303 lines
36 KiB
XML
303 lines
36 KiB
XML
<document id="3B9EF7CE93FCD6F1053FC887877CA6A5" ID-CLB-Dataset="2431" ID-DOI="10.11646/zootaxa.5093.1.1" ID-GBIF-Dataset="0faf68be-3777-4466-8cd5-9eb8ca72a239" ID-ISSN="1175-5326" ID-Zenodo-Dep="5901006" ID-ZooBank="C2F5E368-CB15-4207-9944-E52C190FBF20" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1643101690477" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2022" docId="03C03B6BFF8AFFE5FF7B2F4CFC6BFAEA" docLanguage="en" docName="zootaxa.5093.1.1.pdf" docOrigin="Zootaxa 5093 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euconnus (Euconnus) oscillans Sharp 1886" docType="treatment" docVersion="7" lastPageNumber="20" masterDocId="FFF94313FF9BFFF6FFEC2B57FFE6FF93" masterDocTitle="Euconnus Thomson of Japan: redescriptions of species established by Reitter, Sharp and Franz, new synonyms, and summary of current state of knowledge (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="18" updateTime="1699292035325" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A59CE917ABFBFDFE314BC365DE49D00A">Euconnus Thomson of Japan: redescriptions of species established by Reitter, Sharp and Franz, new synonyms, and summary of current state of knowledge (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="566B9F0665EFD96922397097E1FDAE78">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="66F6E449737A4300648E000E0722DB65">Zootaxa</mods:title>
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<mods:part id="42FFCDD2FCB4CF74631292231C945136">
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<mods:date id="3C3C74B7B375FF42FF33C4D03C602E02">2022</mods:date>
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<mods:number id="887BB28B4E09AFED07C6C96CE8C16BE6">2022-01-25</mods:number>
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<mods:number id="27D25F8B5405FF4957C82558D17FCF50">5093</mods:number>
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<mods:end id="671A0C7B20871417E53AC28668BF03DF">37</mods:end>
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<treatment id="03C03B6BFF8AFFE5FF7B2F4CFC6BFAEA" ID-DOI="http://doi.org/10.5281/zenodo.5901016" ID-GBIF-Taxon="192597247" ID-Zenodo-Dep="5901016" LSID="urn:lsid:plazi:treatment:03C03B6BFF8AFFE5FF7B2F4CFC6BFAEA" httpUri="http://treatment.plazi.org/id/03C03B6BFF8AFFE5FF7B2F4CFC6BFAEA" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
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<subSubSection id="C373D9F6FF8AFFE7FF7B2F4CFD96FBA7" box="[151,624,1050,1077]" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="8BD68A7DFF8AFFE7FF7B2F4CFD96FBA7" blockId="17.[151,624,1050,1077]" box="[151,624,1050,1077]" pageId="17" pageNumber="18">
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<heading id="D09E3D11FF8AFFE7FF7B2F4CFD96FBA7" bold="true" box="[151,624,1050,1077]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
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<taxonomicName id="4C69F1FEFF8AFFE7FF7B2F4CFD96FBA7" ID-CoL="9G95M" authority="Sharp" authorityName="Sharp" authorityYear="1886" box="[151,624,1050,1077]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="oscillans" subGenus="Euconnus">
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<emphasis id="B91D566FFF8AFFE7FF7B2F4CFD96FBA7" bold="true" box="[151,624,1050,1077]" pageId="17" pageNumber="18">
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<emphasis id="B91D566FFF8AFFE7FF7B2F4CFEF5FBA6" bold="true" box="[151,275,1051,1077]" italics="true" pageId="17" pageNumber="18">Euconnus</emphasis>
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(
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<emphasis id="B91D566FFF8AFFE7FEC82F4CFE46FBA6" bold="true" box="[292,416,1051,1077]" italics="true" pageId="17" pageNumber="18">Euconnus</emphasis>
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)
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<emphasis id="B91D566FFF8AFFE7FE5D2F4DFDFDFBA6" bold="true" box="[433,539,1050,1077]" italics="true" pageId="17" pageNumber="18">oscillans</emphasis>
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Sharp
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C373D9F6FF8AFFE7FF2B2F37FEC1FB0F" pageId="17" pageNumber="18" type="reference_group">
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<paragraph id="8BD68A7DFF8AFFE7FF2B2F37FDDCFBE4" blockId="17.[151,570,1119,1180]" box="[199,570,1119,1143]" pageId="17" pageNumber="18">
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<treatmentCitationGroup id="AB79AD53FF8AFFE7FF2B2F37FDDCFBE4" box="[199,570,1119,1143]" pageId="17" pageNumber="18">
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<treatmentCitation id="0AC8AC6CFF8AFFE7FF2B2F37FDD0FBE4" author="Sharp, D." box="[199,566,1119,1143]" page="48" pageId="17" pageNumber="18" year="1886">
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<taxonomicName id="4C69F1FEFF8AFFE7FF2B2F37FDD0FBE4" ID-CoL="9G95M" authority="Sharp, 1886: 48" authorityName="Sharp" authorityPageNumber="48" authorityYear="1886" box="[199,566,1119,1143]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="oscillans">
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<emphasis id="B91D566FFF8AFFE7FF2B2F37FE6DFBE4" box="[199,395,1119,1143]" italics="true" pageId="17" pageNumber="18">Euconnus oscillans</emphasis>
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<bibRefCitation id="EFF8F78CFF8AFFE7FE7E2F08FDD0FBE4" author="Sharp, D." box="[402,566,1119,1143]" pageId="17" pageNumber="18" pagination="46 - 51" refId="ref26383" refString="Sharp, D. (1886) The Scydmaenidae of Japan. Entomologist's Monthly Magazine, 23, 46 - 51." type="journal article" year="1886">Sharp, 1886: 48</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BD68A7DFF8AFFE7FF7B2FD6FEC1FB0F" blockId="17.[151,570,1119,1180]" box="[151,295,1153,1180]" pageId="17" pageNumber="18">
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(
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<figureCitation id="135296F8FF8AFFE7FF732FD6FEF9FB0F" box="[159,287,1153,1180]" captionStart="FIGURES 64–69" captionStartId="18.[151,264,1696,1720]" captionTargetBox="[158,1423,358,1660]" captionTargetId="figure-100@18.[151,1436,345,1673]" captionTargetPageId="18" captionText="FIGURES 64–69. Euconnus oscillans, male. Dorsal habitus (64) and annotated mounting card with historical labels (65) of holotype; aedeagus in ventral (66, 68) and lateral (67, 69) views." figureDoi="http://doi.org/10.5281/zenodo.5901032" httpUri="https://zenodo.org/record/5901032/files/figure.png" pageId="17" pageNumber="18">Figs 64–69</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C373D9F6FF8AFFE7FF7B2F9EFDE5F948" pageId="17" pageNumber="18" type="materials_examined">
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<paragraph id="8BD68A7DFF8AFFE7FF7B2F9EFB06FAB8" blockId="17.[151,1437,1225,2008]" pageId="17" pageNumber="18">
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<emphasis id="B91D566FFF8AFFE7FF7B2F9EFE7FFB70" bold="true" box="[151,409,1225,1251]" pageId="17" pageNumber="18">Type material studied</emphasis>
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.
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<materialsCitation id="3B018020FF8AFFE7FE4A2F9EFB06FAB8" ID-GBIF-Occurrence="3454958304" collectionCode="NHM" collectorName="G. Lewis." country="Japan" location="Japan" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B91D566FFF8AFFE7FE4A2F9EFDF5FB70" bold="true" box="[422,531,1225,1251]" pageId="17" pageNumber="18">
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<typeStatus id="54D234DFFF8AFFE7FE4A2F9EFDF5FB70" box="[422,531,1225,1251]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
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</emphasis>
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:
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<specimenCount id="9D6F41F4FF8AFFE7FDCD2F9DFDD0FB77" box="[545,566,1226,1252]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
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(
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<figureCitation id="135296F8FF8AFFE7FDA82F9EFD70FB77" box="[580,662,1225,1252]" captionStart="FIGURES 64–69" captionStartId="18.[151,264,1696,1720]" captionTargetBox="[158,1423,358,1660]" captionTargetId="figure-100@18.[151,1436,345,1673]" captionTargetPageId="18" captionText="FIGURES 64–69. Euconnus oscillans, male. Dorsal habitus (64) and annotated mounting card with historical labels (65) of holotype; aedeagus in ventral (66, 68) and lateral (67, 69) views." figureDoi="http://doi.org/10.5281/zenodo.5901032" httpUri="https://zenodo.org/record/5901032/files/figure.png" pageId="17" pageNumber="18">Fig. 64</figureCitation>
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), mounted on thick card with annotation: “
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<taxonomicName id="4C69F1FEFF8AFFE7FB872F9EFB3AFB70" box="[1131,1244,1225,1251]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Euconnus</taxonomicName>
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osci- / lans. Type / D.S. / Hitoyoshi. / 3.5.81. Lewis.”, with two labels (
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<figureCitation id="135296F8FF8AFFE7FD192FBAFCACFA9B" box="[757,842,1261,1288]" captionStart="FIGURES 64–69" captionStartId="18.[151,264,1696,1720]" captionTargetBox="[158,1423,358,1660]" captionTargetId="figure-100@18.[151,1436,345,1673]" captionTargetPageId="18" captionText="FIGURES 64–69. Euconnus oscillans, male. Dorsal habitus (64) and annotated mounting card with historical labels (65) of holotype; aedeagus in ventral (66, 68) and lateral (67, 69) views." figureDoi="http://doi.org/10.5281/zenodo.5901032" httpUri="https://zenodo.org/record/5901032/files/figure.png" pageId="17" pageNumber="18">Fig. 65</figureCitation>
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): “
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<collectingCountry id="F37ECAEDFF8AFFE7FC842FBAFC4DFA94" box="[872,939,1261,1287]" name="Japan" pageId="17" pageNumber="18">Japan</collectingCountry>
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/
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<collectorName id="269CEFABFF8AFFE7FC2E2FBAFBD7FA94" box="[962,1073,1261,1287]" pageId="17" pageNumber="18">G. Lewis.</collectorName>
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/ 1910-320” [white with orange horizontal line, printed], and circular label “Type / H.T.” [white with red margin, printed] (
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||
<collectionCode id="ED7812B8FF8AFFE7FB612E46FB33FAB8" box="[1165,1237,1297,1323]" pageId="17" pageNumber="18">NHM</collectionCode>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BD68A7DFF8AFFE7FF2B2E62FEA9FAE0" blockId="17.[151,1437,1225,2008]" pageId="17" pageNumber="18">
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||
<emphasis id="B91D566FFF8AFFE7FF2B2E62FD79FADC" bold="true" box="[199,671,1333,1359]" pageId="17" pageNumber="18">Additional material studied. HONSHU</emphasis>
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:
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<materialsCitation id="3B018020FF8AFFE7FD5C2E62FB8BFAC3" ID-GBIF-Occurrence="3454958326" box="[688,1133,1333,1360]" location="Mt. Kasuga-yama" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Nara">
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||
<emphasis id="B91D566FFF8AFFE7FD5C2E62FCD6FADC" bold="true" box="[688,816,1333,1359]" pageId="17" pageNumber="18">
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||
<collectingRegion id="49AD449FFF8AFFE7FD5C2E62FCD6FADC" box="[688,816,1333,1359]" country="Japan" name="Nara" pageId="17" pageNumber="18">Nara Pref.</collectingRegion>
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</emphasis>
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||
:
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<specimenCount id="9D6F41F4FF8AFFE7FCAD2E61FCB0FAC3" box="[833,854,1334,1360]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
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, near
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<location id="8EB6DCA6FF8AFFE7FC732E61FB8BFAC3" LSID="urn:lsid:plazi:treatment:03C03B6BFF8AFFE5FF7B2F4CFC6BFAEA:8EB6DCA6FF8AFFE7FC732E61FB8BFAC3" box="[927,1133,1334,1360]" name="Mt. Kasuga-yama" pageId="17" pageNumber="18" stateProvince="Nara">Mt. Kasuga-yama</location>
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</materialsCitation>
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,
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<materialsCitation id="3B018020FF8AFFE7FB962E61FEACFAE0" ID-GBIF-Occurrence="3454958316" collectingDate="2001-05-05" collectorName="T. Tsuru" location="City" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Nara">
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<collectingRegion id="49AD449FFF8AFFE7FB962E61FB56FAC3" box="[1146,1200,1334,1360]" country="Japan" name="Nara" pageId="17" pageNumber="18">Nara</collectingRegion>
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<location id="8EB6DCA6FF8AFFE7FB562E62FB0DFADC" LSID="urn:lsid:plazi:treatment:03C03B6BFF8AFFE5FF7B2F4CFC6BFAEA:8EB6DCA6FF8AFFE7FB562E62FB0DFADC" box="[1210,1259,1333,1359]" name="City" pageId="17" pageNumber="18" stateProvince="Nara">City</location>
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,
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||
<date id="FFD7ACBDFF8AFFE7FB152E62FA9FFAC3" box="[1273,1401,1333,1360]" pageId="17" pageNumber="18" value="2001-05-05">
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||
<collectingDate id="EF935555FF8AFFE7FB152E62FA9FFAC3" box="[1273,1401,1333,1360]" pageId="17" pageNumber="18" value="2001-05-05">05.05.2001</collectingDate>
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||
</date>
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||
,
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<collectorName id="269CEFABFF8AFFE7FA6A2E61FF30FAE7" pageId="17" pageNumber="18">T. Tsuru</collectorName>
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leg. (cPJ)
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</materialsCitation>
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||
.
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||
</paragraph>
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<paragraph id="8BD68A7DFF8AFFE7FF2B2E2AFDE5F948" blockId="17.[151,1437,1225,2008]" pageId="17" pageNumber="18">
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||
<emphasis id="B91D566FFF8AFFE7FF2B2E2AFE51FA04" bold="true" box="[199,439,1405,1431]" pageId="17" pageNumber="18">Emended diagnosis.</emphasis>
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Among Japanese species,
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<taxonomicName id="4C69F1FEFF8AFFE7FD042E29FC8BFA04" authorityName="Sharp" authorityYear="1886" box="[744,877,1405,1431]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="oscillans">
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<emphasis id="B91D566FFF8AFFE7FD042E29FC8BFA04" box="[744,877,1405,1431]" italics="true" pageId="17" pageNumber="18">E. oscillans</emphasis>
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||
</taxonomicName>
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can be identified by: small body (BL ~
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||
<quantity id="4C912798FF8AFFE7FADF2E2AFA76FA0B" box="[1331,1424,1405,1432]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="mm" value="1.2">1.2 mm</quantity>
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); conspicuous setal pattern: head covered with very thin and short, moderately dense suberect setae, lacking thick bristles, pronotum densely covered with setae and bristles, and elytra moderately densely covered with suberect setae much longer and thicker than those on head; relatively compact antennae with antennomeres 3–7 each about as long as broad and distinct tetramerous club; head covered densely with fine punctures, pronotum with inconspicuously fine punctures, but elytra densely, deeply, distinctly punctate; pronotum with one pair of shallow lateral antebasal pits, connected by superficial transverse impression covered densely with fine punctures, and with a pair of short but distinct sublateral carinae, broadest indistinctly behind middle; legs in male unmodified; and aedeagus stout, with presumably subtriangular apex of mediodistal region of dorsal plate and a complex set of strongly asymmetrically distributed endophallic sclerites.
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</paragraph>
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</subSubSection>
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<subSubSection id="C373D9F6FF8AFFE5FF2B2DB2FCA5FC86" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="description">
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||
<paragraph id="8BD68A7DFF8AFFE7FF2B2DB2FF0FF8D4" blockId="17.[151,1437,1225,2008]" pageId="17" pageNumber="18">
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<emphasis id="B91D566FFF8AFFE7FF2B2DB2FE93F96C" bold="true" box="[199,373,1765,1791]" pageId="17" pageNumber="18">Redescription.</emphasis>
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Body of male (
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<figureCitation id="135296F8FF8AFFE7FDC72DB2FD64F96C" box="[555,642,1765,1791]" captionStart="FIGURES 64–69" captionStartId="18.[151,264,1696,1720]" captionTargetBox="[158,1423,358,1660]" captionTargetId="figure-100@18.[151,1436,345,1673]" captionTargetPageId="18" captionText="FIGURES 64–69. Euconnus oscillans, male. Dorsal habitus (64) and annotated mounting card with historical labels (65) of holotype; aedeagus in ventral (66, 68) and lateral (67, 69) views." figureDoi="http://doi.org/10.5281/zenodo.5901032" httpUri="https://zenodo.org/record/5901032/files/figure.png" pageId="17" pageNumber="18">Fig. 64</figureCitation>
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||
:
|
||
<typeStatus id="54D234DFFF8AFFE7FD632DB2FD15F893" box="[655,755,1765,1792]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
|
||
) moderately slender, strongly convex, BL
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<quantity id="4C912798FF8AFFE7FB372DB2FAA2F893" box="[1243,1348,1765,1792]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="17" pageNumber="18" unit="mm" value="1.14">1.14 mm</quantity>
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; cuticle glossy, pigmentation light brown with slightly reddish hue, vestiture of setae and bristles slightly lighter than cuticle.
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||
</paragraph>
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<paragraph id="8BD68A7DFF8AFFE4FF2B2C06FB22FED2" blockId="17.[151,1437,1225,2008]" lastBlockId="18.[151,1437,151,321]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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Head elliptical, slightly elongate and broadest at eyes, HL
|
||
<quantity id="4C912798FF8AFFE7FCA52C06FC48F8FF" box="[841,942,1873,1900]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="17" pageNumber="18" unit="mm" value="0.23">0.23 mm</quantity>
|
||
, HW
|
||
<quantity id="4C912798FF8AFFE7FC002C06FBB7F8FF" box="[1004,1105,1873,1900]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="18" unit="mm" value="0.25">0.25 mm</quantity>
|
||
; tempora in dorsal view about twice as long as eyes and weakly converging posterad, weakly but distinctly rounded; vertex and frons confluent, weakly and evenly convex, posterior margin of vertex rounded, posteriorly convex, not bulging posterodorsad; supraantennal tubercles feebly marked; frons between antennae steeply declining; clypeus unmodified. Eyes moderately large and finely faceted, weakly projecting laterad from the head silhouette, in lateral view oval. Head densely covered with fine and superficial punctures, so that surface of vertex and frons appears semi-glossy; setae moderately dense and long, suberect, directed posterad, thick bristles lacking. Antennae slender and moderately compactly assembled, with distinct tetramerous clubs, AnL
|
||
<quantity id="4C912798FF89FFE4FCAF2A55FC4AFE8E" box="[835,940,258,285]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="18" pageNumber="19" unit="mm" value="0.53">0.53 mm</quantity>
|
||
; antennomeres 1–2 distinctly elongate, 3–7 each about as long as brad, 8–10 each distinctly transverse, 11 slightly elongate, as broad as 10.
|
||
</paragraph>
|
||
<caption id="DF16DAF5FF89FFE4FF7B2DF7FCC1F94F" ID-DOI="http://doi.org/10.5281/zenodo.5901032" ID-Zenodo-Dep="5901032" httpUri="https://zenodo.org/record/5901032/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,264,1696,1720]" targetBox="[158,1423,358,1660]" targetPageId="18">
|
||
<paragraph id="8BD68A7DFF89FFE4FF7B2DF7FCC1F94F" blockId="18.[151,1436,1696,1756]" pageId="18" pageNumber="19">
|
||
<emphasis id="B91D566FFF89FFE4FF7B2DF7FEB0F92B" bold="true" box="[151,342,1696,1720]" pageId="18" pageNumber="19">FIGURES 64–69.</emphasis>
|
||
<taxonomicName id="4C69F1FEFF89FFE4FEB22DF6FDC2F92B" authorityName="Sharp" authorityYear="1886" box="[350,548,1696,1720]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="oscillans">
|
||
<emphasis id="B91D566FFF89FFE4FEB22DF6FDC2F92B" box="[350,548,1696,1720]" italics="true" pageId="18" pageNumber="19">Euconnus oscillans</emphasis>
|
||
</taxonomicName>
|
||
, male. Dorsal habitus (64) and annotated mounting card with historical labels (65) of holotype; aedeagus in ventral (66, 68) and lateral (67, 69) views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD68A7DFF89FFE5FF2B2C5CFE45FF45" blockId="18.[151,1437,1803,2009]" lastBlockId="19.[151,1437,151,1401]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
|
||
Pronotum bell-shaped with rounded sides, widest slightly behind middle, but dense vestiture in intact specimens may obscure the shape, PL
|
||
<quantity id="4C912798FF89FFE4FE252C78FDC9F8DA" box="[457,559,1839,1865]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="18" pageNumber="19" unit="mm" value="0.26">0.26 mm</quantity>
|
||
, PW
|
||
<quantity id="4C912798FF89FFE4FD872C78FD34F8DA" box="[619,722,1839,1865]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="18" pageNumber="19" unit="mm" value="0.28">0.28 mm</quantity>
|
||
; anterior margin nearly straight and much shorter than posterior margin, anterior corners indistinct, sides of pronotum strongly rounded and strongly converging anterad; posterior corners strongly obtuse-angled, well-marked; posterior margin weakly arcuate. Pronotal base with shallow transverse impression covered densely with fine and shallow punctures, one pair of lateral antebasal pits is situated within this impression, sublateral carinae sharply marked. Disc covered with inconspicuously fine punctures, densely covered with thin, suberect setae and thick, straight bristles distributed not only on sides, but also on dorsum (in
|
||
<typeStatus id="54D234DFFF88FFE5FAD62BC0FA7AFF22" box="[1338,1436,151,177]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
male partly broken off).
|
||
</paragraph>
|
||
<paragraph id="8BD68A7DFF88FFE5FF2B2B88FEF6FE3E" blockId="19.[151,1437,151,1401]" pageId="19" pageNumber="20">
|
||
Elytra oval, broadest distinctly in front of middle, EL
|
||
<quantity id="4C912798FF88FFE5FCAF2B89FC4AFF6A" box="[835,940,222,249]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="19" pageNumber="20" unit="mm" value="0.65">0.65 mm</quantity>
|
||
, EW
|
||
<quantity id="4C912798FF88FFE5FC1D2B89FBBDFF6A" box="[1009,1115,222,249]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="20" unit="mm" value="0.5">0.50 mm</quantity>
|
||
, EI 1.30; basal impressions shallow but distinct, humeral calli prominent and weakly elongate, each delimited from adscutellar region by round impression; elytral apices separately rounded. Punctures on elytral disc conspicuously large and dense, those on anterior half separated by spaces subequal to their diameters, punctures reducing in diameter and depth toward sides and apices; setae moderately dense and long, but clearly longer than those on head, suberect. Hind wings long, functional.
|
||
</paragraph>
|
||
<paragraph id="8BD68A7DFF88FFE5FF2B2AE0FD3EFE42" blockId="19.[151,1437,151,1401]" box="[199,728,439,465]" pageId="19" pageNumber="20">Legs moderately long and slender, unmodified.</paragraph>
|
||
<paragraph id="8BD68A7DFF88FFE5FF2B2A8CFE20FE66" blockId="19.[151,1437,151,1401]" box="[199,454,475,501]" pageId="19" pageNumber="20">Abdomen unmodified.</paragraph>
|
||
<paragraph id="8BD68A7DFF88FFE5FF2B2AA8FAADFD5E" blockId="19.[151,1437,151,1401]" pageId="19" pageNumber="20">
|
||
Aedeagus (
|
||
<figureCitation id="135296F8FF88FFE5FEAA2AA8FE2FFD89" box="[326,457,511,538]" captionStart="FIGURES 64–69" captionStartId="18.[151,264,1696,1720]" captionTargetBox="[158,1423,358,1660]" captionTargetId="figure-100@18.[151,1436,345,1673]" captionTargetPageId="18" captionText="FIGURES 64–69. Euconnus oscillans, male. Dorsal habitus (64) and annotated mounting card with historical labels (65) of holotype; aedeagus in ventral (66, 68) and lateral (67, 69) views." figureDoi="http://doi.org/10.5281/zenodo.5901032" httpUri="https://zenodo.org/record/5901032/files/figure.png" pageId="19" pageNumber="20">Figs 66–69</figureCitation>
|
||
) in the only known male slightly damaged, with apex of median lobe broken off and proximal half of median lobe distorted; stout, AeL
|
||
<quantity id="4C912798FF88FFE5FD522974FCC5FDAE" box="[702,803,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="19" pageNumber="20" unit="mm" value="0.19">0.19 mm</quantity>
|
||
; median lobe in ventral view broadest in subapical region, ventral apical plate subtriangular with rounded apex, dorsal apical plate longer, presumably with subtriangular median projection; endophallic structures asymmetrical, with several elongate and curved sclerites of various shapes and lengths distributed in sub-median and subapical regions. Parameres slender and long, but not reaching apex of median lobe, in lateral view parameres weakly curved in proximal half, each with 3–4 long subapical setae.
|
||
</paragraph>
|
||
<paragraph id="8BD68A7DFF88FFE5FF2B2980FCA5FC86" blockId="19.[151,1437,151,1401]" pageId="19" pageNumber="20">
|
||
Female. Externally indistinguishable from male. BL
|
||
<quantity id="4C912798FF88FFE5FCF42980FC99FD62" box="[792,895,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="20" unit="mm" value="1.2">1.20 mm</quantity>
|
||
; HL
|
||
<quantity id="4C912798FF88FFE5FC5B2980FBFBFD62" box="[951,1053,727,753]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="19" pageNumber="20" unit="mm" value="0.23">0.23 mm</quantity>
|
||
, HW
|
||
<quantity id="4C912798FF88FFE5FBB32981FB23FD62" box="[1119,1221,726,753]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="20" unit="mm" value="0.25">0.25 mm</quantity>
|
||
, AnL
|
||
<quantity id="4C912798FF88FFE5FAE52981FA97FD62" box="[1289,1393,726,753]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="19" pageNumber="20" unit="mm" value="0.53">0.53 mm</quantity>
|
||
; PL
|
||
<quantity id="4C912798FF88FFE5FF7B29ACFF1BFC86" box="[151,253,763,789]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="19" pageNumber="20" unit="mm" value="0.28">0.28 mm</quantity>
|
||
, PW
|
||
<quantity id="4C912798FF88FFE5FED529ACFE46FC86" box="[313,416,763,789]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="20" unit="mm" value="0.3">0.30 mm</quantity>
|
||
; EL
|
||
<quantity id="4C912798FF88FFE5FE3829ACFDDCFC86" box="[468,570,763,789]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="20" unit="mm" value="0.7">0.70 mm</quantity>
|
||
, EW
|
||
<quantity id="4C912798FF88FFE5FD9B29ADFD3BFC86" box="[631,733,762,789]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="19" pageNumber="20" unit="mm" value="0.53">0.53 mm</quantity>
|
||
, EI 1.33.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C373D9F6FF88FFE5FF2B2849FD9CFCAA" box="[199,634,798,825]" pageId="19" pageNumber="20" type="distribution">
|
||
<paragraph id="8BD68A7DFF88FFE5FF2B2849FD9CFCAA" blockId="19.[151,1437,151,1401]" box="[199,634,798,825]" pageId="19" pageNumber="20">
|
||
<emphasis id="B91D566FFF88FFE5FF2B2849FEBFFCAB" bold="true" box="[199,345,798,824]" pageId="19" pageNumber="20">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F37ECAEDFF88FFE5FE8B2848FE4DFCAA" box="[359,427,799,825]" name="Japan" pageId="19" pageNumber="20">Japan</collectingCountry>
|
||
: Honshu, Kyushu.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C373D9F6FF88FFE5FF2B2814FC6BFAEA" pageId="19" pageNumber="20" type="discussion">
|
||
<paragraph id="8BD68A7DFF88FFE5FF2B2814FD63FBEE" blockId="19.[151,1437,151,1401]" pageId="19" pageNumber="20">
|
||
<emphasis id="B91D566FFF88FFE5FF2B2814FEDDFCCE" bold="true" box="[199,315,835,861]" pageId="19" pageNumber="20">Remarks.</emphasis>
|
||
The aedeagus of the only known male of
|
||
<taxonomicName id="4C69F1FEFF88FFE5FCF32814FC42FCCE" authorityName="Sharp" authorityYear="1886" box="[799,932,835,861]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="oscillans">
|
||
<emphasis id="B91D566FFF88FFE5FCF32814FC42FCCE" box="[799,932,835,861]" italics="true" pageId="19" pageNumber="20">E. oscillans</emphasis>
|
||
</taxonomicName>
|
||
has been partly damaged during a previous, unskillful preparation, and the true shape of the dorsal mediodistal plate remains unknown. It closely resembles the aedeagus illustrated by Hoshina (2020) for
|
||
<taxonomicName id="4C69F1FEFF88FFE5FD5228DCFCC2FC36" box="[702,804,907,933]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="akane">
|
||
<emphasis id="B91D566FFF88FFE5FD5228DCFCC2FC36" box="[702,804,907,933]" italics="true" pageId="19" pageNumber="20">E. akane</emphasis>
|
||
</taxonomicName>
|
||
, based on a male collected on Shikoku. However, the shapes and arrangement of the endophallic sclerites in
|
||
<taxonomicName id="4C69F1FEFF88FFE5FCE028F8FC97FC5A" box="[780,881,943,969]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="akane">
|
||
<emphasis id="B91D566FFF88FFE5FCE028F8FC97FC5A" box="[780,881,943,969]" italics="true" pageId="19" pageNumber="20">E. akane</emphasis>
|
||
</taxonomicName>
|
||
, if illustrated correctly, seem different (although also as strikingly asymmetrical as those in
|
||
<taxonomicName id="4C69F1FEFF88FFE5FD962884FD18FC7E" authorityName="Sharp" authorityYear="1886" box="[634,766,979,1005]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="oscillans">
|
||
<emphasis id="B91D566FFF88FFE5FD962884FD18FC7E" box="[634,766,979,1005]" italics="true" pageId="19" pageNumber="20">E. oscillans</emphasis>
|
||
</taxonomicName>
|
||
). According to Hoshina (2020), adults of
|
||
<taxonomicName id="4C69F1FEFF88FFE5FB392884FADEFC7E" box="[1237,1336,979,1005]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="akane">
|
||
<emphasis id="B91D566FFF88FFE5FB392884FADEFC7E" box="[1237,1336,979,1005]" italics="true" pageId="19" pageNumber="20">E. akane</emphasis>
|
||
</taxonomicName>
|
||
measure
|
||
<quantity id="4C912798FF88FFE5FF7B28A0FEDDFB82" box="[151,315,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.4249999999999998" metricValueMax="1.43" metricValueMin="1.42" pageId="19" pageNumber="20" unit="mm" value="1.4249999999999998" valueMax="1.43" valueMin="1.42">1.42‒1.43 mm</quantity>
|
||
of body length and have inconspicuous punctures on the elytra. In contrast, the male and female of
|
||
<taxonomicName id="4C69F1FEFF88FFE5FF7B2F4CFEFCFBA6" authorityName="Sharp" authorityYear="1886" box="[151,282,1051,1077]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="oscillans">
|
||
<emphasis id="B91D566FFF88FFE5FF7B2F4CFEFCFBA6" box="[151,282,1051,1077]" italics="true" pageId="19" pageNumber="20">E. oscillans</emphasis>
|
||
</taxonomicName>
|
||
examined during the present study measure
|
||
<quantity id="4C912798FF88FFE5FCE42F4CFC4BFBA6" box="[776,941,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" metricValueMax="1.2" metricValueMin="1.14" pageId="19" pageNumber="20" unit="mm" value="1.17" valueMax="1.2" valueMin="1.14">1.14‒1.20 mm</quantity>
|
||
, and have strikingly dense, deep and distinct elytral punctures. Therefore, it seems that these are two distinct species. The conspicuous elytral punctation of
|
||
<taxonomicName id="4C69F1FEFF88FFE5FA682F68FF1DFBEE" authorityName="Sharp" authorityYear="1886" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="oscillans">
|
||
<emphasis id="B91D566FFF88FFE5FA682F68FF1DFBEE" italics="true" pageId="19" pageNumber="20">E. oscillans</emphasis>
|
||
</taxonomicName>
|
||
is unique among Japanese species.
|
||
</paragraph>
|
||
<paragraph id="8BD68A7DFF88FFE5FF2B2FD0FA93FB56" blockId="19.[151,1437,151,1401]" pageId="19" pageNumber="20">
|
||
<taxonomicName id="4C69F1FEFF88FFE5FF2B2FD0FE44FB32" authorityName="Sharp" authorityYear="1886" box="[199,418,1159,1185]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="oscillans">
|
||
<emphasis id="B91D566FFF88FFE5FF2B2FD0FE44FB32" box="[199,418,1159,1185]" italics="true" pageId="19" pageNumber="20">Euconnus oscillans</emphasis>
|
||
</taxonomicName>
|
||
is here placed in
|
||
<taxonomicName id="4C69F1FEFF88FFE5FD802FD0FD3CFB33" box="[620,730,1159,1184]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91D566FFF88FFE5FD802FD0FD3CFB33" box="[620,730,1159,1184]" italics="true" pageId="19" pageNumber="20">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
(s. str.), based on a tetramerous antennal club, pronotum bellshaped, broadest behind middle, lack of the median antebasal pronotal pit and a non-emarginate aedeagal apex.
|
||
</paragraph>
|
||
<paragraph id="8BD68A7DFF88FFE5FF2B2F98FC6BFAEA" blockId="19.[151,1437,151,1401]" pageId="19" pageNumber="20">
|
||
<materialsCitation id="3B018020FF88FFE5FF2B2F98FC6CFAEA" ID-GBIF-Occurrence="3454958306" collectorName="SC Honshu" county="southern Kyushu" location="Mt. Kasuga" municipality="Mt. Kasuga" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Kumamoto" typeStatus="holotype">
|
||
The
|
||
<typeStatus id="54D234DFFF88FFE5FF152F98FEBDFB7A" box="[249,347,1231,1257]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
male of
|
||
<taxonomicName id="4C69F1FEFF88FFE5FE512F98FD70FB7A" authorityName="Sharp" authorityYear="1886" box="[445,662,1231,1257]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="oscillans">
|
||
<emphasis id="B91D566FFF88FFE5FE512F98FD70FB7A" box="[445,662,1231,1257]" italics="true" pageId="19" pageNumber="20">Euconnus oscillans</emphasis>
|
||
</taxonomicName>
|
||
comes from Hitoyoshi,
|
||
<collectingRegion id="49AD449FFF88FFE5FC4F2F98FB7BFB7A" box="[931,1181,1231,1257]" country="Japan" name="Kumamoto" pageId="19" pageNumber="20">Kumamoto Prefecture</collectingRegion>
|
||
,
|
||
<collectingCounty id="62B7F2F1FF88FFE5FB442F98FA8FFB7A" box="[1192,1385,1231,1257]" pageId="19" pageNumber="20">southern Kyushu</collectingCounty>
|
||
, and collecting circumstances remain unknown. The only additional specimen available to my study was found in
|
||
<collectingRegion id="49AD449FFF88FFE5FA8A2FA4FA7AFA9E" box="[1382,1436,1267,1293]" country="Japan" name="Nara" pageId="19" pageNumber="20">Nara</collectingRegion>
|
||
City, (
|
||
<collectingRegion id="49AD449FFF88FFE5FF302E40FE74FAA2" box="[220,402,1303,1329]" country="Japan" name="Nara" pageId="19" pageNumber="20">Nara Prefecture</collectingRegion>
|
||
,
|
||
<collectorName id="269CEFABFF88FFE5FE712E40FDC4FAA1" box="[413,546,1303,1330]" pageId="19" pageNumber="20">SC Honshu</collectorName>
|
||
), nearly
|
||
<quantity id="4C912798FF88FFE5FD682E41FD3FFAA2" box="[644,729,1302,1329]" metricMagnitude="5" metricUnit="m" metricValue="5.5" pageId="19" pageNumber="20" unit="km" value="550.0">550 km</quantity>
|
||
NE of the
|
||
<emphasis id="B91D566FFF88FFE5FCBA2E40FC3CFAA3" box="[854,986,1303,1328]" italics="true" pageId="19" pageNumber="20">terra typica</emphasis>
|
||
. Unfortunately, no additional collecting data are available, except for near
|
||
<collectingMunicipality id="6BB21007FF88FFE5FDF42E6CFD78FAC6" box="[536,670,1339,1365]" pageId="19" pageNumber="20">Mt. Kasuga</collectingMunicipality>
|
||
, Tullgren. This suggests sifted leaf litter, as
|
||
<location id="8EB6DCA6FF88FFE5FB672E6CFAE8FAC6" LSID="urn:lsid:plazi:treatment:03C03B6BFF8AFFE5FF7B2F4CFC6BFAEA:8EB6DCA6FF88FFE5FB672E6CFAE8FAC6" box="[1163,1294,1339,1365]" county="southern Kyushu" municipality="Mt. Kasuga" name="Mt. Kasuga" pageId="19" pageNumber="20" stateProvince="Kumamoto">Mt. Kasuga</location>
|
||
east of
|
||
<collectingRegion id="49AD449FFF88FFE5FA8A2E6CFA7AFAC6" box="[1382,1436,1339,1365]" country="Japan" name="Nara" pageId="19" pageNumber="20">Nara</collectingRegion>
|
||
is covered with and surrounded by the
|
||
<location id="8EB6DCA6FF88FFE5FDA62E08FC6CFAEA" LSID="urn:lsid:plazi:treatment:03C03B6BFF8AFFE5FF7B2F4CFC6BFAEA:8EB6DCA6FF88FFE5FDA62E08FC6CFAEA" box="[586,906,1375,1401]" county="southern Kyushu" municipality="Mt. Kasuga" name="Mt. Kasuga Primeval Forest" pageId="19" pageNumber="20" stateProvince="Kumamoto">Mt. Kasuga Primeval Forest</location>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |