treatments-xml/data/67/6A/87/676A87E0FFB4B15A2392FD5741630B19.xml

<document id="F46C4D44524FF367B1EDE27310DD48B5" ID-CLB-Dataset="22806" ID-DOI="10.11646/zootaxa.4407.1.2" ID-GBIF-Dataset="8e0aac3a-6556-4b96-a67e-fce9f0db0ba9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3766290" ID-ZooBank="B45E9ABD-CB52-4EE7-91D6-783E8E958AC5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587827908598" checkinUser="plazi" docAuthor="Short, Andrew Edward Z. &amp; Girón, Jennifer C." docDate="2018" docId="676A87E0FFB4B15A2392FD5741630B19" docLanguage="en" docName="zootaxa.4407.1.2.pdf.imf" docOrigin="Zootaxa 4407 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helochares (Hydrobaticus) politus Short &amp; Girón 2018, n. sp." docType="treatment" docUuid="1FB1AFED-3D86-48B0-A9B8-6E2E38831DD9" docUuidSource="ZooBank" docVersion="6" lastPageNumber="45" masterDocId="9B53FF98FFA4B14A2305FFE3420C0E4A" masterDocTitle="Review of the Helochares (Hydrobaticus) MacLeay of the New World (Coleoptera: Hydrophilidae: Acidocerinae)" masterLastPageNumber="50" masterPageNumber="29" pageNumber="45" updateTime="1698835862916" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="65DB6C49F20FD1BD7678560D757FD030">Review of the Helochares (Hydrobaticus) MacLeay of the New World (Coleoptera: Hydrophilidae: Acidocerinae)</mods:title>
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<mods:namePart id="25480E96EBAB19FC9D044D1C9F5751E9">Short, Andrew Edward Z.</mods:namePart>
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<mods:namePart id="A9C52A957E002E8BBAE1F2E141609DC1">Girón, Jennifer C.</mods:namePart>
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<treatment id="676A87E0FFB4B15A2392FD5741630B19" ID-DOI="http://doi.org/10.5281/zenodo.3803376" ID-GBIF-Taxon="163743972" ID-Zenodo-Dep="3803376" LSID="urn:lsid:zoobank.org:act:1FB1AFED-3D86-48B0-A9B8-6E2E38831DD9" httpUri="http://treatment.plazi.org/id/676A87E0FFB4B15A2392FD5741630B19" lastPageNumber="45" pageId="16" pageNumber="45">
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<paragraph id="EF7C36F6FFB4B15A2392FD57408D0C84" blockId="16.[151,641,692,751]" box="[151,641,692,718]" pageId="16" pageNumber="45">
<heading id="B434819AFFB4B15A2392FD57408D0C84" bold="true" box="[151,641,692,718]" fontSize="11" level="1" pageId="16" pageNumber="45" reason="1">
<emphasis id="DDB7EAE4FFB4B15A2392FD57408D0C84" bold="true" box="[151,641,692,718]" pageId="16" pageNumber="45">
<taxonomicName id="28C34D75FFB4B15A2392FD57403B0C84" ID-CoL="8Y4DP" authority="Short &amp; Girón, 2018" authorityName="Short &amp; Girón" authorityYear="2018" box="[151,567,692,718]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="politus" status="sp. nov." subGenus="Hydrobaticus">
<emphasis id="DDB7EAE4FFB4B15A2392FD57403B0C84" bold="true" box="[151,567,692,718]" italics="true" pageId="16" pageNumber="45">Helochares (Hydrobaticus) politus</emphasis>
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<taxonomicNameLabel id="C684579FFFB4B15A213AFD57408D0C84" box="[575,641,692,718]" pageId="16" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="A7D9657DFFB4B15A2392FD35438A0CA5" box="[151,390,726,751]" pageId="16" pageNumber="45" type="description">
<paragraph id="EF7C36F6FFB4B15A2392FD35438A0CA5" blockId="16.[151,641,692,751]" box="[151,390,726,751]" pageId="16" pageNumber="45">
<figureCitation id="77F82A73FFB4B15A2392FD3542F60CA5" box="[151,250,726,751]" captionStart="FIGURE 4" captionStartId="10.[151,250,1284,1306]" captionTargetBox="[151,1436,193,1262]" captionTargetId="figure@10.[151,1436,193,1262]" captionTargetPageId="10" captionText="FIGURE 4. Dorsal habitus images of Helochares (Hydrobaticus) spp. (A) H. politus n. sp., (B) H. laevis n. sp." figureDoi="http://doi.org/10.5281/zenodo.3766298" httpUri="https://zenodo.org/record/3766298/files/figure.png" pageId="16" pageNumber="45">Figs. 4A</figureCitation>
, 
<figureCitation id="77F82A73FFB4B15A2200FD3543130CA5" box="[261,287,726,751]" captionStart="FIGURE 5" captionStartId="11.[151,250,1936,1958]" captionTargetBox="[151,1436,493,1914]" captionTargetId="figure@11.[151,1436,493,1914]" captionTargetPageId="11" captionText="FIGURE 5. Heads of Helochares (Hydrobaticus) spp. (A) H. maculicollis (Pennsylvania), (B) H. normatus (California), (C) H. normatus (Arizona), (D) H. nexus n. sp., (E) H. zamora n. sp., (F) H. trujillo n. sp., (G) H. championi, (H) H. sp. A., (I) H. politus n. sp." figureDoi="http://doi.org/10.5281/zenodo.3766300" httpUri="https://zenodo.org/record/3766300/files/figure.png" pageId="16" pageNumber="45">5I</figureCitation>
, 
<figureCitation id="77F82A73FFB4B15A222EFD3443480CA4" box="[299,324,727,750]" captionStart="FIGURE 7" captionStartId="13.[151,250,1200,1222]" captionTargetBox="[347,1229,206,1166]" captionTargetId="figure@13.[340,1247,193,1179]" captionTargetPageId="13" captionText="FIGURE 7. Aedeagi of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. zamora n. sp., (C) H. maculicollis (Florida), (D) H. maculicollis (Kansas), (E) H. maculicollis (Pennsylvania), (F) H. nexus n. sp. (Venezuela), (G) H. nexus n. sp. (Panama), (H) H. nexus n. sp. (Ecuador), (I) H. politus n. sp., (J) H. laevis n. sp., (K) H. championi." figureDoi="http://doi.org/10.5281/zenodo.3766304" httpUri="https://zenodo.org/record/3766304/files/figure.png" pageId="16" pageNumber="45">7I</figureCitation>
, 
<figureCitation id="77F82A73FFB4B15A2255FD34438E0CA5" box="[336,386,727,751]" captionStart="FIGURE 10" captionStartId="18.[151,250,1999,2021]" captionTargetBox="[151,1436,373,1976]" captionTargetId="figure@18.[151,1436,373,1977]" captionTargetPageId="18" captionText="FIGURE 10. Distribution of New World Helochares (Hydrobaticus) spp." figureDoi="http://doi.org/10.5281/zenodo.3766310" httpUri="https://zenodo.org/record/3766310/files/figure.png" pageId="16" pageNumber="45">10B</figureCitation>
.
</paragraph>
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<subSubSection id="A7D9657DFFB4B15A2392FCFD41E90D11" pageId="16" pageNumber="45" type="materials_examined">
<paragraph id="EF7C36F6FFB4B15A2392FCFD41E90D11" blockId="16.[151,1437,798,1363]" pageId="16" pageNumber="45">
<emphasis id="DDB7EAE4FFB4B15A2392FCFD434B0D7D" bold="true" box="[151,327,798,823]" pageId="16" pageNumber="45">Type material.</emphasis>
<materialsCitation id="5FAB3CABFFB4B15A224AFCFD41E90D11" ID-GBIF-Occurrence="2608702033" collectionCode="CAS" collectorName="N. Santa Eulalia &amp; D. E. Breedlove" country="Guatemala" elevation="2865" location="Ixtantan" municipality="San Mateo" pageId="16" pageNumber="45" specimenCount="1" stateProvince="Huehuetenango" typeStatus="holotype">
<emphasis id="DDB7EAE4FFB4B15A224AFCFD40160D7D" bold="true" box="[335,538,798,823]" pageId="16" pageNumber="45">
<typeStatus id="30788854FFB4B15A224AFCFD43B00D7D" box="[335,444,798,823]" pageId="16" pageNumber="45" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
“ 
<collectingCountry id="97D47666FFB4B15A2128FCFC40EE0D7D" box="[557,738,799,823]" name="Guatemala" pageId="16" pageNumber="45">GUATEMALA</collectingCountry>
: Departamen-/to de 
<collectingRegion id="2D07F814FFB4B15A20CFFCFD468C0D7D" box="[970,1152,798,823]" country="Guatemala" name="Huehuetenango" pageId="16" pageNumber="45">Huehuetenango</collectingRegion>
,/ 
<quantity id="283B9B13FFB4B15A2788FCFD46DD0D7D" box="[1165,1233,798,823]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="16" pageNumber="45" unit="km" value="11.0">11 km</quantity>
<collectorName id="42365320FFB4B15A27DCFCFC47980D7D" box="[1241,1428,798,823]" pageId="16" pageNumber="45">N. Santa Eulalia</collectorName>
/ on road to 
<collectingMunicipality id="0F18AC8CFFB4B15A220AFCA0438B0D11" box="[271,391,835,859]" pageId="16" pageNumber="45">San Mateo</collectingMunicipality>
/ 
<location id="EA1C602DFFB4B15A2292FCA143F90D11" LSID="urn:lsid:plazi:treatment:676A87E0FFB4B15A2392FD5741630B19:EA1C602DFFB4B15A2292FCA143F90D11" box="[407,501,834,859]" country="Guatemala" municipality="San Mateo" name="Ixtantan" pageId="16" pageNumber="45" stateProvince="Huehuetenango">Ixtantan</location>
,/ 
<quantity id="283B9B13FFB4B15A2104FCA140430D11" box="[513,591,834,859]" metricMagnitude="3" metricUnit="m" metricValue="2.8649999999999998" pageId="16" pageNumber="45" unit="m" value="2865.0">
<elevation id="64EED1C5FFB4B15A2104FCA140430D11" box="[513,591,834,859]" metricMagnitude="3" metricUnit="m" metricValue="2.8649999999999998" pageId="16" pageNumber="45" unit="m" value="2865.0">2865m</elevation>
</quantity>
5.II&gt;1965/ 
<collectorName id="42365320FFB4B15A21CAFCA141890D11" box="[719,901,834,859]" pageId="16" pageNumber="45">D. E. Breedlove</collectorName>
” (
<collectionCode id="89D2AE33FFB4B15A2098FCA041D50D11" box="[925,985,835,859]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="45" type="Museum">CAS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFB4B15A23C2FC85439E0AD5" pageId="16" pageNumber="45" type="diagnosis">
<paragraph id="EF7C36F6FFB4B15A23C2FC8546320A1D" blockId="16.[151,1437,798,1363]" pageId="16" pageNumber="45">
<emphasis id="DDB7EAE4FFB4B15A23C2FC8543C10D35" bold="true" box="[199,461,870,895]" pageId="16" pageNumber="45">Differential diagnosis.</emphasis>
Body size 
<quantity id="283B9B13FFB4B15A214FFC8540AE0D34" box="[586,674,870,895]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="16" pageNumber="45" unit="mm" value="6.6">6.6 mm</quantity>
. Clypeus dark yellow with the central third darkened to dark brown. Elytra without any detectable serial punctures or striae (
<figureCitation id="77F82A73FFB4B15A202FFC6941850DE9" box="[810,905,906,931]" captionStart="FIGURE 4" captionStartId="10.[151,250,1284,1306]" captionTargetBox="[151,1436,193,1262]" captionTargetId="figure@10.[151,1436,193,1262]" captionTargetPageId="10" captionText="FIGURE 4. Dorsal habitus images of Helochares (Hydrobaticus) spp. (A) H. politus n. sp., (B) H. laevis n. sp." figureDoi="http://doi.org/10.5281/zenodo.3766298" httpUri="https://zenodo.org/record/3766298/files/figure.png" pageId="16" pageNumber="45">Fig. 4A</figureCitation>
). Abdominal ventrites uniformly dark brown; pubescence dense and evenly distributed, the longest hairs shorter than those on the metaventrite. Aedeagus (
<figureCitation id="77F82A73FFB4B15A266BFC4D42A30DA1" captionStart="FIGURE 7" captionStartId="13.[151,250,1200,1222]" captionTargetBox="[347,1229,206,1166]" captionTargetId="figure@13.[340,1247,193,1179]" captionTargetPageId="13" captionText="FIGURE 7. Aedeagi of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. zamora n. sp., (C) H. maculicollis (Florida), (D) H. maculicollis (Kansas), (E) H. maculicollis (Pennsylvania), (F) H. nexus n. sp. (Venezuela), (G) H. nexus n. sp. (Panama), (H) H. nexus n. sp. (Ecuador), (I) H. politus n. sp., (J) H. laevis n. sp., (K) H. championi." figureDoi="http://doi.org/10.5281/zenodo.3766304" httpUri="https://zenodo.org/record/3766304/files/figure.png" pageId="16" pageNumber="45">Fig. 7I</figureCitation>
) 2.5-times longer than its greatest width; apex of parameres rounded, with outer corners widely rounded; inner corners of apex of parameres forming a nearly right angle; lateral outer margins of parameres parallel, slightly constricted near midlength; maximum width of aedeagus at basal half, nearly as wide as maximum width of aedeagus at apical half; median lobe with apex gradually tapering to an acute point.
</paragraph>
<paragraph id="EF7C36F6FFB4B15A23C2FB81439E0AD5" blockId="16.[151,1437,798,1363]" pageId="16" pageNumber="45">
This species is extremely similar to 
<taxonomicName id="28C34D75FFB4B15A215BFB8040CF0A31" authorityName="Short &amp; Girón" authorityYear="2018" box="[606,707,1123,1147]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="DDB7EAE4FFB4B15A215BFB8040CF0A31" box="[606,707,1123,1147]" italics="true" pageId="16" pageNumber="45">H. laevis</emphasis>
</taxonomicName>
, from which it can only be definitively separated by the form of the aedeagus (
<figureCitation id="77F82A73FFB4B15A2233FB6543890AD5" box="[310,389,1158,1183]" captionStart="FIGURE 7" captionStartId="13.[151,250,1200,1222]" captionTargetBox="[347,1229,206,1166]" captionTargetId="figure@13.[340,1247,193,1179]" captionTargetPageId="13" captionText="FIGURE 7. Aedeagi of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. zamora n. sp., (C) H. maculicollis (Florida), (D) H. maculicollis (Kansas), (E) H. maculicollis (Pennsylvania), (F) H. nexus n. sp. (Venezuela), (G) H. nexus n. sp. (Panama), (H) H. nexus n. sp. (Ecuador), (I) H. politus n. sp., (J) H. laevis n. sp., (K) H. championi." figureDoi="http://doi.org/10.5281/zenodo.3766304" httpUri="https://zenodo.org/record/3766304/files/figure.png" pageId="16" pageNumber="45">Fig. 7I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFB4B15A23C2FB49471C0AAD" pageId="16" pageNumber="45" type="description">
<paragraph id="EF7C36F6FFB4B15A23C2FB49471C0AAD" blockId="16.[151,1437,798,1363]" pageId="16" pageNumber="45">
<emphasis id="DDB7EAE4FFB4B15A23C2FB4943550A89" bold="true" box="[199,345,1194,1219]" pageId="16" pageNumber="45">Description.</emphasis>
In addition to differential diagnosis: pronotum dark brown to black in central half (
<figureCitation id="77F82A73FFB4B15A2634FB4947830A89" box="[1329,1423,1194,1219]" captionStart="FIGURE 4" captionStartId="10.[151,250,1284,1306]" captionTargetBox="[151,1436,193,1262]" captionTargetId="figure@10.[151,1436,193,1262]" captionTargetPageId="10" captionText="FIGURE 4. Dorsal habitus images of Helochares (Hydrobaticus) spp. (A) H. politus n. sp., (B) H. laevis n. sp." figureDoi="http://doi.org/10.5281/zenodo.3766298" httpUri="https://zenodo.org/record/3766298/files/figure.png" pageId="16" pageNumber="45">Fig. 4A</figureCitation>
). Ground punctation on head and pronotum moderately impressed, slightly less impressed on the elytra.
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFB4B15A23C2FB11474F0B41" box="[199,1347,1266,1291]" pageId="16" pageNumber="45" type="distribution">
<paragraph id="EF7C36F6FFB4B15A23C2FB11474F0B41" blockId="16.[151,1437,798,1363]" box="[199,1347,1266,1291]" pageId="16" pageNumber="45">
<emphasis id="DDB7EAE4FFB4B15A23C2FB11436C0B41" bold="true" box="[199,352,1266,1291]" pageId="16" pageNumber="45">Distribution.</emphasis>
<collectingCountry id="97D47666FFB4B15A2262FB1143EC0B41" box="[359,480,1266,1291]" name="Guatemala" pageId="16" pageNumber="45">Guatemala</collectingCountry>
(
<figureCitation id="77F82A73FFB4B15A22F5FB11405A0B41" box="[496,598,1266,1291]" captionStart="FIGURE 10" captionStartId="18.[151,250,1999,2021]" captionTargetBox="[151,1436,373,1976]" captionTargetId="figure@18.[151,1436,373,1977]" captionTargetPageId="18" captionText="FIGURE 10. Distribution of New World Helochares (Hydrobaticus) spp." figureDoi="http://doi.org/10.5281/zenodo.3766310" httpUri="https://zenodo.org/record/3766310/files/figure.png" pageId="16" pageNumber="45">Fig. 10B</figureCitation>
). The single specimen was taken at a very high elevation (
<quantity id="283B9B13FFB4B15A27E1FB11473B0B41" box="[1252,1335,1266,1291]" metricMagnitude="3" metricUnit="m" metricValue="2.8649999999999998" pageId="16" pageNumber="45" unit="m" value="2865.0">2865m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFB4B15A23C2FAF547080B65" box="[199,1284,1302,1327]" pageId="16" pageNumber="45" type="etymology">
<paragraph id="EF7C36F6FFB4B15A23C2FAF547080B65" blockId="16.[151,1437,798,1363]" box="[199,1284,1302,1327]" pageId="16" pageNumber="45">
<emphasis id="DDB7EAE4FFB4B15A23C2FAF543420B65" bold="true" box="[199,334,1302,1327]" pageId="16" pageNumber="45">Etymology.</emphasis>
<emphasis id="DDB7EAE4FFB4B15A2251FAF543AF0B64" box="[340,419,1302,1326]" italics="true" pageId="16" pageNumber="45">Politus</emphasis>
(L), after the smooth elytra which lacks any coarse serial punctures or striae.
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFB4B15A23C2FAD941630B19" box="[199,879,1338,1363]" pageId="16" pageNumber="45" type="biology_ecology">
<paragraph id="EF7C36F6FFB4B15A23C2FAD941630B19" blockId="16.[151,1437,798,1363]" box="[199,879,1338,1363]" pageId="16" pageNumber="45">
<emphasis id="DDB7EAE4FFB4B15A23C2FAD9432B0B19" bold="true" box="[199,295,1338,1363]" pageId="16" pageNumber="45">Biology.</emphasis>
Nothing is known about the biology of this species.
</paragraph>
</subSubSection>
</treatment>
</document>