164 lines
18 KiB
XML
164 lines
18 KiB
XML
<document id="07ACFA91158F5AEDFBD4ECE1C3EA25EB" ID-ISSN="2201-4349" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629577487243" checkinUser="felipe" docAuthor="Bradley J Sinclair" docDate="2003" docId="038287C6FF81FFD7FED553AAFE03B79D" docLanguage="en" docName="RecAustMus.55.1.1-44.pdf" docOrigin="Records of the Australian Museum 55" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Ceratomerus bickeli Bradley J Sinclair 2003, n.sp." docType="treatment" docVersion="3" lastPageNumber="19" masterDocId="FFBBFFBEFF93FFC5FF995332FFA7B045" masterDocTitle="Taxonomy, Phylogeny and Zoogeography of the Subfamily Ceratomerinae of Australia (Diptera: Empidoidea)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="19" updateTime="1699079322007" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo id="C8EA65913D13AACC1898D5986DBFA433">
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<mods:title id="D419E1E76F4930C8F4EB5C42F27F7695">Taxonomy, Phylogeny and Zoogeography of the Subfamily Ceratomerinae of Australia (Diptera: Empidoidea)</mods:title>
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<mods:name id="E589F599EE72AEAC05477A83A0912C4E" type="personal">
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<mods:roleTerm id="C09E2E01346117FF0416E3347ECDFFF8">Author</mods:roleTerm>
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<mods:namePart id="373E7B9511233C8624AFDF307A7E1565">Bradley J Sinclair</mods:namePart>
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<mods:typeOfResource id="62BDB1778E9D232D26BAAA99613D302F">text</mods:typeOfResource>
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<mods:title id="7CD48284EE5CC23F87BA8095503394F5">Records of the Australian Museum</mods:title>
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<mods:part id="24BEC5C324C30E49EF6043AE743993BA">
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<mods:date id="E9651D89807CFD974ED38914104BE4B8">2003</mods:date>
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<mods:detail id="E54A785BF5FC9FBAD4CD20F2120A05AE" type="volume">
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<mods:number id="FB3471BB657CF0272CA9622DD333AB0E">55</mods:number>
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<mods:end id="4512C7FA3B43D37E9F521612F5C1656C">44</mods:end>
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<mods:classification id="41FEE3585F6C1487297B3548DA69697C">journal article</mods:classification>
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<mods:identifier id="FB3E5AE7CD4A9FB65B53082B23120FF8" type="ISSN">2201-4349</mods:identifier>
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<treatment id="038287C6FF81FFD7FED553AAFE03B79D" ID-DOI="http://doi.org/10.5281/zenodo.5239777" ID-Zenodo-Dep="5239777" LSID="urn:lsid:plazi:treatment:038287C6FF81FFD7FED553AAFE03B79D" httpUri="http://treatment.plazi.org/id/038287C6FF81FFD7FED553AAFE03B79D" lastPageNumber="19" pageId="18" pageNumber="19">
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<subSubSection id="C331655BFF81FFD7FED553AAFDD3B0F5" box="[332,628,152,176]" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="8B9436D0FF81FFD7FED553AAFDD3B0F5" blockId="18.[332,628,152,176]" box="[332,628,152,176]" pageId="18" pageNumber="19">
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<heading id="D0DC81BCFF81FFD7FED553AAFDD3B0F5" box="[332,628,152,176]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
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<emphasis id="B95FEAC2FF81FFD7FED553AAFDD3B0F5" bold="true" box="[332,628,152,176]" pageId="18" pageNumber="19">
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<taxonomicName id="4C2B4D53FF81FFD7FED553AAFD92B0F5" ID-CoL="SNDB" authority="Bradley J Sinclair, 2003" authorityName="Bradley J Sinclair" authorityYear="2003" box="[332,565,152,176]" class="Insecta" family="Brachystomatidae" genus="Ceratomerus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bickeli" status="sp. nov.">
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<emphasis id="B95FEAC2FF81FFD7FED553AAFD92B0F5" bold="true" box="[332,565,152,176]" italics="true" pageId="18" pageNumber="19">Ceratomerus bickeli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26C57B9FF81FFD7FDA553AAFDD3B0F5" box="[572,628,152,176]" pageId="18" pageNumber="19" rank="species">n.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C331655BFF81FFD7FE0F53F7FD8DB099" box="[406,554,197,220]" pageId="18" pageNumber="19" type="description">
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<paragraph id="8B9436D0FF81FFD7FE0F53F7FD8DB099" blockId="18.[406,554,197,220]" box="[406,554,197,220]" pageId="18" pageNumber="19">
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<figureCitation id="13102A55FF81FFD7FE0F53F7FE53B099" box="[406,500,197,220]" captionStart="Figs" captionStartId="19.[121,166,1887,1908]" captionTargetBox="[155,1360,146,1872]" captionTargetId="figure-10@19.[151,1361,146,1874]" captionTargetPageId="19" captionText="Figs. 44–50. Terminalia of Ceratomerus. 44, C. bickeli, male, lateral view. 45–46, C. bulbosus: 45, male, dorsal view; 46, male, lateral view. 47–50, C. globosus: 47, hypandrium and phallus, lateral view; 48, male, lateral view; 49, female, lateral view; 50, spermatheca. Scale bars = 0.1 mm. Abbreviations: cerc, cercus; gcx apod, gonocoxal apodeme; ej apod, ejaculatory apodeme; epand, epandrium; epand lb, epandrial lobe; hypd, hypandrium; hypd proc, hypandrial process; pgt, postgonite; ph, phallus; S, sternite; sur, surstylus." pageId="18" pageNumber="19">Figs. 44</figureCitation>
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,
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<figureCitation id="13102A55FF81FFD7FD9853F7FD8DB099" box="[513,554,197,220]" captionStart="Figs" captionStartId="34.[265,310,741,762]" captionTargetBox="[234,1399,151,719]" captionTargetId="figure-661@34.[227,1399,151,719]" captionTargetPageId="34" captionText="Figs. 100–101. Known distribution of species of Ceratomerus (Tasmania). 100, Ceratomerus bickeli *, C. bulbosus -, C. hibernus •. 101, Ceratomerus ordinatus." pageId="18" pageNumber="19">100</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C331655BFF81FFD7FF3F53C3FCB5B170" pageId="18" pageNumber="19" type="materials_examined">
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<paragraph id="8B9436D0FF81FFD7FF3F53C3FCB5B170" blockId="18.[166,786,241,309]" pageId="18" pageNumber="19">
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<emphasis id="B95FEAC2FF81FFD7FF3F53C3FE81B141" bold="true" box="[166,294,241,260]" pageId="18" pageNumber="19">Type material</emphasis>
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.
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<materialsCitation id="3B433C8DFF81FFD7FEA853C0FCB5B170" collectionCode="AMS" collectorName="D. Bickel" country="Australia" location="Mt. Wedge" pageId="18" pageNumber="19" specimenCode="K175243" specimenCount="2" typeStatus="holotype">
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<typeStatus id="54908872FF81FFD7FEA853C0FE37B146" box="[305,400,242,261]" pageId="18" pageNumber="19" type="holotype">HOLOTYPE</typeStatus>
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3, “TAS[mania,
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<collectingCountry id="F33C7640FF81FFD7FD8753C0FDD4B140" box="[542,627,242,261]" name="Australia" pageId="18" pageNumber="19">Australia</collectingCountry>
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]: Frodshams Pass/ nr.
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<location id="8EF4600BFF81FFD7FF595238FE95B158" LSID="urn:lsid:plazi:treatment:038287C6FF81FFD7FED553AAFE03B79D:8EF4600BFF81FFD7FF595238FE95B158" box="[192,306,266,285]" country="Australia" name="Mt. Wedge" pageId="18" pageNumber="19">Mt. Wedge</location>
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;4–5.ii./ -1989; rainforest/
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<collectorName id="26DE5306FF81FFD7FDA15238FD2AB158" box="[568,653,266,285]" pageId="18" pageNumber="19">D.Bickel</collectorName>
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;yellowpan”; “
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<typeStatus id="54908872FF81FFD7FF375210FEB0B170" box="[174,279,290,309]" pageId="18" pageNumber="19" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="4C2B4D53FF81FFD7FEB25210FE3CB170" authorityName="Philippi" authorityYear="1865" box="[299,411,290,309]" class="Insecta" family="Brachystomatidae" genus="Ceratomerus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B95FEAC2FF81FFD7FEB25210FE3CB170" box="[299,411,290,309]" italics="true" pageId="18" pageNumber="19">Ceratomerus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C2B4D53FF81FFD7FE3C5210FE79B170" authorityName="Bradley J Sinclair" authorityYear="2003" box="[421,478,290,309]" class="Insecta" family="Brachystomatidae" genus="Ceratomerus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bickeli">
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<emphasis id="B95FEAC2FF81FFD7FE3C5210FE79B170" box="[421,478,290,309]" italics="true" pageId="18" pageNumber="19">bickeli</emphasis>
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</taxonomicName>
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/ Sinclair [red label]”
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<collectionCode id="ED3AAE15FF81FFD7FD105210FD10B170" box="[649,695,290,309]" pageId="18" pageNumber="19">AMS</collectionCode>
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<specimenCode id="DB8D9EABFF81FFD7FD225210FCA9B170" box="[699,782,290,309]" collectionCode="AMS" pageId="18" pageNumber="19">K175243</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C331655BFF81FFD7FF3F527AFE5EB1DF" pageId="18" pageNumber="19" type="diagnosis">
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<paragraph id="8B9436D0FF81FFD7FF3F527AFE5EB1DF" blockId="18.[166,794,328,410]" pageId="18" pageNumber="19">
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<emphasis id="B95FEAC2FF81FFD7FF3F527AFEBFB125" bold="true" box="[166,280,328,352]" pageId="18" pageNumber="19">Diagnosis</emphasis>
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. Recognized by the strong inflexion of R
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<subScript id="17AF3495FF81FFD7FD665267FCBDB121" attach="left" box="[767,794,341,356]" fontSize="6" pageId="18" pageNumber="19">2+3</subScript>
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around the stigma and the very long, slender posterior hypandrial processes and cerci.
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</paragraph>
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</subSubSection>
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<subSubSection id="C331655BFF81FFD7FF3F529CFEDDB701" pageId="18" pageNumber="19" type="description">
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<paragraph id="8B9436D0FF81FFD7FF3F529CFECDB1A7" blockId="18.[166,793,430,482]" pageId="18" pageNumber="19">
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<emphasis id="B95FEAC2FF81FFD7FF3F529CFE8CB183" bold="true" box="[166,299,430,454]" pageId="18" pageNumber="19">Description</emphasis>
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. Wing length 3.7 mm, similar to
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<taxonomicName id="4C2B4D53FF81FFD7FD12529CFCBEB180" authorityName="Bradley J Sinclair" authorityYear="2003" box="[651,793,430,453]" class="Insecta" family="Brachystomatidae" genus="Ceratomerus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="attenuatus">
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<emphasis id="B95FEAC2FF81FFD7FD12529CFCBEB180" box="[651,793,430,453]" italics="true" pageId="18" pageNumber="19">C. attenuatus</emphasis>
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</taxonomicName>
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except as follows:
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</paragraph>
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<paragraph id="8B9436D0FF81FFD7FF3F52C4FE39B2F8" blockId="18.[166,794,502,1816]" pageId="18" pageNumber="19">
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<emphasis id="B95FEAC2FF81FFD7FF3F52C4FF47B24B" bold="true" box="[166,224,502,526]" pageId="18" pageNumber="19">Male</emphasis>
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.
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<emphasis id="B95FEAC2FF81FFD7FF7552C5FE82B24B" box="[236,293,503,526]" italics="true" pageId="18" pageNumber="19">Head</emphasis>
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. Four pairs of vertical setae, shorter than ocellar bristles; postocular bristles short, overlapping less than onethird of eye. Antenna with length of scape slightly less than height of eye; postpedicel length less than height of head; three-segmented stylus longer than postpedicel; segment 8 elongate, 4× as long as wide. Palpus brown, slender, onesixth length of labrum.
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</paragraph>
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<paragraph id="8B9436D0FF81FFD7FF5D51F6FE55B2BD" blockId="18.[166,794,502,1816]" pageId="18" pageNumber="19">
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<emphasis id="B95FEAC2FF81FFD7FF5D51F6FEB5B29E" box="[196,274,708,731]" italics="true" pageId="18" pageNumber="19">Thorax</emphasis>
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. Upper margin of pleura brown, remaining pale brown; postpronotal lobe pale.
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</paragraph>
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<paragraph id="8B9436D0FF81FFD7FF5D51CDFE15B376" blockId="18.[166,794,502,1816]" pageId="18" pageNumber="19">
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<emphasis id="B95FEAC2FF81FFD7FF5D51CDFF5CB353" box="[196,251,767,790]" italics="true" pageId="18" pageNumber="19">Wing</emphasis>
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. R
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<subScript id="17AF3495FF81FFD7FE855038FE91B35C" attach="left" box="[284,310,778,793]" fontSize="6" pageId="18" pageNumber="19">2+3</subScript>
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running parallel and close to costa, strongly deflected around stigma.
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</paragraph>
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<paragraph id="8B9436D0FF81FFD7FF5D500BFE0EB3A6" blockId="18.[166,794,502,1816]" pageId="18" pageNumber="19">
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<emphasis id="B95FEAC2FF81FFD7FF5D500BFF50B315" box="[196,247,825,848]" italics="true" pageId="18" pageNumber="19">Legs</emphasis>
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. Fore and mid coxae and ventral of femora pale yellow, remaining leg segments brown, darker towards tip. Fore femur slightly swollen, with av row of fine setae, decreasing in length apically; basal setae nearly subequal in length to width of femur. First tarsomere greater than half length of fore tibia.
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</paragraph>
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<paragraph id="8B9436D0FF81FFD7FF5D50DBFEE6B4AE" blockId="18.[166,794,502,1816]" pageId="18" pageNumber="19">
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Mid femur more strongly swollen than fore femur, with pv row of short, dark setae and 1–2 long basal setae; basal half with av brush of short dark setae, increasing in length medially. Mid tibia very slightly swollen at mid-length, with av row of short setae; pv surface with row of slender setae, longer than width of tibia; ventroapically with row of short erect setae on apical sixth; 1 ad seta on proximal half. First tarsomere
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<emphasis id="B95FEAC2FF81FFD7FE815784FE8BB48E" bold="true" box="[280,300,1206,1227]" pageId="18" pageNumber="19">
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<superScript id="7C5E9B98FF81FFD7FE815784FE87B481" attach="left" box="[280,288,1206,1220]" fontSize="6" pageId="18" pageNumber="19">4</superScript>
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<subScript id="17AF3495FF81FFD7FEBD578FFE8BB48E" attach="right" box="[292,300,1213,1227]" fontSize="6" pageId="18" pageNumber="19">5</subScript>
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</emphasis>
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length of remaining 4 tarsomeres, with many erect pv setae.
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</paragraph>
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<paragraph id="8B9436D0FF81FFD7FF5D57C3FEF8B5FD" blockId="18.[166,794,502,1816]" pageId="18" pageNumber="19">Hind femur straight with 1 preapical dorsal and 1 ad seta; ventral surface with av row of fine setae, shorter than width of femur. Hind tibia straight, shorter than femur, apex partially dilated, bearing posteroapical comb; 2 erect ad setae at mid-length and 1 subapical ad seta; apical two-thirds with row of long, erect dorsal setae. Hind tarsomeres slightly longer than tibia.</paragraph>
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<paragraph id="8B9436D0FF81FFD7FF5D568DFD9AB5B6" blockId="18.[166,794,502,1816]" pageId="18" pageNumber="19">
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<emphasis id="B95FEAC2FF81FFD7FF5D568DFE8EB593" box="[196,297,1471,1494]" italics="true" pageId="18" pageNumber="19">Abdomen</emphasis>
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pale brown, concolorous with thorax, sclerites weakly sclerotized; T8 not examined.
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</paragraph>
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<paragraph id="8B9436D0FF81FFD7FF5D56CBFEDDB75D" blockId="18.[166,794,502,1816]" pageId="18" pageNumber="19">
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<taxonomicName id="4C2B4D53FF81FFD7FF5D56CBFE9EB655" box="[196,313,1529,1552]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="genus">
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<emphasis id="B95FEAC2FF81FFD7FF5D56CBFE9EB655" box="[196,313,1529,1552]" italics="true" pageId="18" pageNumber="19">Terminalia</emphasis>
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</taxonomicName>
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(based on undissected specimen) (
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<figureCitation id="13102A55FF81FFD7FD2256CBFCA9B655" box="[699,782,1529,1552]" captionStart="Figs" captionStartId="19.[121,166,1887,1908]" captionTargetBox="[155,1360,146,1872]" captionTargetId="figure-10@19.[151,1361,146,1874]" captionTargetPageId="19" captionText="Figs. 44–50. Terminalia of Ceratomerus. 44, C. bickeli, male, lateral view. 45–46, C. bulbosus: 45, male, dorsal view; 46, male, lateral view. 47–50, C. globosus: 47, hypandrium and phallus, lateral view; 48, male, lateral view; 49, female, lateral view; 50, spermatheca. Scale bars = 0.1 mm. Abbreviations: cerc, cercus; gcx apod, gonocoxal apodeme; ej apod, ejaculatory apodeme; epand, epandrium; epand lb, epandrial lobe; hypd, hypandrium; hypd proc, hypandrial process; pgt, postgonite; ph, phallus; S, sternite; sur, surstylus." pageId="18" pageNumber="19">Fig. 44</figureCitation>
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). Hypandrium with pair of long, slender posterior processes, partially twisted; 1 pair of short anterior, hooked, articulated processes arched posteriorly near surstyli; postgonites arched anteriorly, flanking pair of phallic processes. Epandrial lamella greatly inflated laterally, round; fused ventrally to lateral wall of hypandrium. Surstylus fingershaped, shorter than cercus. Cercus with broad base, with long, heavily sclerotized lateral process, similar to posterior hypandrial process.
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</paragraph>
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<paragraph id="8B9436D0FF81FFD7FF3F541FFEDDB701" blockId="18.[166,378,1837,1861]" box="[166,378,1837,1861]" pageId="18" pageNumber="19">
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<emphasis id="B95FEAC2FF81FFD7FF3F541FFF5DB700" bold="true" box="[166,250,1837,1861]" pageId="18" pageNumber="19">Female</emphasis>
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. Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C331655BFF81FFD7FF3F546BFCBDB734" box="[166,794,1881,1905]" pageId="18" pageNumber="19" type="distribution">
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<paragraph id="8B9436D0FF81FFD7FF3F546BFCBDB734" blockId="18.[166,794,1881,1905]" box="[166,794,1881,1905]" pageId="18" pageNumber="19">
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||
<emphasis id="B95FEAC2FF81FFD7FF3F546BFE8BB734" bold="true" box="[166,300,1881,1905]" pageId="18" pageNumber="19">Distribution</emphasis>
|
||
. This species is restricted to
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<collectingRegion id="49EFF832FF81FFD7FDD15468FD0DB734" box="[584,682,1882,1905]" country="Australia" name="Tasmania" pageId="18" pageNumber="19">Tasmania</collectingRegion>
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(
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<figureCitation id="13102A55FF81FFD7FD2F5468FCA9B734" box="[694,782,1882,1905]" captionStart="Figs" captionStartId="34.[265,310,741,762]" captionTargetBox="[234,1399,151,719]" captionTargetId="figure-661@34.[227,1399,151,719]" captionTargetPageId="34" captionText="Figs. 100–101. Known distribution of species of Ceratomerus (Tasmania). 100, Ceratomerus bickeli *, C. bulbosus -, C. hibernus •. 101, Ceratomerus ordinatus." pageId="18" pageNumber="19">Fig. 100</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C331655BFF81FFD7FF3F54B4FE03B79D" pageId="18" pageNumber="19" type="etymology">
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<paragraph id="8B9436D0FF81FFD7FF3F54B4FE03B79D" blockId="18.[166,794,1926,2008]" pageId="18" pageNumber="19">
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<emphasis id="B95FEAC2FF81FFD7FF3F54B4FE87B7DB" bold="true" box="[166,288,1926,1950]" pageId="18" pageNumber="19">Etymology</emphasis>
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. The specific name honours Dr D.J. Bickel, who collected the
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<typeStatus id="54908872FF81FFD7FEA15491FE31B7FF" box="[312,406,1955,1978]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
|
||
of this species and many other rare species from
|
||
<collectingRegion id="49EFF832FF81FFD7FEAF54F3FE07B79D" box="[310,416,1985,2008]" country="Australia" name="Tasmania" pageId="18" pageNumber="19">Tasmania</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |