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<document id="21EB819F00010AF25ABED320256C93B1" ID-CLB-Dataset="268615" ID-DOI="10.1163/18759866-20191416" ID-GBIF-Dataset="6388d839-a72d-4598-a141-053897bfc900" ID-ISSN="1875-9866" ID-Zenodo-Dep="8343061" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_requiresApprovalFor="plazi" IM.tables_requiresApprovalFor="plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1694662606570" checkinUser="felipe" docAuthor="Hyman, Isabel T. &amp; Köhler, Frank" docDate="2019" docId="03F32A40FF9DB149519F9A34F5CAFD62" docLanguage="en" docName="ContribZool.88.4.351-451.pdf" docOrigin="Contributions to Zoology 88 (4)" docSource="http://dx.doi.org/10.1163/18759866-20191416" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.2:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="2" docTitle="Fastosarion hannianus" docType="treatment" docVersion="8" lastPageNumber="423" masterDocId="FFCA5238FFD9B100533A9C10F477FFFB" masterDocTitle="Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)" masterLastPageNumber="451" masterPageNumber="351" pageNumber="419" updateTime="1699651795867" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4E23D0C9A39AFC8ACCFBBCBE2F3A20E3">Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)</mods:title>
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<subSubSection id="C340C8DDFF9DB144519F9A34F02AF9C5" box="[677,1117,1571,1598]" pageId="68" pageNumber="419" type="nomenclature">
<paragraph id="8BE59B56FF9DB144519F9A34F02AF9C5" blockId="68.[677,1117,1571,1598]" box="[677,1117,1571,1598]" pageId="68" pageNumber="419">
<heading id="D0AD2C3AFF9DB144519F9A34F02AF9C5" box="[677,1117,1571,1598]" fontSize="11" level="2" pageId="68" pageNumber="419" reason="2">
<taxonomicName id="4C5AE0D5FF9DB144519F9A34F02AF9C5" ID-CoL="7RBRF" ID-ENA="2658037" authority="(Stanisic, 2010)" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[677,1117,1571,1598]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="68" pageNumber="419" phylum="Mollusca" rank="species" species="hannianus">
<emphasis id="B92E4744FF9DB144519F9A34F02AF9C5" bold="true" box="[677,1117,1571,1598]" pageId="68" pageNumber="419">
<emphasis id="B92E4744FF9DB144519F9A34F7CDF9C5" bold="true" box="[677,954,1571,1598]" italics="true" pageId="68" pageNumber="419">Fastosarion hannianus (</emphasis>
<bibRefCitation id="EFCBE6A7FF9DB14450819A33F024F9C6" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." box="[955,1107,1571,1597]" pageId="68" pageNumber="419" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic, 2010</bibRefCitation>
<emphasis id="B92E4744FF9DB14457699A34F02AF9C5" bold="true" box="[1107,1117,1572,1598]" italics="true" pageId="68" pageNumber="419">)</emphasis>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C340C8DDFF9DB144519F9A57F7BAF999" box="[677,973,1607,1634]" pageId="68" pageNumber="419" type="description">
<paragraph id="8BE59B56FF9DB144519F9A57F7BAF999" blockId="68.[677,1187,1607,1706]" box="[677,973,1607,1634]" pageId="68" pageNumber="419">
<figureCitation id="136187D3FF9DB144519F9A57F682F999" box="[677,757,1607,1634]" captionStart="FIGURE 18" captionStartId="66.[129,202,1582,1603]" captionTargetBox="[129,1187,188,1550]" captionTargetId="figure-8@66.[129,1187,188,1550]" captionTargetPageId="66" captionText="FIGURE 18 Distribution of Clade B species. █ = F.alyssa, ▼ = F.deensis, × = F.ephelis, ♢ = F.hannianus, ▲ = F. insularis, ◻ = F.katatonos, ∆ = F.longimentula, ● = F.mcdonaldi, ♠ = F.minerva, ◆ = F.papillosus, ♣ = F.pustulosus, ♥ = F.rowani, Ǫ = F.sarina, ▽ = F.tuljun." figureDoi="http://doi.org/10.5281/zenodo.8343097" httpUri="https://zenodo.org/record/8343097/files/figure.png" pageId="68" pageNumber="419">Figs. 18</figureCitation>
,
<figureCitation id="136187D3FF9DB14451C49A58F75FF999" box="[766,808,1608,1634]" captionStart="FIGURE 19" captionStartId="67.[98,171,1480,1501]" captionTargetBox="[98,1104,171,1448]" captionTargetId="figure-115@67.[98,1104,170,1448]" captionTargetPageId="67" captionText="FIGURE 19 Shells of Clade B species. A. Fastosarion alyssa (type locality of D.peterbrocki), NMV F230748. B. Fastosarion deensis, MO85834 (holotype). C. Fastosarion ephelis, MO85835 (holotype). D. Fastosarion hannianus, MO56684.E. Fastosarion insularis sp. nov., MO76354. F. Fastosarion katatonos, MO85837 (holotype). G.Fastosarion longimentula, MO65313.H. Fastosarion mcdonaldi, MO50434.I. Fastosarion minerva, MO78358. J.Fastosarion papillosus, MO78896 (Mt Superbus).K.Fastosarion papillosus, MO62397 (Kroombit Tops). L.Fastosarion pustulosus, MO78894 (Kalpowar SF).M. Fastosarion rowani, MO85839 (holotype). N. Fastosarion sarina, MO85840 (holotype).O. Fastosarion tujun (Denham Range), MO76081 (holotype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8343099" httpUri="https://zenodo.org/record/8343099/files/figure.png" pageId="68" pageNumber="419">19D</figureCitation>
,
<figureCitation id="136187D3FF9DB144500B9A58F72DF999" box="[817,858,1608,1634]" captionStart="FIGURE 20" captionStartId="69.[98,171,1087,1108]" captionTargetBox="[98,1157,188,1050]" captionTargetId="figure-74@69.[98,1157,188,1050]" captionTargetPageId="69" captionText="FIGURE 20 Fastosarion live images (Clade B). A-D.Fastosarion alyssa, A:AM C.512537, Mt Etna Caves; B:AM C.512480, Mt Ossa; C:AM C.512579, The Hummock. D: NMV F230748,Brandy Creek (Type locality of F.peterbrocki; A.Moussalli, NMV). E. Fastosarion ephelis (J. Stanisic).F.Fastosarion hannianus (QM image).G. Fastosarion mcdonaldi, Mt Dalrymple (QM image). H. Fastosarion rowani, AM C.512541,Dan Dan NP." figureDoi="http://doi.org/10.5281/zenodo.8343101" httpUri="https://zenodo.org/record/8343101/files/figure.png" pageId="68" pageNumber="419">20F</figureCitation>
,
<figureCitation id="136187D3FF9DB144505E9A58F7DDF999" box="[868,938,1608,1634]" captionStart="FIGURE 22" captionStartId="71.[98,171,1710,1731]" captionTargetBox="[98,887,184,1677]" captionTargetId="figure-39@71.[98,887,184,1677]" captionTargetPageId="71" captionText="FIGURE 22 Spermatophores of Clade B species. A-B. F.alyssa, MO77158.C-D.F.deensis, MO85834.E-F. F.hannianus, MO72277.G-H.F.mcdonaldi, MO50434.I-J. F.minerva, MO78358." figureDoi="http://doi.org/10.5281/zenodo.8343105" httpUri="https://zenodo.org/record/8343105/files/figure.png" pageId="68" pageNumber="419">22E–F</figureCitation>
,
<figureCitation id="136187D3FF9DB14450899A58F7BAF999" box="[947,973,1608,1634]" captionStart="FIGURE 24" captionStartId="74.[129,202,1314,1335]" captionTargetBox="[129,1177,192,1282]" captionTargetId="figure-8@74.[129,1187,188,1282]" captionTargetPageId="74" captionText="FIGURE 24 Genitalia of F.hannianus. A, B. MO72277,Bakers Blue Mountain. C.MO76564, Hann Tableland.A. Reproductive system. Arrow shows the junction of the penis and epiphallus. B–C. Penis interior.Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8343109" httpUri="https://zenodo.org/record/8343109/files/figure.png" pageId="68" pageNumber="419">24</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C340C8DDFF9DB14451F99A7CF050F952" pageId="68" pageNumber="419" type="reference_group">
<paragraph id="8BE59B56FF9DB14451F99A7CF050F952" blockId="68.[677,1187,1607,1706]" pageId="68" pageNumber="419">
<treatmentCitationGroup id="AB4ABC78FF9DB14451F99A7CF050F952" pageId="68" pageNumber="419">
<taxonomicName id="4C5AE0D5FF9DB14451F99A7CF7B5F97E" ID-CoL="ad02c51b-097a-428e-9847-47a314167ba6" authorityName="J. Stanisic" authorityYear="2010" box="[707,962,1643,1669]" class="Gastropoda" family="Helicarionidae" genus="Hymanarion" kingdom="Animalia" order="Stylommatophora" pageId="68" pageNumber="419" phylum="Mollusca" rank="species" species="hanniana">
<emphasis id="B92E4744FF9DB14451F99A7CF7B5F97E" box="[707,962,1643,1669]" italics="true" pageId="68" pageNumber="419">Hymanarion hanniana</emphasis>
</taxonomicName>
[sic!]
<bibRefCitation id="EFCBE6A7FF9DB14457379A7BF0D4F97D" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." box="[1037,1187,1643,1670]" pageId="68" pageNumber="419" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">
Stanisic,
<quantity id="4CA236B3FF9DB14457499A7CF0D4F97D" box="[1139,1187,1644,1670]" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" pageId="68" pageNumber="419" unit="in" value="2010.0">2010</quantity>
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in
<treatmentCitation id="0AFBBD47FF9DB14451FB9A9FF786F952" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." box="[705,1009,1679,1706]" page="318" pageId="68" pageNumber="419" year="2010">
<bibRefCitation id="EFCBE6A7FF9DB14451FB9A9FF786F952" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." box="[705,1009,1679,1706]" pageId="68" pageNumber="419" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al., 2010: 318–319</bibRefCitation>
</treatmentCitation>
, 334.
</treatmentCitationGroup>
</paragraph>
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<caption id="DF25CBDEFF9CB1455358982FF551FB31" ID-DOI="http://doi.org/10.5281/zenodo.8343101" ID-Zenodo-Dep="8343101" httpUri="https://zenodo.org/record/8343101/files/figure.png" pageId="69" pageNumber="420" startId="69.[98,171,1087,1108]" targetBox="[98,1157,188,1050]" targetPageId="69" targetType="figure">
<paragraph id="8BE59B56FF9CB1455358982FF551FB31" blockId="69.[98,1149,1087,1226]" pageId="69" pageNumber="420">
FIGURE 20
<taxonomicName id="4C5AE0D5FF9CB14553E19850F534FBAE" authorityName="Iredale" authorityYear="1933" box="[219,323,1088,1109]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="69" pageNumber="420" phylum="Mollusca" rank="genus">
<emphasis id="B92E4744FF9CB14553E19850F534FBAE" box="[219,323,1088,1109]" italics="true" pageId="69" pageNumber="420">Fastosarion</emphasis>
</taxonomicName>
live images (Clade B). A-D.
<taxonomicName id="4C5AE0D5FF9CB14551049850F695FBAF" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[574,738,1087,1109]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="69" pageNumber="420" phylum="Mollusca" rank="species" species="alyssa">
<emphasis id="B92E4744FF9CB14551049850F695FBAF" box="[574,738,1087,1109]" italics="true" pageId="69" pageNumber="420">Fastosarion alyssa</emphasis>
</taxonomicName>
, A: AM C.512537, Mt Etna Caves; B: AM C.512480, Mt Ossa; C: AM C.512579, The Hummock. D: NMV F230748, Brandy Creek (Type locality of
<taxonomicName id="4C5AE0D5FF9CB14553E1986AF525FB74" box="[219,338,1146,1167]" class="Gastropoda" family="Helicarionidae" genus="Dimidarion" kingdom="Animalia" order="Stylommatophora" pageId="69" pageNumber="420" phylum="Mollusca" rank="species" species="peterbrocki">
<emphasis id="B92E4744FF9CB14553E1986AF525FB74" box="[219,338,1146,1167]" italics="true" pageId="69" pageNumber="420">F.peterbrocki</emphasis>
</taxonomicName>
; A. Moussalli, NMV). E.
<taxonomicName id="4C5AE0D5FF9CB1455117986AF731FB6B" authority="(J. Stanisic)" baseAuthorityName="J. Stanisic" box="[557,838,1146,1168]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="69" pageNumber="420" phylum="Mollusca" rank="species" species="ephelis">
<emphasis id="B92E4744FF9CB1455117986AF6A0FB74" box="[557,727,1146,1167]" italics="true" pageId="69" pageNumber="420">Fastosarion ephelis</emphasis>
(J. Stanisic)
</taxonomicName>
. F.
<taxonomicName id="4C5AE0D5FF9CB1455058986AF047FB74" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[866,1072,1146,1167]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="69" pageNumber="420" phylum="Mollusca" rank="species" species="hannianus">
<emphasis id="B92E4744FF9CB1455058986AF047FB74" box="[866,1072,1146,1167]" italics="true" pageId="69" pageNumber="420">Fastosarion hannianus</emphasis>
</taxonomicName>
(QM image). G.
<taxonomicName id="4C5AE0D5FF9CB14552079888F6E5FB56" authority=", Mt Dalrymple" authorityName="Mt Dalrymple" box="[317,658,1175,1197]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="69" pageNumber="420" phylum="Mollusca" rank="species" species="mcdonaldi">
<emphasis id="B92E4744FF9CB14552079888F67EFB57" box="[317,521,1175,1197]" italics="true" pageId="69" pageNumber="420">Fastosarion mcdonaldi</emphasis>
, Mt Dalrymple
</taxonomicName>
(QM image). H.
<taxonomicName id="4C5AE0D5FF9CB14550139888F063FB56" authority=", AM C." authorityName="AM C." box="[809,1044,1175,1197]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="69" pageNumber="420" phylum="Mollusca" rank="species" species="rowani">
<emphasis id="B92E4744FF9CB14550139888F7A4FB56" box="[809,979,1176,1197]" italics="true" pageId="69" pageNumber="420">Fastosarion rowani</emphasis>
, AM C.
</taxonomicName>
512541, Dan Dan NP.
</paragraph>
</caption>
<subSubSection id="C340C8DDFF9CB1495366994FF5CAFD62" lastPageId="73" lastPageNumber="424" pageId="69" pageNumber="420" type="materials_examined">
<paragraph id="8BE59B56FF9CB1455366994FF547FA82" blockId="69.[92,608,1375,1617]" box="[92,304,1375,1401]" pageId="69" pageNumber="420">Material examined</paragraph>
<paragraph id="8BE59B56FF9CB14553589994F5E5F9F2" blockId="69.[92,608,1375,1617]" pageId="69" pageNumber="420">
<emphasis id="B92E4744FF9CB14553589994F4D6FA66" box="[98,161,1412,1437]" italics="true" pageId="69" pageNumber="420">Types</emphasis>
:
<materialsCitation id="3B32910BFF9CB145538E9993F5E5F9F2" ID-GBIF-Occurrence="4413347336" collectingDate="1989-12-30" elevation="900" latitude="-16.7" location="Bakers Blue Mtn" longLatPrecision="1281" longitude="145.16667" pageId="69" pageNumber="420" specimenCount="1" typeStatus="holotype">
<typeStatus id="54E125F4FF9CB145538E9993F568FA66" box="[180,287,1411,1437]" pageId="69" pageNumber="420" type="holotype">Holotype</typeStatus>
: QMMO48176 (
<location id="8E85CD8DFF9CB14552EF9993F4E4FA39" LSID="urn:lsid:plazi:treatment:03F32A40FF9DB149519F9A34F5CAFD62:8E85CD8DFF9CB14552EF9993F4E4FA39" latitude="-16.7" longLatPrecision="1281" longitude="145.16667" name="Bakers Blue Mtn" pageId="69" pageNumber="420">Bakers Blue Mtn</location>
, c.
<locationDeviation id="9A578274FF9CB145538699B7F5B6FA3A" box="[188,449,1447,1474]" location="Mt Molloy" pageId="69" pageNumber="420">
<quantity id="4CA236B3FF9CB145538699B7F489FA3A" box="[188,254,1447,1473]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="69" pageNumber="420" unit="km" value="17.0">17 km</quantity>
W of Mt Molloy
</locationDeviation>
, NEQ
<geoCoordinate id="EE6EFD91FF9CB145513699B7F62BFA3A" box="[524,604,1447,1473]" degrees="16" direction="south" minutes="42" orientation="latitude" pageId="69" pageNumber="420" precision="925" value="-16.7">16°42’S</geoCoordinate>
,
<geoCoordinate id="EE6EFD91FF9CB145535899DBF4CAFA1D" box="[98,189,1483,1510]" degrees="145" direction="east" minutes="10" orientation="longitude" pageId="69" pageNumber="420" precision="925" value="145.16667">145°10’E</geoCoordinate>
, rainforest,
<quantity id="4CA236B3FF9CB145527299DBF5A5FA1D" box="[328,466,1483,1510]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="69" pageNumber="420" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">
<elevation id="00777C65FF9CB145527299DBF5A5FA1D" box="[328,466,1483,1510]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="69" pageNumber="420" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">800–1000 m</elevation>
</quantity>
, under logs, coll.
<date id="FFE4BD96FF9CB14553AD99FFF56CF9F2" box="[151,283,1519,1546]" pageId="69" pageNumber="420" value="1989-12-30">
<collectingDate id="EFA0447EFF9CB14553AD99FFF56CF9F2" box="[151,283,1519,1546]" pageId="69" pageNumber="420" value="1989-12-30">30 Dec 1989</collectingDate>
</date>
, ANZSES).
</materialsCitation>
</paragraph>
<paragraph id="8BE59B56FF9CB14553BB9A04F637F9D6" blockId="69.[92,608,1375,1617]" box="[129,576,1555,1582]" pageId="69" pageNumber="420">
<materialsCitation id="3B32910BFF9CB14553BB9A04F5E3F9D6" ID-GBIF-Occurrence="4413347366" box="[129,404,1555,1582]" collectionCode="QM" pageId="69" pageNumber="420" specimenCode="MO48167" specimenCount="1" typeStatus="paratype">
<typeStatus id="54E125F4FF9CB14553BB9A04F487F9D5" box="[129,240,1556,1582]" pageId="69" pageNumber="420" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="ED4B0393FF9CB14553C39A03F553F9D6" box="[249,292,1555,1581]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="69" pageNumber="420" type="Museum">QM</collectionCode>
<specimenCode id="DBFC332DFF9CB14552139A03F5E3F9D6" box="[297,404,1555,1581]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="69" pageNumber="420" type="Museum">MO48167</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B32910BFF9CB14552A79A03F64CF9D6" ID-GBIF-Occurrence="4413347375" box="[413,571,1555,1581]" collectionCode="QM" pageId="69" pageNumber="420" specimenCode="MO78907" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED4B0393FF9CB14552A79A03F5BFF9D6" box="[413,456,1555,1581]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="69" pageNumber="420" type="Museum">QM</collectionCode>
<specimenCode id="DBFC332DFF9CB14552F79A03F64CF9D6" box="[461,571,1555,1581]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="69" pageNumber="420" type="Museum">MO78907</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE59B56FF9CB14553BB9A28F5BCF9AA" blockId="69.[92,608,1375,1617]" box="[129,459,1591,1617]" pageId="69" pageNumber="420">
<emphasis id="B92E4744FF9CB14553BB9A28F53BF9AA" box="[129,332,1591,1617]" italics="true" pageId="69" pageNumber="420">Non-type material:</emphasis>
See
<tableCitation id="C6D8AEEDFF9CB14552449A27F5B0F9AA" box="[382,455,1591,1617]" captionStart="TABLE 1" captionStartId="3.[56,116,402,428]" captionTargetBox="[56,1638,477,1528]" captionTargetPageId="3" captionText="TABLE 1 Material examined. Abbreviations: AM, Australian Museum, Sydney; H, holotype; NMV,Museums Victoria, Melbourne; P, Paratype; QM, Queensland Museum, Brisbane; S, syntype; NHRM, Naturhistoriska Riksmuseet, Stockholm, Sweden" httpUri="http://table.plazi.org/id/DF25CBDEFFDAB10353029D82F71DFE37" pageId="69" pageNumber="420" tableUuid="DF25CBDEFFDAB10353029D82F71DFE37">table 1</tableCitation>
.
</paragraph>
<paragraph id="8BE59B56FF9CB14553669A6FF4BEF962" blockId="69.[92,609,1663,1761]" box="[92,201,1663,1689]" pageId="69" pageNumber="420">Diagnosis</paragraph>
<paragraph id="8BE59B56FF9CB14553589AB3F6B6F98E" blockId="69.[92,609,1663,1761]" lastBlockId="69.[646,1157,1375,1760]" pageId="69" pageNumber="420">
<emphasis id="B92E4744FF9CB14553589AB3F526F946" box="[98,337,1699,1725]" italics="true" pageId="69" pageNumber="420">External morphology</emphasis>
: Shell (fig. 19D) small (
<quantity id="4CA236B3FF9CB14553509AD6F4B1F91A" box="[106,198,1734,1761]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="69" pageNumber="420" unit="mm" value="9.6">9.6 mm</quantity>
), golden amber, transparent, 2.1–3.0 whorls, subglobose, thin, last whorl large. Body (fig. 20F)
<quantity id="4CA236B3FF9CB14550109993F7EAFA65" box="[810,925,1411,1438]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="69" pageNumber="420" unit="mm" value="21.0" valueMax="24.0" valueMin="18.0">18–24 mm</quantity>
, dark reddish brown, darker on tail; dark bands on either side of neck. Mantle lobes and shell lappets moderately large, darker in colour, smooth. Tail not keeled, slime network weak. White pigmentation on visceral mass visible through shell.
</paragraph>
<paragraph id="8BE59B56FF9CB148519F9A6FF784F9A9" blockId="69.[646,1157,1375,1760]" lastBlockId="72.[676,1189,1483,1618]" lastPageId="72" lastPageNumber="423" pageId="69" pageNumber="420">
<emphasis id="B92E4744FF9CB145519F9A6FF715F963" box="[677,866,1663,1689]" italics="true" pageId="69" pageNumber="420">Genital anatomy:</emphasis>
Genitalia (figs. 22E–F, 24) with very short vagina; bursa copulatrix very short, duct distinct, bursa spherical. Penis large, cylindrical, internally often with 3–4 short, narrow longitudinal pilasters basally, two narrow longitudinal pilasters proximally, and one stimulator and one rounded pilaster medially; internal wall pustulose distally and longitudinally ridged proximally; 50% of penis contained in penial tunica. Epiphallus approx. equal to penis length; epiphallus arms equal in length; epiphallic caecum short; flagellum moderately long, uncoiled, slender, slightly swollen towards tip. Spermatophore simple (based on a fragment).
</paragraph>
<caption id="DF25CBDEFF9FB14653BB9A7EF664F920" ID-DOI="http://doi.org/10.5281/zenodo.8343103" ID-Zenodo-Dep="8343103" httpUri="https://zenodo.org/record/8343103/files/figure.png" pageId="70" pageNumber="421" startId="70.[129,202,1646,1667]" targetBox="[129,1066,188,1614]" targetPageId="70" targetType="figure">
<paragraph id="8BE59B56FF9FB14653BB9A7EF664F920" blockId="70.[129,1173,1646,1756]" pageId="70" pageNumber="421">
FIGURE 21 Genitalia of
<taxonomicName id="4C5AE0D5FF9FB14652519A7FF5C2F978" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[363,437,1646,1668]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="70" pageNumber="421" phylum="Mollusca" rank="species" species="alyssa">
<emphasis id="B92E4744FF9FB14652519A7FF5C2F978" box="[363,437,1646,1668]" italics="true" pageId="70" pageNumber="421">F. alyssa</emphasis>
</taxonomicName>
. A, D. MO21620, Johanssen’s Caves. B-C, F. MO77188, Lords Table Mountain, Peak Range (type locality of
<taxonomicName id="4C5AE0D5FF9FB14652F69A9CF657F95B" box="[460,544,1675,1697]" class="Gastropoda" family="Helicarionidae" genus="Dimidarion" kingdom="Animalia" order="Stylommatophora" pageId="70" pageNumber="421" phylum="Mollusca" rank="species" species="slatyeri">
<emphasis id="B92E4744FF9FB14652F69A9CF657F95B" box="[460,544,1675,1697]" italics="true" pageId="70" pageNumber="421">F. slatyeri</emphasis>
</taxonomicName>
). D. F230748. Brandy Creek (type locality of
<taxonomicName id="4C5AE0D5FF9FB14650889A9CF05EF95B" box="[946,1065,1675,1697]" class="Gastropoda" family="Helicarionidae" genus="Dimidarion" kingdom="Animalia" order="Stylommatophora" pageId="70" pageNumber="421" phylum="Mollusca" rank="species" species="peterbrocki">
<emphasis id="B92E4744FF9FB14650889A9CF05EF95B" box="[946,1065,1675,1697]" italics="true" pageId="70" pageNumber="421">F.peterbrocki</emphasis>
</taxonomicName>
). A–B: Reproductive system. Arrows show the junction of the penis and epiphallus. C: Spermatophore. D–F: Penis interior. Scale bars 1 mm.
</paragraph>
</caption>
<caption id="DF25CBDEFF9EB14753589ABEF7D2F91A" ID-DOI="http://doi.org/10.5281/zenodo.8343105" ID-Zenodo-Dep="8343105" httpUri="https://zenodo.org/record/8343105/files/figure.png" pageId="71" pageNumber="422" startId="71.[98,171,1710,1731]" targetBox="[98,887,184,1677]" targetPageId="71" targetType="figure">
<paragraph id="8BE59B56FF9EB14753589ABEF7D2F91A" blockId="71.[98,1041,1710,1761]" pageId="71" pageNumber="422">
FIGURE 22 Spermatophores of Clade B species. A-B.
<taxonomicName id="4C5AE0D5FF9EB147516B9ABFF6B0F93F" authority=", MO" authorityName="MO" box="[593,711,1710,1732]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="71" pageNumber="422" phylum="Mollusca" rank="species" species="alyssa">
<emphasis id="B92E4744FF9EB147516B9ABFF6ECF938" box="[593,667,1710,1732]" italics="true" pageId="71" pageNumber="422">F. alyssa</emphasis>
, MO
</taxonomicName>
77158. C-D.
<taxonomicName id="4C5AE0D5FF9EB14750149ABFF7D8F93F" authority=", MO" authorityName="MO" box="[814,943,1710,1732]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="71" pageNumber="422" phylum="Mollusca" rank="species" species="deensis">
<emphasis id="B92E4744FF9EB14750149ABFF7F5F938" box="[814,898,1710,1732]" italics="true" pageId="71" pageNumber="422">F.deensis</emphasis>
, MO
</taxonomicName>
85834. E-F.
<taxonomicName id="4C5AE0D5FF9EB14753E19ADCF50DF91A" authority=", MO" authorityName="MO" box="[219,378,1740,1761]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="71" pageNumber="422" phylum="Mollusca" rank="species" species="hannianus">
<emphasis id="B92E4744FF9EB14753E19ADCF538F91A" box="[219,335,1740,1761]" italics="true" pageId="71" pageNumber="422">F.hannianus</emphasis>
, MO
</taxonomicName>
72277. G-H.
<taxonomicName id="4C5AE0D5FF9EB14752DC9ADCF6F1F91A" authority=", MO" authorityName="MO" box="[486,646,1739,1761]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="71" pageNumber="422" phylum="Mollusca" rank="species" species="mcdonaldi">
<emphasis id="B92E4744FF9EB14752DC9ADCF62EF91A" box="[486,601,1740,1761]" italics="true" pageId="71" pageNumber="422">F.mcdonaldi</emphasis>
, MO
</taxonomicName>
50434. I-J.
<taxonomicName id="4C5AE0D5FF9EB14751D89ADCF71CF91A" authority=", MO" authorityName="MO" box="[738,875,1739,1761]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="71" pageNumber="422" phylum="Mollusca" rank="species" species="minerva">
<emphasis id="B92E4744FF9EB14751D89ADCF748F91A" box="[738,831,1740,1761]" italics="true" pageId="71" pageNumber="422">F. minerva</emphasis>
, MO
</taxonomicName>
78358.
</paragraph>
</caption>
<caption id="DF25CBDEFF91B14853BB9905F61AFA9D" ID-DOI="http://doi.org/10.5281/zenodo.8343107" ID-Zenodo-Dep="8343107" httpUri="https://zenodo.org/record/8343107/files/figure.png" pageId="72" pageNumber="423" startId="72.[129,202,1301,1322]" targetBox="[129,1187,184,1243]" targetPageId="72" targetType="figure">
<paragraph id="8BE59B56FF91B14853BB9905F61AFA9D" blockId="72.[129,1185,1301,1382]" pageId="72" pageNumber="423">
FIGURE 23 Genitalia. A–C.
<taxonomicName id="4C5AE0D5FF91B14852B39906F67DFAD0" authority=", MO" authorityName="MO" box="[393,522,1301,1323]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="72" pageNumber="423" phylum="Mollusca" rank="species" species="deensis">
<emphasis id="B92E4744FF91B14852B39906F5AAFAD1" box="[393,477,1301,1323]" italics="true" pageId="72" pageNumber="423">F.deensis</emphasis>
, MO
</taxonomicName>
85834, Mt Morgan. D-E.
<taxonomicName id="4C5AE0D5FF91B14851DF9906F714FAD0" authority=", MO" authorityName="MO" box="[741,867,1301,1323]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="72" pageNumber="423" phylum="Mollusca" rank="species" species="ephelis">
<emphasis id="B92E4744FF91B14851DF9906F741FAD1" box="[741,822,1301,1323]" italics="true" pageId="72" pageNumber="423">F.ephelis</emphasis>
, MO
</taxonomicName>
85835, Broken River. A, D. Reproductive system. Arrows show the junction of the penis and epiphallus. B, E. Penis interior. F. Spermatophore. Scale bars 1 mm (E), 2 mm (A–D).
</paragraph>
</caption>
<paragraph id="8BE59B56FF91B14851A59A6FF776F962" blockId="72.[671,1187,1663,1761]" box="[671,769,1663,1689]" pageId="72" pageNumber="423">Remarks</paragraph>
<paragraph id="8BE59B56FF91B149519F9AB3F5CAFD62" blockId="72.[671,1187,1663,1761]" lastBlockId="73.[97,611,171,665]" lastPageId="73" lastPageNumber="424" pageId="72" pageNumber="423">
The spelling of the species name is adapted herein to agree with the masculine gender of the genus name.
<taxonomicName id="4C5AE0D5FF90B14952649CBCF616FF3E" box="[350,609,171,197]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="species" species="hannianus">
<emphasis id="B92E4744FF90B14952649CBCF616FF3E" box="[350,609,171,197]" italics="true" pageId="73" pageNumber="424">Fastosarion hannianus</emphasis>
</taxonomicName>
is only known from the Hann Tableland and Bakers Blue Mountain, SW of Mossman in northeastern Qld (fig. 18), where it is found in vine thicket on basalt. This species shares with
<taxonomicName id="4C5AE0D5FF90B14953A69D70F48FFE82" box="[156,248,351,377]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="species" species="alyssa">
<emphasis id="B92E4744FF90B14953A69D70F48FFE82" box="[156,248,351,377]" italics="true" pageId="73" pageNumber="424">F. alyssa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5AE0D5FF90B149523E9D70F50CFE82" authorityName="Hyman &amp; Köhler" authorityYear="2019" box="[260,379,351,377]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="species" species="insularis">
<emphasis id="B92E4744FF90B149523E9D70F50CFE82" box="[260,379,351,377]" italics="true" pageId="73" pageNumber="424">F. insularis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5AE0D5FF90B14952899D70F637FE82" box="[435,576,351,377]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="species" species="mcdonaldi">
<emphasis id="B92E4744FF90B14952899D70F637FE82" box="[435,576,351,377]" italics="true" pageId="73" pageNumber="424">F. mcdonaldi</emphasis>
</taxonomicName>
its small size and low-spired shell no greater than three whorls with, a short vagina and bursa copulatrix, equal length arms of the epiphallus, and a penis with some rounded pilasters proximally. It differs from these species in its more globose shell.
<taxonomicName id="4C5AE0D5FF90B14952729E27F631FDAB" box="[328,582,566,592]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="species" species="hannianus">
<emphasis id="B92E4744FF90B14952729E27F631FDAB" box="[328,582,566,592]" italics="true" pageId="73" pageNumber="424">Fastosarion hannianus</emphasis>
</taxonomicName>
is genetically very distinct from its nearest relatives (
<taxonomicName id="4C5AE0D5FF90B149539C9E6FF576FD63" box="[166,257,638,664]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="species" species="alyssa">
<emphasis id="B92E4744FF90B149539C9E6FF576FD63" box="[166,257,638,664]" italics="true" pageId="73" pageNumber="424">F. alyssa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5AE0D5FF90B149520D9E6FF5DAFD63" authorityName="Hyman &amp; Köhler" authorityYear="2019" box="[311,429,638,664]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="species" species="insularis">
<emphasis id="B92E4744FF90B149520D9E6FF5DAFD63" box="[311,429,638,664]" italics="true" pageId="73" pageNumber="424">F. insularis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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