treatments-xml/data/03/B5/3F/03B53F6A2952D64BFF59CEF880DB3EC6.xml

<document id="3F345956CEBE07AFD273BDA302C795A1" ID-CLB-Dataset="29841" ID-DOI="10.11646/zootaxa.4438.2.6" ID-GBIF-Dataset="fcb2bfd4-caf1-4467-b6c4-a9d5ad013e17" ID-ISSN="1175-5326" ID-Zenodo-Dep="1294774" ID-ZooBank="5D28883B-01C3-426B-9117-9F946D5E86DB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529568931826" checkinUser="plazi" docAuthor="Nguyen, Sang Ngoc, Nguyen, Luan Thanh, Nguyen, Vu Dang Hoang, Orlov, Nikolai L. &amp; Murphy, Robert W." docDate="2018" docId="03B53F6A2952D64BFF59CEF880DB3EC6" docLanguage="en" docName="zootaxa.4438.2.6.pdf.imf" docOrigin="Zootaxa 4438 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphenomorphus yersini Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="323" masterDocId="FF8C47122950D641FFCEC845827F3C43" masterDocTitle="A new skink of the genus Sphenomorphus Fitzinger, 1843 (Squamata: Scincidae) from Hon Ba Nature Reserve, southern Vietnam" masterLastPageNumber="326" masterPageNumber="313" pageNumber="315" updateTime="1698550208823" updateUser="ExternalLinkService">
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<mods:title id="9EAF6D2C2A3C500EBAEC25EEC9F4C9C0">A new skink of the genus Sphenomorphus Fitzinger, 1843 (Squamata: Scincidae) from Hon Ba Nature Reserve, southern Vietnam</mods:title>
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<mods:namePart id="78E6317CE6DC5E67645A4AA2B1495281">Nguyen, Sang Ngoc</mods:namePart>
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<mods:namePart id="23B42AFCDDDF8F889366870EE947B9AE">Nguyen, Luan Thanh</mods:namePart>
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<mods:namePart id="C19D3CC79D94E22E16E4AA1C2A783B89">Orlov, Nikolai L.</mods:namePart>
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<treatment id="03B53F6A2952D64BFF59CEF880DB3EC6" ID-DOI="http://doi.org/10.5281/zenodo.5964244" ID-GBIF-Taxon="146016102" ID-Zenodo-Dep="5964244" LSID="urn:lsid:plazi:treatment:03B53F6A2952D64BFF59CEF880DB3EC6" httpUri="http://treatment.plazi.org/id/03B53F6A2952D64BFF59CEF880DB3EC6" lastPageId="10" lastPageNumber="323" pageId="2" pageNumber="315">
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<emphasis id="B968526E2952D643FF59CEF880643A94" bold="true" box="[151,539,1725,1751]" pageId="2" pageNumber="315">
<taxonomicName id="4C1CF5FF2952D643FF59CEF883C93A94" authority="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy, 2018" authorityName="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy" authorityYear="2018" box="[151,438,1725,1751]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="315" phylum="Chordata" rank="species" species="yersini" status="sp. nov.">
<emphasis id="B968526E2952D643FF59CEF883C93A94" bold="true" box="[151,438,1725,1751]" italics="true" pageId="2" pageNumber="315">Sphenomorphus yersini</emphasis>
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<taxonomicNameLabel id="A25BEF152952D643FE73CEF880643A94" box="[445,539,1725,1751]" pageId="2" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="132792F92952D643FF51CEA583763ABB" box="[159,265,1760,1785]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1671,1693]" captionStartId-1="4.[151,250,1774,1796]" captionStartId-2="5.[151,250,1951,1973]" captionStartId-3="6.[151,250,1668,1690]" captionTargetBox-0="[151,1436,507,1655]" captionTargetBox-1="[172,1431,1011,1758]" captionTargetBox-2="[161,1433,497,1934]" captionTargetBox-3="[151,1436,232,1613]" captionTargetId-0="figure@3.[151,1436,507,1655]" captionTargetId-1="figure@4.[172,1431,1011,1758]" captionTargetId-2="figure@5.[161,1433,497,1934]" captionTargetId-3="figure@6.[151,1436,232,1616]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Holotype of Sphenomorphus yersini sp. nov., ITBCZ 5685. A, dorsolateral view; B, ventral view; C&amp;D, dorsal and lateral views of head; E, lower eyelid scaly; F, two asymmetrical hemipenes in preservation." captionText-1="FIGURE 2. Paratypes of Sphenomorphus yersini sp. nov. A&amp;B, ITBCZ 5684, male; C&amp;D, ITBCZ 5686, female. Scales in lower eyelid can be observed in figures B and D." captionText-2="FIGURE 3. Hemipenes of paratype ITBCZ 5684 of Sphenomorphus yersini sp. nov. A&amp;B, sulcate and asulcate views of the right hemipenis, respectively (fresh); B&amp;C, asulcate and sulcate views of left hemipenis, respectively (fresh); E&amp;F, the two hemipenes in preservation." captionText-3="FIGURE 4. Released referred specimen HBA 44 of Sphenomorphus yersini sp. nov. A–C, living specimen; D–H, midbody scale rows illustrated and checked via detail photos. Photographs detail five parts. Red spots are marks on midbody; blue dots illustrate counting of 33 longitudinal scale rows." httpUri-0="https://zenodo.org/record/3095246/files/figure.png" httpUri-1="https://zenodo.org/record/3095250/files/figure.png" httpUri-2="https://zenodo.org/record/3095256/files/figure.png" httpUri-3="https://zenodo.org/record/3095268/files/figure.png" pageId="2" pageNumber="315">Figs. 1–4</figureCitation>
)
</paragraph>
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<paragraph id="8BA38E7C2952D643FF59CF6D83ED3BCB" blockId="2.[151,1436,1832,2037]" pageId="2" pageNumber="315">
<materialsCitation id="3B7484212952D643FF59CF6D83ED3BCB" ID-GBIF-Occurrence="1851020268" collectingDate="2016-10-16" collectionCode="ITBCZ" collectorName="Hon Ba NR &amp; L. T. Nguyen &amp; V. D. H. Nguyen &amp; S. N. Nguyen" country="Vietnam" elevation="1162" latitude="12.136945" location="Khanh Hoa Province" longLatPrecision="21" longitude="108.96083" pageId="2" pageNumber="315" specimenCode="ITBCZ 5685" specimenCount="1" stateProvince="Khanh Hoa Province" typeStatus="holotype">
<emphasis id="B968526E2952D643FF59CF6D837B3B02" bold="true" box="[151,260,1832,1857]" pageId="2" pageNumber="315">
<typeStatus id="54A730DE2952D643FF59CF6D837B3B02" box="[151,260,1832,1857]" pageId="2" pageNumber="315" type="holotype">Holotype</typeStatus>
</emphasis>
. 
<specimenCode id="DBBA26072952D643FED8CF6D83D13B02" box="[278,430,1832,1857]" collectionCode="ITBCZ" pageId="2" pageNumber="315">ITBCZ 5685</specimenCode>
, adult male, collected from 
<collectorName id="26E9EBAA2952D643FD32CF6C81F03B03" box="[764,911,1833,1856]" pageId="2" pageNumber="315">Hon Ba NR</collectorName>
., 
<collectingRegion id="49D8409E2952D643FC6DCF6D86E53B02" box="[931,1178,1832,1857]" country="Vietnam" name="Khanh Hoa" pageId="2" pageNumber="315">
Khanh 
<collectingRegion id="49D8409E2952D643FC37CF6C86E53B02" box="[1017,1178,1832,1857]" inferredBySuffix="true" pageId="2" pageNumber="315">Hoa Province</collectingRegion>
</collectingRegion>
, 
<collectingCountry id="F30BCEEC2952D643FB67CF6D87743B02" box="[1193,1291,1832,1857]" name="Vietnam" pageId="2" pageNumber="315">Vietnam</collectingCountry>
; coordinates 
<geoCoordinate id="EE28E8BB2952D643FF59CF08836B3B26" box="[151,276,1869,1893]" direction="north" orientation="latitude" pageId="2" pageNumber="315" precision="15" value="12.136945">12°8’13”N</geoCoordinate>
, 
<geoCoordinate id="EE28E8BB2952D643FEEBCF0983C23B26" box="[293,445,1868,1893]" direction="east" orientation="longitude" pageId="2" pageNumber="315" precision="15" value="108.96083">108°57’39”E</geoCoordinate>
; 
<elevation id="0031694F2952D643FE00CF0980A33B26" box="[462,732,1868,1893]" metricMagnitude="3" metricUnit="m" metricValue="1.162" pageId="2" pageNumber="315" unit="m" value="1162.0">
elevation 
<quantity id="4CE423992952D643FD8FCF0980E43B27" box="[577,667,1868,1893]" metricMagnitude="3" metricUnit="m" metricValue="1.162" pageId="2" pageNumber="315" unit="m" value="1162.0">1162 m</quantity>
a.s.l.
</elevation>
by 
<collectorName id="26E9EBAA2952D643FCDFCF0881D83B26" box="[785,935,1869,1893]" pageId="2" pageNumber="315">L.T. Nguyen</collectorName>
, 
<collectorName id="26E9EBAA2952D643FC76CF0886113B26" box="[952,1134,1869,1893]" pageId="2" pageNumber="315">V.D.H. Nguyen</collectorName>
, and 
<collectorName id="26E9EBAA2952D643FB7ACF0887343B26" box="[1204,1355,1869,1893]" pageId="2" pageNumber="315">S.N. Nguyen</collectorName>
on 
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<collectingDate id="EFE651542952D643FA4ECF09834F3BCA" pageId="2" pageNumber="315" value="2016-10-16">16 October 2016</collectingDate>
</date>
(
<figureCitation id="132792F92952D643FEF1CF3583FE3BCB" box="[319,385,1904,1929]" captionStart="FIGURE 1" captionStartId="3.[151,250,1671,1693]" captionTargetBox="[151,1436,507,1655]" captionTargetId="figure@3.[151,1436,507,1655]" captionTargetPageId="3" captionText="FIGURE 1. Holotype of Sphenomorphus yersini sp. nov., ITBCZ 5685. A, dorsolateral view; B, ventral view; C&amp;D, dorsal and lateral views of head; E, lower eyelid scaly; F, two asymmetrical hemipenes in preservation." httpUri="https://zenodo.org/record/3095246/files/figure.png" pageId="2" pageNumber="315">Fig. 1</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA38E7C2952D643FF09CFD1867A3BB6" blockId="2.[151,1436,1832,2037]" pageId="2" pageNumber="315">
<emphasis id="B968526E2952D643FF09CFD183403BEE" bold="true" box="[199,319,1940,1965]" pageId="2" pageNumber="315">
<typeStatus id="54A730DE2952D643FF09CFD183403BEE" box="[199,319,1940,1965]" pageId="2" pageNumber="315" type="paratype">Paratypes</typeStatus>
</emphasis>
. Two specimens, also collected from Hon Ba NR by L.T. Nguyen, V.D.H. Nguyen, and S.N. Nguyen: 
<collectionCode id="ED0D16B92952D643FF31CFFD832F3B93" box="[255,336,1976,2000]" pageId="2" pageNumber="315">ITBCZ</collectionCode>
5686, adult female (
<figureCitation id="132792F92952D643FDF5CFFD80C03B93" box="[571,703,1976,2001]" captionStart="FIGURE 2" captionStartId="4.[151,250,1774,1796]" captionTargetBox="[172,1431,1011,1758]" captionTargetId="figure@4.[172,1431,1011,1758]" captionTargetPageId="4" captionText="FIGURE 2. Paratypes of Sphenomorphus yersini sp. nov. A&amp;B, ITBCZ 5684, male; C&amp;D, ITBCZ 5686, female. Scales in lower eyelid can be observed in figures B and D." httpUri="https://zenodo.org/record/3095250/files/figure.png" pageId="2" pageNumber="315">Fig. 2C&amp;D</figureCitation>
), and 
<collectionCode id="ED0D16B92952D643FCCACFFD812A3B93" box="[772,853,1976,2000]" pageId="2" pageNumber="315">ITBCZ</collectionCode>
5684, adult male (
<figureCitation id="132792F92952D643FBE4CFFD86CA3B93" box="[1066,1205,1976,2001]" captionStart="FIGURE 2" captionStartId="4.[151,250,1774,1796]" captionTargetBox="[172,1431,1011,1758]" captionTargetId="figure@4.[172,1431,1011,1758]" captionTargetPageId="4" captionText="FIGURE 2. Paratypes of Sphenomorphus yersini sp. nov. A&amp;B, ITBCZ 5684, male; C&amp;D, ITBCZ 5686, female. Scales in lower eyelid can be observed in figures B and D." httpUri="https://zenodo.org/record/3095250/files/figure.png" pageId="2" pageNumber="315">Figs. 2A&amp;B</figureCitation>
and 
<figureCitation id="132792F92952D643FB23CFFD86833B93" box="[1261,1276,1976,2000]" captionStart="FIGURE 3" captionStartId="5.[151,250,1951,1973]" captionTargetBox="[161,1433,497,1934]" captionTargetId="figure@5.[161,1433,497,1934]" captionTargetPageId="5" captionText="FIGURE 3. Hemipenes of paratype ITBCZ 5684 of Sphenomorphus yersini sp. nov. A&amp;B, sulcate and asulcate views of the right hemipenis, respectively (fresh); B&amp;C, asulcate and sulcate views of left hemipenis, respectively (fresh); E&amp;F, the two hemipenes in preservation." httpUri="https://zenodo.org/record/3095256/files/figure.png" pageId="2" pageNumber="315">3</figureCitation>
), collected on 
<date id="FFA2A8BC2952D643FF59CF9883293BB6" box="[151,342,2012,2037]" pageId="2" pageNumber="315" value="2016-10-14">14 October 2016</date>
, coordinates 
<geoCoordinate id="EE28E8BB2952D643FE27CF9880183BB6" box="[489,615,2013,2037]" direction="north" orientation="latitude" pageId="2" pageNumber="315" precision="15" value="12.139444">12°8’22”N</geoCoordinate>
, 
<geoCoordinate id="EE28E8BB2952D643FDBCCF9981763BB6" box="[626,777,2012,2037]" direction="east" orientation="longitude" pageId="2" pageNumber="315" precision="15" value="108.96833">108°58’06”E</geoCoordinate>
; elevation 
<quantity id="4CE423992952D643FC4DCF9881B53BB7" box="[899,970,2013,2037]" metricMagnitude="2" metricUnit="m" metricValue="9.32" pageId="2" pageNumber="315" unit="m" value="932.0">932 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="C306DDF72953D642FF09C8D280CC3C96" pageId="3" pageNumber="316" type="description">
<paragraph id="8BA38E7C2953D642FF09C8D280CC3C96" blockId="3.[151,1436,151,464]" pageId="3" pageNumber="316">
<emphasis id="B968526E2953D642FF09C8D283D43CF3" bold="true" box="[199,427,151,176]" pageId="3" pageNumber="316">Referred specimen</emphasis>
. HBA 44 (released, 
<figureCitation id="132792F92953D642FD6BC8D280923CF3" box="[677,749,151,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,1668,1690]" captionTargetBox="[151,1436,232,1613]" captionTargetId="figure@6.[151,1436,232,1616]" captionTargetPageId="6" captionText="FIGURE 4. Released referred specimen HBA 44 of Sphenomorphus yersini sp. nov. A–C, living specimen; D–H, midbody scale rows illustrated and checked via detail photos. Photographs detail five parts. Red spots are marks on midbody; blue dots illustrate counting of 33 longitudinal scale rows." httpUri="https://zenodo.org/record/3095268/files/figure.png" pageId="3" pageNumber="316">Fig. 4</figureCitation>
), adult female, collected on 
<date id="FFA2A8BC2953D642FB8CC8DD87743CF3" box="[1090,1291,151,176]" pageId="3" pageNumber="316" value="2016-10-19">19 October 2016</date>
, coordinates 12°8’22”N, 108°58’06”E; elevation 
<quantity id="4CE423992953D642FDFEC8F980083C97" box="[560,631,188,212]" metricMagnitude="2" metricUnit="m" metricValue="9.32" pageId="3" pageNumber="316" unit="m" value="932.0">932 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="C306DDF72953D642FF09C89A870A3D93" pageId="3" pageNumber="316" type="diagnosis">
<paragraph id="8BA38E7C2953D642FF09C89A870A3D93" blockId="3.[151,1436,151,464]" pageId="3" pageNumber="316">
<emphasis id="B968526E2953D642FF09C89A83453CBB" bold="true" box="[199,314,223,248]" pageId="3" pageNumber="316">Diagnosis</emphasis>
. 
<taxonomicName id="4C1CF5FF2953D642FE87C8A5802E3CB4" authority="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy, 2018" authorityName="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy" authorityYear="2018" box="[329,593,224,248]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="316" phylum="Chordata" rank="species" species="yersini" status="sp. nov.">
<emphasis id="B968526E2953D642FE87C8A5802E3CB4" box="[329,593,224,248]" italics="true" pageId="3" pageNumber="316">Sphenomorphus yersini</emphasis>
</taxonomicName>
<emphasis id="B968526E2953D642FD97C89A80CC3CBB" bold="true" box="[601,691,223,248]" pageId="3" pageNumber="316">
<taxonomicNameLabel id="A25BEF152953D642FD97C89A80CC3CBB" box="[601,691,223,248]" pageId="3" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from all of its congeners by a combination of the following morphological characters: size in adults (SVL) up to 
<quantity id="4CE423992953D642FCA6C94181C43D5F" box="[872,955,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="316" unit="mm" value="56.0">56 mm</quantity>
; TaL/SVL ratio 1.81; toes reach to fingers when limbs adpressed; 32–34 smooth midbody scale rows; 61–69 paravertebral scales; 58–67 ventral scale rows; 112 subcaudal scales; four, rarely five, supraoculars; prefrontals in broad contact with one another; two loreal scales; tympanum deeply sunk; smooth lamellae beneath Finger IV and Toe 
<date id="FFA2A8BC2953D642FC24C93586DB3DCB" box="[1002,1188,367,392]" pageId="3" pageNumber="316" value="2010-04-12" valueMax="2010-04-18" valueMin="2010-04-12">IV 10–12 and 18</date>
–20, respectively; two enlarged precloacal scales; hemipenis smooth, deeply forked, asymmetrical with a long lobe and another short; black and interruptive dorsolateral line; lateral side and lower part of head, neck, and tail orange to red in male.
</paragraph>
</subSubSection>
<caption id="DF63DEF42953D642FF59CEC286173AFF" httpUri="https://zenodo.org/record/3095246/files/figure.png" pageId="3" pageNumber="316" startId="3.[151,250,1671,1693]" targetBox="[151,1436,507,1655]" targetPageId="3">
<paragraph id="8BA38E7C2953D642FF59CEC286173AFF" blockId="3.[151,1436,1671,1724]" pageId="3" pageNumber="316">
<emphasis id="B968526E2953D642FF59CEC2836B3ADF" bold="true" box="[151,276,1671,1693]" pageId="3" pageNumber="316">FIGURE 1.</emphasis>
Holotype of 
<taxonomicName id="4C1CF5FF2953D642FE51CEC280F03ADE" authority="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy, 2018" authorityName="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy" authorityYear="2018" box="[415,655,1671,1693]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="316" phylum="Chordata" rank="species" species="yersini" status="sp. nov.">
<emphasis id="B968526E2953D642FE51CEC280F03ADE" box="[415,655,1671,1693]" italics="true" pageId="3" pageNumber="316">Sphenomorphus yersini</emphasis>
</taxonomicName>
<emphasis id="B968526E2953D642FD56CEC280953ADF" bold="true" box="[664,746,1671,1692]" pageId="3" pageNumber="316">
<taxonomicNameLabel id="A25BEF152953D642FD56CEC280953ADF" box="[664,746,1671,1692]" pageId="3" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ITBCZ 5685. A, dorsolateral view; B, ventral view; C&amp;D, dorsal and lateral views of head; E, lower eyelid scaly; F, two asymmetrical hemipenes in preservation.
</paragraph>
</caption>
<subSubSection id="C306DDF72953D644FF09CEA7836C3D85" lastPageId="5" lastPageNumber="318" pageId="3" pageNumber="316" type="description">
<paragraph id="8BA38E7C2953D642FF09CEA781713B02" blockId="3.[151,1436,1762,2031]" pageId="3" pageNumber="316">
<emphasis id="B968526E2953D642FF09CEA783983AB8" bold="true" box="[199,487,1762,1787]" pageId="3" pageNumber="316">
Description of 
<typeStatus id="54A730DE2953D642FEB7CEA7839D3AB8" box="[377,482,1762,1787]" pageId="3" pageNumber="316" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male; head small and elongate (HW/HL = 0.7); SVL 
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; tail long (TaL/ SVL = 1.81), 
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in length; lower eyelid scaly; tympanum deeply sunk with a prominent oblique edge; limbs pentadactyl, toes reach to fingers when limbs adpressed.
</paragraph>
<paragraph id="8BA38E7C2953D645FF09CF0E83A33DC3" blockId="3.[151,1436,1762,2031]" lastBlockId="4.[151,1437,151,974]" lastPageId="4" lastPageNumber="317" pageId="3" pageNumber="316">Head scalation smooth; rostral convex, distinctly visible from above, in broad contact with frontonasal, which is broader than long; no supranasals; prefrontals in broad contact with one another; 5 supraoculars on left side and 4 on right side; a pair of frontoparietals, shorter than frontal; frontal narrowing posteriorly, longer than wide, bordered laterally by first two supraoculars, anteriorly by prefrontals, and posteriorly by frontoparietals; frontoparietals in contact with the second to fifth supraoculars on left side and the second to fourth supraoculars on right side; parietals in contact posteriorly, behind the interparietal; no nuchal; 7 supralabials on both sides, the fifth and sixth below the eye; 2 loreals, posterior loreal larger than anterior; nostril in center of nasal, which is in contact with the first supralabial, rostral, anterior loreal, and frontonasal; 8 supraciliaries, first largest; 1 anterior temporal, in contact with sixth and seventh supralabials; 2 secondary temporals, lower temporal overlapping upper one and in contact with seventh supralabial, upper temporal larger, in contact with parietal; lower eyelid scaly, 2 scales in center larger than scales in posterior and anterior areas; 6 infralabials, first pair medially in contact with each other; 1 anterior and 2 postmentals.</paragraph>
<paragraph id="8BA38E7C2954D645FF09C9CF865F3DAB" blockId="4.[151,1437,151,974]" pageId="4" pageNumber="317">Dorsal scales smooth, not larger than lateral and ventral scales; 34 midbody scale rows; 61 paravertebral scales; ventral scales smooth, in 60 rows; 112 subcaudal scales, slightly enlarged posteriorly; 11 and 20 smooth lamellae beneath finger IV and toe IV, respectively; 2 enlarged precloacal scales.</paragraph>
<paragraph id="8BA38E7C2954D645FF09C9B683C03ED5" blockId="4.[151,1437,151,974]" pageId="4" pageNumber="317">
<emphasis id="B968526E2954D645FF09C9B6833C3E49" box="[199,323,499,522]" italics="true" pageId="4" pageNumber="317">Hemipenes</emphasis>
. Each hemipenis deeply forked and asymmetrical with two smooth lobes. In fully everted position, the inner lobe is shorter and the outer lobe is much longer. Clear sulcus spermaticus starts from the base, divides into two branches at the fork and extends to the tips of the two lobes; body of the long lobe forming regular transversal shallow grooves and ending with terminal papillae. Short lobe obtuse, not forming regular transversal shallow grooves (
<figureCitation id="132792F92954D645FE93CA3883CE3ED5" box="[349,433,637,662]" captionStart="FIGURE 1" captionStartId="3.[151,250,1671,1693]" captionTargetBox="[151,1436,507,1655]" captionTargetId="figure@3.[151,1436,507,1655]" captionTargetPageId="3" captionText="FIGURE 1. Holotype of Sphenomorphus yersini sp. nov., ITBCZ 5685. A, dorsolateral view; B, ventral view; C&amp;D, dorsal and lateral views of head; E, lower eyelid scaly; F, two asymmetrical hemipenes in preservation." httpUri="https://zenodo.org/record/3095246/files/figure.png" pageId="4" pageNumber="317">Fig. 1F</figureCitation>
).
</paragraph>
<paragraph id="8BA38E7C2954D645FF09CAE583ED3F8D" blockId="4.[151,1437,151,974]" pageId="4" pageNumber="317">
<emphasis id="B968526E2954D645FF09CAE5833D3EFB" box="[199,322,672,696]" italics="true" pageId="4" pageNumber="317">Coloration</emphasis>
. In life, overall dorsal coloration black (Hex #000000) to dark brown (Hex #654321) with two interrupted copper (Hex #B87333) dorsal lines in margins of dorsum extending from neck to base of tail; black and interrupted lateral lines under the copper ones; dorsum with irregular black spots; lateral side and lower part of head, neck, and tail orange (Hex #FF6600) to red (Hex #FE2712); venter from chest to tail base yellow (Hex #FFD300) to cream (Hex #FFFDD0); upper side of limbs black with bright dots. Lateral margins of supralabials and infralabials with black blotches. Free margin of upper eyelid orange and the margin of lower eyelid yellow (Hex #FFD300). Eyes black. In preservation, color fades; orange and yellow disappear; overall dorsal coloration black with two interrupted brighter lines in margins of dorsum extending from neck to base of tail; venter zebra white (Hex #F5F5F5).
</paragraph>
<caption id="DF63DEF42954D645FF59CEAB80F93B60" httpUri="https://zenodo.org/record/3095250/files/figure.png" pageId="4" pageNumber="317" startId="4.[151,250,1774,1796]" targetBox="[172,1431,1011,1758]" targetPageId="4">
<paragraph id="8BA38E7C2954D645FF59CEAB80F93B60" blockId="4.[151,1436,1774,1827]" pageId="4" pageNumber="317">
<emphasis id="B968526E2954D645FF59CEAB836A3B47" bold="true" box="[151,277,1774,1796]" pageId="4" pageNumber="317">FIGURE 2.</emphasis>
Paratypes of 
<taxonomicName id="4C1CF5FF2954D645FE6BCEAB80E93B46" authority="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy, 2018" authorityName="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy" authorityYear="2018" box="[421,662,1774,1797]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="317" phylum="Chordata" rank="species" species="yersini" status="sp. nov.">
<emphasis id="B968526E2954D645FE6BCEAB80E93B46" box="[421,662,1774,1797]" italics="true" pageId="4" pageNumber="317">Sphenomorphus yersini</emphasis>
</taxonomicName>
<emphasis id="B968526E2954D645FD50CEAA808F3B47" bold="true" box="[670,752,1775,1796]" pageId="4" pageNumber="317">
<taxonomicNameLabel id="A25BEF152954D645FD50CEAA808F3B47" box="[670,752,1775,1796]" pageId="4" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A&amp;B, ITBCZ 5684, male; C&amp;D, ITBCZ 5686, female. Scales in lower eyelid can be observed in figures B and D.
</paragraph>
</caption>
<paragraph id="8BA38E7C2954D645FF09CF0C80783BAE" blockId="4.[151,1436,1865,2029]" pageId="4" pageNumber="317">
<emphasis id="B968526E2954D645FF09CF0C83433B21" bold="true" box="[199,316,1865,1890]" pageId="4" pageNumber="317">Variation.</emphasis>
<tableCitation id="C69EBBC72954D645FE82CF0C83D93B21" box="[332,422,1865,1890]" captionStart="TABLE 2" captionStartId="10.[151,239,686,709]" captionText="TABLE 2. Measurements (in mm) and scalation of the types series of Sphenomorphus yersini sp. nov. See Material and Methods for abbreviations. Reg. = regenerated." pageId="4" pageNumber="317">Table 2</tableCitation>
summarizes variation in size and scalation of the 
<typeStatus id="54A730DE2954D645FBD8CF0C86043B21" box="[1046,1147,1865,1890]" pageId="4" pageNumber="317" type="holotype">holotype</typeStatus>
, 
<typeStatus id="54A730DE2954D645FB43CF0F86843B21" box="[1165,1275,1866,1890]" pageId="4" pageNumber="317" type="paratype">paratypes</typeStatus>
, and referred specimen. Supraoculars four in all specimens except for the left side of the 
<typeStatus id="54A730DE2954D645FC15CF2986423BC6" box="[987,1085,1900,1925]" pageId="4" pageNumber="317" type="holotype">holotype</typeStatus>
which bears 5 scales. Midbody scale rows vary from 32 to 34. Paravertebral and ventral scale rows range from 61 to 69 and from 58 to 67, respectively. Hemipenes of the second 
<typeStatus id="54A730DE2954D645FD92CFF780C23B89" box="[604,701,1970,1994]" pageId="4" pageNumber="317" type="paratype">paratype</typeStatus>
ITBCZ 5684 (
<figureCitation id="132792F92954D645FCA3CFF481CD3B89" box="[877,946,1969,1994]" captionStart="FIGURE 3" captionStartId="5.[151,250,1951,1973]" captionTargetBox="[161,1433,497,1934]" captionTargetId="figure@5.[161,1433,497,1934]" captionTargetPageId="5" captionText="FIGURE 3. Hemipenes of paratype ITBCZ 5684 of Sphenomorphus yersini sp. nov. A&amp;B, sulcate and asulcate views of the right hemipenis, respectively (fresh); B&amp;C, asulcate and sulcate views of left hemipenis, respectively (fresh); E&amp;F, the two hemipenes in preservation." httpUri="https://zenodo.org/record/3095256/files/figure.png" pageId="4" pageNumber="317">Fig. 3</figureCitation>
) similar to those of the 
<typeStatus id="54A730DE2954D645FB1FCFF4874C3B89" box="[1233,1331,1969,1994]" pageId="4" pageNumber="317" type="holotype">holotype</typeStatus>
in being deeply forked and asymmetrical.
</paragraph>
<paragraph id="8BA38E7C2955D644FF09C8D283743D54" blockId="5.[151,1436,151,454]" pageId="5" pageNumber="318">
<emphasis id="B968526E2955D644FF09C8D283FB3CF3" bold="true" box="[199,388,151,176]" pageId="5" pageNumber="318">Natural history.</emphasis>
All specimens were collected at night and under rotting leaf layer in evergreen forest (
<figureCitation id="132792F92955D644FA84C8D287F23CF3" box="[1354,1421,151,176]" captionStart="FIGURE 5" captionStartId="7.[151,250,1345,1367]" captionTargetBox="[152,1436,194,1324]" captionTargetId="figure@7.[151,1436,193,1324]" captionTargetPageId="7" captionText="FIGURE 5. Map (left) showing the type locality (red dot) of Sphenomorphus yersini sp. nov. in southern Vietnam and habitat (right) of the new species in Hon Ba Nature Reserve." httpUri="https://zenodo.org/record/3095280/files/figure.png" pageId="5" pageNumber="318">Fig. 5</figureCitation>
), elevations 
<quantity id="4CE423992955D644FEDEC8FE83273C91" box="[272,344,187,211]" metricMagnitude="2" metricUnit="m" metricValue="9.32" pageId="5" pageNumber="318" unit="m" value="932.0">932 m</quantity>
and 
<quantity id="4CE423992955D644FE5FC8FF83993C91" box="[401,486,186,211]" metricMagnitude="3" metricUnit="m" metricValue="1.162" pageId="5" pageNumber="318" unit="m" value="1162.0">1162 m</quantity>
a.s.l., between 19:00–23:30. However, the species was observed being active in the daytime and it may be diurnal. 
<taxonomicName id="4C1CF5FF2955D644FE3AC89880833CB6" authority="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy, 2018" authorityName="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy" authorityYear="2018" box="[500,764,221,245]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="318" phylum="Chordata" rank="species" species="yersini" status="sp. nov.">
<emphasis id="B968526E2955D644FE3AC89880833CB6" box="[500,764,221,245]" italics="true" pageId="5" pageNumber="318">Sphenomorphus yersini</emphasis>
</taxonomicName>
<emphasis id="B968526E2955D644FCCDC89881223CB5" bold="true" box="[771,861,221,246]" pageId="5" pageNumber="318">
<taxonomicNameLabel id="A25BEF152955D644FCCDC89881223CB5" box="[771,861,221,246]" pageId="5" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was observed to be sympatric with other congener, 
<taxonomicName id="4C1CF5FF2955D644FF59C945837B3D5B" baseAuthorityName="Gray" baseAuthorityYear="1853" box="[151,260,256,280]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="318" phylum="Chordata" rank="species" species="indicus">
<emphasis id="B968526E2955D644FF59C945837B3D5B" box="[151,260,256,280]" italics="true" pageId="5" pageNumber="318">S. indicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA38E7C2955D644FF09C967836C3D85" blockId="5.[151,1436,151,454]" pageId="5" pageNumber="318">
<emphasis id="B968526E2955D644FF09C96783C53D78" bold="true" box="[199,442,290,315]" pageId="5" pageNumber="318">Sexual dimorphism.</emphasis>
Males are larger than females (SVL 
<quantity id="4CE423992955D644FCBFC96781BC3D79" box="[881,963,290,315]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="318" unit="mm" value="56.0">56 mm</quantity>
vs. 
<quantity id="4CE423992955D644FC36C96786333D79" box="[1016,1100,290,315]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="318" unit="mm" value="52.0">52 mm</quantity>
, n=4) and have orange (Hex #FF6600) or red (Hex #FE2712) color on lateral sides and lower part of head, neck, and tail. The color on the lateral side in females faded to yellow (Hex #FFD300) or brown (Hex #964B00) with bright spots; lower part of head, limbs, and tail white (Hex #FFFFFF) to zebra white (Hex #F5F5F5); venter from neck to vent yellow (Hex #FCE883).
</paragraph>
</subSubSection>
<caption id="DF63DEF42955D644FF59CFDA83D73BB1" httpUri="https://zenodo.org/record/3095256/files/figure.png" pageId="5" pageNumber="318" startId="5.[151,250,1951,1973]" targetBox="[161,1433,497,1934]" targetPageId="5">
<paragraph id="8BA38E7C2955D644FF59CFDA83D73BB1" blockId="5.[151,1436,1951,2034]" pageId="5" pageNumber="318">
<emphasis id="B968526E2955D644FF59CFDA836C3BF7" bold="true" box="[151,275,1951,1973]" pageId="5" pageNumber="318">FIGURE 3.</emphasis>
Hemipenes of paratype ITBCZ 5684 of 
<taxonomicName id="4C1CF5FF2955D644FD78CFDA81D93BF6" authority="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy, 2018" authorityName="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy" authorityYear="2018" box="[694,934,1951,1973]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="318" phylum="Chordata" rank="species" species="yersini" status="sp. nov.">
<emphasis id="B968526E2955D644FD78CFDA81D93BF6" box="[694,934,1951,1973]" italics="true" pageId="5" pageNumber="318">Sphenomorphus yersini</emphasis>
</taxonomicName>
<emphasis id="B968526E2955D644FC63CFDA81803BF7" bold="true" box="[941,1023,1951,1972]" pageId="5" pageNumber="318">
<taxonomicNameLabel id="A25BEF152955D644FC63CFDA81803BF7" box="[941,1023,1951,1972]" pageId="5" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A&amp;B, sulcate and asulcate views of the right hemipenis, respectively (fresh); B&amp;C, asulcate and sulcate views of left hemipenis, respectively (fresh); E&amp;F, the two hemipenes in preservation.
</paragraph>
</caption>
<caption id="DF63DEF42956D647FF59CEC180FD3A94" httpUri="https://zenodo.org/record/3095268/files/figure.png" pageId="6" pageNumber="319" startId="6.[151,250,1668,1690]" targetBox="[151,1436,232,1613]" targetPageId="6">
<paragraph id="8BA38E7C2956D647FF59CEC180FD3A94" blockId="6.[151,1436,1668,1751]" pageId="6" pageNumber="319">
<emphasis id="B968526E2956D647FF59CEC1836B3ADA" bold="true" box="[151,276,1668,1690]" pageId="6" pageNumber="319">FIGURE 4.</emphasis>
Released referred specimen HBA 44 of 
<taxonomicName id="4C1CF5FF2956D647FD73CEC181D13AD9" authority="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy, 2018" authorityName="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy" authorityYear="2018" box="[701,942,1668,1690]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="319" phylum="Chordata" rank="species" species="yersini" status="sp. nov.">
<emphasis id="B968526E2956D647FD73CEC181D13AD9" box="[701,942,1668,1690]" italics="true" pageId="6" pageNumber="319">Sphenomorphus yersini</emphasis>
</taxonomicName>
<emphasis id="B968526E2956D647FC78CEC186773ADA" bold="true" box="[950,1032,1668,1689]" pageId="6" pageNumber="319">
<taxonomicNameLabel id="A25BEF152956D647FC78CEC186773ADA" box="[950,1032,1668,1689]" pageId="6" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A–C, living specimen; D–H, midbody scale rows illustrated and checked via detail photos. Photographs detail five parts. Red spots are marks on midbody; blue dots illustrate counting of 33 longitudinal scale rows.
</paragraph>
</caption>
<subSubSection id="C306DDF72956D647FF09CEBB83283B78" pageId="6" pageNumber="319" type="distribution">
<paragraph id="8BA38E7C2956D647FF09CEBB83283B78" blockId="6.[151,1437,1790,2031]" pageId="6" pageNumber="319">
<emphasis id="B968526E2956D647FF09CEBB831F3B54" bold="true" box="[199,352,1790,1815]" pageId="6" pageNumber="319">Distribution.</emphasis>
The new species is currently known only from Hon Ba NR, 
<collectingRegion id="49D8409E2956D647FBF9CEBB87523B54" box="[1079,1325,1790,1815]" country="Vietnam" name="Khanh Hoa" pageId="6" pageNumber="319">Khanh Hoa Province</collectingRegion>
, southern 
<collectingCountry id="F30BCEEC2956D647FF59CF6782893B78" box="[151,246,1826,1851]" name="Vietnam" pageId="6" pageNumber="319">Vietnam</collectingCountry>
(
<figureCitation id="132792F92956D647FECBCF6783363B78" box="[261,329,1826,1851]" captionStart="FIGURE 5" captionStartId="7.[151,250,1345,1367]" captionTargetBox="[152,1436,194,1324]" captionTargetId="figure@7.[151,1436,193,1324]" captionTargetPageId="7" captionText="FIGURE 5. Map (left) showing the type locality (red dot) of Sphenomorphus yersini sp. nov. in southern Vietnam and habitat (right) of the new species in Hon Ba Nature Reserve." httpUri="https://zenodo.org/record/3095280/files/figure.png" pageId="6" pageNumber="319">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C306DDF72956D64BFF09CF0380293DEF" lastPageId="10" lastPageNumber="323" pageId="6" pageNumber="319" type="etymology">
<paragraph id="8BA38E7C2956D647FF09CF03805F3BAC" blockId="6.[151,1437,1790,2031]" pageId="6" pageNumber="319">
<emphasis id="B968526E2956D647FF09CF0383393B1C" bold="true" box="[199,326,1862,1887]" pageId="6" pageNumber="319">Etymology</emphasis>
. We name this new species in honor of the famous physician and bacteriologist, Alexandre Yersin (
<date id="FFA2A8BC2956D647FF51CF2F835D3BC0" box="[159,290,1898,1923]" pageId="6" pageNumber="319" value="1863" valueMax="1943">1863–1943</date>
), who discovered the bacterium responsible for bubonic plague. Hon Ba NR associates with the name of Alexandre Yersin who built a research station on the top of the mountain and worked there. Currently, the research station has been reconstructed and opened to visitors. We recommend Yersin’s Forest Skink as the common name of this new species.
</paragraph>
<caption id="DF63DEF42957D646FF59CD0480CE3935" httpUri="https://zenodo.org/record/3095280/files/figure.png" pageId="7" pageNumber="320" startId="7.[151,250,1345,1367]" targetBox="[152,1436,194,1324]" targetPageId="7">
<paragraph id="8BA38E7C2957D646FF59CD0480CE3935" blockId="7.[151,1436,1345,1398]" pageId="7" pageNumber="320">
<emphasis id="B968526E2957D646FF59CD04836C3914" bold="true" box="[151,275,1345,1367]" pageId="7" pageNumber="320">FIGURE 5.</emphasis>
Map (left) showing the type locality (red dot) of 
<taxonomicName id="4C1CF5FF2957D646FCC4CD0481873914" authority="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy, 2018" authorityName="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy" authorityYear="2018" box="[778,1016,1345,1367]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="320" phylum="Chordata" rank="species" species="yersini" status="sp. nov.">
<emphasis id="B968526E2957D646FCC4CD0481873914" box="[778,1016,1345,1367]" italics="true" pageId="7" pageNumber="320">Sphenomorphus yersini</emphasis>
</taxonomicName>
<emphasis id="B968526E2957D646FC31CD07862F3914" bold="true" box="[1023,1104,1346,1367]" pageId="7" pageNumber="320">
<taxonomicNameLabel id="A25BEF152957D646FC31CD07862F3914" box="[1023,1104,1346,1367]" pageId="7" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in southern Vietnam and habitat (right) of the new species in Hon Ba Nature Reserve.
</paragraph>
</caption>
<paragraph id="8BA38E7C2957D64BFF09CDE780293DEF" blockId="7.[151,1437,1442,2007]" lastBlockId="10.[151,1436,151,645]" lastPageId="10" lastPageNumber="323" pageId="7" pageNumber="320">
<emphasis id="B968526E2957D646FF09CDE7831239F8" bold="true" box="[199,365,1442,1467]" pageId="7" pageNumber="320">Comparisons.</emphasis>
<taxonomicName id="4C1CF5FF2957D646FEB4CDE680F739F9" authority="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy, 2018" authorityName="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy" authorityYear="2018" box="[378,648,1443,1467]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="320" phylum="Chordata" rank="species" species="yersini" status="sp. nov.">
<emphasis id="B968526E2957D646FEB4CDE680F739F9" box="[378,648,1443,1467]" italics="true" pageId="7" pageNumber="320">Sphenomorphus yersini</emphasis>
</taxonomicName>
<emphasis id="B968526E2957D646FD5BCDE7808A39F8" bold="true" box="[661,757,1442,1467]" pageId="7" pageNumber="320">
<taxonomicNameLabel id="A25BEF152957D646FD5BCDE7808A39F8" box="[661,757,1442,1467]" pageId="7" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from its congeners in Indochina (
<collectingCountry id="F30BCEEC2957D646FB24CDE7873239F8" box="[1258,1357,1442,1467]" name="Vietnam" pageId="7" pageNumber="320">Vietnam</collectingCountry>
, 
<collectingCountry id="F30BCEEC2957D646FA90CDE687E839F9" box="[1374,1431,1443,1466]" name="Laos" pageId="7" pageNumber="320">Laos</collectingCountry>
, 
<collectingCountry id="F30BCEEC2957D646FF59CD838372399C" box="[151,269,1478,1503]" name="Cambodia" pageId="7" pageNumber="320">Cambodia</collectingCountry>
, 
<collectingCountry id="F30BCEEC2957D646FEECCD8383F7399C" box="[290,392,1478,1503]" name="Thailand" pageId="7" pageNumber="320">Thailand</collectingCountry>
, 
<collectingCountry id="F30BCEEC2957D646FE50CD8D807239A3" box="[414,525,1480,1504]" name="Myanmar" pageId="7" pageNumber="320">Myanmar</collectingCountry>
, Peninsular 
<collectingCountry id="F30BCEEC2957D646FD67CD83816C399C" box="[681,787,1478,1503]" name="Malaysia" pageId="7" pageNumber="320">Malaysia</collectingCountry>
, and southern 
<collectingCountry id="F30BCEEC2957D646FC1DCD838669399C" box="[979,1046,1478,1503]" name="China" pageId="7" pageNumber="320">China</collectingCountry>
[
<collectingRegion id="49D8409E2957D646FBE0CD8286F4399C" box="[1070,1163,1479,1503]" country="China" name="Yunnan" pageId="7" pageNumber="320">Yunnan</collectingRegion>
]) as follows: from 
<taxonomicName id="4C1CF5FF2957D646FA49CD8283D13A40" authority="(Boulenger)" authorityName="Boulenger" baseAuthorityName="Boulenger" baseAuthorityYear="1890" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="320" phylum="Chordata" rank="species" species="anomalopus">
<emphasis id="B968526E2957D646FA49CD82835F3A40" italics="true" pageId="7" pageNumber="320">S. anomalopus</emphasis>
(Boulenger)
</taxonomicName>
by having a smaller size (SVL 
<quantity id="4CE423992957D646FCDFCDAF81F53A41" box="[785,906,1514,1539]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="7" pageNumber="320" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992957D646FC76CDAE86743A40" box="[952,1035,1515,1539]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="320" unit="mm" value="70.0">70 mm</quantity>
), fewer midbody scale rows (32–34 vs. 38) and more lamellae beneath toe IV (18–20 vs. 14); from 
<taxonomicName id="4C1CF5FF2957D646FCB1CE4A86643A64" baseAuthorityName="Eremchenko" baseAuthorityYear="2003" box="[895,1051,1551,1575]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="320" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B968526E2957D646FCB1CE4A86643A64" box="[895,1051,1551,1575]" italics="true" pageId="7" pageNumber="320">S. bacboensis</emphasis>
</taxonomicName>
by having one (vs. two) anterior temporal, more midbody scale rows (32–34 vs. 30–32), and more supralabials (7 vs. 6); from 
<taxonomicName id="4C1CF5FF2957D646FB67CE7687E33A08" authority="Smith" authorityName="Smith" authorityYear="1924" box="[1193,1436,1586,1611]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="320" phylum="Chordata" rank="species" species="cameronicus">
<emphasis id="B968526E2957D646FB67CE76872D3A09" box="[1193,1362,1587,1610]" italics="true" pageId="7" pageNumber="320">S. cameronicus</emphasis>
Smith
</taxonomicName>
by having a smaller size (SVL 
<quantity id="4CE423992957D646FDC8CE1380FD3A2C" box="[518,642,1622,1647]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="7" pageNumber="320" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992957D646FD79CE1281753A2C" box="[695,778,1623,1647]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="320" unit="mm" value="70.0">70 mm</quantity>
) and fewer midbody scale rows (32–34 vs. 38); from 
<taxonomicName id="4C1CF5FF2957D646FA49CE1283153AD0" authority="Boulenger" authorityName="Boulenger" authorityYear="1908" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="320" phylum="Chordata" rank="species" species="cophias">
<emphasis id="B968526E2957D646FA49CE1282913AD0" italics="true" pageId="7" pageNumber="320">S. cophias</emphasis>
Boulenger
</taxonomicName>
by having a larger size (SVL 
<quantity id="4CE423992957D646FD72CE3F814A3AD1" box="[700,821,1658,1683]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="7" pageNumber="320" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992957D646FCADCE3E81C93AD0" box="[867,950,1659,1683]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="320" unit="mm" value="37.0">37 mm</quantity>
), more midbody scale rows (32–34 vs. 24), more lamellae under fourth toe (18–20 vs. 9) and prefrontals in broad contact (vs. separated); from 
<taxonomicName id="4C1CF5FF2957D646FB34CEDA87073AF4" authorityName="Darevsky, Orlov &amp; Ho" authorityYear="2004" box="[1274,1400,1695,1719]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="320" phylum="Chordata" rank="species" species="cryptotis">
<emphasis id="B968526E2957D646FB34CEDA87073AF4" box="[1274,1400,1695,1719]" italics="true" pageId="7" pageNumber="320">S. cryptotis</emphasis>
</taxonomicName>
by having a smaller size (SVL 
<quantity id="4CE423992957D646FE21CE8780113A99" box="[495,622,1730,1755]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="7" pageNumber="320" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992957D646FD67CE8781533A98" box="[681,812,1730,1755]" metricMagnitude="-2" metricUnit="m" metricValue="6.85" metricValueMax="7.9" metricValueMin="5.8" pageId="7" pageNumber="320" unit="mm" value="68.5" valueMax="79.0" valueMin="58.0">58–79 mm</quantity>
), fewer midbody scale rows (32–34 vs. 36–39) and tympanum deeply sunk (vs. superficial); from 
<taxonomicName id="4C1CF5FF2957D646FD69CEA281DE3ABC" authority="Taylor" authorityName="Taylor" authorityYear="1962" box="[679,929,1766,1791]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="320" phylum="Chordata" rank="species" species="grandisonae">
<emphasis id="B968526E2957D646FD69CEA2812F3ABC" box="[679,848,1767,1791]" italics="true" pageId="7" pageNumber="320">S. grandisonae</emphasis>
Taylor
</taxonomicName>
by having a larger size (SVL 
<quantity id="4CE423992957D646FB34CEA3870B3ABC" box="[1274,1396,1766,1791]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="7" pageNumber="320" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992957D646FF59CF4E82953B60" box="[151,234,1803,1827]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="320" unit="mm" value="30.0">30 mm</quantity>
), one (vs. two) anterior temporal, more supralabials (7 vs. 6) and lamellae under toe IV (18–20 vs. 12), and adpressed limbs overlapping (vs. failing to touch); from 
<taxonomicName id="4C1CF5FF2957D646FCD4CF6A81893B04" authority="Cochran" authorityName="Cochran" authorityYear="1927" box="[794,1014,1838,1863]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="320" phylum="Chordata" rank="species" species="helenae">
<emphasis id="B968526E2957D646FCD4CF6A81F13B04" box="[794,910,1839,1863]" italics="true" pageId="7" pageNumber="320">S. helenae</emphasis>
Cochran
</taxonomicName>
by having more midbody scale rows (32–34 vs. 30), prefrontals in broad contact (vs. separated), and presence of an interrupted (vs. uninterrupted) lateral stripe; from 
<taxonomicName id="4C1CF5FF2957D646FEBECF32807D3BCC" baseAuthorityName="Thompson" baseAuthorityYear="1912" box="[368,514,1911,1935]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="320" phylum="Chordata" rank="species" species="incognitus">
<emphasis id="B968526E2957D646FEBECF32807D3BCC" box="[368,514,1911,1935]" italics="true" pageId="7" pageNumber="320">S. incognitus</emphasis>
</taxonomicName>
by having a smaller size (SVL 
<quantity id="4CE423992957D646FCA7CF33819C3BCC" box="[873,995,1910,1935]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="7" pageNumber="320" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992957D646FBDFCF3286E23BCC" box="[1041,1181,1911,1935]" metricMagnitude="-2" metricUnit="m" metricValue="9.15" metricValueMax="10.3" metricValueMin="8.0" pageId="7" pageNumber="320" unit="mm" value="91.5" valueMax="103.0" valueMin="80.0">80–103 mm</quantity>
), fewer midbody scale rows (32–34 vs. 36–40), and one (vs. two) anterior temporal; from 
<taxonomicName id="4C1CF5FF2957D646FC49CFDE818B3BF0" baseAuthorityName="Gray" baseAuthorityYear="1853" box="[903,1012,1947,1971]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="320" phylum="Chordata" rank="species" species="indicus">
<emphasis id="B968526E2957D646FC49CFDE818B3BF0" box="[903,1012,1947,1971]" italics="true" pageId="7" pageNumber="320">S. indicus</emphasis>
</taxonomicName>
by having a smaller size (SVL 
<quantity id="4CE423992957D646FA98CFDE82BC3B94" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="7" pageNumber="320" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992957D646FF3DCFFA83373B94" box="[243,328,1983,2007]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="320" unit="mm" value="90.0">90 mm</quantity>
), prefrontals in broad contact (vs. separated), and asymmetrical and deeply forked hemipenis (vs. symmetrical); from 
<taxonomicName id="4C1CF5FF2958D649FE4FC8DD80CD3CF3" authority="Taylor" authorityName="Taylor" authorityYear="1962" box="[385,690,151,176]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="321" phylum="Chordata" rank="species" species="lineopunctulatus">
<emphasis id="B968526E2958D649FE4FC8DD80223CF3" box="[385,605,152,176]" italics="true" pageId="8" pageNumber="321">S. lineopunctulatus</emphasis>
Taylor
</taxonomicName>
by having a smaller size (SVL 
<quantity id="4CE423992958D649FBF8C8D286CA3CEC" box="[1078,1205,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="8" pageNumber="321" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992958D649FB22C8DD873B3CF3" box="[1260,1348,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="8.4" pageId="8" pageNumber="321" unit="mm" value="84.0">84 mm</quantity>
), fewer midbody scale rows (32–34 vs. 38), fewer paravertebral scale rows (61–69 vs. 76), and prefrontals in broad contact (vs. separated); from 
<taxonomicName id="4C1CF5FF2958D649FE40C8A5805F3CBB" baseAuthorityName="Blyth" baseAuthorityYear="1853" box="[398,544,224,248]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="321" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="B968526E2958D649FE40C8A5805F3CBB" box="[398,544,224,248]" italics="true" pageId="8" pageNumber="321">S. maculatus</emphasis>
</taxonomicName>
by having a smaller size (SVL 
<quantity id="4CE423992958D649FC5AC89A866F3CB4" box="[916,1040,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="8" pageNumber="321" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992958D649FB8CC89A86E73CBB" box="[1090,1176,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="321" unit="mm" value="62.0">62 mm</quantity>
), fewer midbody scale rows (32–34 vs. 38–42), and prefrontals in broad contact (vs. separated); from 
<taxonomicName id="4C1CF5FF2958D649FBE7C94186B13D5F" baseAuthorityName="Doria" baseAuthorityYear="1888" box="[1065,1230,260,284]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="321" phylum="Chordata" rank="species" species="malayanum">
<emphasis id="B968526E2958D649FBE7C94186B13D5F" box="[1065,1230,260,284]" italics="true" pageId="8" pageNumber="321">S. malayanum</emphasis>
</taxonomicName>
by having fewer ventral scales (32–34 vs. 74), fewer paravertebral scales (61–69 vs. 76–80), more lamellae under fourth toe (18–20 vs. 15), and deeply sunk (vs. shallow) tympanum; from 
<taxonomicName id="4C1CF5FF2959D648FCC2CD7781F9390A" authorityName="Taylor" authorityYear="1962" box="[780,902,1330,1353]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="322" phylum="Chordata" rank="species" species="mimicus">
<emphasis id="B968526E2959D648FCC2CD7781F9390A" box="[780,902,1330,1353]" italics="true" pageId="9" pageNumber="322">S. mimicus</emphasis>
</taxonomicName>
by having a larger size (SVL 
<quantity id="4CE423992959D648FB16CD74872D390A" box="[1240,1362,1329,1354]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="9" pageNumber="322" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992959D648FA4ECD7482B8392D" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="9" pageNumber="322" unit="mm" value="36.0">36 mm</quantity>
), more midbody scale rows (32–34 vs. 30) and more lamellae under fourth toe (18–20 vs. 16); from 
<taxonomicName id="4C1CF5FF2959D648FA49CD13831C39D1" authority="(Shreve)" authorityName="Shreve" baseAuthorityName="Shreve" baseAuthorityYear="1940" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="322" phylum="Chordata" rank="species" species="orientale">
<emphasis id="B968526E2959D648FA49CD13828339D2" italics="true" pageId="9" pageNumber="322">S. orientale</emphasis>
(Shreve)
</taxonomicName>
by having more midbody scale rows (32–34 vs. 24–26) and fewer paravertebral scale rows (61– 69 vs. 69–71); and from 
<taxonomicName id="4C1CF5FF2959D648FE66CDDB80B939F5" authority="(Boulenger)" authorityName="Boulenger" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[424,710,1437,1462]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="322" phylum="Chordata" rank="species" species="praesignis">
<emphasis id="B968526E2959D648FE66CDDB804639F6" box="[424,569,1438,1461]" italics="true" pageId="9" pageNumber="322">S. praesignis</emphasis>
(Boulenger)
</taxonomicName>
by having a smaller size (SVL 
<quantity id="4CE423992959D648FBEACDD886E339F6" box="[1060,1180,1437,1462]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="9" pageNumber="322" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992959D648FB06CDDB875A39F5" box="[1224,1317,1438,1462]" metricMagnitude="-1" metricUnit="m" metricValue="1.09" pageId="9" pageNumber="322" unit="mm" value="109.0">109 mm</quantity>
) and more midbody scale rows (32–34 vs. 28). 
<taxonomicName id="4C1CF5FF2959D648FDE5CD84814E399A" authority="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy, 2018" authorityName="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy" authorityYear="2018" box="[555,817,1473,1497]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="322" phylum="Chordata" rank="species" species="yersini" status="sp. nov.">
<emphasis id="B968526E2959D648FDE5CD84814E399A" box="[555,817,1473,1497]" italics="true" pageId="9" pageNumber="322">Sphenomorphus yersini</emphasis>
</taxonomicName>
<emphasis id="B968526E2959D648FCF6CD8481EE3999" bold="true" box="[824,913,1473,1498]" pageId="9" pageNumber="322">
<taxonomicNameLabel id="A25BEF152959D648FCF6CD8481EE3999" box="[824,913,1473,1498]" pageId="9" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="4C1CF5FF2959D648FBEDCD8787053999" authority="(Dumeril &amp; Bibron)" authorityName="Dumeril &amp; Bibron" box="[1059,1402,1473,1498]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="322" phylum="Chordata" rank="species" species="sanctus">
<emphasis id="B968526E2959D648FBEDCD8786EC399A" box="[1059,1171,1474,1497]" italics="true" pageId="9" pageNumber="322">S. sanctus</emphasis>
(Duméril &amp; Bibron)
</taxonomicName>
by having a larger size (SVL 
<quantity id="4CE423992959D648FE75CDA0804C39BE" box="[443,563,1509,1534]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="9" pageNumber="322" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992959D648FDAECDA080A239BD" box="[608,733,1509,1534]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="9" pageNumber="322" unit="mm" value="42.5" valueMax="45.0" valueMin="40.0">40–45 mm</quantity>
), fewer paravertebral scales (61–69 vs. 71), fewer supraoculars (4[5] vs. 5), and fewer lamellae under toe IV (18–20 vs. 26–27); from 
<taxonomicName id="4C1CF5FF2959D648FC79CE4F869C3A61" authority="(Boulenger)" authorityName="Boulenger" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[951,1251,1545,1570]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="322" phylum="Chordata" rank="species" species="scotophilus">
<emphasis id="B968526E2959D648FC79CE4F862C3A62" box="[951,1107,1545,1569]" italics="true" pageId="9" pageNumber="322">S. scotophilus</emphasis>
(Boulenger)
</taxonomicName>
by having more midbody scale rows (32–34 vs. 28–31), fewer supraoculars (4 vs. 5), and fewer lamellae under fourth toe (18–20 vs. 22–23); from 
<taxonomicName id="4C1CF5FF2959D648FEA8CE1780EE3A29" authority="Grismer &amp; Quah" authorityName="Grismer &amp; Quah" authorityYear="2015" box="[358,657,1617,1642]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="322" phylum="Chordata" rank="species" species="senja">
<emphasis id="B968526E2959D648FEA8CE1783BE3A2A" box="[358,449,1618,1641]" italics="true" pageId="9" pageNumber="322">S. senja</emphasis>
Grismer &amp; Quah
</taxonomicName>
by having a smaller size (SVL 
<quantity id="4CE423992959D648FBDECE1486F13A2A" box="[1040,1166,1617,1642]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="9" pageNumber="322" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992959D648FB0ACE14873A3A29" box="[1220,1349,1617,1642]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="9" pageNumber="322" unit="mm" value="62.5" valueMax="65.0" valueMin="60.0">60–65 mm</quantity>
), fewer paravertebral scales (61–69 vs. 72–73), fewer ventral scale rows (60–67 vs. 68), one (vs. two) anterior temporal, more lamellae under toe IV (18–20 vs. 13–17) and prefrontals in broad contact with each other (vs. narrow in contact or slightly separated); from 
<taxonomicName id="4C1CF5FF2959D648FDE1CEFB80F83A96" authorityName="Nguyen, Nguyen, van Devender, Bonkowski &amp; Ziegler" authorityYear="2013" box="[559,647,1725,1749]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="322" phylum="Chordata" rank="species" species="sheai">
<emphasis id="B968526E2959D648FDE1CEFB80F83A96" box="[559,647,1725,1749]" italics="true" pageId="9" pageNumber="322">S. sheai</emphasis>
</taxonomicName>
by having a larger size (SVL 
<quantity id="4CE423992959D648FC2CCEF886223A96" box="[994,1117,1725,1750]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="9" pageNumber="322" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992959D648FB40CEF8869D3A95" box="[1166,1250,1725,1750]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="322" unit="mm" value="35.0">35 mm</quantity>
), more midbody scale rows (32–34 vs. 20), more paravertebral scales (61–69 vs. 53), more supralabials (7 vs. 6), more lamellae under fourth toe (18–20 vs. 6), and adpressed limbs overlapping (vs. separated); from 
<taxonomicName id="4C1CF5FF2959D648FB9CCF4387E33B5D" authority="(Boulenger 1900)" authorityName="Boulenger" authorityYear="1900" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[1106,1436,1797,1822]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="322" phylum="Chordata" rank="species" species="shelfordi">
<emphasis id="B968526E2959D648FB9CCF4386AF3B5E" box="[1106,1232,1797,1821]" italics="true" pageId="9" pageNumber="322">S. shelfordi</emphasis>
(
<bibRefCitation id="EF8DF38D2959D648FB10CF4087EB3B5D" box="[1246,1428,1797,1822]" pageId="9" pageNumber="322" refId="ref6053" refString="Boulenger, G. A. (1900) Description of new reptiles and batrachians from Borneo. Proceedings of the Zoological Society of London, 69, 182 - 187. https: // doi. org / 10.1111 / j. 1096 - 3642.1890. tb 01716. x" type="journal article">Boulenger 1900</bibRefCitation>
)
</taxonomicName>
by having a smaller size (SVL 
<quantity id="4CE423992959D648FDCCCF6C80013B02" box="[514,638,1833,1858]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="9" pageNumber="322" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992959D648FD7FCF6C81783B01" box="[689,775,1833,1858]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="9" pageNumber="322" unit="mm" value="67.0">67 mm</quantity>
), fewer lamellae under toe IV (18–20 vs. 28–29) and the absence of nuchals (vs. presence of a single pair of nuchals); from 
<taxonomicName id="4C1CF5FF2959D648FCB3CF0B81893B26" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[893,1014,1869,1893]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="322" phylum="Chordata" rank="species" species="stellatus">
<emphasis id="B968526E2959D648FCB3CF0B81893B26" box="[893,1014,1869,1893]" italics="true" pageId="9" pageNumber="322">S. stellatus</emphasis>
</taxonomicName>
by having a smaller size (SVL 
<quantity id="4CE423992959D648FA98CF0882BC3BCA" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="9" pageNumber="322" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992959D648FF3ECF37833D3BC9" box="[240,322,1906,1930]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="322" unit="mm" value="80.0">80 mm</quantity>
), more midbody scale rows (32–34 vs. 24) and absence (vs. presence) of two enlarged, broader than long, vertebral scale rows; from 
<taxonomicName id="4C1CF5FF2959D648FDC3CFD3864A3BED" authority="Sumarli, Grismer, Wood, Ahmad" authorityName="Sumarli, Grismer, Wood, Ahmad" box="[525,1077,1941,1966]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="322" phylum="Chordata" rank="species" species="sungaicolus">
<emphasis id="B968526E2959D648FDC3CFD380CE3BEE" box="[525,689,1941,1965]" italics="true" pageId="9" pageNumber="322">S. sungaicolus</emphasis>
Sumarli, Grismer, Wood, Ahmad
</taxonomicName>
, 
<collectingRegion id="49D8409E2959D648FB8CCFD086FF3BED" box="[1090,1152,1941,1966]" country="Philippines" name="Rizal" pageId="9" pageNumber="322">Rizal</collectingRegion>
, Ismail, Izam, Ahmad &amp; Linkem by having a smaller size (SVL 
<quantity id="4CE423992959D648FD96CFFC80AD3B92" box="[600,722,1977,2002]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="9" pageNumber="322" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE423992959D648FCCCCFFC81FF3B91" box="[770,896,1977,2002]" metricMagnitude="-2" metricUnit="m" metricValue="7.85" metricValueMax="9.0" metricValueMin="6.7" pageId="9" pageNumber="322" unit="mm" value="78.5" valueMax="90.0" valueMin="67.0">67–90 mm</quantity>
), fewer midbody scale rows (32–34 vs. 39–44), fewer paravertebral scales (61–69 vs. 72–81) and fewer ventral scale rows (32–34 vs. 74–86); from 
<emphasis id="B968526E2959D648FAF9CF9B87E43BB6" box="[1335,1435,2014,2037]" italics="true" pageId="9" pageNumber="322">S. tarsus</emphasis>
(Smith) by having a smaller size (SVL 
<quantity id="4CE42399295AD64BFD80C8D280B83CEC" box="[590,711,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="10" pageNumber="323" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE42399295AD64BFD3BC8DD81103CF3" box="[757,879,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" metricValueMax="9.2" metricValueMin="9.0" pageId="10" pageNumber="323" unit="mm" value="91.0" valueMax="92.0" valueMin="90.0">90–92 mm</quantity>
) and two loreals (vs. three); from 
<taxonomicName id="4C1CF5FF295AD64BFB20C8DD87E33CF3" baseAuthorityName="Darevsky &amp; Orlov" baseAuthorityYear="2005" box="[1262,1436,152,176]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="323" phylum="Chordata" rank="species" species="tetradactylus">
<emphasis id="B968526E295AD64BFB20C8DD87E33CF3" box="[1262,1436,152,176]" italics="true" pageId="10" pageNumber="323">S. tetradactylus</emphasis>
</taxonomicName>
by having more midbody scale rows (32–34 vs. 20), absence (vs. presence) of external ear opening, and forelimb with five digits (vs. four digits); from 
<taxonomicName id="4C1CF5FF295AD64BFD8DC8A580A23CBB" authorityName="Nguyen, Schmitz, Nguyen, Orlov, Bohme &amp; Ziegler" authorityYear="2011" box="[579,733,224,248]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="323" phylum="Chordata" rank="species" species="tonkinensis">
<emphasis id="B968526E295AD64BFD8DC8A580A23CBB" box="[579,733,224,248]" italics="true" pageId="10" pageNumber="323">S. tonkinensis</emphasis>
</taxonomicName>
by having a larger size (SVL 
<quantity id="4CE42399295AD64BFBFEC89A86D63CB4" box="[1072,1193,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="10" pageNumber="323" unit="mm" value="53.0" valueMax="56.0" valueMin="50.0">50–56 mm</quantity>
vs. 
<quantity id="4CE42399295AD64BFB19C89A872B3CBB" box="[1239,1364,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.9" metricValueMin="3.6" pageId="10" pageNumber="323" unit="mm" value="42.5" valueMax="49.0" valueMin="36.0">36–49 mm</quantity>
), TaL/ SVL ratio 1.80 (vs. 1.32), and one (vs. two) anterior temporal; from 
<taxonomicName id="4C1CF5FF295AD64BFC55C94186613D5F" baseAuthorityName="Bourret" baseAuthorityYear="1939" box="[923,1054,260,284]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="323" phylum="Chordata" rank="species" species="tridigitus">
<emphasis id="B968526E295AD64BFC55C94186613D5F" box="[923,1054,260,284]" italics="true" pageId="10" pageNumber="323">S. tridigitus</emphasis>
</taxonomicName>
by having a larger size (SVL 50– 56 vs. 35), more midbody scale rows (32–34 vs. 20), absence (vs. presence) of external ear opening and forelimb with five digits (vs. three digits); and from 
<taxonomicName id="4C1CF5FF295AD64BFD55C90981433D27" baseAuthorityName="Bourret" baseAuthorityYear="1937" box="[667,828,332,356]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="323" phylum="Chordata" rank="species" species="tritaeniatus">
<emphasis id="B968526E295AD64BFD55C90981433D27" box="[667,828,332,356]" italics="true" pageId="10" pageNumber="323">S. tritaeniatus</emphasis>
</taxonomicName>
by having a larger size (SVL 50–56 vs. 47), fewer midbody scale rows (32–34 vs. 38) and paravertebral scales (61–69 vs. 81), more lamellae under toe IV (18–20 vs. 15) and one anterior temporal (vs. two).
</paragraph>
</subSubSection>
<caption id="DF63DEF42958D649FF59CFD983DA3BAC" httpUri="https://zenodo.org/record/3095282/files/figure.png" pageId="8" pageNumber="321" startId="8.[151,250,1948,1970]" targetBox="[174,1438,334,1932]" targetPageId="8">
<paragraph id="8BA38E7C2958D649FF59CFD983DA3BAC" blockId="8.[151,1436,1948,2031]" pageId="8" pageNumber="321">
<emphasis id="B968526E2958D649FF59CFD9836D3BF2" bold="true" box="[151,274,1948,1970]" pageId="8" pageNumber="321">FIGURE 6.</emphasis>
Examined specimens of 
<taxonomicName id="4C1CF5FF2958D649FDDECFD981583BF1" authorityName="Darevsky &amp; Nguyen" authorityYear="1983" box="[528,807,1948,1970]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="321" phylum="Chordata" rank="species" species="buenloicus">
<emphasis id="B968526E2958D649FDDECFD981583BF1" box="[528,807,1948,1970]" italics="true" pageId="8" pageNumber="321">Sphenomorphus buenloicus</emphasis>
</taxonomicName>
. A–C, holotype, ZISP 19795, hemipenes in C showing forked points near the tips of the organs; D, ITBCZ 5060 (upper) and ITBCZ 4357 (lower); E&amp;F, lateral views of ITBCZ 5060 and ITBCZ 4357, respectively.
</paragraph>
</caption>
<caption id="DF63DEF42959D648FF59CCD687EA38A5" httpUri="https://zenodo.org/record/3095292/files/figure.png" pageId="9" pageNumber="322" startId="9.[151,250,1171,1193]" targetBox="[151,1436,193,1148]" targetPageId="9">
<paragraph id="8BA38E7C2959D648FF59CCD687EA38A5" blockId="9.[151,1436,1171,1254]" pageId="9" pageNumber="322">
<emphasis id="B968526E2959D648FF59CCD6836C38EB" bold="true" box="[151,275,1171,1193]" pageId="9" pageNumber="322">FIGURE 7.</emphasis>
Additional specimen (ITBCZ 6027) of 
<taxonomicName id="4C1CF5FF2959D648FD67CCD681BE38EA" authorityName="Darevsky &amp; Nguyen" authorityYear="1983" box="[681,961,1171,1193]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="322" phylum="Chordata" rank="species" species="buenloicus">
<emphasis id="B968526E2959D648FD67CCD681BE38EA" box="[681,961,1171,1193]" italics="true" pageId="9" pageNumber="322">Sphenomorphus buenloicus</emphasis>
</taxonomicName>
from Tram Lap Forest, Gia Lai Prov. showing the fully everted hemipenes. A, living specimen; B, hemipenes under not in fully everted condition (note that this is similar to the condition of the holotype’s hemipenes in figure 6C); C&amp;D, sulcate and asulcate views of the right hemipenis, respectively.
</paragraph>
</caption>
<subSubSection id="C306DDF7295AD64BFF09C9FD80DB3EC6" pageId="10" pageNumber="323" type="description">
<paragraph id="8BA38E7C295AD64BFF09C9FD80DB3EC6" blockId="10.[151,1436,151,645]" pageId="10" pageNumber="323">
<taxonomicName id="4C1CF5FF295AD64BFF09C9FD83B13D8C" authority="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy, 2018" authorityName="Nguyen &amp; Nguyen &amp; Nguyen &amp; Orlov &amp; Murphy" authorityYear="2018" box="[199,462,440,464]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="323" phylum="Chordata" rank="species" species="yersini" status="sp. nov.">
<emphasis id="B968526E295AD64BFF09C9FD83B13D8C" box="[199,462,440,464]" italics="true" pageId="10" pageNumber="323">Sphenomorphus yersini</emphasis>
</taxonomicName>
<emphasis id="B968526E295AD64BFE1BC9F280503D93" bold="true" box="[469,559,439,464]" pageId="10" pageNumber="323">
<taxonomicNameLabel id="A25BEF15295AD64BFE1BC9F280503D93" box="[469,559,439,464]" pageId="10" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to 
<taxonomicName id="4C1CF5FF295AD64BFD0EC9FD812A3D93" authorityName="Darevsky &amp; Nguyen" authorityYear="1983" box="[704,853,440,464]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="323" phylum="Chordata" rank="species" species="buenloicus">
<emphasis id="B968526E295AD64BFD0EC9FD812A3D93" box="[704,853,440,464]" italics="true" pageId="10" pageNumber="323">S. buenloicus</emphasis>
</taxonomicName>
in many aspects (size, midbody scale rows, number of anterior temporals, lamellae under fourth toe, etc.). However, the new species can be distinguished from 
<taxonomicName id="4C1CF5FF295AD64BFA49C999836F3E5B" authorityName="Darevsky &amp; Nguyen" authorityYear="1983" class="Reptilia" family="Scincidae" genus="Sphenomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="323" phylum="Chordata" rank="species" species="buenloicus">
<emphasis id="B968526E295AD64BFA49C999836F3E5B" italics="true" pageId="10" pageNumber="323">S. buenloicus</emphasis>
</taxonomicName>
by having a relatively longer tail (TaL/SVL ratio 1.8 vs. 1.2), usually more ventral scale rows (58–67 vs. 55–58) and deeply forked hemipenis with two asymmetrical lobes and the outer lobe is much longer than the inner one (vs. hemipenis forked at the middle point of its length with two developing lobes and the outer lobe is slightly longer than the inner one [
<figureCitation id="132792F9295AD64BFDD6CA2980183EC6" box="[536,615,620,645]" captionStart="FIGURE 6" captionStartId="8.[151,250,1948,1970]" captionTargetBox="[174,1438,334,1932]" captionTargetId="figure@8.[174,1438,334,1932]" captionTargetPageId="8" captionText="FIGURE 6. Examined specimens of Sphenomorphus buenloicus. A–C, holotype, ZISP 19795, hemipenes in C showing forked points near the tips of the organs; D, ITBCZ 5060 (upper) and ITBCZ 4357 (lower); E&amp;F, lateral views of ITBCZ 5060 and ITBCZ 4357, respectively." httpUri="https://zenodo.org/record/3095282/files/figure.png" pageId="10" pageNumber="323">Figs. 6</figureCitation>
&amp; 
<figureCitation id="132792F9295AD64BFD4FCA2980EF3EC6" box="[641,656,620,645]" captionStart="FIGURE 7" captionStartId="9.[151,250,1171,1193]" captionTargetBox="[151,1436,193,1148]" captionTargetId="figure@9.[151,1436,193,1149]" captionTargetPageId="9" captionText="FIGURE 7. Additional specimen (ITBCZ 6027) of Sphenomorphus buenloicus from Tram Lap Forest, Gia Lai Prov. showing the fully everted hemipenes. A, living specimen; B, hemipenes under not in fully everted condition (note that this is similar to the condition of the holotype’s hemipenes in figure 6C); C&amp;D, sulcate and asulcate views of the right hemipenis, respectively." httpUri="https://zenodo.org/record/3095292/files/figure.png" pageId="10" pageNumber="323">7</figureCitation>
]).
</paragraph>
</subSubSection>
</treatment>
</document>