89 lines
12 KiB
XML
89 lines
12 KiB
XML
<document id="7CC735A04F8E4F40452D6685EE89E4C4" ENCODING="UTF-8" ID-GBIF-Dataset="62345364-7287-4277-9efa-f628aab56456" ModsDocID="z01551p031" checkinTime="1247324133964" checkinUser="thomas" docAuthor="Jonathan H. Walsh & David A. Ebert" docDate="2007" docId="0D39DACE952DCF7DB177C65CF1966697" docLanguage="en" docName="2007_Walsh_Ebert_gg1.xml" docOrigin="Zootaxa 1551" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:DD00E8E3-2C28-41A8-A401-E0D00DA325B9" docTitle="Squatina Dumeril" docType="treatment" docVersion="12" lastPageNumber="46" masterDocId="B7D8EEA7B6D99A7AE0DA9D9FF27F9CAF" masterDocTitle="A review of the systematics of western North Pacific angel sharks, genus Squatina, with redescriptions of Squatina formosa, S. japonica, and S. nebulosa (Chondrichthyes: Squatiniformes, Squatinidae)." masterLastPageNumber="47" masterPageNumber="31" pageNumber="46" updateTime="1701314684852" updateUser="plazi">
|
|
<mods:mods id="680BC7BF3BE1CDF54E4C0181A465FEA3" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="A57DB6AE3BCA13D7BCA8C3501D5571F6">
|
|
<mods:title id="946F8B2B2C79F30B017F0C9ADFA42649">A review of the systematics of western North Pacific angel sharks, genus Squatina, with redescriptions of Squatina formosa, S. japonica, and S. nebulosa (Chondrichthyes: Squatiniformes, Squatinidae).</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="D648E8BA7E17119FBFCAB589AAC3BE63" type="personal">
|
|
<mods:role id="E55FF00E4042C2E07D002F748F52F1F9">
|
|
<mods:roleTerm id="38F9AD1B8FAF70E5915A649BB5938ED8">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="8F238077FA20D40F8960385A757C22F4">Jonathan H. Walsh</mods:namePart>
|
|
</mods:name>
|
|
<mods:name id="9952E32D4BC957B2D2C9AD89E5C14616" type="personal">
|
|
<mods:role id="34FEFB3F9FC11430FF1003E89177D72A">
|
|
<mods:roleTerm id="8DB3366A7B37B8261E4790C296394669">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="023987E6316C9383B4F7B23D06E632C2">David A. Ebert</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource id="6AD1F546CCB3B8AC9826FFEB11693663">text</mods:typeOfResource>
|
|
<mods:relatedItem id="6ABBD5A23F00D0F18AD3C69F101FBB33" type="host">
|
|
<mods:titleInfo id="CB6985B4AAC2D30AD075F09C61C2159B">
|
|
<mods:title id="7C245A9D2B0B435C085AB6939CFFF0F9">Zootaxa</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="B5014C62A04707358DE7D91477124E8C">
|
|
<mods:date id="E8692870AD0B1D464A21AD647F14F4D1">2007</mods:date>
|
|
<mods:detail id="A2A11BA93CB2BF075AFBFBD8A1CB327D" type="volume">
|
|
<mods:number id="32F70ACCA3D487CE354A89274438459B">1551</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="FE9E7F35A8D73FC3EE6A2755A5288C88" unit="page">
|
|
<mods:start id="A955E5568916D1FD2AEC562D2085E3BD">31</mods:start>
|
|
<mods:end id="D45B79575D69C66DC7D4CE66E13638AD">47</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location id="105385BC170A5169B44ACDE888B76726">
|
|
<mods:url id="C31A8716AC18ABF899B99ED755CAEBFC">http://www.zoobank.org/urn:lsid:zoobank.org:pub:DD00E8E3-2C28-41A8-A401-E0D00DA325B9</mods:url>
|
|
</mods:location>
|
|
<mods:classification id="2EB25EFCF141F12EE0CA9E495DE8D4E4">journal article</mods:classification>
|
|
<mods:identifier id="7AC5FFEF0C502E4459AFFADD7F4C9AD4" type="Plazi-Custom">z01551p031</mods:identifier>
|
|
<mods:identifier id="9E5BE2DA619BAC344EAD7E90CBBA4148" type="ZooBank">DD00E8E3-2C28-41A8-A401-E0D00DA325B9</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="0D39DACE952DCF7DB177C65CF1966697" ID-DOI="http://doi.org/10.5281/zenodo.6242940" ID-GBIF-Taxon="100121801" ID-Zenodo-Dep="6242940" LSID="urn:lsid:plazi:treatment:0D39DACE952DCF7DB177C65CF1966697" httpUri="http://treatment.plazi.org/id/0D39DACE952DCF7DB177C65CF1966697" lastPageNumber="46" pageNumber="46">
|
|
<subSubSection id="ED71E2DF3B41F8DD0553DEBD74EEC702" type="nomenclature">
|
|
<paragraph id="2073667C78DC0D801C0E53A54E6F27BE" pageNumber="46">
|
|
[[ Genus
|
|
<taxonomicName id="5CC0DBB78851C070787C5058265B2B26" ID-CoL="7LRS" ID-ENA="55141" LSID-ZBK="urn:lsid:zoobank.org:act:2B9CCE5C-316C-44E4-B124-7DF961210153" class="Elasmobranchii" family="Squatinidae" genus="Squatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Squatina Risso 1810:45" order="Squatiniformes" pageNumber="46" phylum="Chordata" rank="genus">Squatina Dumeril</taxonomicName>
|
|
]]
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="4FF42161C7EED62653B4988A348CBF05" type="discussion">
|
|
<paragraph id="3E6435B761CBF7A982E6AA44270A9493" pageNumber="46">
|
|
The main confusion among WNP squatinids lies in differentiating
|
|
<taxonomicName id="0A5930C1C069D0AAA32F65FE1BF8CDE5" LSID-ZBK="urn:lsid:zoobank.org:act:C995E0F2-9BA7-4F6B-9726-E20E986E0BFE" class="Elasmobranchii" family="Squatinidae" genus="Squatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Squatina formosa Shen & Ting 1972:21" order="Squatiniformes" pageNumber="46" phylum="Chordata" rank="species" species="formosa">S. formosa</taxonomicName>
|
|
and
|
|
<taxonomicName id="AFD1D766AD271D86389031EE2BD225CD" LSID-ZBK="urn:lsid:zoobank.org:act:D454401C-992B-4434-8434-FA522035C589" class="Elasmobranchii" family="Squatinidae" genus="Squatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Squatina nebulosa Regan 1906:439" order="Squatiniformes" pageNumber="46" phylum="Chordata" rank="species" species="nebulosa">S. nebulosa</taxonomicName>
|
|
from one another. The main character used in the past to differentiate these two species was the number of dermal lobes at the corners of the mouth (Chen 1963, Nakabo 2002, Compagno 2005a). Based upon our examination, this character does not appear to be robust in separating these species. Although all
|
|
<taxonomicName id="81DC5C36358C8E48ACF3CD8A40FB1735" LSID-ZBK="urn:lsid:zoobank.org:act:C995E0F2-9BA7-4F6B-9726-E20E986E0BFE" class="Elasmobranchii" family="Squatinidae" genus="Squatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Squatina formosa Shen & Ting 1972:21" order="Squatiniformes" pageNumber="46" phylum="Chordata" rank="species" species="formosa">S. formosa</taxonomicName>
|
|
individuals studied had one dermal lobe,
|
|
<taxonomicName id="B68F22E6B462A3975C424A3C39ADC8EA" LSID-ZBK="urn:lsid:zoobank.org:act:D454401C-992B-4434-8434-FA522035C589" class="Elasmobranchii" family="Squatinidae" genus="Squatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Squatina nebulosa Regan 1906:439" order="Squatiniformes" pageNumber="46" phylum="Chordata" rank="species" species="nebulosa">S. nebulosa</taxonomicName>
|
|
specimens had one or two lobes, invalidating this character as a method of separating the two species. Additionally, the number of dermal lobes was often difficult to determine since differentiation between lobes is often vague and these lobes are susceptible to damage during capture and or preservation of specimens. A combination of characters should be used to identify individuals of these two species, since the differences are subtle and a great deal of overlap exists among many characters. The best characters available, based on our findings, appear to be the shape of the caudal pelvic and dorsal fins, the ratio of standardized pelvic girdle distance with the standardized head length, and the shape and height of the upper lip arch. The upper lip arch appears to be of particular value as a field character. It is a feature that is not prone to damage and is easy to assess quickly in the field. More specimens are needed to assess the utility of this character but it appears to be useful for all WNP squatinids.
|
|
</paragraph>
|
|
<paragraph id="0E849B809BA8EAA01A9BDE52912AC2D7" pageNumber="46">Based on the key characters presented herein, we provide the following key to the WNP squatinids:</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="A4F58A46DC573CE8E18A87160D53B1BE" type="key">
|
|
<paragraph id="3565514C08FBE9229166AF252D86BE5E" pageNumber="46">
|
|
Key to Western North Pacific
|
|
<taxonomicName id="63E210C4901EE9EAF3B54DF257D63D09" LSID-ZBK="urn:lsid:zoobank.org:act:2B9CCE5C-316C-44E4-B124-7DF961210153" class="Elasmobranchii" family="Squatinidae" genus="Squatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Squatina Risso 1810:45" order="Squatiniformes" pageNumber="46" phylum="Chordata" rank="genus">Squatina</taxonomicName>
|
|
Species
|
|
</paragraph>
|
|
<paragraph id="D4D8B2FDAB8B8F7C6BAD35820196BABB" pageNumber="46">1. Pelvic fin tips do not extend to origin of first dorsal fin..............................................................................2.</paragraph>
|
|
<paragraph id="2178391BE160415E01C6835A5038AC98" pageNumber="46">- Pelvic fin tips extend to or surpass origin of first dorsal fin........................................................................3.</paragraph>
|
|
<paragraph id="F5B6DD7A33FC3CFDB75F69A4EF7F46E7" pageNumber="46">
|
|
2. A prominent row of thorn-like denticles extending from mid-back to caudal peduncle; no distinct ocelli on posterior lobes of the pectoral fins................................................................................................
|
|
<taxonomicName id="E4D5E0CE5DB4763E4F77C57E1A074079" LSID-ZBK="urn:lsid:zoobank.org:act:605F995D-6CF7-494C-AD85-65000563A021" class="Elasmobranchii" family="Squatinidae" genus="Squatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Squatina japonica Bleeker 1858:40" order="Squatiniformes" pageNumber="46" phylum="Chordata" rank="species" species="japonica">S. japonica</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="C19016F3B56EFF05F97C378C111F878C" pageNumber="46">
|
|
- No row of thorn-like denticles extending from the mid-back to the caudal peduncle, distinct paired ocelli on the posterior lobes of the pectoral fins.........................................................................
|
|
<taxonomicName id="86CD886F98FA9AAB8B2CB3B480F92D1C" LSID-ZBK="urn:lsid:zoobank.org:act:A0160AC7-329C-46D5-9BEA-7DC2BACCF091" class="Elasmobranchii" family="Squatinidae" genus="Squatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Squatina tergocellatoides Chen 1963:98" order="Squatiniformes" pageNumber="46" phylum="Chordata" rank="species" species="tergocellatoides">S. tergocellatoides</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="1DECBDAE7A55626D368DBD0D8E290BAD" pageNumber="46">
|
|
3. Upper lip arch semi-circular in shape (>1.5% TL in height); dorsals are lobed with a curvilinear anterior margin; pelvic girdle distance 1.4 times or less head length; caudal fin is lobed, especially dorsally, with a curvilinear postventral caudal margin...........................................................................................
|
|
<taxonomicName id="8FA588EBECB19FCEE8F2C30A8982EE84" LSID-ZBK="urn:lsid:zoobank.org:act:C995E0F2-9BA7-4F6B-9726-E20E986E0BFE" class="Elasmobranchii" family="Squatinidae" genus="Squatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Squatina formosa Shen & Ting 1972:21" order="Squatiniformes" pageNumber="46" phylum="Chordata" rank="species" species="formosa">S. formosa</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="6C7D63D24E5541EEC01E54D2873B3E57" pageNumber="46">
|
|
- Upper lip arch is not semi-circular in shape (<1.5% TL in height); dorsals are not lobed (angular) without a curvilinear anterior margin (straight); pelvic girdle distance greater than 1.4 times head length; caudal fin is not lobed (angular), especially dorsally, without a curvilinear postventral caudal margin (straight) ... .....................................................................................................................................................
|
|
<taxonomicName id="79CCB5D4EC5692CE42741FCD5672786E" LSID-ZBK="urn:lsid:zoobank.org:act:D454401C-992B-4434-8434-FA522035C589" class="Elasmobranchii" family="Squatinidae" genus="Squatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Squatina nebulosa Regan 1906:439" order="Squatiniformes" pageNumber="46" phylum="Chordata" rank="species" species="nebulosa">S. nebulosa</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |