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<mods:title id="FCD71F63AD914A8EEC8DD8B67ABED45E">Ultrastructure of attachment specializations of hexapods (Arthropoda): evolutionary patterns inferred from a revised ordinal phylogeny</mods:title>
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<mods:namePart id="65923983972A7AB8398920B13A7D48E0">R. G. Beutel</mods:namePart>
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<mods:namePart id="75677943228C8281B1E9A8747D70A22E">S. N. Gorb</mods:namePart>
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24
<taxonomicName id="4C3D69CC2F618E18029AFF7FFEA8FF6E" authorityName="Latreille" authorityYear="1810" box="[176,297,190,212]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteroptera" pageId="5" pageNumber="182" phylum="Arthropoda" rank="order">Heteroptera</taxonomicName>
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<paragraph id="8B82124F2F618E1802BAFF1AFDA4FE39" blockId="5.[144,781,190,387]" pageId="5" pageNumber="182">
The tarsus is maximally three­segmented. An arolium is present in slaterellines (
<taxonomicName id="4C3D69CC2F618E1803A6FF39FE78FEB4" authorityName="Schilling" authorityYear="1829" box="[396,505,248,270]" class="Insecta" family="Lygaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="182" phylum="Arthropoda" rank="family">Lygaeidae</taxonomicName>
), an arolium and smooth euplantulae in mirines (
<taxonomicName id="4C3D69CC2F618E180354FED4FE55FE91" box="[382,468,277,299]" class="Insecta" family="Miridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="182" phylum="Arthropoda" rank="family">Miridae</taxonomicName>
), smooth pulvilli are present in members of
<taxonomicName id="4C3D69CC2F618E180303FEF2FE09FEF3" authorityName="" authorityYear="1815" box="[297,392,307,329]" class="Insecta" family="Coreidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="182" phylum="Arthropoda" rank="family">Coreidae</taxonomicName>
,
<taxonomicName id="4C3D69CC2F618E18038AFEF2FDACFEF3" authorityName="Leach" authorityYear="1815" box="[416,557,307,329]" class="Insecta" family="Pentatomidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="182" phylum="Arthropoda" rank="family">Pentatomidae</taxonomicName>
and
<taxonomicName id="4C3D69CC2F618E180051FEF2FC8CFEF3" box="[635,781,307,329]" class="Insecta" family="Pyrrhocoridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="182" phylum="Arthropoda" rank="family">Pyrrhocoridae</taxonomicName>
(
<figureCitation id="13060ECA2F618E1802BDFE91FF66FEDC" box="[151,231,336,358]" captionStart="Fig. 3" captionStartId="3.[1132,1165,405,424]" captionTargetBox="[149,976,192,769]" captionTargetId="figure@3.[144,979,190,771]" captionTargetPageId="3" captionText="Fig. 3. Diversity of the leg attachment devices (gray­colored areas) in hexapods. (a) Arolium (smooth). (b) Pulvilli (smooth or hairy). (c) Empodial pulvillus (ep) (hairy). (d) Hairy adhesive soles of tarsomeres. (e) Eversible pretarsal bladder (smooth). (f) Eversible structure between tibia and tarsus (smooth). (g) Fossula spongiosa (hairy). (h) Euplantulae (eu) and claw pad (cp) (both smooth). (i) Tarsal thorns transformed into adhesive structures (th), claw pad (cp) (both smooth). (j) Adhesive claw setae" httpUri="https://zenodo.org/record/2360333/files/figure.png" pageId="5" pageNumber="182">Figs 3b</figureCitation>
,
<figureCitation id="13060ECA2F618E1802D9FE91FE95FEDC" box="[243,276,336,358]" captionStart="Fig. 5" captionStartId="6.[182,215,1652,1671]" captionTargetBox="[212,1447,190,1619]" captionTargetId="figure@6.[212,1447,190,1619]" captionTargetPageId="6" captionText="Fig. 5. Details of hexapod tarsi and pretarsi. (A) Coreus marginatus (Heteroptera), hindleg, pretarsus, ventro­lateral aspect. (B) Cantharis fusca (Coleoptera, Polyphaga), hindleg, first and second tarsomeres, ventral aspect. (C, D, F) Priacma serrata (Coleoptera, Archostemata), midleg, tarsus, ventral aspect. (E, I) Sialis lutaria (Megaloptera), midleg, tarsus, ventral aspect. (G) Rhagonycha fulva (Coleoptera, Polyphaga), midleg, tarsus, ventral aspect. (H) Cantharis fusca (Coleoptera, Polyphaga), foreleg, tarsus, adhesive setae. HS, hairy soles; PU, pulvilli; TAR, tarsal segments; UN, claw" httpUri="https://zenodo.org/record/2360347/files/figure.png" pageId="5" pageNumber="182">5A</figureCitation>
), and smooth pulvilli and a fossula spongiosa in reduviids (
<bibRefCitation id="EFAC6FBE2F618E1802D6FEACFE36FE39" author="Carver, M. &amp; Gross, G. F. &amp; Woodward, T. E." box="[252,439,365,387]" editor="Ithaca, NY" journalOrPublisher="Cornell University Press" pageId="5" pageNumber="182" pagination="429 - 510" refId="ref11486" refString="Carver, M.; Gross, G. F.; Woodward, T. E., 1991: Hemiptera. In: CSIRO (ed.), The Insects of Australia. Ithaca, NY: Cornell University Press, pp. 429 - 510." title="Hemiptera" type="book chapter" volumeTitle="CSIRO" year="1991">Carver et al. 1991</bibRefCitation>
) (
<figureCitation id="13060ECA2F618E1803E7FEACFD98FE39" box="[461,537,365,387]" captionStart="Fig. 3" captionStartId="3.[1132,1165,405,424]" captionTargetBox="[149,976,192,769]" captionTargetId="figure@3.[144,979,190,771]" captionTargetPageId="3" captionText="Fig. 3. Diversity of the leg attachment devices (gray­colored areas) in hexapods. (a) Arolium (smooth). (b) Pulvilli (smooth or hairy). (c) Empodial pulvillus (ep) (hairy). (d) Hairy adhesive soles of tarsomeres. (e) Eversible pretarsal bladder (smooth). (f) Eversible structure between tibia and tarsus (smooth). (g) Fossula spongiosa (hairy). (h) Euplantulae (eu) and claw pad (cp) (both smooth). (i) Tarsal thorns transformed into adhesive structures (th), claw pad (cp) (both smooth). (j) Adhesive claw setae" httpUri="https://zenodo.org/record/2360333/files/figure.png" pageId="5" pageNumber="182">Fig. 3g</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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