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<document id="8D58AC1C396177A8BB26C314B0B3B64B" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629615386486" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2009" docId="1D214E2BF164FFE0DE8277F22F85FB2D" docLanguage="en" docName="zt02038p119.pdf" docOrigin="Zootaxa 2038" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Parhydraena asperita Perkins 2009, new species" docType="treatment" docVersion="3" lastPageNumber="26" masterDocId="E1183653F17CFFF9DE1575472E27FFB2" masterDocTitle="Revisions of the genera Parhydraena Orchymont, Protozantaena Perkins, Decarthrocerus Orchymont, and Parhydraenopsis nomen novum, aquatic and humicolous beetles from Africa and Madagascar, and comparative morphology of the tribe Parhydraenini (Coleoptera: Hydraenidae)" masterLastPageNumber="119" masterPageNumber="1" pageNumber="25" updateTime="1699083612395" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="C4660A7383619AF8AE364CB97C3C3081" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="59D661946A207F0EA56160F9972E13BC">
<mods:title id="16802A471175660A985235218501C90D">Revisions of the genera Parhydraena Orchymont, Protozantaena Perkins, Decarthrocerus Orchymont, and Parhydraenopsis nomen novum, aquatic and humicolous beetles from Africa and Madagascar, and comparative morphology of the tribe Parhydraenini (Coleoptera: Hydraenidae)</mods:title>
</mods:titleInfo>
<mods:name id="42EB5DF9144493F2185008FE6311A1E7" type="personal">
<mods:role id="78A031E93239E5F14FEB97F1147E0710">
<mods:roleTerm id="AB8759772174453E2DAFD4D91DBB040D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="643D399773008472356EB8FEF2509905">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B34EF5ADBA307E5FB6A810668C8CC948">text</mods:typeOfResource>
<mods:relatedItem id="9CB1445A503E5DBD92165821A41430E3" type="host">
<mods:titleInfo id="9A0488ED1EBCCD444B6FF585BD93080B">
<mods:title id="595DC487D4C06FF1DFE247699B01DF33">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0DA99030CBD969A8BCD754D449410B1A">
<mods:date id="7CFE775CAB2DF0623E0226AB2504AC30">2009</mods:date>
<mods:detail id="7848B8A78856D6974CD22C57B63A790D" type="pubDate">
<mods:number id="8F39A8C6DEE2BFA116B765762E7DF7ED">2009-03-16</mods:number>
</mods:detail>
<mods:detail id="A279B1F03879373A52ED6CFCAC389F58" type="volume">
<mods:number id="6D583047C92535513BA4EC6D21876A7C">2038</mods:number>
</mods:detail>
<mods:extent id="3FA039F39433AFEA31BF7007E2FEC9D1" unit="page">
<mods:start id="E73CB805E7A3BCB3FD940F056FDC7D17">1</mods:start>
<mods:end id="38D2135A1D2E6C84C247DD60FA63CFBF">119</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BD851EF89949FB0004BCA6A39445E553">journal article</mods:classification>
<mods:identifier id="CB7BD7BAF1B124FF9A4303C8A8703F6D" type="ISSN">1175­5334</mods:identifier>
</mods:mods>
<treatment id="1D214E2BF164FFE0DE8277F22F85FB2D" ID-DOI="http://doi.org/10.5281/zenodo.5321730" ID-Zenodo-Dep="5321730" LSID="urn:lsid:plazi:treatment:1D214E2BF164FFE0DE8277F22F85FB2D" httpUri="http://treatment.plazi.org/id/1D214E2BF164FFE0DE8277F22F85FB2D" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<subSubSection id="DD92ACB6F164FFE1DE8277F22C14FD7D" box="[151,563,693,719]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="9537FF3DF164FFE1DE8277F22C14FD7D" blockId="24.[151,563,693,754]" box="[151,563,693,719]" pageId="24" pageNumber="25">
<heading id="CE7F4851F164FFE1DE8277F22C14FD7D" bold="true" box="[151,563,693,719]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="A7FC232FF164FFE1DE8277F22C14FD7D" bold="true" box="[151,563,693,719]" pageId="24" pageNumber="25">
<taxonomicName id="528884BEF164FFE1DE8277F22FBBFD7D" ID-CoL="4DLYJ" authority="Perkins, 2009" authorityName="Perkins" authorityYear="2009" box="[151,412,693,719]" class="Insecta" family="Hydraenidae" genus="Parhydraena" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="asperita" status="sp. nov.">
<emphasis id="A7FC232FF164FFE1DE8277F22FBBFD7D" bold="true" box="[151,412,693,719]" italics="true" pageId="24" pageNumber="25">Parhydraena asperita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCCF9E54F164FFE1DFB677F22C14FD7D" box="[419,563,693,719]" pageId="24" pageNumber="25" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DD92ACB6F164FFE1DE82779F2FD5FD40" box="[151,498,728,754]" pageId="24" pageNumber="25" type="description">
<paragraph id="9537FF3DF164FFE1DE82779F2FD5FD40" blockId="24.[151,563,693,754]" box="[151,498,728,754]" pageId="24" pageNumber="25">
(
<figureCitation id="0DB3E3B8F164FFE1DE8A779F2ED5FD40" box="[159,242,728,754]" captionStart="FIGURE 2" captionStartId="59.[151,255,1718,1742]" captionTargetBox="[151,1431,207,1658]" captionTargetId="figure-28@59.[151,1436,194,1694]" captionTargetPageId="59" captionText="FIGURE 2. Parhydraena asperita and Parhydraena brahma, scanning electron micrographs of ventral surface." pageId="24" pageNumber="25">Figs. 2</figureCitation>
,
<figureCitation id="0DB3E3B8F164FFE1DEEA779F2F28FD40" box="[255,271,728,754]" captionStart="FIGURES 4–8" captionStartId="61.[151,269,1669,1693]" captionTargetBox="[157,1430,212,1611]" captionTargetId="figure-41@61.[151,1436,194,1646]" captionTargetPageId="61" captionText="FIGURES 4–8. Antennae (hydrofuge hair omitted). 4. Decarthrocerus jeanneli. 5. Discozantaena genuvela. 6. Parhydraena asperita. 7. Parhydraena brunovacca. 8. Protozantaena labrata." pageId="24" pageNumber="25">6</figureCitation>
,
<figureCitation id="0DB3E3B8F164FFE1DF09779F2F1CFD40" box="[284,315,728,754]" captionStart="FIGURES 12–13" captionStartId="63.[151,269,1730,1754]" captionTargetBox="[294,1281,208,1689]" captionTargetId="figure-30@63.[268,1319,194,1706]" captionTargetPageId="63" captionText="FIGURES 12–13. Elytron punctation pattern, and microsculpture detail. 12. Discozantaena sepiola. 13. Parhydraena asperita." pageId="24" pageNumber="25">13</figureCitation>
,
<figureCitation id="0DB3E3B8F164FFE1DF5D779F2F4FFD40" box="[328,360,728,754]" captionStart="FIGURES 30–33" captionStartId="69.[151,269,1692,1716]" captionTargetBox="[151,1429,220,1646]" captionTargetId="figure-41@69.[151,1436,194,1668]" captionTargetPageId="69" captionText="FIGURES 30–33. Terminal abdominal segments, and spermathecae. 30. Parhydraenopsis cooperi, female. 31. Parhydraenopsis cooperi, male. 32. Parhydraena asperita, male. 33. Pneuminion velamen." pageId="24" pageNumber="25">32</figureCitation>
,
<figureCitation id="0DB3E3B8F164FFE1DF61779F2FB3FD40" box="[372,404,728,754]" captionStart="FIGURE 50" captionStartId="82.[151,255,1730,1754]" captionTargetBox="[225,1353,194,1706]" captionTargetId="figure-25@82.[206,1381,194,1706]" captionTargetPageId="82" captionText="FIGURE 50. Parhydraena lancicula, dorsal and lateral habitus of holotype." pageId="24" pageNumber="25">50</figureCitation>
,
<figureCitation id="0DB3E3B8F164FFE1DFB5779F2FE7FD40" box="[416,448,728,754]" captionStart="FIGURES 52–53" captionStartId="84.[151,269,1730,1754]" captionTargetBox="[246,1333,222,1670]" captionTargetId="figure-31@84.[228,1360,194,1706]" captionTargetPageId="84" captionText="FIGURES 52–53. Parhydraena, aedeagi of holotypes. 52. P. lancicula. 53. P. namaqua." pageId="24" pageNumber="25">52</figureCitation>
,
<figureCitation id="0DB3E3B8F164FFE1DFD9779F2FCDFD40" box="[460,490,728,754]" captionStart="FIGURES 96–97" captionStartId="116.[151,269,1730,1754]" captionTargetBox="[218,1369,194,1702]" captionTargetId="figure-21@116.[218,1369,194,1706]" captionTargetPageId="116" captionText="FIGURES 96–97. Geographical distributions of Parhydraena species." pageId="24" pageNumber="25">96</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DD92ACB6F164FFE1DE8276622EDDFB29" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="9537FF3DF164FFE1DE8276622EDDFB29" blockId="24.[151,1436,805,2030]" pageId="24" pageNumber="25">
<emphasis id="A7FC232FF164FFE1DE8276622F72FC8D" bold="true" box="[151,341,805,831]" pageId="24" pageNumber="25">Type material.</emphasis>
<materialsCitation id="25E0F560F164FFE1DF7176622F90FC3E" collectingDate="1981-12-12" collectingDateMax="1981-12-30" collectingDateMin="1981-12-12" collectorName="S. &amp; J. Peck. Deposited" country="South Africa" county="Knysna" elevation="450" latitude="-34.05" location="Podocarpus forest" longLatPrecision="1201" longitude="23.05" municipality="Diepwalle" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<typeStatus id="4A33419FF164FFE1DF7176622FF6FC8D" box="[356,465,805,831]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="A7FC232FF164FFE1DC5176622A1BFC8D" bold="true" box="[580,1084,805,831]" pageId="24" pageNumber="25">
<collectingCountry id="ED9FBFADF164FFE1DC5176622CD6FC8D" box="[580,753,805,831]" name="South Africa" pageId="24" pageNumber="25">South Africa</collectingCountry>
:
<collectingRegion id="574C31DFF164FFE1DD1376622A1BFC8D" box="[774,1084,805,831]" country="South Africa" name="Western Cape" pageId="24" pageNumber="25">Western Cape Province</collectingRegion>
</emphasis>
,
<collectingCounty id="7C5687B1F164FFE1DA4676622A88FC8D" box="[1107,1199,805,831]" pageId="24" pageNumber="25">Knysna</collectingCounty>
,
<collectingMunicipality id="75536547F164FFE1DAD676622B19FC8D" box="[1219,1342,805,831]" pageId="24" pageNumber="25">Diepwalle</collectingMunicipality>
, mixed
<location id="9057A9E6F164FFE1DE82760B2F57FCD4" LSID="urn:lsid:plazi:treatment:1D214E2BF164FFE0DE8277F22F85FB2D:9057A9E6F164FFE1DE82760B2F57FCD4" box="[151,368,844,870]" country="South Africa" county="Knysna" latitude="-34.05" longLatPrecision="1201" longitude="23.05" municipality="Diepwalle" name="Podocarpus forest" pageId="24" pageNumber="25" stateProvince="Western Cape">Podocarpus forest</location>
,
<collectingMethod id="4CC9872AF164FFE1DF94760B2CACFCD4" box="[385,651,844,870]" pageId="24" pageNumber="25">window-malaise traps</collectingMethod>
, elev.
<quantity id="527052D8F164FFE1DCC8760B2D09FCD4" box="[733,814,844,870]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="24" pageNumber="25" unit="m" value="450.0">
<elevation id="1EA5180EF164FFE1DCC8760B2D09FCD4" box="[733,814,844,870]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="24" pageNumber="25" unit="m" value="450.0">450 m</elevation>
</quantity>
,
<geoCoordinate id="F0BC99FAF164FFE1DD2A760B2D81FCD4" box="[831,934,844,870]" degrees="34" direction="south" minutes="3" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="-34.05">34° 3' S</geoCoordinate>
,
<geoCoordinate id="F0BC99FAF164FFE1DDAD760B2A06FCD4" box="[952,1057,844,870]" degrees="23" direction="east" minutes="3" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="23.05">23° 3' E</geoCoordinate>
,
<date id="E136D9FDF164FFE1DA27760B2AC2FCD4" box="[1074,1253,844,870]" pageId="24" pageNumber="25" value="1981-12-12" valueMax="1981-12-30" valueMin="1981-12-12">
<collectingDate id="F1722015F164FFE1DA27760B2AC2FCD4" box="[1074,1253,844,870]" pageId="24" pageNumber="25" value="1981-12-12" valueMax="1981-12-30" valueMin="1981-12-12">12–30.xii.1981</collectingDate>
</date>
,
<collectorName id="387D9AEBF164FFE1DAE2760B2F29FC3E" pageId="24" pageNumber="25">S. &amp; J. Peck. Deposited</collectorName>
in the TMSA
</materialsCitation>
.
<materialsCitation id="25E0F560F164FFE1DFD176352BB0FC01" collectingDate="1981-12-05" collectingDateMax="1981-12-30" collectingDateMin="1981-12-05" collectionCode="MCZ" collectorName="S. &amp; J. Peck" country="South Africa" county="Stormsrivier" latitude="-33.983334" location="Podocarpus forest" longLatPrecision="1202" longitude="23.866667" municipality="Goesabos" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
<emphasis id="A7FC232FF164FFE1DFD176352A55FC3E" bold="true" box="[452,1138,882,908]" pageId="24" pageNumber="25">
<typeStatus id="4A33419FF164FFE1DFD176352C66FC3E" box="[452,577,882,908]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
(121):
<collectingCountry id="ED9FBFADF164FFE1DC8976352D66FC3E" box="[668,833,882,908]" name="South Africa" pageId="24" pageNumber="25">South Africa</collectingCountry>
:
<collectingRegion id="574C31DFF164FFE1DD4576352A55FC3E" box="[848,1138,882,908]" country="South Africa" name="Eastern Cape" pageId="24" pageNumber="25">Eastern Cape Province</collectingRegion>
</emphasis>
,
<collectingCounty id="7C5687B1F164FFE1DA9776352B31FC3E" box="[1154,1302,882,908]" pageId="24" pageNumber="25">Stormsrivier</collectingCounty>
,
<collectingMunicipality id="75536547F164FFE1DB3676352BBFFC3E" box="[1315,1432,882,908]" pageId="24" pageNumber="25">Goesabos</collectingMunicipality>
, mixed
<location id="9057A9E6F164FFE1DEFC76DE2FE7FC01" LSID="urn:lsid:plazi:treatment:1D214E2BF164FFE0DE8277F22F85FB2D:9057A9E6F164FFE1DEFC76DE2FE7FC01" box="[233,448,921,947]" country="South Africa" county="Stormsrivier" latitude="-33.983334" longLatPrecision="1202" longitude="23.866667" municipality="Goesabos" name="Podocarpus forest" pageId="24" pageNumber="25" stateProvince="Eastern Cape">Podocarpus forest</location>
,
<collectingMethod id="4CC9872AF164FFE1DFD976DE2CF5FC01" box="[460,722,921,947]" pageId="24" pageNumber="25">window-malaise traps</collectingMethod>
,
<geoCoordinate id="F0BC99FAF164FFE1DCCA76DE2D69FC01" box="[735,846,921,947]" degrees="33" direction="south" minutes="59" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="-33.983334">33° 59' S</geoCoordinate>
,
<geoCoordinate id="F0BC99FAF164FFE1DD4976DE2DEAFC01" box="[860,973,921,947]" degrees="23" direction="east" minutes="52" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="23.866667">23° 52' E</geoCoordinate>
,
<date id="E136D9FDF164FFE1DDCF76DE2AA7FC01" box="[986,1152,921,947]" pageId="24" pageNumber="25" value="1981-12-05" valueMax="1981-12-30" valueMin="1981-12-05">
<collectingDate id="F1722015F164FFE1DDCF76DE2AA7FC01" box="[986,1152,921,947]" pageId="24" pageNumber="25" value="1981-12-05" valueMax="1981-12-30" valueMin="1981-12-05">5–30.xii.1981</collectingDate>
</date>
,
<collectorName id="387D9AEBF164FFE1DA9876DE2B06FC01" box="[1165,1313,921,947]" pageId="24" pageNumber="25">S. &amp; J. Peck</collectorName>
(8
<collectionCode id="F39967F8F164FFE1DB5E76DE2BB7FC01" box="[1355,1424,921,947]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="24" pageNumber="25" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="25E0F560F164FFE1DE8276872FF9FBB2" collectingDate="1981-12-06" collectingDateMax="1981-12-30" collectingDateMin="1981-12-06" collectionCode="FMNH, TMSA" collectorName="S. &amp; J. Peck" country="South Africa" county="Stormsrivier" latitude="-33.983334" location="Podocarpus forest" longLatPrecision="1202" longitude="23.866667" municipality="Skuinsbos" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
<collectingCounty id="7C5687B1F164FFE1DE8276872F0CFC68" box="[151,299,960,986]" pageId="24" pageNumber="25">Stormsrivier</collectingCounty>
,
<collectingMunicipality id="75536547F164FFE1DF2276872F95FC68" box="[311,434,960,986]" pageId="24" pageNumber="25">Skuinsbos</collectingMunicipality>
, mixed
<location id="9057A9E6F164FFE1DC0576872CC1FC68" LSID="urn:lsid:plazi:treatment:1D214E2BF164FFE0DE8277F22F85FB2D:9057A9E6F164FFE1DC0576872CC1FC68" box="[528,742,960,986]" country="South Africa" county="Stormsrivier" latitude="-33.983334" longLatPrecision="1202" longitude="23.866667" municipality="Skuinsbos" name="Podocarpus forest" pageId="24" pageNumber="25" stateProvince="Eastern Cape">Podocarpus forest</location>
,
<collectingMethod id="4CC9872AF164FFE1DCE776872DB3FC68" box="[754,916,960,986]" pageId="24" pageNumber="25">window traps</collectingMethod>
,
<geoCoordinate id="F0BC99FAF164FFE1DDB476872A29FC68" box="[929,1038,960,986]" degrees="33" direction="south" minutes="59" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="-33.983334">33° 59' S</geoCoordinate>
,
<geoCoordinate id="F0BC99FAF164FFE1DA0976872AACFC68" box="[1052,1163,960,986]" degrees="23" direction="east" minutes="52" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="23.866667">23° 52' E</geoCoordinate>
,
<date id="E136D9FDF164FFE1DA8D76872B1AFC68" box="[1176,1341,960,986]" pageId="24" pageNumber="25" value="1981-12-06" valueMax="1981-12-30" valueMin="1981-12-06">
<collectingDate id="F1722015F164FFE1DA8D76872B1AFC68" box="[1176,1341,960,986]" pageId="24" pageNumber="25" value="1981-12-06" valueMax="1981-12-30" valueMin="1981-12-06">6–30.xii.1981</collectingDate>
</date>
,
<collectorName id="387D9AEBF164FFE1DB5E76872EF7FBB2" pageId="24" pageNumber="25">S. &amp; J. Peck</collectorName>
(1
<collectionCode id="F39967F8F164FFE1DEE276A12F68FBB2" box="[247,335,998,1024]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="24" pageNumber="25" type="Museum">FMNH</collectionCode>
, 34
<collectionCode id="F39967F8F164FFE1DF6A76A12FF1FBB2" box="[383,470,998,1024]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="24" pageNumber="25">TMSA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="25E0F560F164FFE1DFFF76A12FBBFB95" collectingDate="1997-03-07" collectionCode="NMW" collectorName="Hess &amp; Heckes" country="South Africa" latitude="-34.083332" location="Tsitsikamma Mts." longLatPrecision="1201" longitude="23.35" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
<location id="9057A9E6F164FFE1DFFF76A12C99FBB2" LSID="urn:lsid:plazi:treatment:1D214E2BF164FFE0DE8277F22F85FB2D:9057A9E6F164FFE1DFFF76A12C99FBB2" box="[490,702,998,1024]" country="South Africa" latitude="-34.083332" longLatPrecision="1201" longitude="23.35" name="Tsitsikamma Mts." pageId="24" pageNumber="25" stateProvince="Eastern Cape">Tsitsikamma Mts.</location>
,
<locationDeviation id="8485E61FF164FFE1DCDF76A12DD6FBB2" box="[714,1009,998,1024]" location="Plettenberg Bay" pageId="24" pageNumber="25">
<quantity id="527052D8F164FFE1DCDF76A12D33FBB2" box="[714,788,998,1024]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="25" unit="km" value="40.0">40 km</quantity>
E Plettenberg Bay
</locationDeviation>
,
<geoCoordinate id="F0BC99FAF164FFE1DDEB76A12A7CFBB2" box="[1022,1115,998,1024]" degrees="34" direction="south" minutes="5" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="-34.083332">34° 5' S</geoCoordinate>
,
<geoCoordinate id="F0BC99FAF164FFE1DA7D76A12AF0FBB2" box="[1128,1239,998,1024]" degrees="23" direction="east" minutes="21" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="23.35">23° 21' E</geoCoordinate>
,
<date id="E136D9FDF164FFE1DAF176A12B71FBB2" box="[1252,1366,998,1024]" pageId="24" pageNumber="25" value="1997-03-07">
<collectingDate id="F1722015F164FFE1DAF176A12B71FBB2" box="[1252,1366,998,1024]" pageId="24" pageNumber="25" value="1997-03-07">7.iii.1997</collectingDate>
</date>
,
<collectorName id="387D9AEBF164FFE1DB7676A12BBBFBB2" box="[1379,1436,998,1024]" pageId="24" pageNumber="25">Hess</collectorName>
&amp;
<collectorName id="387D9AEBF164FFE1DEAE714A2F36FB95" box="[187,273,1037,1063]" pageId="24" pageNumber="25">Heckes</collectorName>
(3
<collectionCode id="F39967F8F164FFE1DF51714A2FB3FB95" box="[324,404,1037,1063]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="24" pageNumber="25">NMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="25E0F560F164FFE1DFBB714A2B6AFBFC" collectingDate="1981-12-12" collectingDateMax="1981-12-30" collectingDateMin="1981-12-12" collectionCode="TMSA, FMNH" collectorName="S. &amp; J. Peck" country="South Africa" county="Knysna" elevation="450" latitude="-34.05" location="Podocarpus forest" longLatPrecision="1201" longitude="23.05" municipality="Diepwalle" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype">
<emphasis id="A7FC232FF164FFE1DFBB714A2CC7FB95" bold="true" box="[430,736,1037,1063]" pageId="24" pageNumber="25">
<collectingRegion id="574C31DFF164FFE1DFBB714A2CC7FB95" box="[430,736,1037,1063]" country="South Africa" name="Western Cape" pageId="24" pageNumber="25">Western Cape Province</collectingRegion>
</emphasis>
,
<collectingCounty id="7C5687B1F164FFE1DCE0714A2D76FB95" box="[757,849,1037,1063]" pageId="24" pageNumber="25">Knysna</collectingCounty>
,
<collectingMunicipality id="75536547F164FFE1DD76714A2DF9FB95" box="[867,990,1037,1063]" pageId="24" pageNumber="25">Diepwalle</collectingMunicipality>
, mixed
<location id="9057A9E6F164FFE1DA53714A2B05FB95" LSID="urn:lsid:plazi:treatment:1D214E2BF164FFE0DE8277F22F85FB2D:9057A9E6F164FFE1DA53714A2B05FB95" box="[1094,1314,1037,1063]" country="South Africa" county="Knysna" latitude="-34.05" longLatPrecision="1201" longitude="23.05" municipality="Diepwalle" name="Podocarpus forest" pageId="24" pageNumber="25" stateProvince="Western Cape">Podocarpus forest</location>
,
<collectingMethod id="4CC9872AF164FFE1DB21714A2F10FBFC" pageId="24" pageNumber="25">windowmalaise traps</collectingMethod>
, elev.
<quantity id="527052D8F164FFE1DF9271732FF0FBFC" box="[391,471,1076,1102]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="24" pageNumber="25" unit="m" value="450.0">
<elevation id="1EA5180EF164FFE1DF9271732FF0FBFC" box="[391,471,1076,1102]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="24" pageNumber="25" unit="m" value="450.0">450 m</elevation>
</quantity>
,
<geoCoordinate id="F0BC99FAF164FFE1DFF271732C6BFBFC" box="[487,588,1076,1102]" degrees="34" direction="south" minutes="3" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="-34.05">34° 3' S</geoCoordinate>
,
<geoCoordinate id="F0BC99FAF164FFE1DC4971732CE4FBFC" box="[604,707,1076,1102]" degrees="23" direction="east" minutes="3" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="23.05">23° 3' E</geoCoordinate>
,
<date id="E136D9FDF164FFE1DCC671732DA1FBFC" box="[723,902,1076,1102]" pageId="24" pageNumber="25" value="1981-12-12" valueMax="1981-12-30" valueMin="1981-12-12">
<collectingDate id="F1722015F164FFE1DCC671732DA1FBFC" box="[723,902,1076,1102]" pageId="24" pageNumber="25" value="1981-12-12" valueMax="1981-12-30" valueMin="1981-12-12">12–30.xii.1981</collectingDate>
</date>
,
<collectorName id="387D9AEBF164FFE1DD8371732A16FBFC" box="[918,1073,1076,1102]" pageId="24" pageNumber="25">S. &amp; J. Peck</collectorName>
(65
<collectionCode id="F39967F8F164FFE1DA7B71732AE5FBFC" box="[1134,1218,1076,1102]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="24" pageNumber="25">TMSA</collectionCode>
, 7
<collectionCode id="F39967F8F164FFE1DAFE71732B62FBFC" box="[1259,1349,1076,1102]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="24" pageNumber="25" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="25E0F560F164FFE1DB4871732ED2FB29" collectingDate="1992-11-21" collectionCode="TMSA" collectorName="Endrody-Younga" country="South Africa" latitude="-33.616665" location="Kammanasieberge" longLatPrecision="1204" longitude="22.55" municipality="Little Karroo" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype">
<collectingMunicipality id="75536547F164FFE1DB4871732ECAFBC6" pageId="24" pageNumber="25">Little Karroo</collectingMunicipality>
,
<location id="9057A9E6F164FFE1DEEA711D2FFCFBC6" LSID="urn:lsid:plazi:treatment:1D214E2BF164FFE0DE8277F22F85FB2D:9057A9E6F164FFE1DEEA711D2FFCFBC6" box="[255,475,1114,1140]" country="South Africa" latitude="-33.616665" longLatPrecision="1204" longitude="22.55" municipality="Little Karroo" name="Kammanasieberge" pageId="24" pageNumber="25" stateProvince="Western Cape">Kammanasieberge</location>
, river stones,
<geoCoordinate id="F0BC99FAF164FFE1DC87711D2D2BFBC6" box="[658,780,1114,1140]" degrees="33" direction="south" minutes="37" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="-33.616665">33° 37' S</geoCoordinate>
,
<geoCoordinate id="F0BC99FAF164FFE1DD35711D2DBBFBC6" box="[800,924,1114,1140]" degrees="22" direction="east" minutes="33" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="22.55">22° 33' E</geoCoordinate>
,
<date id="E136D9FDF164FFE1DDBA711D2A08FBC6" box="[943,1071,1114,1140]" pageId="24" pageNumber="25" value="1992-11-21">
<collectingDate id="F1722015F164FFE1DDBA711D2A08FBC6" box="[943,1071,1114,1140]" pageId="24" pageNumber="25" value="1992-11-21">21.xi.1992</collectingDate>
</date>
,
<collectorName id="387D9AEBF164FFE1DA57711D2B2DFBC6" box="[1090,1290,1114,1140]" pageId="24" pageNumber="25">Endrödy-Younga</collectorName>
(#2930) (3
<collectionCode id="F39967F8F164FFE1DE8271C62EC9FB29" box="[151,238,1153,1179]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="24" pageNumber="25">TMSA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DD92ACB6F164FFE1DED371E02CDEFAEF" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="9537FF3DF164FFE1DED371E02CDEFAEF" blockId="24.[151,1436,805,2030]" pageId="24" pageNumber="25">
<emphasis id="A7FC232FF164FFE1DED371E02FFFFB73" bold="true" box="[198,472,1191,1217]" pageId="24" pageNumber="25">Differential diagnosis.</emphasis>
Recognized among members of the genus by the combination of the unilinear row of large granules on the elytra series and intervals, the comparatively long maxillary palpi (palpus length/head width ca. 0.89), and the comparatively large body size (length ca.
<quantity id="527052D8F164FFE1DDE271B22A4BFABD" box="[1015,1132,1269,1295]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" pageId="24" pageNumber="25" unit="mm" value="1.93">1.93 mm</quantity>
); males are additionally recognized by the modified 6th abdominal ventrite (
<figureCitation id="0DB3E3B8F164FFE1DCEE705B2D69FA84" box="[763,846,1308,1334]" captionStart="FIGURE 2" captionStartId="59.[151,255,1718,1742]" captionTargetBox="[151,1431,207,1658]" captionTargetId="figure-28@59.[151,1436,194,1694]" captionTargetPageId="59" captionText="FIGURE 2. Parhydraena asperita and Parhydraena brahma, scanning electron micrographs of ventral surface." pageId="24" pageNumber="25">Figs. 2</figureCitation>
,
<figureCitation id="0DB3E3B8F164FFE1DD4E705B2D5BFA84" box="[859,892,1308,1334]" captionStart="FIGURE 47" captionStartId="80.[151,255,1730,1754]" captionTargetBox="[256,1332,213,1689]" captionTargetId="figure-27@80.[236,1352,194,1706]" captionTargetPageId="80" captionText="FIGURE 47. Parhydraena asperita, dorsal, lateral and ventral habitus of holotype." pageId="24" pageNumber="25">47</figureCitation>
). The aedeagus, while distinct, might possibly be compared with that of
<taxonomicName id="528884BEF164FFE1DFD670042C6AFAEE" authorityName="P. D. Perkins &amp; J. Balfour-Browne" authorityYear="1994" box="[451,589,1347,1372]" class="Insecta" family="Hydraenidae" genus="Parhydraena" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lancicula">
<emphasis id="A7FC232FF164FFE1DFD670042C6AFAEE" box="[451,589,1347,1372]" italics="true" pageId="24" pageNumber="25">P. lancicula</emphasis>
</taxonomicName>
(
<figureCitation id="0DB3E3B8F164FFE1DC4870042C98FAEF" box="[605,703,1347,1373]" captionStart="FIGURES 48–49" captionStartId="81.[151,269,1730,1754]" captionTargetBox="[313,1259,209,1684]" captionTargetId="figure-29@81.[298,1288,194,1706]" captionTargetPageId="81" captionText="FIGURES 48–49. Parhydraena, aedeagi of holotypes. 48. P. ancylis. 49. P. asperita." pageId="24" pageNumber="25">Figs. 49</figureCitation>
,
<figureCitation id="0DB3E3B8F164FFE1DCDE70042CCBFAEF" box="[715,748,1347,1373]" captionStart="FIGURES 52–53" captionStartId="84.[151,269,1730,1754]" captionTargetBox="[246,1333,222,1670]" captionTargetId="figure-31@84.[228,1360,194,1706]" captionTargetPageId="84" captionText="FIGURES 52–53. Parhydraena, aedeagi of holotypes. 52. P. lancicula. 53. P. namaqua." pageId="24" pageNumber="25">52</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DD92ACB6F164FFE0DED3702F2A7AFB99" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
<paragraph id="9537FF3DF164FFE1DED3702F2C81FA45" blockId="24.[151,1436,805,2030]" pageId="24" pageNumber="25">
<emphasis id="A7FC232FF164FFE1DED3702F2F79FA30" bold="true" box="[198,350,1384,1410]" pageId="24" pageNumber="25">Description.</emphasis>
Size (length/width, mm)
<typeStatus id="4A33419FF164FFE1DC9E702E2CD1FA31" box="[651,758,1385,1411]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
: body (length to elytral apices) 1.93/0.83; head 0.31/0.51; pronotum 0.45/0.68, PA 0.54, PB 0.58; elytra 1.20 /0.83. Head piceous; pronotum dark brown to piceous, except margined laterally with light brown; elytra dark brown; legs brown to dark brown; maxillary palpi brown, distal 1/3 of last mere slightly darker.
</paragraph>
<paragraph id="9537FF3DF164FFE1DED373432ACEF8B4" blockId="24.[151,1436,805,2030]" pageId="24" pageNumber="25">Head with eyes moderately large, in dorsal aspect 9 convex facets in longest series. Ocelli distinct round small shining, each located at base of shallow oblique interocular fovea. Frons microreticulate, dull, disc indistinctly finely punctate, ca. 0.5xef, interstices ca. 2–4xpd; area between interocular foveae and eye microreticulate, dull. Clypeus dull, markedly microreticulate, except anterior margin shining. Labrum dull, apicomedially emarginate to form rounded lobe on each side; laterally fringed with short arcuate setae. Maxillary palpi elongate, combined lengths of meres slightly less than width of head (ratio ca. 31/35); ratios of meres 2–4 ca. 12/7/15. Mentum and submentum sparsely finely punctulate, microreticulate.</paragraph>
<paragraph id="9537FF3DF164FFE0DED372542A56FF6A" blockId="24.[151,1436,805,2030]" lastBlockId="25.[151,1437,152,1183]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">Pronotum with anterior margin weakly arcuate, except median 1/3 straight and with extremely narrow hyaline border. Anterior angles obtuse, widely rounded. Sides rounded, widest just before middle, behind middle gradually attenuate, very weakly sinuate. Posterior angle obtusangulate. Disc microreticulate, dull, impressions very shallow, broadly U-shaped, punctation on disc fine and dense, many punctures confluent in impressions, sparser, 0.5–2xpd, on low relief separating impressions; shallow wide impression at each posterior angle. Sculpture laterally subrugose, dull. Each puncture with anterior margin extended posteriorly as narrow ridge, dividing puncture into a pair of two smaller punctures each of which smaller than eye-facet, short recumbent or decumbent seta arising from posterior extreme of dividing ridge.</paragraph>
<paragraph id="9537FF3DF165FFE0DED375A22CB7FE28" blockId="25.[151,1437,152,1183]" pageId="25" pageNumber="26">Elytra dull, elongate-ovate, sides weakly arcuate, margins distinct from shoulder for 4/5 of length, sutural angle more acute in female than in male. Ten-seriate punctate, but punctures obsolete, granule of each &quot;puncture&quot; large, serial granules closely spaced, very similar in size and spacing to unilinear row of granules on each interval. Each granule at its posterior side with short recumbent seta, setae of serial and interval granules very similar in length and density.</paragraph>
<paragraph id="9537FF3DF165FFE0DED374E12ACBFCAE" blockId="25.[151,1437,152,1183]" pageId="25" pageNumber="26">Metaventrite with shallow median elongate oval impression in basal 1/2, midlongitudinal shining carina between anterior extreme of oval impression and mesoventral intercoxal process. Metaventrite and abdominal ventrites 1–4 hydrofuge pubescent. Ventrite 5, in male, hydrofuge pubescent in basal 1/3 laterally, posterior margin of pubescent area produced in middle to 3/4 length of ventrite, in female basal 1/2 of ventrite hydrofuge pubescent; shining part of ventrite 5 very sparsely pubescent in both sexes. Ventrite 6 very weakly microreticulate, shining, in male middle part markedly raised and apicomedially emarginate, about 3 or 4 setae on each side of emargination; in female shape simple, with transverse band of sparse setae across posterior 1/2. Male: last ventrite asymmetrical, apex lobiform and setose; last tergite with tuft of setae on each side of apicomedian notch into which fits lobe of last ventrite. Female: last ventrite sparsely pubescent, margin rounded; last tergite with small tuft of setae on each side of apicomedian emargination.</paragraph>
<paragraph id="9537FF3DF165FFE0DED3766E2C2EFC22" blockId="25.[151,1437,152,1183]" pageId="25" pageNumber="26">Legs moderately long and slender, combined lengths of metatibia and metatarsus greater than width of head (ratio ca. 51/35), ratio of lengths of metatibia/metatarsus ca. 31/20. Basal 3 pro- and mesotarsomeres, in male, with pad of suction setae.</paragraph>
<paragraph id="9537FF3DF165FFE0DED376DA2D5FFC05" blockId="25.[151,1437,152,1183]" box="[198,888,925,951]" pageId="25" pageNumber="26">
Wings fully developed in
<typeStatus id="4A33419FF165FFE0DFE076DA2C7CFC05" box="[501,603,925,951]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
and all dissected males.
</paragraph>
<paragraph id="9537FF3DF165FFE0DED376832A7AFB99" blockId="25.[151,1437,152,1183]" pageId="25" pageNumber="26">
Aedeagus length ca.
<quantity id="527052D8F165FFE0DFD476832C08FC6C" box="[449,559,964,990]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="25" pageNumber="26" unit="mm" value="0.27">0.27 mm</quantity>
; main-piece, in left lateral view, with two large pointed processes; mainpiece wider in lateral view than ventral view, with finely strigose surface over much of ventral and lateral surface; distal piece produced in large lobe on right side, pointed at apex (
<figureCitation id="0DB3E3B8F165FFE0DDED71562A77FB99" box="[1016,1104,1041,1067]" captionStart="FIGURES 48–49" captionStartId="81.[151,269,1730,1754]" captionTargetBox="[313,1259,209,1684]" captionTargetId="figure-29@81.[298,1288,194,1706]" captionTargetPageId="81" captionText="FIGURES 48–49. Parhydraena, aedeagi of holotypes. 48. P. ancylis. 49. P. asperita." pageId="25" pageNumber="26">Fig. 49</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DD92ACB6F165FFE0DED371702AD6FBE0" box="[198,1265,1079,1106]" pageId="25" pageNumber="26" type="etymology">
<paragraph id="9537FF3DF165FFE0DED371702AD6FBE0" blockId="25.[151,1437,152,1183]" box="[198,1265,1079,1106]" pageId="25" pageNumber="26">
<emphasis id="A7FC232FF165FFE0DED371702F74FBE3" bold="true" box="[198,339,1079,1105]" pageId="25" pageNumber="26">Etymology.</emphasis>
Named in reference to the asperite dorsum, with comparatively large granules.
</paragraph>
</subSubSection>
<subSubSection id="DD92ACB6F165FFE0DED371192F85FB2D" pageId="25" pageNumber="26" type="distribution">
<paragraph id="9537FF3DF165FFE0DED371192F85FB2D" blockId="25.[151,1437,152,1183]" pageId="25" pageNumber="26">
<emphasis id="A7FC232FF165FFE0DED371192F40FBCA" bold="true" box="[198,359,1118,1144]" pageId="25" pageNumber="26">Distribution.</emphasis>
Currently known from three localities near Knysna and one in the Little Karroo, in southern
<collectingCountry id="ED9FBFADF165FFE0DE8271C22F09FB2D" box="[151,302,1157,1183]" name="South Africa" pageId="25" pageNumber="26">South Africa</collectingCountry>
(
<figureCitation id="0DB3E3B8F165FFE0DF2871C22FB2FB2D" box="[317,405,1157,1183]" captionStart="FIGURES 94–95" captionStartId="115.[151,269,1730,1754]" captionTargetBox="[216,1371,194,1703]" captionTargetId="figure-20@115.[216,1371,194,1706]" captionTargetPageId="115" captionText="FIGURES 94–95. Geographical distributions of Parhydraena species." pageId="25" pageNumber="26">Fig. 95</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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