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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1030.62544" ID-GBIF-Dataset="91fec855-7fa9-4ad6-84d7-9b9d35e23e8b" ID-PMC="PMC8060247" ID-Pensoft-Pub="1313-2970-1030-1" ID-Pensoft-UUID="875199397EEE5F7898EA1DB25DA62D25" ID-PubMed="33958904" ModsDocID="1313-2970-1030-1" checkinTime="1618371561543" checkinUser="pensoft" docAuthor="Means, Jackson C., Hennen, Derek A. &amp; Marek, Paul E." docDate="2021" docId="11ACA8B4D4585F9BBD8EB3A44CF71419" docLanguage="en" docName="ZooKeys 1030: 1-180" docOrigin="ZooKeys 1030" docPubDate="2021-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1030.62544" docTitle="Nannaria hokie Means, Hennen &amp; Marek 2021" docType="treatment" docVersion="5" id="875199397EEE5F7898EA1DB25DA62D25" lastPageNumber="1" masterDocId="875199397EEE5F7898EA1DB25DA62D25" masterDocTitle="A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1673961214978" updateUser="julia">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Means, Jackson C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7377-0696</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
<mods:nameIdentifier type="email">mjacks4@vt.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hennen, Derek A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7005-1151</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marek, Paul E.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1030</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>180</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1030.62544</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1030.62544</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1030-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">875199397EEE5F7898EA1DB25DA62D25</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="181004761" LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419" httpUri="http://treatment.plazi.org/id/11ACA8B4D4585F9BBD8EB3A44CF71419" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="86Y4H" LSID="11ACA8B4-D458-5F9B-BD8E-B3A44CF71419" authority="Means, Hennen &amp; Marek, 2021" authorityName="Means, Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria hokie" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hokie">Nannaria hokie Means, Hennen &amp; Marek, 2021</taxonomicName>
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Nannaria hokie Means, Hennen &amp; Marek in Means et al. 2021, paratype ♂ (VTEC, MPE 00880) left gonopod. A anterior view; red arrow indicates hood-like flange partially obscuring view of projected solenomere B medial view C posterior view; red triangle indicates medial flange. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure35" httpUri="https://binary.pensoft.net/fig/531138" pageId="0" pageNumber="1">Figs 35</figureCitation>
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Nannaria hokie Means, Hennen &amp; Marek in Means et al. 2021, holotype ♂ (VTEC, MPE 04803) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure36" httpUri="https://binary.pensoft.net/fig/531139" pageId="0" pageNumber="1">, 36</figureCitation>
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Nannaria hokie Means, Hennen &amp; Marek, 2021 variation A Floyd County, Virginia non-type ♂ (VMNH, NAN 0003; red arrow indicates reduced lateral and medial flanges) B Pulaski County, Virginia non-type ♂ (VTEC, MPE 00253); red triangle indicates prefemoral process lacking prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure37" httpUri="https://binary.pensoft.net/fig/531140" pageId="0" pageNumber="1">, 37 Vernacular name: &quot;The Virginia Tech Twisted-Claw Millipede&quot;</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Means, Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria hokie" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hokie">Nannaria hokie</taxonomicName>
Means, Hennen &amp; Marek, in
<bibRefCitation DOI="https://doi.org/10.1093/isd/ixab003" author="Means, JC" journalOrPublisher="Insect Systematics and Diversity" pageId="0" pageNumber="1" pagination="1 - 26" refId="B52" refString="Means, JC, Hennen, DA, Tanabe, T, Marek, PE, 2021. Phylogenetic systematics of the millipede family Xystodesmidae. Insect Systematics and Diversity 5: 1 - 26, DOI: https://doi.org/10.1093/isd/ixab003" title="Phylogenetic systematics of the millipede family Xystodesmidae." url="https://doi.org/10.1093/isd/ixab003" volume="5" year="2021">Means et al. 2021</bibRefCitation>
: 4, 5, 7, 10, 17-19, 24, S70.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355557" accessionNumber="MPE04803" collectingDate="2019-04-17" collectorName="P. Marek" country="United States" county="Montgomery County" elevation="637" latitude="37.2216" location="Virginia Tech" longLatPrecision="6" longitude="-80.4159" municipality="Blacksburg" specimenCount="♂" stateProvince="Virginia" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
</emphasis>
<collectingCountry name="United States of America">United States</collectingCountry>
-
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
</emphasis>
•
<specimenCount>♂</specimenCount>
;
<collectingCounty>Montgomery County</collectingCounty>
,
<collectingMunicipality>Blacksburg</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:FF026762E86E804543138512112A4A19" country="United States" county="Montgomery County" latitude="37.2216" longLatPrecision="6" longitude="-80.4159" municipality="Blacksburg" name="Virginia Tech" stateProvince="Virginia">Virginia Tech</location>
campus,
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:E2FED81D960FD39D0135AFBE14FFE657" country="United States" county="Montgomery County" latitude="37.2216" longLatPrecision="6" longitude="-80.4159" municipality="Blacksburg" name="Stadium Woods" stateProvince="Virginia">Stadium Woods</location>
;
<geoCoordinate degrees="37.2216" direction="north" orientation="latitude" precision="5" value="37.2216">37.2216°N</geoCoordinate>
, -
<geoCoordinate degrees="80.4159" direction="west" orientation="longitude" precision="5" value="-80.4159">80.4159°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.37" unit="m" value="637.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.37" unit="m" value="637.0">637 m</elevation>
</quantity>
;
<collectingDate value="2019-04-17">17 Apr. 2019</collectingDate>
; hand collected;
<collectorName>P. Marek</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE04803">MPE04803</accessionNumber>
.
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355508" accessionNumber="MPE00880" collectingDate="2015-10-13" collectorName="D. Hennen" country="United States" county="Montgomery County" elevation="625" latitude="37.225" location="Virginia Tech" longLatPrecision="6" longitude="-80.4276" municipality="Blacksburg" specimenCount="5" specimenCount-male="5" stateProvince="Virginia" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<collectingCountry name="United States of America">United States</collectingCountry>
-
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
</emphasis>
•
<specimenCount type="male">5 ♂♂</specimenCount>
;
<collectingCounty>Montgomery County</collectingCounty>
,
<collectingMunicipality>Blacksburg</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:41F99E17649DEF9C96125181F91A3CAA" country="United States" county="Montgomery County" latitude="37.225" longLatPrecision="6" longitude="-80.4276" municipality="Blacksburg" name="Virginia Tech" stateProvince="Virginia">Virginia Tech</location>
campus, south side of
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:2509E487C6696C73F9FEA21B60681F3A" country="United States" county="Montgomery County" latitude="37.225" longLatPrecision="6" longitude="-80.4276" municipality="Blacksburg" name="Duck Pond" stateProvince="Virginia">Duck Pond</location>
;
<geoCoordinate degrees="37.2250" direction="north" orientation="latitude" precision="5" value="37.225">37.2250°N</geoCoordinate>
, -
<geoCoordinate degrees="80.4276" direction="west" orientation="longitude" precision="5" value="-80.4276">80.4276°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.25" unit="m" value="625.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.25" unit="m" value="625.0">625 m</elevation>
</quantity>
;
<collectingDate value="2015-10-13">13 Oct. 2015</collectingDate>
; hand collected;
<collectorName>D. Hennen</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE00880">MPE00880</accessionNumber>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3071355756" accessionNumber="MPE00884" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="United States" specimenCount="1" stateProvince="Virginia" typeStatus="paratype">
882, 883, VMNH,
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE00884">MPE00884</accessionNumber>
</materialsCitation>
, 886 •
<materialsCitation ID-GBIF-Occurrence="3071355316" accessionNumber="MPE00881" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="United States" specimenCount="2" specimenCount-female="2" stateProvince="Virginia" typeStatus="paratype">
<specimenCount type="female">2 ♀♀</specimenCount>
; same collection data as preceding; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE00881">MPE00881</accessionNumber>
</materialsCitation>
, 885.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355606" accessionNumber="MPE00266, MPE03464, MPE04998, MPE00227, MPE03829, MPE03831, MPE05027, MPE03808, MPE03705" collectingDate="1956-10-28" collectingDateMax="2020-03-13" collectingDateMin="1956-10-28" collectorName="R. Hoffman, Knight, Golden Hills Disc Golf, J. Means, D. Hennen, P. Marek, Blacksburg, Virginia Tech, G. Schiermeyer, F. Vasquez, I. Huerta, Gateway Trail, E. Francis, K. Lawler, N. Zegler, C. Hall, Fairview Ave., Airport, Fairview Rds., G. Scheirmeyer, Rt." country="United States" county="Floyd County" elevation="564" latitude="37.1849" location="Montgomery County" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" specimenCount="54" specimenCount-female="5" specimenCount-male="49" stateProvince="Virginia">
<collectingCountry name="United States of America">United States</collectingCountry>
-
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
</emphasis>
•
<specimenCount type="male">4 ♂♂</specimenCount>
;
<collectingCounty>Floyd County</collectingCounty>
, along
<collectingMunicipality>Goose Creek</collectingMunicipality>
, ca.
<locationDeviation location="Simpsons">
<quantity metricMagnitude="3" metricUnit="m" metricValue="4.828032" unit="mi" value="3.0">3 mi.</quantity>
N of Simpsons
</locationDeviation>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.6388" unit="ft" value="1850.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.6388" unit="ft" value="1850.0">1850 ft.</elevation>
</quantity>
;
<geoCoordinate degrees="37.0815" direction="north" orientation="latitude" precision="5" value="37.0815">37.0815°N</geoCoordinate>
, -
<geoCoordinate degrees="80.2055" direction="west" orientation="longitude" precision="5" value="-80.2055">80.2055°W</geoCoordinate>
;
<collectingDate value="1971-10-09">9 Oct. 1971</collectingDate>
;
<collectorName>R. Hoffman</collectorName>
,
<collectorName>Knight</collectorName>
leg.; VMNH NAN0005, 0244 •
<specimenCount type="male">10 ♂♂</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:20917D541CB86C33F74E6BB8B2C15ABA" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="Montgomery County" stateProvince="Virginia">Montgomery County</location>
,
<collectorName>Golden Hills Disc Golf</collectorName>
course, in gully;
<geoCoordinate degrees="37.1729" direction="north" orientation="latitude" precision="5" value="37.1729">37.1729°N</geoCoordinate>
, -
<geoCoordinate degrees="80.4078" direction="west" orientation="longitude" precision="5" value="-80.4078">80.4078°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.279999999999999" unit="m" value="628.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.279999999999999" unit="m" value="628.0">628 m</elevation>
</quantity>
;
<collectingDate value="2014-11-08">8 Nov. 2014</collectingDate>
; hand collected;
<collectorName>J. Means</collectorName>
,
<collectorName>D. Hennen</collectorName>
,
<collectorName>P. Marek</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE00266">MPE00266</accessionNumber>
, 275, 280, 283-89 •
<specimenCount type="male">5 ♂♂</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:D214C8B68FBF64BDE9E491F2E205A82B" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="Montgomery County" stateProvince="Virginia">Montgomery County</location>
,
<collectorName>Blacksburg</collectorName>
,
<collectorName>Virginia Tech</collectorName>
, patch of woods
<locationDeviation firstLocation="Grove Ln." secondLocation="Duck Pond">between Grove Ln. and Duck Pond</locationDeviation>
;
<geoCoordinate degrees="37.2247" direction="north" orientation="latitude" precision="5" value="37.2247">37.2247°N</geoCoordinate>
, -
<geoCoordinate degrees="80.4260" direction="west" orientation="longitude" precision="5" value="-80.426">80.4260°W</geoCoordinate>
, elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.279999999999999" unit="m" value="628.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.279999999999999" unit="m" value="628.0">628 m</elevation>
</quantity>
;
<collectingDate value="2017-11-09">9 Nov. 2017</collectingDate>
; hand collected;
<collectorName>J. Means</collectorName>
,
<collectorName>D. Hennen</collectorName>
,
<collectorName>G. Schiermeyer</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03464">MPE03464</accessionNumber>
-68 •
<specimenCount type="male">1 ♂</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:F7667C1FBE1EE55F907427B2881FACFD" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="Montgomery County" stateProvince="Virginia">Montgomery County</location>
,
<collectorName>Blacksburg</collectorName>
, end of
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:05A542C55F31B22EF548E53BE9DF63D1" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="Valley View Drive" stateProvince="Virginia">Valley View Drive</location>
near quarry;
<geoCoordinate degrees="37.2218" direction="north" orientation="latitude" precision="5" value="37.2218">37.2218°N</geoCoordinate>
, -
<geoCoordinate degrees="80.3894" direction="west" orientation="longitude" precision="5" value="-80.3894">80.3894°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.709999999999999" unit="m" value="671.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.709999999999999" unit="m" value="671.0">671 m</elevation>
</quantity>
;
<collectingDate value="2019-09-13">13 Sep. 2019</collectingDate>
; hand collected;
<collectorName>J. Means</collectorName>
,
<collectorName>D. Hennen</collectorName>
,
<collectorName>P. Marek</collectorName>
,
<collectorName>F. Vasquez</collectorName>
,
<collectorName>I. Huerta</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE04998">MPE04998</accessionNumber>
•
<specimenCount type="male">2 ♂♂</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:3CDE480D916FBE9C12870F77A80239A4" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="Montgomery County" stateProvince="Virginia">Montgomery County</location>
,
<collectorName>Gateway Trail</collectorName>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.04672" unit="mi" value="0.5">0.5 mi</quantity>
in from main trail by stream;
<geoCoordinate degrees="37.2506" direction="north" orientation="latitude" precision="5" value="37.2506">37.2506°N</geoCoordinate>
, -
<geoCoordinate degrees="80.4610" direction="west" orientation="longitude" precision="5" value="-80.461">80.4610°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.42" unit="m" value="642.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.42" unit="m" value="642.0">642 m</elevation>
</quantity>
;
<collectingDate value="2014-09-26">26 Sep. 2014</collectingDate>
; hand collected;
<collectorName>J. Means</collectorName>
,
<collectorName>P. Marek</collectorName>
,
<collectorName>E. Francis</collectorName>
,
<collectorName>K. Lawler</collectorName>
,
<collectorName>N. Zegler</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE00227">MPE00227</accessionNumber>
, 229 •
<specimenCount type="male">4 ♂♂</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:4758C63B42BE3256296669E76C1729AD" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="Montgomery County" stateProvince="Virginia">Montgomery County</location>
,
<collectorName>Blacksburg</collectorName>
, 803
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:22D5A4FB093422936E580135E6863F03" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="Airport Rd." stateProvince="Virginia">Airport Rd.</location>
;
<geoCoordinate degrees="37.2125" direction="north" orientation="latitude" precision="5" value="37.2125">37.2125°N</geoCoordinate>
, -
<geoCoordinate degrees="80.4053" direction="west" orientation="longitude" precision="5" value="-80.4053">80.4053°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.81" unit="m" value="581.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.81" unit="m" value="581.0">581 m</elevation>
</quantity>
;
<collectingDate value="2018-03-03">3 Mar. 2018</collectingDate>
; hand collected;
<collectorName>J. Means</collectorName>
,
<collectorName>P. Marek</collectorName>
,
<collectorName>G. Schiermeyer</collectorName>
,
<collectorName>C. Hall</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03829">MPE03829</accessionNumber>
, 3830, 3833, 3835 •
<specimenCount type="female">4 ♀♀</specimenCount>
; same collection data as preceding; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03831">MPE03831</accessionNumber>
, 3832, 3834, 3836 •
<specimenCount type="male">1 ♂</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:80511AF8E50C836AA29EF01C9557C511" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="Montgomery County" stateProvince="Virginia">Montgomery County</location>
,
<collectorName>Blacksburg</collectorName>
, 505
<collectorName>Fairview Ave.</collectorName>
, ex puddle of water at base of driveway in neighborhood;
<geoCoordinate degrees="37.212" direction="north" orientation="latitude" precision="55" value="37.212">37.212°N</geoCoordinate>
, -
<geoCoordinate degrees="80.40663" direction="west" orientation="longitude" precision="1" value="-80.40663">80.40663°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.4" unit="m" value="640.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.4" unit="m" value="640.0">640 m</elevation>
</quantity>
;
<collectingDate value="2020-03-13">13 Mar. 2020</collectingDate>
; hand collected;
<collectorName>P. Marek</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE05027">MPE05027</accessionNumber>
•
<specimenCount type="male">1 ♂</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:0934FB1910EBAC455A16097CFBC07B3B" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="Montgomery County" stateProvince="Virginia">Montgomery County</location>
,
<collectorName>Blacksburg</collectorName>
, jct.
<collectorName>Airport</collectorName>
and
<collectorName>Fairview Rds.</collectorName>
;
<geoCoordinate degrees="37.2122" direction="north" orientation="latitude" precision="5" value="37.2122">37.2122°N</geoCoordinate>
, -
<geoCoordinate degrees="80.4060" direction="west" orientation="longitude" precision="5" value="-80.406">80.4060°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.34" unit="m" value="634.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.34" unit="m" value="634.0">634 m</elevation>
</quantity>
;
<collectingDate value="2018-02-19">19 Feb. 2018</collectingDate>
; hand collected;
<collectorName>P. Marek</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03808">MPE03808</accessionNumber>
•
<specimenCount type="female">1 ♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:32F9A31E69B6F9944A2933CA5A910344" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="Montgomery County" stateProvince="Virginia">Montgomery County</location>
,
<collectorName>Blacksburg</collectorName>
,
<collectorName>Virginia Tech</collectorName>
campus, southern end of
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:7CBE66AB138B9D34152C3B3F99F67D49" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="Duck Pond" stateProvince="Virginia">Duck Pond</location>
;
<geoCoordinate degrees="37.2251" direction="north" orientation="latitude" precision="5" value="37.2251">37.2251°N</geoCoordinate>
, -
<geoCoordinate degrees="80.4272" direction="west" orientation="longitude" precision="5" value="-80.4272">80.4272°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.2299999999999995" unit="m" value="623.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.2299999999999995" unit="m" value="623.0">623 m</elevation>
</quantity>
;
<collectingDate value="2017-06-08">8 June 2017</collectingDate>
;
<collectorName>G. Scheirmeyer</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03705">MPE03705</accessionNumber>
•
<specimenCount type="male">1 ♂</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:1C54CEFED5B18E5693A7F16DED3FD770" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="Montgomery County" stateProvince="Virginia">Montgomery County</location>
,
<collectorName>Blacksburg</collectorName>
;
<geoCoordinate degrees="37.1849" direction="north" orientation="latitude" precision="5" value="37.1849">37.1849°N</geoCoordinate>
, -
<geoCoordinate degrees="80.4087" direction="west" orientation="longitude" precision="5" value="-80.4087">80.4087°W</geoCoordinate>
;
<collectingDate value="1956-10-28">28 Oct. 1956</collectingDate>
;
<collectorName>R. Hoffman</collectorName>
leg.; VMNH NAN0234 •
<specimenCount type="male">20 ♂♂</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:42AA1CE23385C0BDA3439A1CC999C293" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="Montgomery County" stateProvince="Virginia">Montgomery County</location>
,
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:39B6279B8CB55AA5015AE6A110E2808A" country="United States" county="Floyd County" latitude="37.1849" longLatPrecision="6" longitude="-80.4087" municipality="Goose Creek" name="south Blacksburg" stateProvince="Virginia">south Blacksburg</location>
, nr jct.
<collectorName>Rt.</collectorName>
775 and
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355391" accessionNumber="MPE00253, MPE00262" collectingDate="1950-10" collectingDateMax="2014-12-27" collectingDateMin="1950-10" collectorName="R. Hoffman, Blacksburg, Trillium Vale, Highland Farm, J. Means, K. Lawler, Iron Mtn., For, Suppl." country="US" county="Montgomery County" elevation="574" latitude="36.71" location="Pulaski County" longLatPrecision="6" longitude="-81.6188" municipality="Montgomery County" specimenCount="16" specimenCount-female="3" specimenCount-male="13" stateProvince="Virginia">
<collectingCountry name="United States of America">US</collectingCountry>
460, under stones in grassy field;
<geoCoordinate degrees="37.1849" direction="north" orientation="latitude" precision="5" value="37.1849">37.1849°N</geoCoordinate>
, -
<geoCoordinate degrees="80.4087" direction="west" orientation="longitude" precision="5" value="-80.4087">80.4087°W</geoCoordinate>
;
<collectingDate value="1956-11-03">3 Nov. 1956</collectingDate>
;
<collectorName>R. Hoffman</collectorName>
leg.; VMNH NAN0320, 321 •
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Montgomery County</collectingCounty>
;
<collectingDate value="1950-10">Oct. 1950</collectingDate>
;
<collectorName>R. Hoffman</collectorName>
leg.; VMNH NAN0235 •
<specimenCount type="male">5 ♂♂</specimenCount>
;
<collectingMunicipality>Montgomery County</collectingMunicipality>
,
<collectorName>Blacksburg</collectorName>
,
<collectorName>Trillium Vale</collectorName>
;
<geoCoordinate degrees="37.2294" direction="north" orientation="latitude" precision="5" value="37.2294">37.2294°N</geoCoordinate>
, -
<geoCoordinate degrees="80.4141" direction="west" orientation="longitude" precision="5" value="-80.4141">80.4141°W</geoCoordinate>
;
<collectingDate value="1950-10-10">10 Oct. 1950</collectingDate>
;
<collectorName>R. Hoffman</collectorName>
leg.; VMNH NAN0243 •
<specimenCount type="male">6 ♂♂</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:4626592B72E266DD7BC5CABFBB94DB70" country="US" county="Montgomery County" latitude="36.71" longLatPrecision="6" longitude="-81.6188" municipality="Montgomery County" name="Pulaski County" stateProvince="Virginia">Pulaski County</location>
,
<collectorName>Highland Farm</collectorName>
, bottom of hill in dry stream bed;
<geoCoordinate degrees="37.1775" direction="north" orientation="latitude" precision="5" value="37.1775">37.1775°N</geoCoordinate>
, -
<geoCoordinate degrees="80.6456" direction="west" orientation="longitude" precision="5" value="-80.6456">80.6456°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.74" unit="m" value="574.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.74" unit="m" value="574.0">574 m</elevation>
</quantity>
;
<collectingDate value="2014-12-27">27 Dec. 2014</collectingDate>
; hand collected;
<collectorName>J. Means</collectorName>
,
<collectorName>K. Lawler</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE00253">MPE00253</accessionNumber>
-55, 259-61 •
<specimenCount type="female">3 ♀♀</specimenCount>
; same collection data as preceding;
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE00262">MPE00262</accessionNumber>
-64 •
<specimenCount type="male">1 ♂</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:11ACA8B4D4585F9BBD8EB3A44CF71419:6D6F7A40503D12B6D222CFE7C31AF5C0" country="US" county="Montgomery County" latitude="36.71" longLatPrecision="6" longitude="-81.6188" municipality="Montgomery County" name="Smyth County" stateProvince="Virginia">Smyth County</location>
,
<locationDeviation location="Chilhowie">
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.1265408" unit="mi" value="7.0">7 mi</quantity>
SE of Chilhowie
</locationDeviation>
, NW slope of
<collectorName>Iron Mtn.</collectorName>
;
<geoCoordinate degrees="36.7100" direction="north" orientation="latitude" precision="5" value="36.71">36.7100°N</geoCoordinate>
, -
<geoCoordinate degrees="81.6188" direction="west" orientation="longitude" precision="5" value="-81.6188">81.6188°W</geoCoordinate>
;
<collectingDate value="1964-05-04">4 May 1964</collectingDate>
;
<collectorName>R. Hoffman</collectorName>
leg.; VMNH NAN0003.
<collectorName>For</collectorName>
detailed collection data see
<collectorName>Suppl.</collectorName>
material 7
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adult males of
<taxonomicName authorityName="Means, Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria hokie" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hokie">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria hokie</emphasis>
</taxonomicName>
are distinct from other
<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
and the sympatric
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. ericacea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="ericacea">
<emphasis italics="true" pageId="0" pageNumber="1">N. ericacea</emphasis>
</taxonomicName>
, based on the following combination of characters:
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
.
</emphasis>
Gonopodal acropodite long and curving ventromedially before apex, not medially as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. castanea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="1">N. castanea</emphasis>
</taxonomicName>
or straight as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. ericacea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="ericacea">
<emphasis italics="true" pageId="0" pageNumber="1">N. ericacea</emphasis>
</taxonomicName>
. Distal zone bent ventroposteriorly, with laminate flange encircling tip forming a hood-like structure around dorso-posteriorly projected. Acropodite solenomere laminate, partially obscuring solenomere when viewed laterally (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Nannaria hokie Means, Hennen &amp; Marek in Means et al. 2021, paratype ♂ (VTEC, MPE 00880) left gonopod. A anterior view; red arrow indicates hood-like flange partially obscuring view of projected solenomere B medial view C posterior view; red triangle indicates medial flange. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure35" httpUri="https://binary.pensoft.net/fig/531138" pageId="0" pageNumber="1">35A</figureCitation>
, red arrow). Distal zone lacking flat laminate flange, at 90° angle to solenomere as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. castanea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="1">N. castanea</emphasis>
</taxonomicName>
or laminate, sinuous, directed ventro-cephalically as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. ericacea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="ericacea">
<emphasis italics="true" pageId="0" pageNumber="1">N. ericacea</emphasis>
</taxonomicName>
. Acropodite with small medial flange near apex (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Nannaria hokie Means, Hennen &amp; Marek in Means et al. 2021, paratype ♂ (VTEC, MPE 00880) left gonopod. A anterior view; red arrow indicates hood-like flange partially obscuring view of projected solenomere B medial view C posterior view; red triangle indicates medial flange. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure35" httpUri="https://binary.pensoft.net/fig/531138" pageId="0" pageNumber="1">35C</figureCitation>
, red triangle), not with lateral projection as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. ericacea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="ericacea">
<emphasis italics="true" pageId="0" pageNumber="1">N. ericacea</emphasis>
</taxonomicName>
. Telopodite basal zone height ca. 1/3 length of acropodite, not ca. 1/2 length as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. castanea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="1">N. castanea</emphasis>
</taxonomicName>
, or &lt;1/5 as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. ericacea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="ericacea">
<emphasis italics="true" pageId="0" pageNumber="1">N. ericacea</emphasis>
</taxonomicName>
. Prefemoral process small, thin, curving medially, arising from top of projected acuminate prefemoral spine, not large, laminate as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. ericacea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="ericacea">
<emphasis italics="true" pageId="0" pageNumber="1">N. ericacea</emphasis>
</taxonomicName>
or straight as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. castanea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="1">N. castanea</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
.
</emphasis>
Tergites with orange paranotal spots (Fig.
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Nannaria hokie Means, Hennen &amp; Marek in Means et al. 2021, holotype ♂ (VTEC, MPE 04803) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure36" httpUri="https://binary.pensoft.net/fig/531139" pageId="0" pageNumber="1">36</figureCitation>
). Dark brown background. Dorsum of collum smooth with orange margin.
</paragraph>
<caption doi="10.3897/zookeys.1030.62544.figure35" httpUri="https://binary.pensoft.net/fig/531138" pageId="0" pageNumber="1" start="Figure 35" startId="F35">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 35.</emphasis>
<taxonomicName authorityName="Means, Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria hokie" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hokie">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria hokie</emphasis>
</taxonomicName>
Means, Hennen &amp; Marek in
<bibRefCitation DOI="https://doi.org/10.1093/isd/ixab003" author="Means, JC" journalOrPublisher="Insect Systematics and Diversity" pageId="0" pageNumber="1" pagination="1 - 26" refId="B52" refString="Means, JC, Hennen, DA, Tanabe, T, Marek, PE, 2021. Phylogenetic systematics of the millipede family Xystodesmidae. Insect Systematics and Diversity 5: 1 - 26, DOI: https://doi.org/10.1093/isd/ixab003" title="Phylogenetic systematics of the millipede family Xystodesmidae." url="https://doi.org/10.1093/isd/ixab003" volume="5" year="2021">Means et al. 2021</bibRefCitation>
, paratype ♂ (VTEC, MPE00880) left gonopod.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
anterior view; red arrow indicates hood-like flange partially obscuring view of projected solenomere
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
medial view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
posterior view; red triangle indicates medial flange. Scale bar: 0.5 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1030.62544.figure36" httpUri="https://binary.pensoft.net/fig/531139" pageId="0" pageNumber="1" start="Figure 36" startId="F36">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 36.</emphasis>
<taxonomicName authorityName="Means, Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria hokie" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hokie">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria hokie</emphasis>
</taxonomicName>
Means, Hennen &amp; Marek in
<bibRefCitation DOI="https://doi.org/10.1093/isd/ixab003" author="Means, JC" journalOrPublisher="Insect Systematics and Diversity" pageId="0" pageNumber="1" pagination="1 - 26" refId="B52" refString="Means, JC, Hennen, DA, Tanabe, T, Marek, PE, 2021. Phylogenetic systematics of the millipede family Xystodesmidae. Insect Systematics and Diversity 5: 1 - 26, DOI: https://doi.org/10.1093/isd/ixab003" title="Phylogenetic systematics of the millipede family Xystodesmidae." url="https://doi.org/10.1093/isd/ixab003" volume="5" year="2021">Means et al. 2021</bibRefCitation>
, holotype ♂ (VTEC, MPE04803) coloration. Scale bar: 4.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="1">♂ paratype (VTEC, MPE00880): BL = 18.0, CW = 2.9, IW = 2.3, ISW = 0.8, B11W = 3.5, B11H = 2.2; ♀ paratype (VTEC, MPE00881): BL = 22.9, CW = 3.1, IW = 1.6, ISW = 0.8, B11W = 3.9, B11H = 2.4.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph pageId="0" pageNumber="1">
Little morphological variation exists between individuals collected from Montgomery County, Virginia; however, populations in Floyd and Pulaski counties, Virginia display a level of variation that suggests a closer comparison of these populations may be warranted. Both the Floyd and Pulaski County specimens have a much reduced acropodite
<normalizedToken originalValue="“hood”">&quot;hood&quot;</normalizedToken>
(Fig.
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Nannaria hokie Means, Hennen &amp; Marek, 2021 variation A Floyd County, Virginia non-type ♂ (VMNH, NAN 0003; red arrow indicates reduced lateral and medial flanges) B Pulaski County, Virginia non-type ♂ (VTEC, MPE 00253); red triangle indicates prefemoral process lacking prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure37" httpUri="https://binary.pensoft.net/fig/531140" pageId="0" pageNumber="1">37A</figureCitation>
, red arrow), and the Pulaski County specimens have highly variable prefemoral processes, including forms with and without a prefemoral spine (Fig.
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Nannaria hokie Means, Hennen &amp; Marek, 2021 variation A Floyd County, Virginia non-type ♂ (VMNH, NAN 0003; red arrow indicates reduced lateral and medial flanges) B Pulaski County, Virginia non-type ♂ (VTEC, MPE 00253); red triangle indicates prefemoral process lacking prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure37" httpUri="https://binary.pensoft.net/fig/531140" pageId="0" pageNumber="1">37B</figureCitation>
, red triangle).
</paragraph>
<caption doi="10.3897/zookeys.1030.62544.figure37" httpUri="https://binary.pensoft.net/fig/531140" pageId="0" pageNumber="1" start="Figure 37" startId="F37">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 37.</emphasis>
<taxonomicName authorityName="Means, Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria hokie" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hokie">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria hokie</emphasis>
</taxonomicName>
Means, Hennen &amp; Marek, 2021 variation
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Floyd County, Virginia non-type ♂ (VMNH, NAN0003; red arrow indicates reduced lateral and medial flanges)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
Pulaski County, Virginia non-type ♂ (VTEC, MPE00253); red triangle indicates prefemoral process lacking prefemoral spine. Scale bar: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Known only from southwestern Virginia, primarily in the vicinity of Blacksburg, though a single specimen collected from Smyth County, Virginia extends the distribution of the species considerably (Virginia: Floyd, Montgomery, Pulaski, and Smyth counties; Suppl. material 7, Fig.
<figureCitation captionStart="Figure 126" captionStartId="F126" captionText="Figure 126. Distribution map of Nannaria minor species group collection localities in the central Appalachians." figureDoi="10.3897/zookeys.1030.62544.figure126" httpUri="https://binary.pensoft.net/fig/531229" pageId="0" pageNumber="1">126</figureCitation>
). Distribution area: 1,668 km2; status: SRE.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Individuals of
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. hokie" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="hokie">
<emphasis italics="true" pageId="0" pageNumber="1">N. hokie</emphasis>
</taxonomicName>
have been collected from mesic hardwood forests, composed of maple, oak, tuliptree, beech, spicebush, and witch hazel, under deciduous leaf litter, logs, and fallen bark. The Pulaski County specimens were all collected from a 1 m2 section of a dry creek, taken from under 1-2 cm hard packed soil on the bank of the creek.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation DOI="https://doi.org/10.1093/isd/ixab003" author="Means, JC" journalOrPublisher="Insect Systematics and Diversity" pageId="0" pageNumber="1" pagination="1 - 26" refId="B52" refString="Means, JC, Hennen, DA, Tanabe, T, Marek, PE, 2021. Phylogenetic systematics of the millipede family Xystodesmidae. Insect Systematics and Diversity 5: 1 - 26, DOI: https://doi.org/10.1093/isd/ixab003" title="Phylogenetic systematics of the millipede family Xystodesmidae." url="https://doi.org/10.1093/isd/ixab003" volume="5" year="2021">Means et al. (2021)</bibRefCitation>
named
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. hokie" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="hokie">
<emphasis italics="true" pageId="0" pageNumber="1">N. hokie</emphasis>
</taxonomicName>
for its type locality, the campus of Virginia Tech. A
<emphasis italics="true" pageId="0" pageNumber="1">hokie</emphasis>
is a popular term for a member of the Virginia Tech community.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
United States, Virginia, Montgomery County, Blacksburg, Virginia Tech campus, Stadium Woods,
<geoCoordinate degrees="37.2216" direction="north" orientation="latitude" precision="5" value="37.2216">37.2216°N</geoCoordinate>
, -
<geoCoordinate degrees="80.4159" direction="west" orientation="longitude" precision="5" value="-80.4159">80.4159°W</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="notes">
<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph pageId="0" pageNumber="1">
In the original publication
<bibRefCitation DOI="https://doi.org/10.1093/isd/ixab003" author="Means, JC" journalOrPublisher="Insect Systematics and Diversity" pageId="0" pageNumber="1" pagination="1 - 26" refId="B52" refString="Means, JC, Hennen, DA, Tanabe, T, Marek, PE, 2021. Phylogenetic systematics of the millipede family Xystodesmidae. Insect Systematics and Diversity 5: 1 - 26, DOI: https://doi.org/10.1093/isd/ixab003" title="Phylogenetic systematics of the millipede family Xystodesmidae." url="https://doi.org/10.1093/isd/ixab003" volume="5" year="2021">Means et al. (2021</bibRefCitation>
: 17) designated a male holotype (VTEC MPE04803), five male paratypes (VTEC MPE00880, 882, 883; VMNH MPE00884, 886) and two female paratypes (VTEC MPE00881, 885) from the multiple locations on the campus of Virginia Tech collected by the authors from various dates in 2015 and 2019.
</paragraph>
</subSubSection>
</treatment>
</document>
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