227 lines
33 KiB
XML
227 lines
33 KiB
XML
<document ID-CLB-Dataset="21464" ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="EE46802C29515364BE0153EA8BC41EEE" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole antranohofa Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="269DB81E-05EC-47B5-9DCC-F99B6E4E5007" docUuidSource="ZooBank" docVersion="6" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1732651982216" updateUser="ExternalLinkService">
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<mods:titleInfo id="2541392D8383186B738F440DEB9CF87B">
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<mods:title id="4755EAD19734D8F008E442F7B7946DEC">Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name id="19A21B13DE4692B2F706598AC12C1426" type="personal">
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<mods:role id="25C6557852175743487948885BEBEAF3">
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<mods:roleTerm id="BE69EBC222B38C537656E2DEE590C855">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="13CDAE59486353B57CA45BBDFAE02D24">Salata, Sebastian</mods:namePart>
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<mods:affiliation id="9567428CC2464B51A7847A46E5D7C31B">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="4068D029E38F28B87ED65151F57B424D" type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier id="C08BBC0BF827AA578F229A2A8AB39618" type="email">sdsalata@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="F58669A118C0D6A00FE71948E8C73BF2" type="personal">
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</mods:role>
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<mods:namePart id="F60E093CF6B8D9A2944CB63B98B59F2A">Fisher, Brian L.</mods:namePart>
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<mods:affiliation id="5F4DE701EE62892DAF322911D876DC66">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="F5E28F3FCC0E3BD13EBCA22F6AD9DDDC" type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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<mods:typeOfResource id="1DD8E5CEC340C82486D9671F64E46182">text</mods:typeOfResource>
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<mods:title id="2F646A0684582759C2E1878138E3E7A8">ZooKeys</mods:title>
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<mods:part id="28DC57705EDDF76531AA940E71182CBC">
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<mods:date id="2300F4FB94DCA35D1236BCA236A252FE">2020</mods:date>
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<mods:detail id="1AE566FDCBD33095B70950A590D7129E" type="volume">
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<mods:number id="BD2DF0495CD8FF1033F88F539A06FCF7">949</mods:number>
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<mods:start id="5BD4C8A34B88328EF40EBE91FD6F34D0">1</mods:start>
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<mods:url id="E5938F168B8241315D80B6D4CEF82B56">http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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<mods:classification id="0AC876479C96440DDFA7119A7B34BBD9">journal article</mods:classification>
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<mods:identifier id="72D63912BE7149E2CEACED680327749B" type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<mods:identifier id="3FDA31C61C0A526D2B193E6E327A8915" type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
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<mods:identifier id="F3797BBB88C80176EA9642192DCCBE33" type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
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<treatment id="EE46802C29515364BE0153EA8BC41EEE" ID-GBIF-Taxon="165090158" LSID="urn:lsid:zoobank.org:act:269DB81E-05EC-47B5-9DCC-F99B6E4E5007" httpUri="http://treatment.plazi.org/id/EE46802C29515364BE0153EA8BC41EEE" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="97D62B3028A54378D4B47B36FFF48A4B" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="999516C92117141D47DE46F69B7D33F5" pageId="0" pageNumber="1">
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<taxonomicName id="56C8161668CC79D8047A11184C9EB4C4" ID-CoL="CMKVC" LSID="EE46802C-2951-5364-BE01-53EA8BC41EEE" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole antranohofa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="antranohofa" status="sp. nov.">Pheidole antranohofa</taxonomicName>
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<taxonomicNameLabel id="2FD9F7D328808BDA146C690BB2D555AE" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="EA8F7E3BD79EB912E307383BC93034C5" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">Figs 25A-F</figureCitation>
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<figureCitation id="B89809F51C0319DF5631AEB1E8FD223A" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">, 63G</figureCitation>
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<figureCitation id="5D089BC89100C5E3FD9487DCE9413AB4" captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Distribution. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T)." figureDoi="10.3897/zookeys.949.51269.figure65" httpUri="https://binary.pensoft.net/fig/432234" pageId="0" pageNumber="1">, 65G</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D8440855978FDAF85690EF87B1DE12A9" pageId="0" pageNumber="1" type="type material">
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<paragraph id="9861BB864CE281C9479334B44EE6CCC6" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="616414985E675CC1EE39C7072B84D0CD" pageId="0" pageNumber="1">
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<emphasis id="39DABA0B3093014C12EB39CCACEA9759" bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Antsiranana; Parc National de Marojejy, Antranohofa, 26.6 km 31°NNE Andapa, 10.7 km 318°NW Manantenina;
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<geoCoordinate id="8C8981CFABF224265728BC93028C4C86" degrees="14.44333" direction="south" orientation="latitude" precision="1" value="-14.44333">-14.44333</geoCoordinate>
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,
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<geoCoordinate id="D3EFC52FC8B58442C2E21CEFAB788263" degrees="49.74333" direction="east" orientation="longitude" precision="1" value="49.74333">49.74333</geoCoordinate>
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; alt. 1325 m; 20 Nov 2003; B. L. Fisher et al. leg.; montane rainforest, ex root mat, ground layer; BLF09196; CASENT0923295, top specimen on the pin (CASC).
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<emphasis id="FD38651F1FAD7C6A638EEF5D55ABA94A" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes.</emphasis>
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• 3w., 2s; same data as for holotype; CASENT0923294, CASENT0499827CASENT0499829 (CASC, MHNG, PBZT).
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</paragraph>
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<caption id="6D0366C87E6F8DF16BB9D32A3839466A" doi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1" start="Figure 25" startId="F25">
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<paragraph id="44594A0CD8AA7DD4B3F87D64986ED556" pageId="0" pageNumber="1">
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<emphasis id="8F6C2F8249441B61B1312988C9E2EA27" bold="true" pageId="0" pageNumber="1">Figure 25.</emphasis>
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<taxonomicName id="774FA285AC66F21B476656AE3C746098" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole antranohofa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="antranohofa">
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<emphasis id="ABB433B0B82643494DBCDF395826435B" italics="true" pageId="0" pageNumber="1">Pheidole antranohofa</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis id="D92797EDF28D1C7553F67F6FE9D145D5" bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis id="1007E3AD11D04AAA82041A26AE10C74F" bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis id="8721286028CE52BD34A7A2AEA50E1DCC" bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0923294) and full-face view (
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<emphasis id="477CCF4F122F6201EC9949CC47EBA12C" bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis id="B59D7B644343B19389BD64C8C04BB5D8" bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis id="D2147C15FA21FE703279ADF46E11F487" bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0923295).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="EC47DA46B9174E11C8A6BA1D6BE01870" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="BAC1DE516969852291791996BB95AA2D" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="3280F780EFC8A1534009F8ECF8361CDD" pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis id="6506DE9D34FEDA5E244EDA66E6FE7A45" bold="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: •1w., 1q.; Parc National de Marojejy, Antranohofa, 26.6 km 31°NNE Andapa, 10.7 km 318°NW Manantenina;
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<geoCoordinate id="C31246CDD1911CDC6BEE3B32514E4EB7" degrees="14.44333" direction="south" orientation="latitude" precision="1" value="-14.44333">-14.44333</geoCoordinate>
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,
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<geoCoordinate id="EBCB91E8BD359C6F985F66DE6E04029C" degrees="49.74333" direction="east" orientation="longitude" precision="1" value="49.74333">49.74333</geoCoordinate>
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; alt. 1325 m; 20 Nov 2003; B. L. Fisher et al. leg.; montane rainforest, ex rotten tree stump; BLF09202 (CASC). •1w., 1s.; Parc National de Marojejy, Antranohofa, 26.6 km 31°NNE Andapa, 10.7 km 318°NW Manantenina;
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<geoCoordinate id="8669C2B8947299FD56BA627F1BF24EDB" degrees="14.44333" direction="south" orientation="latitude" precision="1" value="-14.44333">-14.44333</geoCoordinate>
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,
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<geoCoordinate id="D6E1A310F25E3BDFE4CEF29B129BA8ED" degrees="49.74333" direction="east" orientation="longitude" precision="1" value="49.74333">49.74333</geoCoordinate>
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; alt. 1325 m; 20 Nov 2003; B. L. Fisher et al. leg.; montane rainforest, ex rotten log; BLF09215 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="DDE11324788F3641EE6F280E474F5725" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="9E6ED14DDF4BDCD1C80B6C66B70A7993" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="6590380D00618226B5FAB4E1E5949EAB" pageId="0" pageNumber="1">
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Moderately large species.
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<emphasis id="EB03BF4F947D340060757264AEF1BF47" bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Head in full-face view sub-oval, widening posteriorly, with anterior and posterior sides convex, in lateral view sub-oval; ventral and dorsal faces convex; sides of the head with dense, short, suberect pilosity; anterior and medial parts of frons with moderately dense and thick, longitudinal and sometimes interrupted rugae with distinctly rugofoveolate interspaces; posterolateral sides with longitudinally irregular and thick rugae with distinctly rugofoveolate interspaces, area posterolateral from eyes without smooth notches; scape, when laid back, exceeding the midlength of head by approximately one-fifth of its length; inner hypostomal teeth distinct, moderately long, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth lobe-like, wider and higher than inner hypostomal teeth, apex directed outward; inner and outer hypostomal teeth closely spaced and connected by indistinct concavity; promesonotum predominantly rugofoveolate, with additional sparse, thin, transverse rugae on pronotum; propodeum with fine rugofoveolae; gaster smooth; body yellowish orange.
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<emphasis id="F4D8312EA2FD0DFE945B1CF7DB055789" bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Head foveolate with additional indistinct rugulae on frons and vertex, area posterolateral from eyes with reduced sculpture, on posterior part predominantly smooth; scape, when laid back, surpassing the posterior head margin by two-fifths of its length; promesonotum moderately high and short; promesonotal groove present; propodeal spines indistinct, triangular; mesosoma smooth and shiny, only dorsum with few transverse, indistinct rugulae and propodeum with indistinct and sparse foveolae; body dark yellow.
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</paragraph>
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</subSubSection>
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<subSubSection id="BA1F7F97EE775B811C701E8DF9406C12" pageId="0" pageNumber="1" type="description">
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<paragraph id="BD2AFC8D35D8610CD558D716E8932432" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="9BD7AB2847EED55BA02DBDAE3A4FBF55" pageId="0" pageNumber="1">
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<emphasis id="0CF75D6F0715A11FFB3EA52D21EC5FE6" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis id="82EFEC9FD3B7DF0314BFC921884473C5" italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 4): HL: 1.18-1.23 (1.21); HW: 1.21-1.27 (1.24); SL: 0.67-0.7 (0.69); EL: 0.14-0.16 (0.15); WL: 1.1-1.13 (1.12); PSL: 0.2-0.21 (0.2); MTL: 0.7-0.73 (0.72); PNW: 0.54-0.59 (0.57); PTW: 0.15-0.17 (0.16); PPW: 0.33-0.35 (0.34); CI: 96.7-97.8 (97.3); SI: 54.1-57.8 (55.4); PSLI: 16.3-17.7 (16.9); PPI: 43.4-50.5 (46.6); PNI: 43.9-46.9 (45.6); MTI: 56.7-58.8 (57.6).
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</paragraph>
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<paragraph id="8C43C3E6A31EBEBC191AE10E64E9A283" pageId="0" pageNumber="1">
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<emphasis id="C901E9D815AA4C06F5D82A2B13C6714A" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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In full-face view sub-oval, widening posteriorly, with anterior and posterior sides convex (Fig.
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<figureCitation id="15BD6CA3BCA5AD739969C1E3D0C95DF6" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth invisible. Sides of the head with dense, short, suberect pilosity; whole head with relatively dense, short, decumbent to erect pilosity. Anterior and medial parts of frons with moderately dense and thick, longitudinal and sometimes interrupted rugae with distinctly rugofoveolate interspaces; posterolateral sides with longitudinally irregular and thick rugae with distinctly rugofoveolate interspaces. Occipital lobes with thick, sparse, irregular rugae, interspaces smooth or with fine, indistinct rugofoveolae. Gena with sparse and thick, longitudinal rugae and smooth interspaces. Area posterolateral from eyes with thin, longitudinal rugae with distinctly rugofoveolate interspaces. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, wide and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by approximately one-fifth of its length; pilosity decumbent to suberect (Fig.
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<figureCitation id="26C15C209254DC92363C079E1F2537C5" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25B, D</figureCitation>
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). Inner hypostomal teeth distinct, moderate, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth lobe-like, wider and higher than inner hypostomal teeth, apex directed outward; inner and outer hypostomal teeth closely spaced and connected by indistinct concavity (Fig.
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<figureCitation id="A1F2B86D86460C351F3A7E95CBC15EE4" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">63G</figureCitation>
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).
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<emphasis id="8636F6A8D29D48234C3107CF7F5CD550" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum short, angular, and moderately high, posterior mesonotum moderately steep, mesonotal process indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines moderate, thin, with acute apex; humeral area laterally weakly produced (Fig.
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|
<figureCitation id="CD4F72875836D9D120B5310202E238FE" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25D</figureCitation>
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). Surface shiny, promesonotum predominantly rugofoveolate, with additional sparse, thin, transverse rugae on pronotum; propodeum with fine rugofoveolae. Pilosity sparse, moderately long, and erect (Fig.
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|
<figureCitation id="534E9A7FCA30FD9B6ADE13119AF893D7" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25D, F</figureCitation>
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).
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<emphasis id="745D1FB1A32574AA98458C78B66C5B32" bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
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Shiny with fine and sparse rugofoveolae; node smooth to indistinctly rugulose, low, triangular, with rounded apex, in rear view node dorsoventrally convex; pilosity moderately sparse and erect (Fig.
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|
<figureCitation id="90CD5C7C20E0239353660E5A0BFE93BA" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25D, F</figureCitation>
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).
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<emphasis id="D0C458EC9CFF9ED36A19BC409E7A6C17" bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
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Shiny, with fine and sparse rugulae; in dorsal view postpetiole oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse, and erect (Fig.
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<figureCitation id="530D07F8EF20C9066135545564FB9E9C" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25D, F</figureCitation>
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).
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<emphasis id="650B601235603FF20CFF13E5134CE1D6" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
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|
Shiny and smooth; pilosity sparse, long, and erect (Fig.
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|
<figureCitation id="8F289D6A28EB3065A21368E998EB72B0" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25D, F</figureCitation>
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).
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<emphasis id="235CA8CA17D6F368118923DB8120B70F" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
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Yellowish orange; mandibles and anterior part of head slightly darker (Fig.
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|
<figureCitation id="00FCE8BCC97501349A797373330666C3" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25D, F</figureCitation>
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).
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</paragraph>
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<paragraph id="280EBD77EF2A056B0A4780696CF9E3A9" pageId="0" pageNumber="1">
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|
<emphasis id="344B8DC1C6533B93C2A00F7896A26EB6" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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|
Measurements (
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|
<emphasis id="2D6206736F30D3ED7EFC0C73E3BACEE5" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
|
= 9): HL: 0.57-0.64 (0.59); HW: 0.47-0.53 (0.5); SL: 0.59-0.66 (0.62); EL: 0.09-0.11 (0.1); WL: 0.7-0.79 (0.74); PSL: 0.06-0.08 (0.07); MTL: 0.48-0.56 (0.52); PNW: 0.3-0.38 (0.33); PTW: 0.07-0.1 (0.09); PPW: 0.13-0.16 (0.14); CI: 112.2-122.3 (119.5); SI: 121.6-130.3 (125.5); PSLI: 10.8-12.6 (11.8); PPI: 51.1-70.3 (61.2); PNI: 64.0-70.2 (66.2); MTI: 101.9-108.0 (104.8).
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|
</paragraph>
|
|
<paragraph id="7C61FFDB3B851CD9C97CA3625165EEB0" pageId="0" pageNumber="1">
|
|
<emphasis id="41FC3B4417AFE1A29661FAAF504C3BEE" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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|
Cephalic margin indistinctly convex (Fig.
|
|
<figureCitation id="9EB51C3CD76E0F3709B9778E6F8A2737" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25A</figureCitation>
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|
). Pilosity relatively sparse, long, and subdecumbent. Head foveolate with additional indistinct rugulae on frons and vertex, area posterolateral from eyes with reduced sculpture and partially smooth; antennal sockets with few thick, curved outward rugae and foveolate interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity dense, subdecumbent to erect (Fig.
|
|
<figureCitation id="742D4B804DF39B2133158FCD85A46BFD" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25A, C</figureCitation>
|
|
).
|
|
<emphasis id="2BC26538631C4840BDC76ED7B2795CF9" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
|
In lateral view, promesonotum moderately high, short, arched; promesonotal groove present, distinct; metanotal groove present and distinct; propodeal spines indistinct, triangular, apex acute (Fig.
|
|
<figureCitation id="7FC095D57381C5D268D3EE7D5F4E4839" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25C</figureCitation>
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). Sculpture smooth and shiny, only dorsum with few transverse, indistinct rugulae and propodeum with indistinct and sparse foveolae. Pilosity sparse, moderately long, and erect (Fig.
|
|
<figureCitation id="1A5FAEA4FF7D5167E598B1DC94B37D75" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25C, E</figureCitation>
|
|
).
|
|
<emphasis id="02EB74FE3E796E96FB062602558BC27D" bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
|
|
Peduncle with ventral face relatively straight; node low, knobbed, and small; with few short, erect setae (Fig.
|
|
<figureCitation id="8776CCC663B5994C2A3E4F773E0E792C" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25C, E</figureCitation>
|
|
).
|
|
<emphasis id="C510B7A27BF6A56FD3F02B0C7C89A75B" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
|
With sparse, erect pilosity (Fig.
|
|
<figureCitation id="AF98A0ADEF236309CD58DFAFF84BC97A" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25C, E</figureCitation>
|
|
).
|
|
<emphasis id="7D4C37E50BB4A8349A0B162B14F4D55A" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
|
Dark yellow (Fig.
|
|
<figureCitation id="7BDA22A6BA67C747504F174251031DB1" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pheidole antranohofa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923294) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923295)." figureDoi="10.3897/zookeys.949.51269.figure25" httpUri="https://binary.pensoft.net/fig/432194" pageId="0" pageNumber="1">25C, E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="8F29638D723C4BFB3DB3BFBF1CA6C048" pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph id="8AF786EA2F17E43E2E4456FC7EFC9AE2" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph id="17C7C198FE6B473F2E9A0C49961E042F" pageId="0" pageNumber="1">From the type locality.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="25B9A224E38EBFD3D3A7682B428DC493" pageId="0" pageNumber="1" type="biology_ecology">
|
|
<paragraph id="8D398167D385C1E30D42ED3D38E87700" pageId="0" pageNumber="1">Biology.</paragraph>
|
|
<paragraph id="972F9311BEF6C56C8BD1DF397DA5A685" pageId="0" pageNumber="1">The species was collected at 1325 m in elevation, in montane rainforest. Nests were located in rotten logs, root mat and rotten tree stump.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="447DBFD2BCE57B3D0AF11948B45C956D" pageId="0" pageNumber="1" type="comments">
|
|
<paragraph id="A2B74BDF68F20ACE6D390A3170A9A9E8" pageId="0" pageNumber="1">Comments.</paragraph>
|
|
<paragraph id="7A6B0E0A5049CBF437848205C2D33396" pageId="0" pageNumber="1">
|
|
<taxonomicName id="8A17A71F63EFF33C3A6C673ED7411ECC" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole antranohofa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="antranohofa">
|
|
<emphasis id="2974A4AED87DDA3DA45DFA5A8EA89C6A" italics="true" pageId="0" pageNumber="1">Pheidole antranohofa</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is known only from its type locality: Parc National de Marojejy in Antsiranana. Morphologically it is most similar to
|
|
<taxonomicName id="383AC082861FBD2CDFF0597AE601E395" lsidName="P. sikorae" pageId="0" pageNumber="1" rank="species" species="sikorae">
|
|
<emphasis id="56CD8DC9747AB4D36D325C3782A63DEE" italics="true" pageId="0" pageNumber="1">P. sikorae</emphasis>
|
|
</taxonomicName>
|
|
Forel which is widely distributed from
|
|
<normalizedToken id="F717453D2FD9284D82AFB1FA3A282249" originalValue="Forêt">Foret</normalizedToken>
|
|
Ambohidena in Toamasina to Andrambovato in Fianarantsoa. Major workers of
|
|
<taxonomicName id="E3AEDA7CE90F04DDA8C8DB7FF98175C5" lsidName="P. antranohofa" pageId="0" pageNumber="1" rank="species" species="antranohofa">
|
|
<emphasis id="378B19BA47F5B51EAF20B04594D86C6C" italics="true" pageId="0" pageNumber="1">P. antranohofa</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be distinguished based on developed sculpture of mesosoma, absence of impressed and smooth concavity placed lateral to antennal socket and tentorial pit and distinctly rugofoveolate interspaces on frons. However, minors of
|
|
<taxonomicName id="3B5EC6E5F72A366AAFC4A279D946551E" lsidName="P. antranohofa" pageId="0" pageNumber="1" rank="species" species="antranohofa">
|
|
<emphasis id="660A51BE55BC30076F5C4DCD43C85903" italics="true" pageId="0" pageNumber="1">P. antranohofa</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName id="865DCDC282879DA4995CB3BED966DB2B" lsidName="P. sikorae" pageId="0" pageNumber="1" rank="species" species="sikorae">
|
|
<emphasis id="06761698B0F87398716591276AFB6168" italics="true" pageId="0" pageNumber="1">P. sikorae</emphasis>
|
|
</taxonomicName>
|
|
are indistinguishable.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |