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<document id="4FE5617DE17624DECB0F12465ED80D6A" ID-CLB-Dataset="2431" ID-DOI="10.11646/zootaxa.5093.1.1" ID-GBIF-Dataset="0faf68be-3777-4466-8cd5-9eb8ca72a239" ID-ISSN="1175-5326" ID-Zenodo-Dep="5901006" ID-ZooBank="C2F5E368-CB15-4207-9944-E52C190FBF20" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1643101690477" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2022" docId="03C03B6BFF8EFFE1FF7B29E4FAACF92D" docLanguage="en" docName="zootaxa.5093.1.1.pdf" docOrigin="Zootaxa 5093 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euconnus (Psomophus) debilis" docType="treatment" docVersion="9" lastPageNumber="24" masterDocId="FFF94313FF9BFFF6FFEC2B57FFE6FF93" masterDocTitle="Euconnus Thomson of Japan: redescriptions of species established by Reitter, Sharp and Franz, new synonyms, and summary of current state of knowledge (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="22" updateTime="1699292035325" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B58D58D5D421FDA7AAEC0C9899429AC8">Euconnus Thomson of Japan: redescriptions of species established by Reitter, Sharp and Franz, new synonyms, and summary of current state of knowledge (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<treatment id="03C03B6BFF8EFFE1FF7B29E4FAACF92D" ID-DOI="http://doi.org/10.5281/zenodo.5902688" ID-GBIF-Taxon="192597246" ID-Zenodo-Dep="5902688" LSID="urn:lsid:plazi:treatment:03C03B6BFF8EFFE1FF7B29E4FAACF92D" httpUri="http://treatment.plazi.org/id/03C03B6BFF8EFFE1FF7B29E4FAACF92D" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
<subSubSection id="C373D9F6FF8EFFE3FF7B29E4FD91FD5F" box="[151,631,690,717]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BD68A7DFF8EFFE3FF7B29E4FD91FD5F" blockId="21.[151,631,690,717]" box="[151,631,690,717]" pageId="21" pageNumber="22">
<heading id="D09E3D11FF8EFFE3FF7B29E4FD91FD5F" bold="true" box="[151,631,690,717]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<taxonomicName id="4C69F1FEFF8EFFE3FF7B29E4FD91FD5F" ID-CoL="9G256" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[151,631,690,717]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="debilis" subGenus="Psomophus">
<emphasis id="B91D566FFF8EFFE3FF7B29E4FD91FD5F" bold="true" box="[151,631,690,717]" pageId="21" pageNumber="22">
<emphasis id="B91D566FFF8EFFE3FF7B29E4FEF5FD5E" bold="true" box="[151,275,691,717]" italics="true" pageId="21" pageNumber="22">Euconnus</emphasis>
(
<emphasis id="B91D566FFF8EFFE3FEC829E5FE56FD5E" bold="true" box="[292,432,690,717]" italics="true" pageId="21" pageNumber="22">Psomophus</emphasis>
)
<emphasis id="B91D566FFF8EFFE3FE2D29E5FDE9FD5E" bold="true" box="[449,527,690,717]" italics="true" pageId="21" pageNumber="22">debilis</emphasis>
(Sharp)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373D9F6FF8EFFE3FF2B29A0FEC1FCE2" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="8BD68A7DFF8EFFE3FF2B29A0FBEFFC9C" blockId="21.[151,1033,759,881]" box="[199,1033,759,783]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB79AD53FF8EFFE3FF2B29A0FBEFFC9C" box="[199,1033,759,783]" pageId="21" pageNumber="22">
<treatmentCitation id="0AC8AC6CFF8EFFE3FF2B29A0FDA3FC9C" author="Sharp, D." box="[199,581,759,783]" page="127" pageId="21" pageNumber="22" year="1874">
<taxonomicName id="4C69F1FEFF8EFFE3FF2B29A0FDA3FC9C" authority="Sharp, 1874: 127" authorityName="Sharp" authorityPageNumber="127" authorityYear="1874" box="[199,581,759,783]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B91D566FFF8EFFE3FF2B29A0FE69FC9C" box="[199,399,759,783]" italics="true" pageId="21" pageNumber="22">Scydmaenus debilis</emphasis>
<bibRefCitation id="EFF8F78CFF8EFFE3FE7929A0FDA3FC9C" author="Sharp, D." box="[405,581,759,783]" pageId="21" pageNumber="22" pagination="105 - 130" refId="ref26336" refString="Sharp, D. (1874) The Pselaphidae and Scydmaenidae of Japan. Transactions of the Entomological Society of London, 1, 105 - 130. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00160. x" type="journal article" year="1874">Sharp, 1874: 127</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Aedeagus illustrated in
<treatmentCitation id="0AC8AC6CFF8EFFE3FCD229A0FBE2FC9C" author="Kurbatov, S." box="[830,1028,759,783]" page="29" pageId="21" pageNumber="22" year="2006">
<bibRefCitation id="EFF8F78CFF8EFFE3FCD229A0FBE2FC9C" author="Kurbatov, S." box="[830,1028,759,783]" pageId="21" pageNumber="22" pagination="25 - 31" refId="ref26039" refString="Kurbatov, S. (2006) Description de quelques Scydmenides et Pselaphines nouveaux russes (Coleoptera: Scydmaenidae et Staphylinidae, Pselaphinae). Russian Entomological Journal, 15 (1), 25 - 31." type="journal article" year="2006">Kurbatov, 2006: 29</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD68A7DFF8EFFE3FF2B2840FCC7FCBD" blockId="21.[151,1033,759,881]" box="[199,801,790,814]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB79AD53FF8EFFE3FF2B2840FCC7FCBD" box="[199,801,790,814]" pageId="21" pageNumber="22">
<taxonomicName id="4C69F1FEFF8EFFE3FF2B2840FD92FCBD" ID-CoL="9G256" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[199,628,790,814]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="debilis" subGenus="Microscydmus">
<emphasis id="B91D566FFF8EFFE3FF2B2840FECDFCBD" box="[199,299,791,814]" italics="true" pageId="21" pageNumber="22">Euconnus</emphasis>
(
<emphasis id="B91D566FFF8EFFE3FED62841FE2BFCBD" box="[314,461,790,814]" italics="true" pageId="21" pageNumber="22">Microscydmus</emphasis>
)
<emphasis id="B91D566FFF8EFFE3FE372841FDF9FCBD" box="[475,543,790,814]" italics="true" pageId="21" pageNumber="22">debilis</emphasis>
(Sharp)
</taxonomicName>
;
<treatmentCitation id="0AC8AC6CFF8EFFE3FD932841FCFBFCBD" author="Csiki, E." box="[639,797,790,814]" page="55" pageId="21" pageNumber="22" year="1919">
<bibRefCitation id="EFF8F78CFF8EFFE3FD932841FCFBFCBD" author="Csiki, E." box="[639,797,790,814]" pageId="21" pageNumber="22" pagination="106" refId="ref24424" refString="Csiki, E. (1919) Pars 70: Scydmaenidae. In: Schenkling, S. &amp; Junk, W. (Eds.), Coleopterorum Catalogus. Vol. 7. W. Junk, Berlin, pp. 106." type="book chapter" year="1919">Csiki, 1919: 55</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD68A7DFF8EFFE3FF2B2862FCCAFCDE" blockId="21.[151,1033,759,881]" box="[199,812,820,845]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB79AD53FF8EFFE3FF2B2862FCCAFCDE" box="[199,812,820,845]" pageId="21" pageNumber="22">
<taxonomicName id="4C69F1FEFF8EFFE3FF2B2862FDB0FCDF" ID-CoL="9G256" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[199,598,820,845]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="debilis" subGenus="Psomophus">
<emphasis id="B91D566FFF8EFFE3FF2B2862FECDFCDF" box="[199,299,821,844]" italics="true" pageId="21" pageNumber="22">Euconnus</emphasis>
(
<emphasis id="B91D566FFF8EFFE3FED62863FE49FCDF" box="[314,431,820,844]" italics="true" pageId="21" pageNumber="22">Psomophus</emphasis>
)
<emphasis id="B91D566FFF8EFFE3FE512863FDE7FCDF" box="[445,513,820,844]" italics="true" pageId="21" pageNumber="22">debilis</emphasis>
(Sharp)
</taxonomicName>
;
<treatmentCitation id="0AC8AC6CFF8EFFE3FD8D2863FCC1FCDE" author="Kurbatov, S." box="[609,807,820,845]" page="27" pageId="21" pageNumber="22" year="2006">
<bibRefCitation id="EFF8F78CFF8EFFE3FD8D2863FCC1FCDE" author="Kurbatov, S." box="[609,807,820,845]" pageId="21" pageNumber="22" pagination="25 - 31" refId="ref26039" refString="Kurbatov, S. (2006) Description de quelques Scydmenides et Pselaphines nouveaux russes (Coleoptera: Scydmaenidae et Staphylinidae, Pselaphinae). Russian Entomological Journal, 15 (1), 25 - 31." type="journal article" year="2006">Kurbatov, 2006: 27</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD68A7DFF8EFFE3FF7B2800FEC1FCE2" blockId="21.[151,1033,759,881]" box="[151,295,855,881]" pageId="21" pageNumber="22">
(
<figureCitation id="135296F8FF8EFFE3FF732800FEC6FCE2" box="[159,288,855,881]" captionStart="FIGURES 7278" captionStartId="22.[151,264,1857,1881]" captionTargetBox="[161,1421,381,1826]" captionTargetId="figure-101@22.[151,1436,381,1834]" captionTargetPageId="22" captionText="FIGURES 7278. Euconnus debilis, male. Dorsal habitus (72) and historical labels (73) of lectotype of Scydmaenus debilis; fore tibia and tarsus of male in lateral view (74); aedeagus in ventral (75, 77) and lateral (76, 78) views." figureDoi="http://doi.org/10.5281/zenodo.5901036" httpUri="https://zenodo.org/record/5901036/files/figure.png" pageId="21" pageNumber="22">Figs 7278</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C373D9F6FF8EFFE3FF2B28C8FE6CF966" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8BD68A7DFF8EFFE3FF2B28C8FBC8FB02" blockId="21.[151,1437,926,2033]" pageId="21" pageNumber="22">
<emphasis id="B91D566FFF8EFFE3FF2B28C8FE37FC2B" bold="true" box="[199,465,926,953]" pageId="21" pageNumber="22">Type material studied</emphasis>
.
<materialsCitation id="3B018020FF8EFFE3FE0E28C8FE3FFBFE" ID-GBIF-Occurrence="3454958311" collectionCode="NHM" collectorName="G. Lewis." country="Japan" determinerName="S. Kurbatov" location="Japan" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="B91D566FFF8EFFE3FE0E28C8FB72FC2B" bold="true" box="[482,1172,926,953]" pageId="21" pageNumber="22">
<typeStatus id="54D234DFFF8EFFE3FE0E28C8FDBEFC2A" box="[482,600,927,953]" pageId="21" pageNumber="22" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="4C69F1FEFF8EFFE3FD6F28C9FC5DFC2B" authority="Sharp" authorityName="Sharp" box="[643,955,926,953]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B91D566FFF8EFFE3FD6F28C9FC8EFC2A" bold="true" box="[643,872,926,953]" italics="true" pageId="21" pageNumber="22">Scydmaenus debilis</emphasis>
Sharp
</taxonomicName>
(here designated)
</emphasis>
:
<specimenCount id="9D6F41F4FF8EFFE3FB4A28C8FB5DFC2A" box="[1190,1211,927,953]" pageId="21" pageNumber="22" type="male"></specimenCount>
(
<figureCitation id="135296F8FF8EFFE3FB2128C8FAC5FC2A" box="[1229,1315,927,953]" captionStart="FIGURES 7278" captionStartId="22.[151,264,1857,1881]" captionTargetBox="[161,1421,381,1826]" captionTargetId="figure-101@22.[151,1436,381,1834]" captionTargetPageId="22" captionText="FIGURES 7278. Euconnus debilis, male. Dorsal habitus (72) and historical labels (73) of lectotype of Scydmaenus debilis; fore tibia and tarsus of male in lateral view (74); aedeagus in ventral (75, 77) and lateral (76, 78) views." figureDoi="http://doi.org/10.5281/zenodo.5901036" httpUri="https://zenodo.org/record/5901036/files/figure.png" pageId="21" pageNumber="22">Fig. 72</figureCitation>
), mounted on simple and not annotated card, with six labels (
<figureCitation id="135296F8FF8EFFE3FD362894FCD7FC4E" box="[730,817,963,989]" captionStart="FIGURES 7278" captionStartId="22.[151,264,1857,1881]" captionTargetBox="[161,1421,381,1826]" captionTargetId="figure-101@22.[151,1436,381,1834]" captionTargetPageId="22" captionText="FIGURES 7278. Euconnus debilis, male. Dorsal habitus (72) and historical labels (73) of lectotype of Scydmaenus debilis; fore tibia and tarsus of male in lateral view (74); aedeagus in ventral (75, 77) and lateral (76, 78) views." figureDoi="http://doi.org/10.5281/zenodo.5901036" httpUri="https://zenodo.org/record/5901036/files/figure.png" pageId="21" pageNumber="22">Fig. 73</figureCitation>
): “Sharp Coll. / 1905-313.” [darkened white, printed], “
<taxonomicName id="4C69F1FEFF8EFFE3FF4D28B0FED7FB92" authorityName="Latreille" authorityYear="1802" box="[161,305,999,1025]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ debilis / Type. D.S.” [white with black frame, handwritten and printed], circular label “Type / H.T.” [white with red margin, printed], circular label “SYN- / TYPE” [white with blue margin, printed], “
<collectingCountry id="F37ECAEDFF8EFFE3FAA52F5CFA6AFBB6" box="[1353,1420,1035,1061]" name="Japan" pageId="21" pageNumber="22">Japan</collectingCountry>
/
<collectorName id="269CEFABFF8EFFE3FF7B2F78FEEEFBDA" box="[151,264,1071,1097]" pageId="21" pageNumber="22">G. Lewis.</collectorName>
” [white with orange horizontal line, printed], and “
<taxonomicName id="4C69F1FEFF8EFFE3FCA72F78FC5AFBDA" box="[843,956,1071,1097]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">Euconnus</taxonomicName>
/
<taxonomicName id="4C69F1FEFF8EFFE3FC382F78FB9CFBD9" authority="(Sharp)" baseAuthorityName="Sharp" box="[980,1146,1071,1098]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="debilis">debilis (Sharp)</taxonomicName>
/ det.
<determinerName id="7E0A66B9FF8EFFE3FB2F2F78FAAAFBDA" box="[1219,1356,1071,1097]" pageId="21" pageNumber="22">S. Kurbatov</determinerName>
, 1994” [white, handwritten] (
<collectionCode id="ED7812B8FF8EFFE3FE662F04FE34FBFE" box="[394,466,1107,1133]" pageId="21" pageNumber="22">NHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B018020FF8EFFE3FE0F2F04FBCCFB02" ID-GBIF-Occurrence="3454958324" collectionCode="NHM" country="Japan" pageId="21" pageNumber="22" specimenCount="3" specimenCount-male="2" typeStatus="paralectotype">
<emphasis id="B91D566FFF8EFFE3FE0F2F04FD14FBFF" bold="true" box="[483,754,1106,1133]" pageId="21" pageNumber="22">
<typeStatus id="54D234DFFF8EFFE3FE0F2F04FD75FBFE" box="[483,659,1107,1133]" pageId="21" pageNumber="22" type="paralectotype">Paralectotypes</typeStatus>
(3 exx):
</emphasis>
<specimenCount id="9D6F41F4FF8EFFE3FD152F04FCD1FBFE" box="[761,823,1107,1133]" pageId="21" pageNumber="22" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="9D6F41F4FF8EFFE3FC812F04FC45FBFE" box="[877,931,1107,1133]" pageId="21" pageNumber="22" type="generic">1 ex.</specimenCount>
of unknown sex (with legs bent under body), same labels as for
<typeStatus id="54D234DFFF8EFFE3FE8A2F20FE2BFB02" box="[358,461,1143,1169]" pageId="21" pageNumber="22" type="lectotype">lectotype</typeStatus>
except without the circular red-margined one (
<collectionCode id="ED7812B8FF8EFFE3FC372F20FBC5FB02" box="[987,1059,1143,1169]" pageId="21" pageNumber="22">NHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD68A7DFF8EFFE3FF2B2FCDFA7AFAD6" blockId="21.[151,1437,926,2033]" pageId="21" pageNumber="22">
<emphasis id="B91D566FFF8EFFE3FF2B2FCDFD7AFB27" bold="true" box="[199,668,1178,1204]" pageId="21" pageNumber="22">Additional material studied. HONSHU</emphasis>
:
<materialsCitation id="3B018020FF8EFFE3FD402FCDFDBBFB4A" ID-GBIF-Occurrence="3454958305" collectingDate="2003-10-18" collectorName="S. Nomura" county="Numayama Moor" elevation="700" location="Aizu" municipality="Shimago-machi" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Fukushima">
<emphasis id="B91D566FFF8EFFE3FD402FCDFC90FB27" bold="true" box="[684,886,1178,1204]" pageId="21" pageNumber="22">
<collectingRegion id="49AD449FFF8EFFE3FD402FCDFC90FB27" box="[684,886,1178,1204]" country="Japan" name="Fukushima" pageId="21" pageNumber="22">Fukushima Pref.</collectingRegion>
</emphasis>
:
<specimenCount id="9D6F41F4FF8EFFE3FC6A2FCCFC7BFB26" box="[902,925,1179,1205]" pageId="21" pageNumber="22" type="male"></specimenCount>
,
<collectingCounty id="62B7F2F1FF8EFFE3FC462FCCFB97FB26" box="[938,1137,1179,1205]" pageId="21" pageNumber="22">Numayama Moor</collectingCounty>
(
<quantity id="4C912798FF8EFFE3FB6E2FCCFB29FB26" box="[1154,1231,1179,1205]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="21" pageNumber="22" unit="m" value="700.0">
<elevation id="00446D4EFF8EFFE3FB6E2FCCFB29FB26" box="[1154,1231,1179,1205]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="21" pageNumber="22" unit="m" value="700.0">700 m</elevation>
</quantity>
),
<collectingMunicipality id="6BB21007FF8EFFE3FB082FCCFA7EFB26" box="[1252,1432,1179,1205]" pageId="21" pageNumber="22">Shimagô-machi</collectingMunicipality>
,
<location id="8EB6DCA6FF8EFFE3FF7B2FE8FF36FB4A" LSID="urn:lsid:plazi:treatment:03C03B6BFF8EFFE1FF7B29E4FAACF92D:8EB6DCA6FF8EFFE3FF7B2FE8FF36FB4A" box="[151,208,1215,1241]" county="Numayama Moor" municipality="Shimago-machi" name="Aizu" pageId="21" pageNumber="22" stateProvince="Fukushima">Aizu</location>
,
<date id="FFD7ACBDFF8EFFE3FF312FE8FEB8FB4A" box="[221,350,1215,1241]" pageId="21" pageNumber="22" value="2003-10-18">
<collectingDate id="EF935555FF8EFFE3FF312FE8FEB8FB4A" box="[221,350,1215,1241]" pageId="21" pageNumber="22" value="2003-10-18">18.10.2003</collectingDate>
</date>
,
<collectorName id="269CEFABFF8EFFE3FE872FE8FE00FB4A" box="[363,486,1215,1241]" pageId="21" pageNumber="22">S. Nomura</collectorName>
leg. (cPJ)
</materialsCitation>
;
<materialsCitation id="3B018020FF8EFFE3FD862FE9FED6FB6E" ID-GBIF-Occurrence="3454958321" collectingDate="1987-06-27" collectionCode="NSMT" collectorName="S. Nomura" location="Geihoku-cho" municipality="Yawatabara" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" stateProvince="Hiroshima">
<emphasis id="B91D566FFF8EFFE3FD862FE9FCCAFB4B" bold="true" box="[618,812,1214,1240]" pageId="21" pageNumber="22">
<collectingRegion id="49AD449FFF8EFFE3FD862FE9FCCAFB4B" box="[618,812,1214,1240]" country="Japan" name="Hiroshima" pageId="21" pageNumber="22">Hiroshima Pref.</collectingRegion>
</emphasis>
:
<specimenCount id="9D6F41F4FF8EFFE3FCD02FE8FCB7FB4A" box="[828,849,1215,1241]" pageId="21" pageNumber="22" type="female"></specimenCount>
,
<collectingMunicipality id="6BB21007FF8EFFE3FCB32FE8FC03FB4A" box="[863,997,1215,1241]" pageId="21" pageNumber="22">Yawatabara</collectingMunicipality>
,
<location id="8EB6DCA6FF8EFFE3FC1E2FE8FB61FB4A" LSID="urn:lsid:plazi:treatment:03C03B6BFF8EFFE1FF7B29E4FAACF92D:8EB6DCA6FF8EFFE3FC1E2FE8FB61FB4A" box="[1010,1159,1215,1241]" municipality="Yawatabara" name="Geihoku-cho" pageId="21" pageNumber="22" stateProvince="Hiroshima">Geihoku-chô</location>
,
<date id="FFD7ACBDFF8EFFE3FB782FE8FAF2FB4A" box="[1172,1300,1215,1241]" pageId="21" pageNumber="22" value="1987-06-27">
<collectingDate id="EF935555FF8EFFE3FB782FE8FAF2FB4A" box="[1172,1300,1215,1241]" pageId="21" pageNumber="22" value="1987-06-27">27.06.1987</collectingDate>
</date>
,
<collectorName id="269CEFABFF8EFFE3FACD2FE8FA7AFB4A" box="[1313,1436,1215,1241]" pageId="21" pageNumber="22">S. Nomura</collectorName>
leg. (
<collectionCode id="ED7812B8FF8EFFE3FF382FB4FECEFB6E" box="[212,296,1251,1277]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="21" pageNumber="22">NSMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B018020FF8EFFE3FEAD2FB4FA7EFB6E" ID-GBIF-Occurrence="3454958323" box="[321,1432,1250,1279]" collectingDate="1994-06-05" collectionCode="NSMT" collectorName="S. Nomura" location="Mt. Ohto" municipality="Hoihoi-dani" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Wakayama">
<emphasis id="B91D566FFF8EFFE3FEAD2FB4FDE8FB6F" bold="true" box="[321,526,1250,1277]" pageId="21" pageNumber="22">
<collectingRegion id="49AD449FFF8EFFE3FEAD2FB4FDE8FB6F" box="[321,526,1250,1277]" country="Japan" name="Wakayama" pageId="21" pageNumber="22">Wakayama Pref.</collectingRegion>
</emphasis>
:
<emphasis id="B91D566FFF8EFFE3FDCE2FB3FDD1FB6C" bold="true" box="[546,567,1252,1279]" pageId="21" pageNumber="22">
<specimenCount id="9D6F41F4FF8EFFE3FDCE2FB3FDD1FB6C" box="[546,567,1252,1279]" pageId="21" pageNumber="22" type="male"></specimenCount>
</emphasis>
,
<collectingMunicipality id="6BB21007FF8EFFE3FDA62FB4FD37FB6E" box="[586,721,1251,1277]" pageId="21" pageNumber="22">Hoihoi-dani</collectingMunicipality>
(riverside),
<location id="8EB6DCA6FF8EFFE3FC892FB4FC35FB6E" LSID="urn:lsid:plazi:treatment:03C03B6BFF8EFFE1FF7B29E4FAACF92D:8EB6DCA6FF8EFFE3FC892FB4FC35FB6E" box="[869,979,1251,1277]" municipality="Hoihoi-dani" name="Mt. Ohto" pageId="21" pageNumber="22" stateProvince="Wakayama">Mt. Ohto</location>
,
<date id="FFD7ACBDFF8EFFE3FC0F2FB5FB82FB6E" box="[995,1124,1250,1277]" pageId="21" pageNumber="22" value="1994-06-05">
<collectingDate id="EF935555FF8EFFE3FC0F2FB5FB82FB6E" box="[995,1124,1250,1277]" pageId="21" pageNumber="22" value="1994-06-05">06.05.1994</collectingDate>
</date>
,
<collectorName id="269CEFABFF8EFFE3FB982FB4FB15FB6E" box="[1140,1267,1251,1277]" pageId="21" pageNumber="22">S. Nomura</collectorName>
leg. (
<collectionCode id="ED7812B8FF8EFFE3FAD02FB4FA76FB6E" box="[1340,1424,1251,1277]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="21" pageNumber="22">NSMT</collectionCode>
)
</materialsCitation>
;
<emphasis id="B91D566FFF8EFFE3FF7B2E51FEFBFAB3" bold="true" box="[151,285,1286,1312]" pageId="21" pageNumber="22">SHIKOKU</emphasis>
:
<materialsCitation id="3B018020FF8EFFE3FEC72E51FAE4FAB2" ID-GBIF-Occurrence="3454958309" box="[299,1282,1286,1313]" collectingDate="2002-05-05" collectionCode="EUMJ" collectorName="T. Kurihara" location="Matsuyama-shi" municipality="Syukuno-machi" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Ehime">
<emphasis id="B91D566FFF8EFFE3FEC72E51FE5FFAB3" bold="true" box="[299,441,1286,1312]" pageId="21" pageNumber="22">
<collectingRegion id="49AD449FFF8EFFE3FEC72E51FE5FFAB3" box="[299,441,1286,1312]" country="Japan" name="Ehime" pageId="21" pageNumber="22">Ehime Pref.</collectingRegion>
</emphasis>
:
<specimenCount id="9D6F41F4FF8EFFE3FE2B2E50FE38FAB2" box="[455,478,1287,1313]" pageId="21" pageNumber="22" type="male"></specimenCount>
,
<collectingMunicipality id="6BB21007FF8EFFE3FE062E50FD78FAB2" box="[490,670,1287,1313]" pageId="21" pageNumber="22">Syukuno-machi</collectingMunicipality>
,
<location id="8EB6DCA6FF8EFFE3FD442E50FCBEFAB2" LSID="urn:lsid:plazi:treatment:03C03B6BFF8EFFE1FF7B29E4FAACF92D:8EB6DCA6FF8EFFE3FD442E50FCBEFAB2" box="[680,856,1287,1313]" municipality="Syukuno-machi" name="Matsuyama-shi" pageId="21" pageNumber="22" stateProvince="Ehime">Matsuyama-shi</location>
,
<date id="FFD7ACBDFF8EFFE3FC8E2E51FC05FAB2" box="[866,995,1286,1313]" pageId="21" pageNumber="22" value="2002-05-05">
<collectingDate id="EF935555FF8EFFE3FC8E2E51FC05FAB2" box="[866,995,1286,1313]" pageId="21" pageNumber="22" value="2002-05-05">05.05.2002</collectingDate>
</date>
,
<collectorName id="269CEFABFF8EFFE3FC012E50FB8BFAB2" box="[1005,1133,1287,1313]" pageId="21" pageNumber="22">T. Kurihara</collectorName>
leg. (
<collectionCode id="ED7812B8FF8EFFE3FB472E50FB1CFAB2" box="[1195,1274,1287,1313]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="21" pageNumber="22">EUMJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B018020FF8EFFE3FAE12E51FA7EFAD6" ID-GBIF-Occurrence="3454958333" collectingDate="1970-08-18" collectionCode="EUMJ" collectorName="Ken Ito &amp; S. Hisamatsu" location="Tosa City" municipality="Usa" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Kochi">
<emphasis id="B91D566FFF8EFFE3FAE12E51FA73FAB3" bold="true" box="[1293,1429,1286,1312]" pageId="21" pageNumber="22">
<collectingRegion id="49AD449FFF8EFFE3FAE12E51FA73FAB3" box="[1293,1429,1286,1312]" country="Japan" name="Kochi" pageId="21" pageNumber="22">Kôchi Pref.</collectingRegion>
</emphasis>
:
<specimenCount id="9D6F41F4FF8EFFE3FF7B2E7CFF48FAD6" box="[151,174,1323,1349]" pageId="21" pageNumber="22" type="male"></specimenCount>
,
<collectingMunicipality id="6BB21007FF8EFFE3FF552E7CFF01FAD6" box="[185,231,1323,1349]" pageId="21" pageNumber="22">Usa</collectingMunicipality>
,
<location id="8EB6DCA6FF8EFFE3FF1D2E7CFEBAFAD6" LSID="urn:lsid:plazi:treatment:03C03B6BFF8EFFE1FF7B29E4FAACF92D:8EB6DCA6FF8EFFE3FF1D2E7CFEBAFAD6" box="[241,348,1323,1349]" municipality="Usa" name="Tosa City" pageId="21" pageNumber="22" stateProvince="Kochi">Tosa City</location>
, 16
<date id="FFD7ACBDFF8EFFE3FE632E7CFDF7FAD6" box="[399,529,1323,1349]" pageId="21" pageNumber="22" value="1970-08-18">
<collectingDate id="EF935555FF8EFFE3FE632E7CFDF7FAD6" box="[399,529,1323,1349]" pageId="21" pageNumber="22" value="1970-08-18">18.08.1970</collectingDate>
</date>
,
<collectorName id="269CEFABFF8EFFE3FDF72E7CFD89FAD6" box="[539,623,1323,1349]" pageId="21" pageNumber="22">Ken Itô</collectorName>
leg. {correctly identified as
<taxonomicName id="4C69F1FEFF8EFFE3FC452E7CFBF4FAD6" box="[937,1042,1323,1349]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B91D566FFF8EFFE3FC452E7CFBF4FAD6" box="[937,1042,1323,1349]" italics="true" pageId="21" pageNumber="22">E. debilis</emphasis>
</taxonomicName>
by
<collectorName id="269CEFABFF8EFFE3FBD52E7CFB2AFAD6" box="[1081,1228,1323,1349]" pageId="21" pageNumber="22">S. Hisamatsu</collectorName>
in 1984} (
<collectionCode id="ED7812B8FF8EFFE3FAAD2E7CFA76FAD6" box="[1345,1424,1323,1349]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="21" pageNumber="22">EUMJ</collectionCode>
)
</materialsCitation>
;
</paragraph>
<paragraph id="8BD68A7DFF8EFFE3FF7B2E19FDB3FA22" blockId="21.[151,1437,926,2033]" pageId="21" pageNumber="22">
<materialsCitation id="3B018020FF8EFFE3FF7B2E19FC84FAFA" ID-GBIF-Occurrence="3454958331" box="[151,866,1358,1385]" collectingDate="1965-09-25" collectionCode="EUMJ" collectorName="M. Sakai" location="Akui" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Tokushima">
<emphasis id="B91D566FFF8EFFE3FF7B2E19FEB8FAFB" bold="true" box="[151,350,1358,1384]" pageId="21" pageNumber="22">
<collectingRegion id="49AD449FFF8EFFE3FF7B2E19FEB8FAFB" box="[151,350,1358,1384]" country="Japan" name="Tokushima" pageId="21" pageNumber="22">Tokushima Pref.</collectingRegion>
</emphasis>
:
<specimenCount id="9D6F41F4FF8EFFE3FE812E18FE62FAFA" box="[365,388,1359,1385]" pageId="21" pageNumber="22" type="male"></specimenCount>
,
<location id="8EB6DCA6FF8EFFE3FE7C2E18FE2DFAFA" LSID="urn:lsid:plazi:treatment:03C03B6BFF8EFFE1FF7B29E4FAACF92D:8EB6DCA6FF8EFFE3FE7C2E18FE2DFAFA" box="[400,459,1359,1385]" name="Akui" pageId="21" pageNumber="22" stateProvince="Tokushima">Akui</location>
,
<date id="FFD7ACBDFF8EFFE3FE3B2E19FDB1FAFA" box="[471,599,1358,1385]" pageId="21" pageNumber="22" value="1965-09-25">
<collectingDate id="EF935555FF8EFFE3FE3B2E19FDB1FAFA" box="[471,599,1358,1385]" pageId="21" pageNumber="22" value="1965-09-25">25.09.1965</collectingDate>
</date>
,
<collectorName id="269CEFABFF8EFFE3FD882E18FD2FFAFA" box="[612,713,1359,1385]" pageId="21" pageNumber="22">M. Sakai</collectorName>
leg. (
<collectionCode id="ED7812B8FF8EFFE3FCE62E18FCBFFAFA" box="[778,857,1359,1385]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="21" pageNumber="22">EUMJ</collectionCode>
)
</materialsCitation>
.
<emphasis id="B91D566FFF8EFFE3FC812E19FBEAFAFB" bold="true" box="[877,1036,1358,1384]" pageId="21" pageNumber="22">RUYUKYUS</emphasis>
:
<materialsCitation id="3B018020FF8EFFE3FBF72E19FF17FA22" ID-GBIF-Occurrence="3454958332" collectingDate="1992-03-23" collectionCode="NSMT" collectorName="S. Nomura" county="Tokara Islands" location="Oh-ike" municipality="Takarajima Island" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Kagoshima">
<emphasis id="B91D566FFF8EFFE3FBF72E19FB03FAFB" bold="true" box="[1051,1253,1358,1384]" pageId="21" pageNumber="22">
<collectingRegion id="49AD449FFF8EFFE3FBF72E19FB03FAFB" box="[1051,1253,1358,1384]" country="Japan" name="Kagoshima" pageId="21" pageNumber="22">Kagoshima Pref.</collectingRegion>
</emphasis>
: 19 exx,
<collectingCounty id="62B7F2F1FF8EFFE3FAA32E18FF0FFA1E" pageId="21" pageNumber="22">Tokara Islands</collectingCounty>
,
<collectingMunicipality id="6BB21007FF8EFFE3FF1F2E24FE58FA1E" box="[243,446,1395,1421]" pageId="21" pageNumber="22">Takarajima Island</collectingMunicipality>
,
<location id="8EB6DCA6FF8EFFE3FE252E24FDFCFA1E" LSID="urn:lsid:plazi:treatment:03C03B6BFF8EFFE1FF7B29E4FAACF92D:8EB6DCA6FF8EFFE3FE252E24FDFCFA1E" box="[457,538,1395,1421]" county="Tokara Islands" municipality="Takarajima Island" name="Oh-ike" pageId="21" pageNumber="22" stateProvince="Kagoshima">Oh-ike</location>
, 20-
<date id="FFD7ACBDFF8EFFE3FDAB2E24FD2CFA1E" box="[583,714,1395,1421]" pageId="21" pageNumber="22" value="1992-03-23">
<collectingDate id="EF935555FF8EFFE3FDAB2E24FD2CFA1E" box="[583,714,1395,1421]" pageId="21" pageNumber="22" value="1992-03-23">23.03.1992</collectingDate>
</date>
,
<collectorName id="269CEFABFF8EFFE3FD392E24FCA8FA1E" box="[725,846,1395,1421]" pageId="21" pageNumber="22">S. Nomura</collectorName>
leg. (
<collectionCode id="ED7812B8FF8EFFE3FC672E24FC3DFA1E" box="[907,987,1395,1421]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="21" pageNumber="22">NSMT</collectionCode>
, cPJ).
<collectorName id="269CEFABFF8EFFE3FBCE2E24FB57FA1E" box="[1058,1201,1395,1421]" pageId="21" pageNumber="22">Additionally</collectorName>
,
<specimenCount id="9D6F41F4FF8EFFE3FB572E24FA86FA1E" box="[1211,1376,1395,1421]" pageId="21" pageNumber="22" type="generic">two specimens</specimenCount>
from
<collectorName id="269CEFABFF8EFFE3FF7B2EC0FF17FA22" box="[151,241,1431,1457]" pageId="21" pageNumber="22">Kyushu</collectorName>
</materialsCitation>
,
<materialsCitation id="3B018020FF8EFFE3FF102EC0FDB7FA22" ID-GBIF-Occurrence="3454958327" box="[252,593,1431,1457]" collectionCode="NSMT" location="Saga Pref." pageId="21" pageNumber="22" specimenCount="1" stateProvince="Saga">
<collectingRegion id="49AD449FFF8EFFE3FF102EC0FE96FA22" box="[252,368,1431,1457]" country="Japan" name="Saga" pageId="21" pageNumber="22">Saga Pref.</collectingRegion>
were seen (
<collectionCode id="ED7812B8FF8EFFE3FE182EC0FDAEFA22" box="[500,584,1431,1457]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="21" pageNumber="22">NSMT</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD68A7DFF8EFFE3FF2B2EECFE6CF966" blockId="21.[151,1437,926,2033]" pageId="21" pageNumber="22">
<emphasis id="B91D566FFF8EFFE3FF2B2EECFE5CFA47" bold="true" box="[199,442,1466,1493]" pageId="21" pageNumber="22">Emended diagnosis.</emphasis>
Small-bodied
<taxonomicName id="4C69F1FEFF8EFFE3FD8B2EECFD0EFA46" box="[615,744,1467,1493]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="24" phylum="Arthropoda" rank="subGenus" subGenus="Psomophus">
<emphasis id="B91D566FFF8EFFE3FD8B2EECFD0EFA46" box="[615,744,1467,1493]" italics="true" pageId="21" pageNumber="22">Psomophus</emphasis>
</taxonomicName>
(Bl ~
<quantity id="4C912798FF8EFFE3FCDD2EECFC6BFA46" box="[817,909,1467,1493]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="22" unit="mm" value="1.2">1.2 mm</quantity>
) with slender antennal club, antennomere 11 slightly less than twice as long as broad; pronotum broadest behind middle, with one pair of small lateral antebasal pits connected by distinct transverse groove, and with short but distinct sublateral carinae; punctures on head, pronotum and elytra inconspicuous; protibiae in male with strongly bent apical regions; aedeagus in ventral view pear-shaped, broadest in sub-basal region, with broadly rounded sides and distinctly delimited apical region composed of shorter ventral apical plate with short, median subtriangular distal projection, and much longer and slenderer dorsal apical plate with sides rounded and strongly converging distad toward subtriangular and blunt apex; endophallus with conspicuous submedian elongate oval median structure; parameres in lateral view almost evenly curved, each with two long apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C373D9F6FF8EFFE1FF2B2DA9FE8CFC5A" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="description">
<paragraph id="8BD68A7DFF8EFFE3FF2B2DA9FC07F8F2" blockId="21.[151,1437,926,2033]" pageId="21" pageNumber="22">
<emphasis id="B91D566FFF8EFFE3FF2B2DA9FE93F88B" bold="true" box="[199,373,1790,1816]" pageId="21" pageNumber="22">Redescription.</emphasis>
Body of male (
<figureCitation id="135296F8FF8EFFE3FDDC2DA8FD6EF88A" box="[560,648,1791,1817]" captionStart="FIGURES 7278" captionStartId="22.[151,264,1857,1881]" captionTargetBox="[161,1421,381,1826]" captionTargetId="figure-101@22.[151,1436,381,1834]" captionTargetPageId="22" captionText="FIGURES 7278. Euconnus debilis, male. Dorsal habitus (72) and historical labels (73) of lectotype of Scydmaenus debilis; fore tibia and tarsus of male in lateral view (74); aedeagus in ventral (75, 77) and lateral (76, 78) views." figureDoi="http://doi.org/10.5281/zenodo.5901036" httpUri="https://zenodo.org/record/5901036/files/figure.png" pageId="21" pageNumber="22">Fig. 72</figureCitation>
:
<typeStatus id="54D234DFFF8EFFE3FD7A2DA8FD19F889" box="[662,767,1791,1818]" pageId="21" pageNumber="22" type="lectotype">lectotype</typeStatus>
) moderately slender, strongly convex, BL
<quantity id="4C912798FF8EFFE3FB032DA8FA7EF88A" box="[1263,1432,1791,1817]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" metricValueMax="1.2" metricValueMin="1.16" pageId="21" pageNumber="22" unit="mm" value="1.18" valueMax="1.2" valueMin="1.16">1.161.20 mm</quantity>
; cuticle glossy, pigmentation light brown, in fully pigmented specimens antennal club slightly to distinctly darker than rest of body, vestiture of setae and bristles slightly lighter than cuticle.
</paragraph>
<paragraph id="8BD68A7DFF8EFFE0FF2B2C3CFF5CFEF6" blockId="21.[151,1437,926,2033]" lastBlockId="22.[151,1436,151,357]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
Head rhomboidal, as long as broad or indistinctly transverse, broadest at eyes, HL
<quantity id="4C912798FF8EFFE3FB932C3DFAC0F816" box="[1151,1318,1898,1925]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="21" pageNumber="22" unit="mm" value="0.24" valueMax="0.25" valueMin="0.23">0.230.25 mm</quantity>
, HW
<quantity id="4C912798FF8EFFE3FA872C3DFF20F83A" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="22" unit="mm" value="0.25">0.25 mm</quantity>
; tempora in dorsal view about 1.5 times as long as eyes and strongly converging posterad, weakly rounded or nearly straight; vertex and frons confluent, weakly and evenly convex, posterior margin of vertex rounded, posteriorly convex, not bulging posterodorsad; supraantennal tubercles feebly marked; frons between antennae steeply declining; clypeus unmodified. Eyes moderately large and finely faceted, weakly projecting laterad from the head silhouette, in lateral view oval. Head with fine and inconspicuous punctures; setae moderately dense and long, suberect, directed posterad, thick bristles distributed on tempora and relatively sparse. Antennae slender and moderately compactly assembled, with distinct trimerous clubs, AnL
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; antennomeres 12 distinctly elongate, 310 each about as long as brad, 11 slightly less than twice as long as broad, indistinctly broader than 10.
</paragraph>
<caption id="DF16DAF5FF8DFFE0FF7B2C16FB54F8EE" ID-DOI="http://doi.org/10.5281/zenodo.5901036" ID-Zenodo-Dep="5901036" httpUri="https://zenodo.org/record/5901036/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,264,1857,1881]" targetBox="[161,1421,381,1826]" targetPageId="22">
<paragraph id="8BD68A7DFF8DFFE0FF7B2C16FB54F8EE" blockId="22.[151,1436,1857,1917]" pageId="22" pageNumber="23">
<emphasis id="B91D566FFF8DFFE0FF7B2C16FEB0F8CA" bold="true" box="[151,342,1857,1881]" pageId="22" pageNumber="23">FIGURES 7278.</emphasis>
<taxonomicName id="4C69F1FEFF8DFFE0FEB12C15FDEBF8CA" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[349,525,1857,1881]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B91D566FFF8DFFE0FEB12C15FDEBF8CA" box="[349,525,1857,1881]" italics="true" pageId="22" pageNumber="23">Euconnus debilis</emphasis>
</taxonomicName>
, male. Dorsal habitus (72) and historical labels (73) of lectotype of
<taxonomicName id="4C69F1FEFF8DFFE0FB212C16FA70F8CA" box="[1229,1430,1857,1881]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B91D566FFF8DFFE0FB212C16FA70F8CA" box="[1229,1430,1857,1881]" italics="true" pageId="22" pageNumber="23">Scydmaenus debilis</emphasis>
</taxonomicName>
; fore tibia and tarsus of male in lateral view (74); aedeagus in ventral (75, 77) and lateral (76, 78) views.
</paragraph>
</caption>
<paragraph id="8BD68A7DFF8DFFE1FF2B2CFBFB62FE19" blockId="22.[151,1436,1963,2026]" lastBlockId="23.[151,1437,151,1726]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
Pronotum bell-shaped with rounded sides, in most specimens widest slightly behind middle, but dense vestiture in intact specimens may obscure the shape, which is slightly variable, PL
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, PW
<quantity id="4C912798FF8DFFE0FB022C98FA7EF879" box="[1262,1432,1999,2026]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="22" pageNumber="23" unit="mm" value="0.28" valueMax="0.3" valueMin="0.26">0.260.30 mm</quantity>
; anterior margin nearly straight and much shorter than posterior margin, anterior corners indistinct, sides of pronotum strongly rounded in anterior half or 1/3 and strongly converging anterad, in posterior half weakly rounded and slightly converging posterad or slightly sinuate; posterior corners strongly obtuse-angled, well-marked; posterior margin weakly arcuate. Pronotal base with distinct transverse groove connecting one pair of small lateral pits, in some specimens groove slightly shallower at middle, sublateral carinae sharply marked. Disc covered with inconspicuously fine punctures, densely covered with thin, suberect setae and thick, straight bristles distributed not only on sides, but also on dorsum (setae and bristles easily brake off during preparation!).
</paragraph>
<paragraph id="8BD68A7DFF8CFFE1FF2B2AC4FBDFFDAE" blockId="23.[151,1437,151,1726]" pageId="23" pageNumber="24">
Elytra oval, broadest distinctly in front of middle, EL
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, EW
<quantity id="4C912798FF8CFFE1FBE32AC5FB53FE3E" box="[1039,1205,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.1" metricValueMin="4.5" pageId="23" pageNumber="24" unit="mm" value="0.48" valueMax="0.51" valueMin="0.45">0.450.51 mm</quantity>
, EI 1.271.42; basal impressions shallow but distinct, humeral calli prominent and weakly elongate, each delimited from adscutellar region by elongate impression; elytral apices separately rounded. Punctures on elytral disc fine, inconspicuous; setae moderately sparse and long, but much longer and thicker than those on head and pronotum, suberect, setae clearly longer than distances between their insertions. Hind wings long, functional.
</paragraph>
<paragraph id="8BD68A7DFF8CFFE1FF2B2910FA7FFDF2" blockId="23.[151,1437,151,1726]" box="[199,1433,583,609]" pageId="23" pageNumber="24">
Legs moderately long and slender, protibiae (
<figureCitation id="135296F8FF8CFFE1FD2E2910FCF2FDF2" box="[706,788,583,609]" captionStart="FIGURES 7278" captionStartId="22.[151,264,1857,1881]" captionTargetBox="[161,1421,381,1826]" captionTargetId="figure-101@22.[151,1436,381,1834]" captionTargetPageId="22" captionText="FIGURES 7278. Euconnus debilis, male. Dorsal habitus (72) and historical labels (73) of lectotype of Scydmaenus debilis; fore tibia and tarsus of male in lateral view (74); aedeagus in ventral (75, 77) and lateral (76, 78) views." figureDoi="http://doi.org/10.5281/zenodo.5901036" httpUri="https://zenodo.org/record/5901036/files/figure.png" pageId="23" pageNumber="24">Fig. 74</figureCitation>
) modified, with short apical portion strongly bent mesad.
</paragraph>
<paragraph id="8BD68A7DFF8CFFE1FF2B293CFE20FD16" blockId="23.[151,1437,151,1726]" box="[199,454,619,645]" pageId="23" pageNumber="24">Abdomen unmodified.</paragraph>
<paragraph id="8BD68A7DFF8CFFE1FF2B29D8FBDCFCCE" blockId="23.[151,1437,151,1726]" pageId="23" pageNumber="24">
Aedeagus (
<figureCitation id="135296F8FF8CFFE1FEAB29D8FE2CFD3A" box="[327,458,654,681]" captionStart="FIGURES 7278" captionStartId="22.[151,264,1857,1881]" captionTargetBox="[161,1421,381,1826]" captionTargetId="figure-101@22.[151,1436,381,1834]" captionTargetPageId="22" captionText="FIGURES 7278. Euconnus debilis, male. Dorsal habitus (72) and historical labels (73) of lectotype of Scydmaenus debilis; fore tibia and tarsus of male in lateral view (74); aedeagus in ventral (75, 77) and lateral (76, 78) views." figureDoi="http://doi.org/10.5281/zenodo.5901036" httpUri="https://zenodo.org/record/5901036/files/figure.png" pageId="23" pageNumber="24">Figs 7578</figureCitation>
) moderately elongate, AeL
<quantity id="4C912798FF8CFFE1FCE729D8FC52FD3A" box="[779,948,655,681]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" metricValueMax="2.6" metricValueMin="2.0" pageId="23" pageNumber="24" unit="mm" value="0.23" valueMax="0.26" valueMin="0.2">0.200.26 mm</quantity>
; median lobe in ventral view pear-shaped, broadest in sub-basal region, with broadly rounded sides and distinctly delimited apical region composed of shorter and subtrapezoidal ventral apical plate bearing short median subtriangular distal projection, and much longer and slenderer dorsal apical plate with sides rounded and strongly converging distad toward subtriangular and blunt apex; endophallus with conspicuous submedian elongate oval median structure; parameres slender, not reaching apex of median lobe, in lateral view almost evenly curved, each with two long apical setae.
</paragraph>
<paragraph id="8BD68A7DFF8CFFE1FF2B2830FE8CFC5A" blockId="23.[151,1437,151,1726]" pageId="23" pageNumber="24">
Female. Externally similar to male but with protibiae very weakly curved; winged. BL
<quantity id="4C912798FF8CFFE1FB7C2830FAD3FC12" box="[1168,1333,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="1.165" metricValueMax="1.2" metricValueMin="1.13" pageId="23" pageNumber="24" unit="mm" value="1.165" valueMax="1.2" valueMin="1.13">1.131.20 mm</quantity>
; HL
<quantity id="4C912798FF8CFFE1FA872830FF23FC36" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="23" pageNumber="24" unit="mm" value="0.23">0.23 mm</quantity>
, HW
<quantity id="4C912798FF8CFFE1FEEB28DDFE8BFC36" box="[263,365,906,933]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="24" unit="mm" value="0.25">0.25 mm</quantity>
, AnL
<quantity id="4C912798FF8CFFE1FE5E28DDFDBEFC36" box="[434,600,906,933]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="23" pageNumber="24" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.500.55 mm</quantity>
; PL
<quantity id="4C912798FF8CFFE1FD6728DCFCD6FC36" box="[651,816,907,933]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="23" pageNumber="24" unit="mm" value="0.29" valueMax="0.3" valueMin="0.28">0.280.30 mm</quantity>
, PW
<quantity id="4C912798FF8CFFE1FC8128DCFBF2FC36" box="[877,1044,907,933]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="23" pageNumber="24" unit="mm" value="0.29" valueMax="0.3" valueMin="0.28">0.280.30 mm</quantity>
; EL
<quantity id="4C912798FF8CFFE1FBA428DCFB08FC36" box="[1096,1262,907,933]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="6.8" metricValueMin="6.3" pageId="23" pageNumber="24" unit="mm" value="0.655" valueMax="0.68" valueMin="0.63">0.630.68 mm</quantity>
, EW
<quantity id="4C912798FF8CFFE1FAC028DDFF23FC5A" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="23" pageNumber="24" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.480.50 mm</quantity>
, EI 1.251.37.
</paragraph>
</subSubSection>
<subSubSection id="C373D9F6FF8CFFE1FF2B2885FBF5FC7D" box="[199,1043,978,1006]" pageId="23" pageNumber="24" type="distribution">
<paragraph id="8BD68A7DFF8CFFE1FF2B2885FBF5FC7D" blockId="23.[151,1437,151,1726]" box="[199,1043,978,1006]" pageId="23" pageNumber="24">
<emphasis id="B91D566FFF8CFFE1FF2B2885FEBFFC7F" bold="true" box="[199,345,978,1004]" pageId="23" pageNumber="24">Distribution</emphasis>
.
<collectingCountry id="F37ECAEDFF8CFFE1FE8B2884FE4DFC7E" box="[359,427,979,1005]" name="Japan" pageId="23" pageNumber="24">Japan</collectingCountry>
: Honshu, Kyushu, Shikoku, Ryukyus (Tokara Islands).
</paragraph>
</subSubSection>
<subSubSection id="C373D9F6FF8CFFE1FF2B28A0FAACF92D" pageId="23" pageNumber="24" type="discussion">
<paragraph id="8BD68A7DFF8CFFE1FF2B28A0FAABFAC6" blockId="23.[151,1437,151,1726]" pageId="23" pageNumber="24">
<emphasis id="B91D566FFF8CFFE1FF2B28A0FEDDFB82" bold="true" box="[199,315,1015,1041]" pageId="23" pageNumber="24">Remarks.</emphasis>
Among described Japanese
<taxonomicName id="4C69F1FEFF8CFFE1FD7C28A0FC86FB82" box="[656,864,1015,1041]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91D566FFF8CFFE1FD7C28A0FD18FB83" box="[656,766,1015,1040]" italics="true" pageId="23" pageNumber="24">Euconnus</emphasis>
species
</taxonomicName>
,
<taxonomicName id="4C69F1FEFF8CFFE1FC9C28A0FC06FB82" box="[880,992,1015,1041]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B91D566FFF8CFFE1FC9C28A0FC06FB82" box="[880,992,1015,1041]" italics="true" pageId="23" pageNumber="24">E. debilis</emphasis>
</taxonomicName>
is the only member of the subgenus
<taxonomicName id="4C69F1FEFF8CFFE1FF7B2F4CFEFEFBA6" box="[151,280,1051,1077]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="subGenus" subGenus="Psomophus">
<emphasis id="B91D566FFF8CFFE1FF7B2F4CFEFEFBA6" box="[151,280,1051,1077]" italics="true" pageId="23" pageNumber="24">Psomophus</emphasis>
</taxonomicName>
and can be distinguished by the subgeneric feature: the trimerous antennal club. As discussed by
<bibRefCitation id="EFF8F78CFF8CFFE1FF7B2F68FE9FFBCA" author="Jaloszynski, P." box="[151,377,1087,1113]" pageId="23" pageNumber="24" pagination="565 - 578" refId="ref25664" refString="Jaloszynski, P. (2017 c) Taxonomy of &quot; Euconnus complex &quot;. Part XII. Synonymy circle of Psomophus, Spanioconnus, Scydmaenites and Xestophus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4272 (4), 565 - 578. https: // doi. org / 10.11646 / zootaxa. 4272.4.5" type="journal article" year="2017">Jałoszyński (2017c)</bibRefCitation>
,
<taxonomicName id="4C69F1FEFF8CFFE1FE682F68FDE3FBCA" box="[388,517,1087,1113]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="subGenus" subGenus="Psomophus">
<emphasis id="B91D566FFF8CFFE1FE682F68FDE3FBCA" box="[388,517,1087,1113]" italics="true" pageId="23" pageNumber="24">Psomophus</emphasis>
</taxonomicName>
is a somehow problematic subgenus and it may in future be merged with the large and morphologically diverse
<taxonomicName id="4C69F1FEFF8CFFE1FE332F34FDABFBEF" box="[479,589,1123,1148]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B91D566FFF8CFFE1FE332F34FDABFBEF" box="[479,589,1123,1148]" italics="true" pageId="23" pageNumber="24">Euconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EEB14FF8CFFE1FDB82F34FD69FBEE" box="[596,655,1123,1149]" pageId="23" pageNumber="24" sensu="stricto">s. str.</taxonomicNameLabel>
Additional findings in Japanese fauna support this view (Jałoszyński, in preparation), as there are more (new) species that inhabit Japanese Islands and have distinctly or indistinctly trimerous clubs, and diverse remaining characters. This problem will be treated separately. Here, however, it should be noted that there are at least 45 undescribed species in
<collectingCountry id="F37ECAEDFF8CFFE1FCF72F98FCBDFB7A" box="[795,859,1231,1257]" name="Japan" pageId="23" pageNumber="24">Japan</collectingCountry>
that highly resemble
<taxonomicName id="4C69F1FEFF8CFFE1FBA02F98FB50FB7A" box="[1100,1206,1231,1257]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B91D566FFF8CFFE1FBA02F98FB50FB7A" box="[1100,1206,1231,1257]" italics="true" pageId="23" pageNumber="24">E. debilis</emphasis>
</taxonomicName>
, and at least three of them have males with the protibiae modified in a very similar way as those in
<taxonomicName id="4C69F1FEFF8CFFE1FC0E2FA4FBACFA9E" box="[994,1098,1267,1293]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B91D566FFF8CFFE1FC0E2FA4FBACFA9E" box="[994,1098,1267,1293]" italics="true" pageId="23" pageNumber="24">E. debilis</emphasis>
</taxonomicName>
. Moreover, their distributional ranges overlap. For this reason, in order to identify any male of
<taxonomicName id="4C69F1FEFF8CFFE1FC8C2E40FC28FAA3" box="[864,974,1303,1328]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B91D566FFF8CFFE1FC8C2E40FC28FAA3" box="[864,974,1303,1328]" italics="true" pageId="23" pageNumber="24">Euconnus</emphasis>
</taxonomicName>
with body form similar to that illustrated in
<figureCitation id="135296F8FF8CFFE1FF582E6CFEE0FAC6" box="[180,262,1339,1365]" captionStart="FIGURES 7278" captionStartId="22.[151,264,1857,1881]" captionTargetBox="[161,1421,381,1826]" captionTargetId="figure-101@22.[151,1436,381,1834]" captionTargetPageId="22" captionText="FIGURES 7278. Euconnus debilis, male. Dorsal habitus (72) and historical labels (73) of lectotype of Scydmaenus debilis; fore tibia and tarsus of male in lateral view (74); aedeagus in ventral (75, 77) and lateral (76, 78) views." figureDoi="http://doi.org/10.5281/zenodo.5901036" httpUri="https://zenodo.org/record/5901036/files/figure.png" pageId="23" pageNumber="24">Fig. 72</figureCitation>
, trimerous antennal club and bent apices of profemora, examination of aedeagus is indispensable.
</paragraph>
<paragraph id="8BD68A7DFF8CFFE1FF2B2E08FAACF92D" blockId="23.[151,1437,151,1726]" pageId="23" pageNumber="24">
<taxonomicName id="4C69F1FEFF8CFFE1FF2B2E08FE6FFAEA" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[199,393,1375,1401]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B91D566FFF8CFFE1FF2B2E08FE6FFAEA" box="[199,393,1375,1401]" italics="true" pageId="23" pageNumber="24">Euconnus debilis</emphasis>
</taxonomicName>
has a broad distribution, currently known localities range from
<collectingRegion id="49AD449FFF8CFFE1FBB12E08FAB2FAEA" box="[1117,1364,1375,1401]" country="Japan" name="Fukushima" pageId="23" pageNumber="24">Fukushima Prefecture</collectingRegion>
in the CN region of Honshu, through Shikoku and Kyushu and the south-western Tokara Islands. Aedeagi of specimens distributed on different islands do not show any differences that could be attributed to separate species or subspecies. However, external morphology of this species is somewhat variable, and although the body form, setal pattern and distribution of punctures are constant, adults of
<taxonomicName id="4C69F1FEFF8CFFE1FD582EB8FCF9F99A" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[692,799,1519,1545]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B91D566FFF8CFFE1FD582EB8FCF9F99A" box="[692,799,1519,1545]" italics="true" pageId="23" pageNumber="24">E. debilis</emphasis>
</taxonomicName>
can have noticeably different proportions of body parts, best visible in the elytral index, which in the small studied sample ranges 1.271.42 among males and 1.251.37 among females. In fully pigmented specimens the antennal club is slightly to distinctly darker than the remaining portion of antenna, but most commonly the antennae are uniformly light brown. There is another, externally similar, and yet undescribed
<taxonomicName id="4C69F1FEFF8CFFE1FE6D2D28FDE4F90A" box="[385,514,1663,1689]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="subGenus" subGenus="Psomophus">
<emphasis id="B91D566FFF8CFFE1FE6D2D28FDE4F90A" box="[385,514,1663,1689]" italics="true" pageId="23" pageNumber="24">Psomophus</emphasis>
</taxonomicName>
broadly distributed in the Ryukyus, from Tokara Islands to Okinawa-jima, which can be easily confused with
<taxonomicName id="4C69F1FEFF8CFFE1FE382DF4FDD8F92E" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[468,574,1699,1725]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B91D566FFF8CFFE1FE382DF4FDD8F92E" box="[468,574,1699,1725]" italics="true" pageId="23" pageNumber="24">E. debilis</emphasis>
</taxonomicName>
, but clearly differs in the male genitalia (Jałoszyński, in preparation).
</paragraph>
</subSubSection>
</treatment>
</document>