162 lines
19 KiB
XML
162 lines
19 KiB
XML
<document id="9CF83C2D6201EE5F6A6720B9B488AC20" ID-DOI="10.5281/zenodo.207073" ID-GBIF-Dataset="a3815507-6100-464e-aaf6-60d6820a1eec" ID-ISSN="1175-5326" ID-Zenodo-Dep="207073" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460228776049" checkinUser="plazi" docAuthor="Torres-Carvajal, Omar, Etheridge, Richard & Queiroz, Kevin De" docDate="2011" docId="038287FDFF99606469838D84D056FAC7" docLanguage="en" docName="zt02752p044.pdf" docOrigin="Zootaxa 2752" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Morunasaurus groi Dunn 1933" docType="treatment" docVersion="9" lastPageNumber="36" masterDocId="FFBBFF85FFBB604769148B6ED174FF93" masterDocTitle="A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="35" updateTime="1698238838839" updateUser="plazi">
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<mods:title id="FD4D403F6D43F49D3C2AAC3EF415A071">A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)</mods:title>
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<mods:namePart id="9B3C6EF4E4740A2E4B4EA28D1EE790D4">Torres-Carvajal, Omar</mods:namePart>
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<mods:namePart id="6836BE66FE7EB8449613C17713E08F6C">Etheridge, Richard</mods:namePart>
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<mods:namePart id="CCC0265E9077E70B8B8F0E5570909573">Queiroz, Kevin De</mods:namePart>
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<mods:title id="5F48CFB0F3A7B93853DD42868A3D1753">Zootaxa</mods:title>
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<mods:date id="6B4D70D95D251D047051B17A562A323E">2011</mods:date>
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<mods:detail id="394A19C7E34E87DD13537F8AC217F66B" type="volume">
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<mods:number id="BB41065343358963BEDC8C9F9C26362E">2752</mods:number>
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<mods:identifier id="BD33C555FB54869943D06A550EBAA61C" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038287FDFF99606469838D84D056FAC7" ID-DOI="http://doi.org/10.5281/zenodo.6184985" ID-GBIF-Taxon="119387055" ID-Zenodo-Dep="6184985" LSID="urn:lsid:plazi:treatment:038287FDFF99606469838D84D056FAC7" httpUri="http://treatment.plazi.org/id/038287FDFF99606469838D84D056FAC7" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
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<subSubSection id="C3316560FF99606569838D84D3D9F8F6" pageId="34" pageNumber="35" type="nomenclature">
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<paragraph id="8B9436EBFF99606569838D84D367F897" blockId="34.[151,531,1770,1796]" box="[151,531,1770,1796]" pageId="34" pageNumber="35">
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<heading id="D0DC8187FF99606569838D84D367F897" bold="true" box="[151,531,1770,1796]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
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<taxonomicName id="4C2B4D68FF99606569838D84D367F897" authority="Dunn 1933" authorityName="Dunn" authorityYear="1933" box="[151,531,1770,1796]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="groi">
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<emphasis id="B95FEAF9FF99606569838D84D367F897" bold="true" box="[151,531,1770,1796]" pageId="34" pageNumber="35">
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<emphasis id="B95FEAF9FF99606569838D84D0F0F897" bold="true" box="[151,388,1770,1796]" italics="true" pageId="34" pageNumber="35">Morunasaurus groi</emphasis>
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<bibRefCitation id="EFBA4B1AFF996065689F8D84D367F897" author="Dunn" box="[395,531,1770,1796]" pageId="34" pageNumber="39" refString="Dunn, E. R. (1933) Amphibians and reptiles from El Valle de Anton, Panama. Occasional Papers of the Boston Society of Natural History, 8, 65 - 79." type="journal article" year="1933">Dunn 1933</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B9436EBFF99606569838C5FD3D9F8F6" blockId="34.[151,685,1841,1893]" pageId="34" pageNumber="35">Proposed standard English name: Gro’s manticores Proposed standard Spanish name: mantícoras de Gro</paragraph>
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</subSubSection>
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<subSubSection id="C3316560FF99606569838CE7D3F4F82E" pageId="34" pageNumber="35" type="reference_group">
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<paragraph id="8B9436EBFF99606569838CE7D3F4F82E" blockId="34.[151,1436,1927,1981]" pageId="34" pageNumber="35">
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<taxonomicName id="4C2B4D68FF99606569838CE7D372F80E" authority="Dunn (1933:76)" authorityName="Dunn" authorityPageNumber="76" authorityYear="1933" box="[151,518,1927,1950]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="groi">
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<emphasis id="B95FEAF9FF99606569838CE7D02FF80D" box="[151,347,1929,1950]" italics="true" pageId="34" pageNumber="35">Morunasaurus groi</emphasis>
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<bibRefCitation id="EFBA4B1AFF99606568758CE6D372F80E" author="Dunn" box="[353,518,1927,1949]" pageId="34" pageNumber="39" refString="Dunn, E. R. (1933) Amphibians and reptiles from El Valle de Anton, Panama. Occasional Papers of the Boston Society of Natural History, 8, 65 - 79." type="journal article" year="1933">Dunn (1933:76)</bibRefCitation>
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</taxonomicName>
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.
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<typeStatus id="54908849FF9960656B1B8CE9D305F80E" box="[527,625,1927,1949]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
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: MCZ 34875, from “El Valle de Anton [
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<geoCoordinate id="EE1F502CFF9960656D178CE9D538F80E" box="[1027,1100,1927,1949]" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="8.6">8º36'N</geoCoordinate>
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,
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<geoCoordinate id="EE1F502CFF9960656D428CE9D5D0F80E" box="[1110,1188,1927,1949]" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="925" value="-80.11667">80º7'W</geoCoordinate>
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], [Provincia Cocle]
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<collectingCountry id="F33C767BFF9960656C798CE6D181F82F" name="Panama" pageId="34" pageNumber="35">Panama</collectingCountry>
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”;
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<bibRefCitation id="EFBA4B1AFF996065681E8CC9D309F82E" author="Peters" box="[266,637,1959,1981]" pageId="34" pageNumber="40" refString="Peters, J. A. & Donoso-Barros, R. (1970) Catalogue of the neotropical Squamata: Part II. Lizards and amphisbaenians. United States National Museum Bulletin, 297, 1 - 293." type="journal article" year="1970">Peters & Donoso-Barros (1970:204)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3316560FF98606469838BF9D433FEAC" pageId="35" pageNumber="36" type="diagnosis">
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<paragraph id="8B9436EBFF98606469838BF9D433FEAC" blockId="35.[151,1437,151,1364]" pageId="35" pageNumber="36">
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<emphasis id="B95FEAF9FF98606469838BF9D065FF23" bold="true" box="[151,273,151,176]" pageId="35" pageNumber="36">Diagnosis.</emphasis>
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This species can be distinguished from other species of
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<taxonomicName id="4C2B4D68FF9860646A8C8BF7D549FF23" box="[920,1085,153,176]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="36" phylum="Chordata" rank="genus">
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<emphasis id="B95FEAF9FF9860646A8C8BF7D549FF23" box="[920,1085,153,176]" italics="true" pageId="35" pageNumber="36">Morunasaurus</emphasis>
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</taxonomicName>
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by lacking an enlarged row of vertebral scales, and by having the caudal whorls of spines separated by three transverse rows of scales ventrally and four transverse rows of scales dorsally (versus two and three, respectively, in other
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<taxonomicName id="4C2B4D68FF9860646D8A8B8FD437FF6B" box="[1182,1347,225,248]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="36" phylum="Chordata" rank="genus">
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<emphasis id="B95FEAF9FF9860646D8A8B8FD437FF6B" box="[1182,1347,225,248]" italics="true" pageId="35" pageNumber="36">Morunasaurus</emphasis>
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</taxonomicName>
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).
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<bibRefCitation id="EFBA4B1AFF9860646C4A8B8ED195FE8F" author="Dunn" pageId="35" pageNumber="39" refString="Dunn, E. R. (1933) Amphibians and reptiles from El Valle de Anton, Panama. Occasional Papers of the Boston Society of Natural History, 8, 65 - 79." type="journal article" year="1933">Dunn (1933)</bibRefCitation>
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mentioned that
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<taxonomicName id="4C2B4D68FF986064688A8A6BD080FE8F" box="[414,500,261,284]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="groi">
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<emphasis id="B95FEAF9FF986064688A8A6BD080FE8F" box="[414,500,261,284]" italics="true" pageId="35" pageNumber="36">M. groi</emphasis>
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</taxonomicName>
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also differs from
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<taxonomicName id="4C2B4D68FF9860646BD18A6BD22CFE8F" box="[709,856,260,284]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="annularis">
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<emphasis id="B95FEAF9FF9860646BD18A6BD22CFE8F" box="[709,856,260,284]" italics="true" pageId="35" pageNumber="36">M. annularis</emphasis>
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</taxonomicName>
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in having a circular (presumably in cross section) rather than compressed tail; however, we find that the tail in both species is nearly circular in cross section.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3316560FF98606469D18A25D05AFB57" pageId="35" pageNumber="36" type="description">
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<paragraph id="8B9436EBFF98606469D18A25D2CCFC87" blockId="35.[151,1437,151,1364]" pageId="35" pageNumber="36">
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<emphasis id="B95FEAF9FF98606469D18A25D023FEF7" bold="true" box="[197,343,331,356]" pageId="35" pageNumber="36">Description.</emphasis>
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(1) dorsal head scales smooth, not projecting dorsally, some occipitals granular or subconical; (2) posterior superciliaries not enlarged relative to adjacent scales; (3) scales on lateral edge of skull roof just posterior to superciliaries not projecting; (4) pretympanic scales similar in size; (5) gular scales granular or flat and smooth, separated by skin, not projecting ventrally; (6) dorsal neck scales heterogeneous in size, granular or large and conical; lateral neck scales granular, homogeneous in size; (7) vertebrals not forming middorsal row of enlarged scales; (8) nuchal region without middorsal crest; (9) dorsals smooth and heterogeneous in size, with largest scales more projecting and abundant on posterior half of body; (10) discontinuous longitudinal row of raised, enlarged scales between dorsals and flank scales present; (11) scales on flanks heterogeneous in size, mostly granular with a few scattered enlarged scales; (12) ventrals smooth; (13) fore limb scales smooth or slightly keeled dorsally and ventrally; (14) hind limb scales smooth or slightly keeled dorsally and ventrally; scattered conical, sharply pointed, enlarged scales present dorsally; scales of pes heterogeneous in size; (15) caudals heterogeneous, with scales making up the posteriormost whorl of each segment modified as conspicuous spines; (16) tail nearly circular in cross section. Meristic and morphometric characters are presented in
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<tableCitation id="C6A90350FF9860646A4A8992D2C6FC87" box="[862,946,764,788]" captionStart="TABLE 1" captionStartId="6.[151,235,1304,1325]" captionTargetBox="[151,1436,1377,1892]" captionTargetPageId="6" captionText="TABLE 1. Summary of counts and measurements (mm) for species of Hoplocercinae. Range (first line) and mean ± standard" httpUri="http://table.plazi.org/id/DF546663FFBD604169838E76D4E8FABD" pageId="35" pageNumber="36" tableUuid="DF546663FFBD604169838E76D4E8FABD">Table 1</tableCitation>
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.
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</paragraph>
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<paragraph id="8B9436EBFF98606469D1884ED0E0FC54" blockId="35.[151,1437,151,1364]" pageId="35" pageNumber="36">
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<emphasis id="B95FEAF9FF98606469D1884ED0F2FCAB" box="[197,390,800,824]" italics="true" pageId="35" pageNumber="36">Coloration in life</emphasis>
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. Dorsal background reddish brown with dark brown transverse bands that reach the middle of the flanks to break up into small dark spots ventrally; head and neck reddish; white band bordered with dark brown on both sides extends from shoulder to scapular region; chin and infralabial region scarlet red in adult males; gular region grayish brown; pectoral region pale chrome orange; venter dirty white in males and yellow in females (
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<bibRefCitation id="EFBA4B1AFF986064698A88C1D0F3FC54" author="Corredor" box="[158,391,943,968]" pageId="35" pageNumber="39" refString="Corredor, V., Rengifo, J. M. & Ayala, S. C. (1985) Discovery of Morunasaurus groi Dunn (Sauria, Iguanidae) in northwestern Colombia. Journal of Herpetology, 19, 162 - 164." type="journal article" year="1985">
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Corredor
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<emphasis id="B95FEAF9FF986064681888DFD030FC5B" box="[268,324,944,968]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
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1985
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</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8B9436EBFF98606469D188BAD05AFB57" blockId="35.[151,1437,151,1364]" pageId="35" pageNumber="36">
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<emphasis id="B95FEAF9FF98606469D188BAD0F0FC7F" bold="true" box="[197,388,979,1004]" pageId="35" pageNumber="36">Natural history.</emphasis>
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<bibRefCitation id="EFBA4B1AFF986064689888BBD368FC7F" author="Dunn" box="[396,540,980,1004]" pageId="35" pageNumber="39" refString="Dunn, E. R. (1933) Amphibians and reptiles from El Valle de Anton, Panama. Occasional Papers of the Boston Society of Natural History, 8, 65 - 79." type="journal article" year="1933">Dunn (1933)</bibRefCitation>
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reported that lizards of this species seem to be diurnal and live under boulders on a mountainside. All specimens were seen within two feet of their burrows, which were short and shallow and into which the lizards retreated when approached. A female and a juvenile were found in the same burrow (
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<bibRefCitation id="EFBA4B1AFF9860646C488F72D1A7FBCB" author="Dunn" pageId="35" pageNumber="39" refString="Dunn, E. R. (1933) Amphibians and reptiles from El Valle de Anton, Panama. Occasional Papers of the Boston Society of Natural History, 8, 65 - 79." type="journal article" year="1933">Dunn 1933</bibRefCitation>
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).
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<bibRefCitation id="EFBA4B1AFF98606469F38F2ED094FBCB" box="[231,480,1088,1112]" pageId="35" pageNumber="39" refString="Corredor, V., Rengifo, J. M. & Ayala, S. C. (1985) Discovery of Morunasaurus groi Dunn (Sauria, Iguanidae) in northwestern Colombia. Journal of Herpetology, 19, 162 - 164." type="journal article">
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Corredor
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<emphasis id="B95FEAF9FF98606468408F2FD0FAFBCB" box="[340,398,1088,1112]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
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(1985)
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</bibRefCitation>
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collected specimens of this species in an area characterized by low rolling hills and sandy, well-drained soil surrounded by dense, moist forest; the lizards were active during the day on the forest floor, in or near open burrows located under logs, with no surrounding vegetation. Clutch size is probably one egg (
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<bibRefCitation id="EFBA4B1AFF986064698A8FC2D055FB57" author="Dunn" box="[158,289,1196,1220]" pageId="35" pageNumber="39" refString="Dunn, E. R. (1933) Amphibians and reptiles from El Valle de Anton, Panama. Occasional Papers of the Boston Society of Natural History, 8, 65 - 79." type="journal article" year="1933">Dunn 1933</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3316560FF98606469D18FA1D056FAC7" pageId="35" pageNumber="36" type="distribution">
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<paragraph id="8B9436EBFF98606469D18FA1D056FAC7" blockId="35.[151,1437,151,1364]" pageId="35" pageNumber="36">
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<emphasis id="B95FEAF9FF98606469D18FA1D02AFB7B" bold="true" box="[197,350,1231,1256]" pageId="35" pageNumber="36">Distribution.</emphasis>
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<taxonomicName id="4C2B4D68FF98606468708FBFD34AFB7B" box="[356,574,1233,1256]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="groi">
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<emphasis id="B95FEAF9FF98606468708FBFD34AFB7B" box="[356,574,1233,1256]" italics="true" pageId="35" pageNumber="36">Morunasaurus groi</emphasis>
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</taxonomicName>
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occurs in
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<collectingCountry id="F33C767BFF9860646BA58FBFD27EFB7B" box="[689,778,1233,1256]" name="Panama" pageId="35" pageNumber="36">Panama</collectingCountry>
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and the western slopes of the western Andean cordillera in northern
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<collectingCountry id="F33C767BFF98606469EA8F9AD018FA9F" box="[254,364,1268,1292]" name="Colombia" pageId="35" pageNumber="36">Colombia</collectingCountry>
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at elevations between
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<quantity id="4CD39B0EFF9860646B678F9AD380FA9F" box="[627,756,1268,1292]" metricMagnitude="2" metricUnit="m" metricValue="7.525" metricValueMax="8.05" metricValueMin="7.0" pageId="35" pageNumber="36" unit="m" value="752.5" valueMax="805.0" valueMin="700.0">700–805 m</quantity>
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(
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<figureCitation id="13102A6EFF9860646A118F9AD22CFA9F" box="[773,856,1268,1292]" captionStart="FIGURE 11" captionStartId="34.[151,250,1664,1687]" captionTargetBox="[159,1434,217,1627]" captionTargetId="figure@34.[151,1436,195,1643]" captionTargetPageId="34" captionText="FIGURE 11. Distribution of Morunasaurus annularis (triangles), M. groi (squares), and M. peruvianus (dots)." httpUri="https://zenodo.org/record/207088/files/figure.png" pageId="35" pageNumber="36">Fig. 11</figureCitation>
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). This species is known to occur in sympatry with
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<taxonomicName id="4C2B4D68FF98606469838E76D0EAFAA3" box="[151,414,1304,1328]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="heterolepis">
|
||
<emphasis id="B95FEAF9FF98606469838E76D0EAFAA3" box="[151,414,1304,1328]" italics="true" pageId="35" pageNumber="36">Enyalioides heterolepis</emphasis>
|
||
</taxonomicName>
|
||
in northwestern
|
||
<collectingCountry id="F33C767BFF9860646B4B8E76D3A4FAA3" box="[607,720,1304,1328]" name="Colombia" pageId="35" pageNumber="36">Colombia</collectingCountry>
|
||
, and it is very likely that the two species are sympatric in
|
||
<collectingCountry id="F33C767BFF9860646C7D8E77D1B2FAC7" name="Panama" pageId="35" pageNumber="36">Panama</collectingCountry>
|
||
as well.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |