC2E36EC660DFFB4FF3FFCDAFE8D4810" blockId="19.[151,1437,151,1841]" pageId="19" pageNumber="478">
<taxonomicName id="0B914D6F660DFFB4FF3FFCDAFE13484C" authority="Axillary" authorityName="Axillary" box="[199,454,805,830]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="19" pageNumber="478" phylum="Nematoda" rank="genus">
<emphasis id="FEE5EAFE660DFFB4FF3FFCDAFE83484C" box="[199,342,805,830]" italics="true" pageId="19" pageNumber="478">Gnathosoma</emphasis>
: Axillary
</taxonomicName>
saccule present. Palpal eupathidium 
<emphasis id="FEE5EAFE660DFFB4FC82FCD9FC7C484F" box="[890,937,806,829]" italics="true" pageId="19" pageNumber="478">acm</emphasis>
attached to solenidion along distal half of solenidial length.
</paragraph>
<paragraph id="CC2E36EC660DFFB4FF3FFC91FE434F2F" blockId="19.[151,1437,151,1841]" pageId="19" pageNumber="478">
<emphasis id="FEE5EAFE660DFFB4FF3FFC91FF2948F7" box="[199,252,878,901]" italics="true" pageId="19" pageNumber="478">Legs</emphasis>
: Setae and solenidia as for tritonymph of 
<taxonomicName id="0B914D6F660DFFB4FD23FC92FBCA48F4" box="[731,1055,877,902]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="19" pageNumber="478" phylum="Arthropoda" rank="species" species="alcescampestris">
<emphasis id="FEE5EAFE660DFFB4FD23FC92FBCA48F4" box="[731,1055,877,902]" italics="true" pageId="19" pageNumber="478">Tectoribates alcescampestris</emphasis>
</taxonomicName>
(
<tableCitation id="81130357660DFFB4FBD6FC92FB5648F4" box="[1070,1155,877,902]" captionStart="TABLE 1" captionStartId="9.[151,239,1132,1155]" captionTargetBox="[159,1399,1215,1903]" captionTargetPageId="9" captionText="TABLE 1. Setiform organs in deutonymph, tritonymph and adult Tectoribates alcescampestris sp. nov. Setae (Roman) and solenidia (Greek) are indicated; parentheses indicate pairs of setae." httpUri="http://table.plazi.org/id/98EE66646617FFAEFF6FFB93FC584FD1" pageId="19" pageNumber="478" tableUuid="98EE66646617FFAEFF6FFB93FC584FD1">Table 1</tableCitation>
). Proral setae of tarsus I appear eupathidial. Subunguinal seta of tarsus I of normal form and inserted proximal to antelateral pair as in adult. Porose areas present on femora I to IV. Distal setae on tarsi I–IV blunt distally. Setae 
<emphasis id="FEE5EAFE660DFFB4FB9BFC4AFBA448BC" box="[1123,1137,949,974]" italics="true" pageId="19" pageNumber="478">d</emphasis>
on femora I to IV barbed, phylliform (
<figureCitation id="54AA2A69660DFFB4FEE6FC26FE534880" box="[286,390,985,1010]" captionStart="FIGURES 10 A – D" captionStartId="17.[151,264,1889,1911]" captionTargetBox="[220,1350,193,1842]" captionTargetId="figure@17.[220,1351,193,1843]" captionTargetPageId="17" captionText="FIGURES 10 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of tritonymph from Alberta, Canada; A, habitus, dorsolateral aspect, with insert of cerotegumental platelets bottom right; B, habitus, ventrolateral aspect, insert of opisthonotal gland opening upper left; C, palptarsus, antiaxial aspect; D, detail of legs I and II, antiaxial aspect, with arrow to phylliform seta d on femur III and coupled seta d on tibia I. Scale bar: A, B = 50 µm, C, = 1 µm, D = 10 µm." httpUri="https://zenodo.org/record/247821/files/figure.png" pageId="19" pageNumber="478">Fig. 10D</figureCitation>
). Tibiae I and II and genu I with seta 
<emphasis id="FEE5EAFE660DFFB4FCDFFC26FCED4880" box="[807,824,985,1010]" italics="true" pageId="19" pageNumber="478">l’</emphasis>
barbed, tapered, partially phylliform, 
<emphasis id="FEE5EAFE660DFFB4FB1AFC26FB2F4880" box="[1250,1274,985,1010]" italics="true" pageId="19" pageNumber="478">l”</emphasis>
more spinose. Solenidion and coupled seta 
<emphasis id="FEE5EAFE660DFFB4FE12FC02FE2D4F64" box="[490,504,1021,1046]" italics="true" pageId="19" pageNumber="478">d</emphasis>
on tibia I on anterodorsal tubercle (
<figureCitation id="54AA2A69660DFFB4FC5EFC02FBC64F64" box="[934,1043,1021,1046]" captionStart="FIGURES 10 A – D" captionStartId="17.[151,264,1889,1911]" captionTargetBox="[220,1350,193,1842]" captionTargetId="figure@17.[220,1351,193,1843]" captionTargetPageId="17" captionText="FIGURES 10 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of tritonymph from Alberta, Canada; A, habitus, dorsolateral aspect, with insert of cerotegumental platelets bottom right; B, habitus, ventrolateral aspect, insert of opisthonotal gland opening upper left; C, palptarsus, antiaxial aspect; D, detail of legs I and II, antiaxial aspect, with arrow to phylliform seta d on femur III and coupled seta d on tibia I. Scale bar: A, B = 50 µm, C, = 1 µm, D = 10 µm." httpUri="https://zenodo.org/record/247821/files/figure.png" pageId="19" pageNumber="478">Fig. 10D</figureCitation>
). Solenidion 
<emphasis id="FEE5EAFE660DFFB4FB57FC01FB6B4F67" box="[1199,1214,1022,1045]" italics="true" pageId="19" pageNumber="478">φ</emphasis>
on tibia II and III subequal in length to seta 
<emphasis id="FEE5EAFE660DFFB4FE3EFBDEFE014F48" box="[454,468,1057,1082]" italics="true" pageId="19" pageNumber="478">d</emphasis>
; solenidion 
<emphasis id="FEE5EAFE660DFFB4FD9BFBDDFDA74F4B" box="[611,626,1058,1081]" italics="true" pageId="19" pageNumber="478">φ</emphasis>
on tibia 
<date id="B82F102C660DFFB4FD24FBDDFCC44F48" box="[732,785,1057,1082]" pageId="19" pageNumber="478">IV 4</date>
/5 length seta 
<emphasis id="FEE5EAFE660DFFB4FC4CFBDEFC174F48" box="[948,962,1057,1082]" italics="true" pageId="19" pageNumber="478">d</emphasis>
. Solenidia 
<emphasis id="FEE5EAFE660DFFB4FBBDFBDDFB874F48" box="[1093,1106,1058,1082]" italics="true" pageId="19" pageNumber="478">δ</emphasis>
on genua I, 
<date id="B82F102C660DFFB4FB1EFBDDFAB24F48" box="[1254,1383,1057,1082]" pageId="19" pageNumber="478">II and III 4</date>
/5 to subequal length seta 
<emphasis id="FEE5EAFE660DFFB4FE79FBBAFE5A4F2C" box="[385,399,1093,1118]" italics="true" pageId="19" pageNumber="478">d</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="848B6567660DFFB4FF3FFB96FE104FD4" pageId="19" pageNumber="478" type="etymology">
<paragraph id="CC2E36EC660DFFB4FF3FFB96FE104FD4" blockId="19.[151,1437,151,1841]" pageId="19" pageNumber="478">
<emphasis id="FEE5EAFE660DFFB4FF3FFB96FE934FF0" bold="true" box="[199,326,1129,1154]" pageId="19" pageNumber="478">Etymology</emphasis>
. The specific epithet 
<taxonomicName id="0B914D6F660DFFB4FDB0FB96FD764FF0" box="[584,675,1129,1154]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="19" pageNumber="478" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="FEE5EAFE660DFFB4FDB0FB96FD764FF0" box="[584,675,1129,1154]" italics="true" pageId="19" pageNumber="478">borealis</emphasis>
</taxonomicName>
is from the Greek meaning ‘northern’ and refers to the northern distribution of this species.
</paragraph>
</subSubSection>
<subSubSection id="848B6567660DFFB4FF3FFB4EFE774C43" pageId="19" pageNumber="478" type="discussion">
<paragraph id="CC2E36EC660DFFB4FF3FFB4EFB934E0F" blockId="19.[151,1437,151,1841]" pageId="19" pageNumber="478">
<emphasis id="FEE5EAFE660DFFB4FF3FFB4EFEE14FB8" bold="true" box="[199,308,1201,1226]" pageId="19" pageNumber="478">Remarks</emphasis>
. This species is similar to two Mongolian species, 
<taxonomicName id="0B914D6F660DFFB4FC72FB4EFB4B4FB8" box="[906,1182,1201,1226]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="19" pageNumber="478" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="FEE5EAFE660DFFB4FC72FB4EFB4B4FB8" box="[906,1182,1201,1226]" italics="true" pageId="19" pageNumber="478">Tectoribates mongolicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="0B914D6F660DFFB4FB22FB4DFAA04FB8" box="[1242,1397,1201,1226]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="19" pageNumber="478" phylum="Arthropoda" rank="species" species="deserticola">
<emphasis id="FEE5EAFE660DFFB4FB22FB4DFAA04FB8" box="[1242,1397,1201,1226]" italics="true" pageId="19" pageNumber="478">T. deserticola</emphasis>
</taxonomicName>
, in having a notogaster with the hexagonal pattern anteriorly and porose areas. It differs from both species in interlamellar setae reaching the tip of the lamella (reaching at most to insertion of seta 
<emphasis id="FEE5EAFE660DFFB4FBA7FB06FBA64E60" box="[1119,1139,1273,1298]" italics="true" pageId="19" pageNumber="478">le</emphasis>
in the Mongolian species) and in well developed medial and lateral dens on the lamellar cusp (medial and lateral dens short and weakly developed in the Mongolian species). It differs from 
<taxonomicName id="0B914D6F660DFFB4FD08FABDFC574E28" box="[752,898,1345,1370]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="19" pageNumber="478" phylum="Arthropoda" rank="species" species="mongolica">
<emphasis id="FEE5EAFE660DFFB4FD08FABDFC574E28" box="[752,898,1345,1370]" italics="true" pageId="19" pageNumber="478">T. mongolica</emphasis>
</taxonomicName>
in having shorter notogastral setae and from 
<taxonomicName id="0B914D6F660DFFB4FA7FFABDFEC14E0C" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="19" pageNumber="478" phylum="Arthropoda" rank="species" species="deserticola">
<emphasis id="FEE5EAFE660DFFB4FA7FFABDFEC14E0C" italics="true" pageId="19" pageNumber="478">T. deserticola</emphasis>
</taxonomicName>
in having elongate, club-like, rather than short, globular bothridial setae.
</paragraph>
<paragraph id="CC2E36EC660DFFB4FF3FFA76FEA84E9F" blockId="19.[151,1437,151,1841]" pageId="19" pageNumber="478">
The tritonymph of this species can be easily distinguished from that of 
<taxonomicName id="0B914D6F660DFFB4FC18FA76FAF64ED0" box="[992,1315,1417,1442]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="19" pageNumber="478" phylum="Arthropoda" rank="species" species="alcescampestris">
<emphasis id="FEE5EAFE660DFFB4FC18FA76FAF64ED0" box="[992,1315,1417,1442]" italics="true" pageId="19" pageNumber="478">Tectoribates alcescampestris</emphasis>
</taxonomicName>
, described above, by the phylliform shape of its setae 
<emphasis id="FEE5EAFE660DFFB4FD7EFA50FD494EB8" box="[646,668,1455,1482]" italics="true" pageId="19" pageNumber="478">p2</emphasis>
and 
<emphasis id="FEE5EAFE660DFFB4FD2EFA51FD394EB8" box="[726,748,1454,1482]" italics="true" pageId="19" pageNumber="478">p3</emphasis>
(vs. barbed, tapered) and phylliform seta 
<emphasis id="FEE5EAFE660DFFB4FB34FA52FB0F4EB4" box="[1228,1242,1453,1478]" italics="true" pageId="19" pageNumber="478">d</emphasis>
on femur IV (vs barbed and tapered).
</paragraph>
<paragraph id="CC2E36EC660DFFB4FF3FFA07FDBD4DD3" blockId="19.[151,1437,151,1841]" pageId="19" pageNumber="478">
<taxonomicName id="0B914D6F660DFFB4FF3FFA07FE6D4D63" box="[199,440,1528,1553]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="19" pageNumber="478" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="FEE5EAFE660DFFB4FF3FFA07FE6D4D63" box="[199,440,1528,1553]" italics="true" pageId="19" pageNumber="478">Tectoribates borealis</emphasis>
</taxonomicName>
has been found in open habitats, primarily native prairie, badlands and pastures in southern Alberta, but we have one specimen from a bog in Wood Buffalo National Park, in the extreme north of the province. Gravid females carry up to 3 large eggs. Gut contents are composed primarily of dark brown fungal hyphae and spores. The association of immatures with adults was addressed under the Discussion following the generic diagnosis (See above, Remark 4).
</paragraph>
<paragraph id="CC2E36EC660DFFB4FF3FF953FE774C43" blockId="19.[151,1437,151,1841]" pageId="19" pageNumber="478">
Three male specimens almost identical to the description given above are known from Florida, 
<collectingCountry id="B486767C660DFFB4FB11F952FAF44DB7" box="[1257,1313,1709,1733]" name="United States of America" pageId="19" pageNumber="478">USA</collectingCountry>
(housed in the RNC). They differ in that the medial dens of the lamellar cusp is subequal to the lateral dens. We hesitate to consider these specimens members of 
<taxonomicName id="0B914D6F660DFFB4FDBFF90BFCE64C7F" box="[583,819,1780,1805]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="19" pageNumber="478" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="FEE5EAFE660DFFB4FDBFF90BFCE64C7F" box="[583,819,1780,1805]" italics="true" pageId="19" pageNumber="478">Tectoribates borealis</emphasis>
</taxonomicName>
, until there are more records of the genus known from Eastern North 
<collectingCountry id="B486767C660DFFB4FEC2F8E7FE4B4C43" box="[314,414,1816,1841]" name="United States of America" pageId="19" pageNumber="478">America</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>