176 lines
18 KiB
XML
176 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1171.104142" ID-Pensoft-Pub="1313-2970-1171-1" ID-Pensoft-UUID="7559C2D42DE85144AEFFC98A16172F97" ID-ZooBank="267D0D4559CA4A18A08034768E652607" ModsDocID="1313-2970-1171-1" checkinTime="1689969829564" checkinUser="pensoft" docAuthor="Short, Andrew Edward Z. & Giron, Jennifer C." docDate="2023" docId="8BAAC0443CFD550BAFE4AA4E9F00AB80" docLanguage="en" docName="ZooKeys 1171: 1-112" docOrigin="ZooKeys 1171" docPubDate="2023-07-20" docSource="http://dx.doi.org/10.3897/zookeys.1171.104142" docTitle="Novochares spangleri Short & Girón 2023, sp. nov." docType="treatment" docUuid="59377017-2936-4FA3-9B74-154A5D09E5E5" docUuidSource="ZooBank" docVersion="1" id="7559C2D42DE85144AEFFC98A16172F97" lastPageNumber="1" masterDocId="7559C2D42DE85144AEFFC98A16172F97" masterDocTitle="Revision of the Neotropical water scavenger beetle genus Novochares Giron & Short (Coleoptera, Hydrophilidae, Acidocerinae)" masterLastPageNumber="112" masterPageNumber="1" pageNumber="1" updateTime="1689969829564" updateUser="pensoft">
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<mods:titleInfo id="98A65203632ACC967AB0057A15D40FC9">
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<mods:title id="91641D05E20A6F8A40BD6802CB85178E">Revision of the Neotropical water scavenger beetle genus Novochares Giron & Short (Coleoptera, Hydrophilidae, Acidocerinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="47C224E739764A1115C16D8B15A7D5C4" type="personal">
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<mods:role id="5FE643E2661CE0CC040AEBEDDF2489F5">
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<mods:roleTerm id="A6CE491D648E84BCB8D4DDF7A88BCBBC">Author</mods:roleTerm>
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<mods:namePart id="F31F5E426AF703F4A6126D223D8B69FF">Short, Andrew Edward Z.</mods:namePart>
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<mods:nameIdentifier id="DAFDB31FE5F59D78AE0F6A8C57AA5652" type="ORCID">https://orcid.org/0000-0002-7467-7116</mods:nameIdentifier>
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<mods:affiliation id="08576D7FF911C83F0EEDB59B3C56F989">Department of Entomology & Nematology, University of Florida, Gainesville, FL, 32611, USA</mods:affiliation>
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<mods:nameIdentifier id="F853BD871899A98A3B58D4AB4D981ECB" type="email">aezshort@ku.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name id="1DBFE1518FC5AB00ECE352A556233AC3" type="personal">
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<mods:roleTerm id="F8D963A04ED7F7EDB804E7821A3BFF9E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="66E35AAF2D1DB42D3328890DD65D4175">Giron, Jennifer C.</mods:namePart>
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<mods:nameIdentifier id="D55295CDBF87F36EF160A3A5A116FF88" type="ORCID">https://orcid.org/0000-0002-0851-6883</mods:nameIdentifier>
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<mods:affiliation id="F579C696D5B05FBF257A73318741C65F">Natural Science Research Laboratory, Museum of Texas Tech University, Lubbock, TX 79409, USA</mods:affiliation>
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<mods:typeOfResource id="4156E0544A3477F0F2FF0D9D62AC0C6A">text</mods:typeOfResource>
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<mods:titleInfo id="EC4AB5D8A59FCF3EE589939F8BA0BFE5">
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<mods:title id="2CD324465FE3D60E0BE20C1356C23EF5">ZooKeys</mods:title>
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<mods:part id="ED10EA89CCB6D0DAB0A292B02E3C39D7">
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<mods:date id="ABEA7925C18544C569F0F3CE6CF4682D">2023</mods:date>
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<mods:number id="77A5DBE260176B16F465E2EF6A2D4C7D">2023-07-20</mods:number>
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<mods:number id="E685A06D5FA2DE8122EB4F6D32BC5987">1171</mods:number>
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<mods:url id="212F422E5639F33683F4708DDA7A361A">http://dx.doi.org/10.3897/zookeys.1171.104142</mods:url>
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<mods:classification id="1BA07EBC88F35E6BC66B215475E8818A">journal article</mods:classification>
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<mods:identifier id="D2511A19AF06BFC7A69FE5A96AE833EF" type="DOI">http://dx.doi.org/10.3897/zookeys.1171.104142</mods:identifier>
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<mods:identifier id="0724312FE8C6941FA274101648F94C71" type="Pensoft-Pub">1313-2970-1171-1</mods:identifier>
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<treatment id="8BAAC0443CFD550BAFE4AA4E9F00AB80" LSID="urn:lsid:zoobank.org:act:59377017-2936-4FA3-9B74-154A5D09E5E5" httpUri="http://treatment.plazi.org/id/8BAAC0443CFD550BAFE4AA4E9F00AB80" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="4BA45046E9B0CC6DACE1722BCB6CEE12" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="A83F6122BBDF49AE051087BF2B29126C" pageId="0" pageNumber="1">
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<taxonomicName id="5C2CD24EB4B17B993185050055A363AA" LSID="https://zoobank.org/59377017-2936-4FA3-9B74-154A5D09E5E5" authority="Short & Girón, 2023" authorityName="Short & Girón" authorityYear="2023" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares spangleri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="spangleri" status="sp. nov.">Novochares spangleri</taxonomicName>
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<taxonomicNameLabel id="240DEA68D16069496B47AF7746A437DB" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="F5660866DEE7DE381005DEC895B4D885" pageId="0" pageNumber="1" type="description">
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<paragraph id="4962A3D7EFB447AFF1DDF4A5AAFF11A3" pageId="0" pageNumber="1">
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<figureCitation id="EF3FC5F1B436ACDDC1D2F87C95A41C4C" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Aedeagi of the Novochares punctatostriatus species group A-D N. dentatus E, F N. spangleri G, H N. geminus I N. triangularis J, K N. pertusus L-Q N. punctatostriatus A-C Venezuela D Ecuador L-N Suriname O Peru P, Q Brazil. A, E, G, I, J, L, O, P dorsal view H, M, Q ventral view C, F, N oblique view K lateral view." figureDoi="10.3897/zookeys.1171.104142.figure15" httpUri="https://binary.pensoft.net/fig/881182" pageId="0" pageNumber="1">Figs 15E-F</figureCitation>
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<figureCitation id="B6885D92A5664CF9EA4B3275999A3DDC" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of Novochares punctatostriatus species group A N. punctatostriatus (red), N. pertusus (yellow) B N. dentatus (red), N. triangularis (yellow), N. spangleri (blue), N. geminus (green)." figureDoi="10.3897/zookeys.1171.104142.figure16" httpUri="https://binary.pensoft.net/fig/881183" pageId="0" pageNumber="1">, 16B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="74F47ED8E93D1153CA50972CD8E59EEA" pageId="0" pageNumber="1" type="type material">
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<paragraph id="1E60FEC1AEBA3FA57CB521849D954988" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="6C4CC1AE6331312DC07605D10531B13A" pageId="0" pageNumber="1">
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<emphasis id="25BFBF2F57E1B18BEC887B734723617B" bold="true" pageId="0" pageNumber="1">
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<emphasis id="FB20F848294223CEF46396CD9B967DB9" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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(male)
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</emphasis>
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: "PERU: Cusco: Paucartambo/ -
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<geoCoordinate id="67A2CD0F0660FF3EF9D95142A3103DC2" degrees="12.91411" direction="south" orientation="latitude" precision="1" value="-12.91411">12.91411S</geoCoordinate>
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, -
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<geoCoordinate id="186D2BED3CD9036D39D6BA990C750E49" degrees="71.37492" direction="west" orientation="longitude" precision="1" value="-71.37492">71.37492W</geoCoordinate>
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, 585m/ c. 3km NE of Pilcopata; 30.v.2022/ grassy pools/ditch along road/ PE22-0530-01C; leg. Short et al." (MHNSM).
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<emphasis id="05A305D171CD1C42FDC55AE4F46C9209" bold="true" pageId="0" pageNumber="1">
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<emphasis id="FF4D67EC5DB0E40C5EFD77EE84BBDDEA" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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(6 exs.): Peru: Cusco
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</emphasis>
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: same data as holotype (1, SEMC); same data except mountain side pools and ditches, PE22-0530-01B (2, SEMC).
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<emphasis id="F0E15F181E0687E63B0D63FF8680233B" bold="true" pageId="0" pageNumber="1">Madre de Dios</emphasis>
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: Villa Carmen Biological Station (ca. 2 km N of Pilcopata), North of Rio
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<normalizedToken id="C4CA606289856D0663C1C300909D4FC6" originalValue="Piñipiñi">Pinipini</normalizedToken>
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, 26.v.2022, leg. Short et al., small muddy pools in landslide, PE22-0526-02B (1, SEMC); same data except 28.v.2022, detrital pools formed by seeps, PE22-0526-02F (2, MHNSM, SEMC).
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</paragraph>
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</subSubSection>
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<subSubSection id="59B225D82F96D74342E186C021A73C1F" pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph id="35081822BB9D6DFBC9BA8D819F813D3E" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph id="2FE27FA6A84EAE4A928143D653FAFD2F" pageId="0" pageNumber="1">
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Among members of the
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<taxonomicName id="AECC9DF2FFA0A08A47555E8B833E0041" authorityName="Short & Girón" authorityYear="2023" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares punctatostriatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="punctatostriatus">
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<emphasis id="5C88C0087150111664B333114295EE77" italics="true" pageId="0" pageNumber="1">Novochares punctatostriatus</emphasis>
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</taxonomicName>
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species group,
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<taxonomicName id="D45DD23383A1BA8FEA60A5447903E9A8" lsidName="N. spangleri" pageId="0" pageNumber="1" rank="species" species="spangleri">
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<emphasis id="F73C53031D879F4BE70638EBE9FD0ECB" italics="true" pageId="0" pageNumber="1">N. spangleri</emphasis>
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</taxonomicName>
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is one of three species that lack distinct rows of elytral serial punctures, the others being
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<taxonomicName id="FB6DEC2FF96D9402FF9D97C92EEBD6E1" lsidName="N. pertusus" pageId="0" pageNumber="1" rank="species" species="pertusus">
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<emphasis id="88D972FA9E15F804E67F0E06951C8D2F" italics="true" pageId="0" pageNumber="1">N. pertusus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="5A0D9F2B9F2B6FB63CD0699DA6BCB54F" lsidName="N. triangularis" pageId="0" pageNumber="1" rank="species" species="triangularis">
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<emphasis id="38C6E6AC9CD7D8747ADA725634A01E75" italics="true" pageId="0" pageNumber="1">N. triangularis</emphasis>
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</taxonomicName>
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. It is most similar to
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<taxonomicName id="1276EA2D8100B20738A0D617765802A7" lsidName="N. triangularis" pageId="0" pageNumber="1" rank="species" species="triangularis">
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<emphasis id="53D0FA819C84C0A5B88CF24438863E8D" italics="true" pageId="0" pageNumber="1">N. triangularis</emphasis>
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</taxonomicName>
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, to which the aedeagal form is very similar, though it can be distinguished by the apices of the forked projection of the dorsal plate of the median lobe being slightly more swollen, with the outer margins of the parameres weakly curved (apices of the forked projection of the dorsal plate of the median lobe being not as swollen and the outer margins of the parameres strongly curved in
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<taxonomicName id="AF1F4B1063B887394E50329573093777" lsidName="N. triangularis" pageId="0" pageNumber="1" rank="species" species="triangularis">
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<emphasis id="0706EB55201AF79BFF44EC9A842A14C4" italics="true" pageId="0" pageNumber="1">N. triangularis</emphasis>
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</taxonomicName>
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; compare Fig.
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<figureCitation id="8F92224FCF4148710D5C79792F4DA109" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Aedeagi of the Novochares punctatostriatus species group A-D N. dentatus E, F N. spangleri G, H N. geminus I N. triangularis J, K N. pertusus L-Q N. punctatostriatus A-C Venezuela D Ecuador L-N Suriname O Peru P, Q Brazil. A, E, G, I, J, L, O, P dorsal view H, M, Q ventral view C, F, N oblique view K lateral view." figureDoi="10.3897/zookeys.1171.104142.figure15" httpUri="https://binary.pensoft.net/fig/881182" pageId="0" pageNumber="1">15E</figureCitation>
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vs. Fig.
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<figureCitation id="B3D6E2934C6E62DBE67B81B2C9C89AA8" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Aedeagi of the Novochares punctatostriatus species group A-D N. dentatus E, F N. spangleri G, H N. geminus I N. triangularis J, K N. pertusus L-Q N. punctatostriatus A-C Venezuela D Ecuador L-N Suriname O Peru P, Q Brazil. A, E, G, I, J, L, O, P dorsal view H, M, Q ventral view C, F, N oblique view K lateral view." figureDoi="10.3897/zookeys.1171.104142.figure15" httpUri="https://binary.pensoft.net/fig/881182" pageId="0" pageNumber="1">15I</figureCitation>
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). The elytral ground punctation is extremely fine and almost not noticeable in
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<taxonomicName id="DB99B1A5D7969C4FD512306D98D6A006" lsidName="N. spangleri" pageId="0" pageNumber="1" rank="species" species="spangleri">
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<emphasis id="44973301DAF1A94104593CBAD4BEC352" italics="true" pageId="0" pageNumber="1">N. spangleri</emphasis>
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</taxonomicName>
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, while it is distinctly coarser in
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<taxonomicName id="F6CE085EC351FE8370829297EF89A63B" lsidName="N. triangularis" pageId="0" pageNumber="1" rank="species" species="triangularis">
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<emphasis id="CB655997E66F9A5A42A80ED8692601C9" italics="true" pageId="0" pageNumber="1">N. triangularis</emphasis>
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</taxonomicName>
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. Additionally,
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<taxonomicName id="AFB95D1E19B09F800E698E39F8D26DD4" lsidName="N. triangularis" pageId="0" pageNumber="1" rank="species" species="triangularis">
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<emphasis id="E828C0EA948BD3172F707FFD5DCCC11B" italics="true" pageId="0" pageNumber="1">N. triangularis</emphasis>
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</taxonomicName>
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has only very narrow paler margins of the pronotum, these margins are much more broadly pale in
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<taxonomicName id="E7BCA1444EF70F6F73F62CD0008B3883" lsidName="N. spangleri" pageId="0" pageNumber="1" rank="species" species="spangleri">
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<emphasis id="B16BC0A7C7C7685049045F1678B434C0" italics="true" pageId="0" pageNumber="1">N. spangleri</emphasis>
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</taxonomicName>
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. See diagnosis of
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<taxonomicName id="126CD9C919184860A4D64FD4446D2979" lsidName="N. pertusus" pageId="0" pageNumber="1" rank="species" species="pertusus">
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<emphasis id="15084C524E0F5F6D51EF6C0509D583DD" italics="true" pageId="0" pageNumber="1">N. pertusus</emphasis>
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</taxonomicName>
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for separation from that species.
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</paragraph>
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</subSubSection>
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<subSubSection id="BB839C9EA4A55510CFE80414C3A3F801" pageId="0" pageNumber="1" type="description">
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<paragraph id="6A21E0CC06DEC7268768B96893D5CA6D" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="D17F0A52A6CDCBFBADC3AF0CFFA6C036" pageId="0" pageNumber="1">
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Body length 5.6-6.0 mm.
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<emphasis id="A443B14FE64770B17AA3CA82D3C78043" bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
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: Dorsal surface of head, labrum, pronotum, and elytra dark brown, gradually paler towards margins.
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<emphasis id="79995825F415BD3D2DFE56129C96399C" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
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: Maxillary palps slightly shorter than width of head, uniformly yellow in color.
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<emphasis id="AAAD5434423A9E1F63D568AAF4D79D5C" bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
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: Ground punctation on pronotum and elytra relatively dense and very shallowly impressed. Elytra without rows of serial punctures, except for rows of very weak serial punctures along lateral regions of each. Prosternum medially convex. Posterior elevation of mesoventrite transverse, blunt, and low.
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<emphasis id="7A8C4CFDDAB5DBAE4511476564B980CC" bold="true" italics="true" pageId="0" pageNumber="1">Aedeagus</emphasis>
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: (Fig.
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<figureCitation id="5E5BAE6AD22EE4FB3DA0B3CA1143A392" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Aedeagi of the Novochares punctatostriatus species group A-D N. dentatus E, F N. spangleri G, H N. geminus I N. triangularis J, K N. pertusus L-Q N. punctatostriatus A-C Venezuela D Ecuador L-N Suriname O Peru P, Q Brazil. A, E, G, I, J, L, O, P dorsal view H, M, Q ventral view C, F, N oblique view K lateral view." figureDoi="10.3897/zookeys.1171.104142.figure15" httpUri="https://binary.pensoft.net/fig/881182" pageId="0" pageNumber="1">15E, F</figureCitation>
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) lateral projection on apical region of outer margin of each paramere rounded to weakly pointed; at closest point, dorsal inner margins of parameres separated by distance slightly greater than greatest width of a paramere; dorsal plate of median lobe with small denticle on each lateral margin, proximal to base of fork; arms of dorsal plate of median lobe distally weakly converging; each arm slightly wider at base and widened on apical region, apically oblique with inner margin extending beyond outer margin; notch between arms at base 1/2 as wide as base of an arm; ventral plate of median lobe moderately sclerotized, at widest point as wide as dorsal plate of median lobe, apically acuminate, apex extending beyond base of fork, not reaching apex of arms of dorsal plate; basal piece 0.4
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<normalizedToken id="E80AC47D5AF47BE70AE2B8D6F64246FA" originalValue="×">x</normalizedToken>
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length of a paramere.
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</paragraph>
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</subSubSection>
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<subSubSection id="37A22D81730B0EBEA79FAE67D24DE871" pageId="0" pageNumber="1" type="etymology">
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<paragraph id="09490C59810D36EA267F2FB77151954D" pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph id="F34BB465FEF97F02B137FD585554BDCA" pageId="0" pageNumber="1">Named after Paul J. Spangler, longtime curator at the US National Museum of Natural History, Smithsonian Institution, and specialist on aquatic beetles, who collected and sorted a lot of the specimens included in this contribution.</paragraph>
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</subSubSection>
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<subSubSection id="91D0C6ED67FEFBB04A5A4A4B713C75CC" pageId="0" pageNumber="1" type="distribution">
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<paragraph id="C14C0CE4C3202D07E6DC2F9F4BFCDD41" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="AD4FA8AC84EEC3B211C13ED41A3E80FA" pageId="0" pageNumber="1">
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Peru (Fig.
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<figureCitation id="EBB4545CABAE7CD44DD1CF94AE585688" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of Novochares punctatostriatus species group A N. punctatostriatus (red), N. pertusus (yellow) B N. dentatus (red), N. triangularis (yellow), N. spangleri (blue), N. geminus (green)." figureDoi="10.3897/zookeys.1171.104142.figure16" httpUri="https://binary.pensoft.net/fig/881183" pageId="0" pageNumber="1">16B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3EEE77521D7878CC63F06EC61286975D" pageId="0" pageNumber="1" type="habitat">
|
||
<paragraph id="5BA5A62B12D3133E61AEE11F58CD45BE" pageId="0" pageNumber="1">Habitat.</paragraph>
|
||
<paragraph id="B6E6B07143CEECF809297FAF8AFC3440" pageId="0" pageNumber="1">This species was collected primarily in forested riparian habitats.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |