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<document id="A861F391882D07262B34478BEE490FD0" ID-DOI="10.11646/zootaxa.4006.2.2" ID-GBIF-Dataset="de6a87e5-efb4-47a3-88ab-8897fba99bf7" ID-ISSN="1175-5326" ID-Zenodo-Dep="254116" ID-ZooBank="B7A45EAC-D817-47BF-A284-731FEF2B8F8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461206532985" checkinUser="plazi" docAuthor="Van Dam, Matthew H. &amp; OBrien, Charles W." docDate="2015" docId="C15787EFE93DFFF6FF08F98B8293FD7F" docLanguage="en" docName="zt04006p284.pdf" docOrigin="Zootaxa 4006 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Miloderes amargosensis Van Dam and O'Brien, sp. nov." docType="treatment" docVersion="7" lastPageNumber="272" masterDocId="3D6EFF97E925FFEFFF9FFFC5803CFFD7" masterDocTitle="Review of the genus Miloderes Casey, 1888 (Coleoptera: Curculionidae: Entiminae), with desciptions of three new species" masterLastPageNumber="284" masterPageNumber="247" pageNumber="271" updateTime="1698601344551" updateUser="plazi">
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<mods:title id="C410360C9AD1A1B5A0C86616689C523C">Review of the genus Miloderes Casey, 1888 (Coleoptera: Curculionidae: Entiminae), with desciptions of three new species</mods:title>
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<mods:namePart id="E0F3E221B3DB4B13379D649E80311FCA">Van Dam, Matthew H.</mods:namePart>
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<mods:namePart id="0B6078155037CEAC659FF2CF3ED88091">OBrien, Charles W.</mods:namePart>
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<mods:title id="3485974A0EB91F938E4264F7E7E7FBA9">Zootaxa</mods:title>
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<mods:date id="10FE39DD4360CF6BE6C1CC2BA1AA806E">2015</mods:date>
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<mods:number id="D3C0405FF9FF00BDD17DC3DE89827B19">4006</mods:number>
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<mods:identifier id="6337EBC26466C3A52799C2D97331E8DB" type="DOI">10.11646/zootaxa.4006.2.2</mods:identifier>
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<treatment id="C15787EFE93DFFF6FF08F98B8293FD7F" ID-DOI="http://doi.org/10.5281/zenodo.5661864" ID-GBIF-Taxon="119630087" ID-Zenodo-Dep="5661864" LSID="urn:lsid:plazi:treatment:C15787EFE93DFFF6FF08F98B8293FD7F" httpUri="http://treatment.plazi.org/id/C15787EFE93DFFF6FF08F98B8293FD7F" lastPageId="25" lastPageNumber="272" pageId="24" pageNumber="271">
<subSubSection id="01E46572E93DFFF7FF08F98B814CF95D" pageId="24" pageNumber="271" type="nomenclature">
<paragraph id="494136F9E93DFFF7FF08F98B8374F9BF" blockId="24.[151,840,1614,1674]" box="[151,840,1614,1640]" pageId="24" pageNumber="271">
<heading id="12098195E93DFFF7FF08F98B8374F9BF" bold="true" box="[151,840,1614,1640]" fontSize="11" level="1" pageId="24" pageNumber="271" reason="1">
<emphasis id="7B8AEAEBE93DFFF7FF08F98B8374F9BF" bold="true" box="[151,840,1614,1640]" pageId="24" pageNumber="271">
<taxonomicName id="8EFE4D7AE93DFFF7FF08F98B82E1F9BF" ID-CoL="43DK2" authority="Van Dam and O'Brien" authorityName="Van Dam and O'Brien" box="[151,733,1614,1640]" class="Insecta" family="Curculionidae" genus="Miloderes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="271" phylum="Arthropoda" rank="species" species="amargosensis" status="sp. nov.">
<emphasis id="7B8AEAEBE93DFFF7FF08F98B8187F9BF" bold="true" box="[151,443,1614,1640]" italics="true" pageId="24" pageNumber="271">Miloderes amargosensis</emphasis>
Van Dam and OBrien
</taxonomicName>
,
<taxonomicNameLabel id="60B95790E93DFFF7FD75F98B8374F9BF" box="[746,840,1614,1640]" pageId="24" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="494136F9E93DFFF7FF08F9B4814CF95D" blockId="24.[151,840,1614,1674]" box="[151,368,1649,1674]" pageId="24" pageNumber="271">
(
<figureCitation id="D1C52A7CE93DFFF7FF00F9B4813BF95D" box="[159,263,1649,1674]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="4.[151,249,1880,1902]" captionStartId-1="5.[151,249,1884,1906]" captionStartId-2="6.[151,249,1958,1980]" captionStartId-3="7.[151,249,1953,1975]" captionStartId-4="8.[151,249,1964,1986]" captionTargetBox-0="[152,1433,195,1855]" captionTargetBox-1="[151,1436,193,1862]" captionTargetBox-2="[201,1380,193,1925]" captionTargetBox-3="[185,1390,193,1931]" captionTargetBox-4="[197,1380,193,1931]" captionTargetId-0="figure@4.[152,1434,195,1855]" captionTargetId-1="figure@5.[151,1436,193,1862]" captionTargetId-2="figure@6.[201,1380,193,1927]" captionTargetId-3="figure@7.[185,1390,193,1932]" captionTargetId-4="figure@8.[197,1380,193,1932]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 1. Miloderes dorsal habitus; A. M. viridis, B. M. ubehebensis sp. nov., C. M. panamintensis sp. nov., D. M. nelsoni, E. M. tingi, F. M. amargosensis sp. nov., G. M. mercuryensis, H. M. setosus." captionText-1="FIGURE 2. Miloderes lateral habitus; A. M. viridis, B. M. ubehebensis sp. nov., C. M. panamintensis sp. nov., D. M. nelsoni, E. M. tingi, F. M. amargosensis sp. nov., G. M. mercuryensis, H. M. setosus." captionText-2="FIGURE 3. Miloderes front tibia dorsal view; A. M. viridis, B. M. ubehebensis sp. nov., C. M. panamintensis sp. nov., D. M. nelsoni, E. M. tingi, F. M. amargosensis sp. nov., G. M. mercuryensis, H. M. setosus." captionText-3="FIGURE 4. Miloderes hind tibia lateral view; A. M. viridis, B. M. ubehebensis sp. nov., C. M. panamintensis sp. nov., D. M. nelsoni, E. M. tingi, F. M. amargosensis sp. nov., G. M. mercuryensis, H. M. setosus." captionText-4="FIGURE 5. Miloderes head anterior view; A. M. viridis, B. M. ubehebensis sp. nov., C. M. panamintensis sp. nov., D. M. nelsoni, E. M. tingi, F. M. amargosensis sp. nov." httpUri-0="https://zenodo.org/record/254117/files/figure.png" httpUri-1="https://zenodo.org/record/254118/files/figure.png" httpUri-2="https://zenodo.org/record/254119/files/figure.png" httpUri-3="https://zenodo.org/record/254121/files/figure.png" httpUri-4="https://zenodo.org/record/254122/files/figure.png" pageId="24" pageNumber="271">Figs. 15</figureCitation>
F, 8B, D)
</paragraph>
</subSubSection>
<subSubSection id="01E46572E93DFFF7FF08F97C81FEF87D" pageId="24" pageNumber="271" type="description">
<paragraph id="494136F9E93DFFF7FF08F97C81FEF87D" blockId="24.[151,1437,1721,2034]" pageId="24" pageNumber="271">
<emphasis id="7B8AEAEBE93DFFF7FF08F97C814DF905" bold="true" box="[151,369,1721,1746]" pageId="24" pageNumber="271">Description. Body</emphasis>
clothed in long dense setae.
<emphasis id="7B8AEAEBE93DFFF7FD5CF97C833FF905" bold="true" box="[707,771,1721,1746]" pageId="24" pageNumber="271">Head</emphasis>
with scales on frons convex.
<emphasis id="7B8AEAEBE93DFFF7FBC0F97C84E4F905" bold="true" box="[1119,1240,1721,1746]" pageId="24" pageNumber="271">Pronotum</emphasis>
with one central vitta, lateral margins strongly convex.
<emphasis id="7B8AEAEBE93DFFF7FDD8F91882AFF921" bold="true" box="[583,659,1757,1782]" pageId="24" pageNumber="271">Elytra</emphasis>
with blue pattern, gray and brown pattern to blue and brown pattern.
<emphasis id="7B8AEAEBE93DFFF7FF08F8C480F3F8CD" bold="true" box="[151,207,1793,1818]" pageId="24" pageNumber="271">Legs</emphasis>
with front tibial shape flat expanded, tibial brush elongate; hind tibial apex truncate.
<emphasis id="7B8AEAEBE93DFFF7FB36F8C4857CF8CD" bold="true" box="[1193,1344,1793,1818]" pageId="24" pageNumber="271">Median lobe</emphasis>
curved.
<emphasis id="7B8AEAEBE93DFFF7FF08F8E08112F8E9" bold="true" box="[151,302,1829,1854]" pageId="24" pageNumber="271">Endophallus</emphasis>
with basal lobe not recurved and distended, dorsal lobe not present, distal projection absent, endophallus terminal not recurved, median ventral lobe absent, middle thick, ventral apical lobe absent, subapical lobe resting on posterior dorsal surface, transfer apparatus not sclerotized or absent.
<emphasis id="7B8AEAEBE93DFFF7FBDBF8A884DDF851" bold="true" box="[1092,1249,1901,1926]" pageId="24" pageNumber="271">Spermatheca</emphasis>
with basal lobes of relatively equal lengths.
</paragraph>
</subSubSection>
<subSubSection id="01E46572E93DFFF6FF58F8708198FE7B" lastPageId="25" lastPageNumber="272" pageId="24" pageNumber="271" type="materials_examined">
<paragraph id="494136F9E93DFFF6FF58F8708198FE7B" blockId="24.[151,1437,1721,2034]" lastBlockId="25.[151,1437,151,680]" lastPageId="25" lastPageNumber="272" pageId="24" pageNumber="271">
<typeStatus id="9645885BE93DFFF7FF58F8708176F819" box="[199,330,1973,1998]" pageId="24" pageNumber="271" type="holotype">
<emphasis id="7B8AEAEBE93DFFF7FF58F8708176F819" bold="true" box="[199,330,1973,1998]" pageId="24" pageNumber="271">HOLOTYPE</emphasis>
</typeStatus>
, (male):
<emphasis id="7B8AEAEBE93DFFF7FE2FF8708278F819" bold="true" box="[432,580,1973,1998]" pageId="24" pageNumber="271">NV: Nye Co.</emphasis>
sand dunes in Amargosa Desert,
<date id="3D401039E93DFFF7FC29F870847BF819" box="[950,1095,1973,1998]" pageId="24" pageNumber="271" value="1973-04-06">6 April 1973</date>
, Derham Giuliani coll. (
<collectionCode id="2FEFAE3CE93DFFF7FACBF87385ACF819" LSID="urn:lsid:biocol.org:col:34700" box="[1364,1424,1974,1998]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="24" pageNumber="271">CAS</collectionCode>
).
<emphasis id="7B8AEAEBE93DFFF7FF58F81C8328F825" bold="true" box="[199,788,2009,2034]" pageId="24" pageNumber="271">
Specimens examined.
<typeStatus id="9645885BE93DFFF7FE4CF81C8261F825" box="[467,605,2009,2034]" pageId="24" pageNumber="271" type="paratype">PARATYPES</typeStatus>
: NV: Nye Co.:
</emphasis>
sand dunes in Amargosa Desert,
<date id="3D401039E93DFFF7FB03F81C850FF825" box="[1180,1331,2009,2034]" pageId="24" pageNumber="271" value="1973-04-06">6 April 1973</date>
, Derham Giuliani coll. (5
<collectionCode id="2FEFAE3CE93CFFF6FED1FF5D81BAFF67" LSID="urn:lsid:biocol.org:col:34700" box="[334,390,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="25" pageNumber="272">CAS</collectionCode>
) [
<geoCoordinate id="2CCA503EE93CFFF6FE04FF528210FF67" box="[411,556,151,176]" direction="north" orientation="latitude" pageId="25" pageNumber="272" precision="1" value="36.64197">N36.641971</geoCoordinate>
,
<geoCoordinate id="2CCA503EE93CFFF6FDA9FF5282D8FF67" box="[566,740,151,176]" direction="west" orientation="longitude" pageId="25" pageNumber="272" precision="1" value="-116.569664">W-116.569667</geoCoordinate>
]; Amargosa Desert, “Big Dune”,
<date id="3D401039E93CFFF6FBCAFF5D84E3FF67" box="[1109,1247,152,176]" pageId="25" pageNumber="272" value="1973-05-01">1 May 1973</date>
, R. McPeak coll. (1
<collectionCode id="2FEFAE3CE93CFFF6FF2AFF7980CCFF03" LSID="urn:lsid:biocol.org:col:34700" box="[181,240,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="25" pageNumber="272">CAS</collectionCode>
); Big Dune,
<date id="3D401039E93CFFF6FEE0FF79821CFF02" box="[383,544,188,213]" pageId="25" pageNumber="272" value="1975-04-28">28 April 1975</date>
, F.G. Andrews and A.R. Hardy coll. (9
<collectionCode id="2FEFAE3CE93CFFF6FC43FF798416FF03" box="[988,1066,188,212]" pageId="25" pageNumber="272">CDFA</collectionCode>
);
<date id="3D401039E93CFFF6FBA2FF79853FFF03" box="[1085,1283,188,213]" pageId="25" pageNumber="272" value="1982-04-29" valueMax="1982-04-30" valueMin="1982-04-29">29-30 April 1982</date>
, L.G. Bezark, cereal bowl trap (1
<collectionCode id="2FEFAE3CE93CFFF6FE1CFF2581E0FF2F" box="[387,476,224,248]" pageId="25" pageNumber="272">CWOB</collectionCode>
); same location:
<date id="3D401039E93CFFF6FD39FF1A837DFF2F" box="[678,833,223,248]" pageId="25" pageNumber="272" value="1973-04-06">6 April 1973</date>
, Derham Giuliani coll. (11
<collectionCode id="2FEFAE3CE93CFFF6FB15FF2584E4FF2F" box="[1162,1240,224,248]" pageId="25" pageNumber="272">CDFA</collectionCode>
);
<date id="3D401039E93CFFF6FB70FF1A85A4FF2F" box="[1263,1432,223,248]" pageId="25" pageNumber="272" value="1974-04-16">16 April 1974</date>
, Derham Giuliani coll. (25
<collectionCode id="2FEFAE3CE93CFFF6FE46FEC18219FECB" box="[473,549,260,284]" pageId="25" pageNumber="272">CDFA</collectionCode>
, 2
<collectionCode id="2FEFAE3CE93CFFF6FDCDFEC18296FECB" box="[594,682,260,284]" pageId="25" pageNumber="272">CMNC</collectionCode>
); E Lathrop,
<date id="3D401039E93CFFF6FCD0FEC183C7FECB" box="[847,1019,260,285]" pageId="25" pageNumber="272" value="1974-04-15">15 April 1974</date>
, Derham Giuliani coll. (2
<collectionCode id="2FEFAE3CE93CFFF6FADCFEC185ADFECB" box="[1347,1425,260,284]" pageId="25" pageNumber="272">CDFA</collectionCode>
); Amargosa Desert,
<date id="3D401039E93CFFF6FEF3FEE281D0FE97" box="[364,492,295,320]" pageId="25" pageNumber="272" value="1973-04">April 1973</date>
, Derham Giuliani coll. (2
<collectionCode id="2FEFAE3CE93CFFF6FC82FEED8357FE97" box="[797,875,296,320]" pageId="25" pageNumber="272">CDFA</collectionCode>
);
<quantity id="8E069B1CE93CFFF6FC1FFEED8384FE97" box="[896,952,295,320]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="25" pageNumber="272" unit="mi" value="7.0">7 mi</quantity>
. E Lathrop, Amargosa River Sand Dunes, under
<taxonomicName id="8EFE4D7AE93CFFF6FF42FE888116FEB3" box="[221,298,333,356]" class="Magnoliopsida" family="Zygophyllaceae" genus="Larrea" kingdom="Plantae" order="Zygophyllales" pageId="25" pageNumber="272" phylum="Tracheophyta" rank="genus">
<emphasis id="7B8AEAEBE93CFFF6FF42FE888116FEB3" box="[221,298,333,356]" italics="true" pageId="25" pageNumber="272">Larrea</emphasis>
</taxonomicName>
,
<date id="3D401039E93CFFF6FEA7FE89823DFEB3" box="[312,513,332,357]" pageId="25" pageNumber="272" value="1980-02-24">24 February 1980</date>
, D.G. Kissinger coll. (1
<collectionCode id="2FEFAE3CE93CFFF6FC92FE898367FEB3" box="[781,859,332,356]" pageId="25" pageNumber="272">CDFA</collectionCode>
); Big Dune, E Lathrop,
<date id="3D401039E93CFFF6FBFAFE89850BFEB2" box="[1125,1335,332,357]" pageId="25" pageNumber="272" value="1985-03-21" valueMax="1985-03-22" valueMin="1985-03-21">21-22 March 1985</date>
, Derham Giuliani coll. (12
<collectionCode id="2FEFAE3CE93CFFF6FEF6FEB5818BFE5F" box="[361,439,368,392]" pageId="25" pageNumber="272">CDFA</collectionCode>
); E. side Big Dune,
<specimenCount id="5FF8FD70E93CFFF6FD36FEAA82D1FE50" box="[681,749,367,392]" pageId="25" pageNumber="272" type="generic">
<quantity id="8E069B1CE93CFFF6FD36FEAA82D5FE50" box="[681,745,367,392]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="25" pageNumber="272" unit="m" value="75.0">75 m</quantity>
.
</specimenCount>
Sand Dune-Creosote Scrub,
<date id="3D401039E93CFFF6FBDEFEAA853FFE5F" box="[1089,1283,367,392]" pageId="25" pageNumber="272" value="1978-04-16" valueMax="1978-04-17" valueMin="1978-04-16">16-17April 1978</date>
, sifting, Rolf Aalbu Coll. (1
<collectionCode id="2FEFAE3CE93CFFF6FEA0FE5181A4FE7B" box="[319,408,404,428]" pageId="25" pageNumber="272">CWOB</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="01E46572E93CFFF6FF58FE728519FE07" box="[199,1317,439,464]" pageId="25" pageNumber="272" type="distribution">
<paragraph id="494136F9E93CFFF6FF58FE728519FE07" blockId="25.[151,1437,151,680]" box="[199,1317,439,464]" pageId="25" pageNumber="272">
<emphasis id="7B8AEAEBE93CFFF6FF58FE72815CFE07" bold="true" box="[199,352,439,464]" pageId="25" pageNumber="272">Distribution.</emphasis>
(
<figureCitation id="D1C52A7CE93CFFF6FEF0FE7281FDFE07" box="[367,449,439,464]" captionStart="FIGURE 19" captionStartId="25.[151,249,1295,1317]" captionTargetBox="[150,1433,740,1271]" captionTargetId="figure@25.[150,1436,740,1274]" captionTargetPageId="25" captionText="FIGURE 19. Map of Miloderes occurrence data: M. setosus dark blue dots, M. tingi light blue dots, M. viridis green dots, M. panamintensis sp. nov. orange dots, M. mercuryensis yellow dots, M. amargosensis sp. nov. light green dot, M. nelsoni white dots, M. ubehebensis sp. nov. pink dot." httpUri="https://zenodo.org/record/254136/files/figure.png" pageId="25" pageNumber="272">Fig. 19</figureCitation>
) This species is endemic to the “Big Dune” of the Amargosa Valley, Nevada.
</paragraph>
</subSubSection>
<subSubSection id="01E46572E93CFFF6FF58FE198293FD7F" pageId="25" pageNumber="272" type="etymology">
<paragraph id="494136F9E93CFFF6FF58FE198162FDCF" blockId="25.[151,1437,151,680]" pageId="25" pageNumber="272">
<emphasis id="7B8AEAEBE93CFFF6FF58FE198172FE22" bold="true" box="[199,334,476,501]" pageId="25" pageNumber="272">Etymology.</emphasis>
This epithet is designated to reflect the geographic location of this species found only in the Amargosa Valley.
</paragraph>
<paragraph id="494136F9E93CFFF6FF58FDE18293FD7F" blockId="25.[151,1437,151,680]" pageId="25" pageNumber="272">
<emphasis id="7B8AEAEBE93CFFF6FF58FDE18327FDEA" bold="true" box="[199,795,548,573]" pageId="25" pageNumber="272">Intra- and interspecific morphological variation.</emphasis>
<taxonomicName id="8EFE4D7AE93CFFF6FCBAFDE18408FDEB" box="[805,1076,548,572]" class="Insecta" family="Curculionidae" genus="Miloderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="272" phylum="Arthropoda" rank="species" species="amargosensis">
<emphasis id="7B8AEAEBE93CFFF6FCBAFDE18408FDEB" box="[805,1076,548,572]" italics="true" pageId="25" pageNumber="272">Miloderes amargosensis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="8EFE4D7AE93CFFF6FA8CFDE08552FDEB" box="[1299,1390,549,572]" class="Insecta" family="Curculionidae" genus="Miloderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="272" phylum="Arthropoda" rank="species" species="tingi">
<emphasis id="7B8AEAEBE93CFFF6FA8CFDE08552FDEB" box="[1299,1390,549,572]" italics="true" pageId="25" pageNumber="272">M. tingi</emphasis>
</taxonomicName>
but differs markedly in the endophallus lacking a ventrobasal lobe and the pronotum having a dark central vitta. The shape of the rostrum is also squarer than in
<taxonomicName id="8EFE4D7AE93CFFF6FDF0FDA882F9FD53" box="[623,709,621,644]" class="Insecta" family="Curculionidae" genus="Miloderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="272" phylum="Arthropoda" rank="species" species="tingi">
<emphasis id="7B8AEAEBE93CFFF6FDF0FDA882F9FD53" box="[623,709,621,644]" italics="true" pageId="25" pageNumber="272">M. tingi</emphasis>
</taxonomicName>
. Little color variation is seen in this species with some individuals having a more blue gray color pattern to brown.
</paragraph>
</subSubSection>
</treatment>
</document>