321 lines
37 KiB
XML
321 lines
37 KiB
XML
<document id="F154B64CDB15DC8638BD703288F232FE" ID-DOI="10.1163/18759866-20191416" ID-ISSN="1875-9866" ID-Zenodo-Dep="8343061" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1694662606570" checkinUser="felipe" docAuthor="Hyman, Isabel T. & Köhler, Frank" docDate="2019" docId="03F32A40FF98B14451BB9D4FF4AEFC4C" docLanguage="en" docName="ContribZool.88.4.351-451.pdf" docOrigin="Contributions to Zoology 88 (4)" docSource="http://dx.doi.org/10.1163/18759866-20191416" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.2:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="2" docTitle="Fastosarion deensis Hyman & Köhler 2019, sp. nov." docType="treatment" docVersion="5" lastPageNumber="419" masterDocId="FFCA5238FFD9B100533A9C10F477FFFB" masterDocTitle="Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)" masterLastPageNumber="451" masterPageNumber="351" pageNumber="416" updateTime="1695298548213" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CDC0531EED83FAC19588895044C40044">Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)</mods:title>
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<mods:namePart id="608526ED1A94C4F7CD0EBF51B603F0BF">Hyman, Isabel T.</mods:namePart>
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<mods:namePart id="AEC47E438DF6DB16536E6C565B345E01">Köhler, Frank</mods:namePart>
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<mods:typeOfResource id="0D196B92CB66933252466ED3DDB55CF1">text</mods:typeOfResource>
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<mods:title id="560016E29543FDC50BE3E620B8BAF4E4">Contributions to Zoology</mods:title>
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<mods:part id="C91F6454888B4D86F8E7B28AE660E040">
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<mods:date id="562263702C1B95E842DDD573E61E33D3">2019</mods:date>
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<mods:number id="C2D7877A0967A3D775F05F20CA554E70">2019-10-03</mods:number>
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<mods:number id="1844E45A335DB808915ECE1C8C9AA082">88</mods:number>
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<mods:url id="FC8A5F0AD382714535A39316C4849BFE">http://dx.doi.org/10.1163/18759866-20191416</mods:url>
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<mods:classification id="8617C4407754F23285672D89E3566D21">journal article</mods:classification>
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<mods:identifier id="66C202E03CE55F7B5B856B29B05D660E" type="DOI">10.1163/18759866-20191416</mods:identifier>
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<treatment id="03F32A40FF98B14451BB9D4FF4AEFC4C" LSID="urn:lsid:plazi:treatment:03F32A40FF98B14451BB9D4FF4AEFC4C" httpUri="http://treatment.plazi.org/id/03F32A40FF98B14451BB9D4FF4AEFC4C" lastPageId="68" lastPageNumber="419" pageId="65" pageNumber="416">
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<subSubSection id="C340C8DDFF98B14151BB9D4FF7CAFE82" box="[641,957,350,377]" pageId="65" pageNumber="416" type="nomenclature">
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<paragraph id="8BE59B56FF98B14151BB9D4FF7CAFE82" blockId="65.[641,957,350,377]" box="[641,957,350,377]" pageId="65" pageNumber="416">
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<heading id="D0AD2C3AFF98B14151BB9D4FF7CAFE82" box="[641,957,350,377]" fontSize="11" level="2" pageId="65" pageNumber="416" reason="2">
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<emphasis id="B92E4744FF98B14151BB9D4FF7CAFE82" bold="true" box="[641,957,350,377]" pageId="65" pageNumber="416">
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<taxonomicName id="4C5AE0D5FF98B14151BB9D4FF713FE82" ID-CoL="7RBR8" ID-ENA="2658016" authority="Hyman & Köhler, 2019" authorityName="Hyman & Köhler" authorityYear="2019" box="[641,868,350,377]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="65" pageNumber="416" phylum="Mollusca" rank="species" species="deensis" status="sp. nov.">
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<emphasis id="B92E4744FF98B14151BB9D4FF713FE82" bold="true" box="[641,868,350,377]" italics="true" pageId="65" pageNumber="416">Fastosarion deensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21DFA3FFF98B14150539D4FF7CAFE82" box="[873,957,351,377]" pageId="65" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C340C8DDFF98B14451BC9D93F4AEFC4C" lastPageId="68" lastPageNumber="419" pageId="65" pageNumber="416" type="description">
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<paragraph id="8BE59B56FF98B14151BC9D93F7C6FE66" blockId="65.[646,945,387,413]" box="[646,945,387,413]" pageId="65" pageNumber="416">
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<figureCitation id="136187D3FF98B14151BC9D93F6A1FE66" box="[646,726,387,413]" captionStart="FIGURE 18" captionStartId="66.[129,202,1582,1603]" captionTargetBox="[129,1187,188,1550]" captionTargetId="figure-8@66.[129,1187,188,1550]" captionTargetPageId="66" captionText="FIGURE 18 Distribution of Clade B species. █ = F.alyssa, ▼ = F.deensis, × = F.ephelis, ♢ = F.hannianus, ▲ = F. insularis, ◻ = F.katatonos, ∆ = F.longimentula, ● = F.mcdonaldi, ♠ = F.minerva, ◆ = F.papillosus, ♣ = F.pustulosus, ♥ = F.rowani, Ǫ = F.sarina, ▽ = F.tuljun." figureDoi="http://doi.org/10.5281/zenodo.8343097" httpUri="https://zenodo.org/record/8343097/files/figure.png" pageId="65" pageNumber="416">Figs. 18</figureCitation>
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,
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<figureCitation id="136187D3FF98B14151DA9D93F770FE66" box="[736,775,387,413]" captionStart="FIGURE 19" captionStartId="67.[98,171,1480,1501]" captionTargetBox="[98,1104,171,1448]" captionTargetId="figure-115@67.[98,1104,170,1448]" captionTargetPageId="67" captionText="FIGURE 19 Shells of Clade B species. A. Fastosarion alyssa (type locality of D.peterbrocki), NMV F230748. B. Fastosarion deensis, MO85834 (holotype). C. Fastosarion ephelis, MO85835 (holotype). D. Fastosarion hannianus, MO56684.E. Fastosarion insularis sp. nov., MO76354. F. Fastosarion katatonos, MO85837 (holotype). G.Fastosarion longimentula, MO65313.H. Fastosarion mcdonaldi, MO50434.I. Fastosarion minerva, MO78358. J.Fastosarion papillosus, MO78896 (Mt Superbus).K.Fastosarion papillosus, MO62397 (Kroombit Tops). L.Fastosarion pustulosus, MO78894 (Kalpowar SF).M. Fastosarion rowani, MO85839 (holotype). N. Fastosarion sarina, MO85840 (holotype).O. Fastosarion tujun (Denham Range), MO76081 (holotype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8343099" httpUri="https://zenodo.org/record/8343099/files/figure.png" pageId="65" pageNumber="416">19B</figureCitation>
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,
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<figureCitation id="136187D3FF98B141502B9D93F72AFE66" box="[785,861,387,413]" captionStart="FIGURE 22" captionStartId="71.[98,171,1710,1731]" captionTargetBox="[98,887,184,1677]" captionTargetId="figure-39@71.[98,887,184,1677]" captionTargetPageId="71" captionText="FIGURE 22 Spermatophores of Clade B species. A-B. F.alyssa, MO77158.C-D.F.deensis, MO85834.E-F. F.hannianus, MO72277.G-H.F.mcdonaldi, MO50434.I-J. F.minerva, MO78358." figureDoi="http://doi.org/10.5281/zenodo.8343105" httpUri="https://zenodo.org/record/8343105/files/figure.png" pageId="65" pageNumber="416">22C–D</figureCitation>
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,
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<figureCitation id="136187D3FF98B141505D9D93F7C6FE66" box="[871,945,387,413]" captionStart="FIGURE 23" captionStartId="72.[129,202,1301,1322]" captionTargetBox="[129,1187,184,1243]" captionTargetId="figure-144@72.[129,1187,184,1243]" captionTargetPageId="72" captionText="FIGURE 23 Genitalia.A–C.F.deensis, MO85834, Mt Morgan. D-E.F.ephelis, MO85835, Broken River.A, D. Reproductive system.Arrows show the junction of the penis and epiphallus.B, E.Penis interior.F.Spermatophore.Scale bars 1 mm (E), 2 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.8343107" httpUri="https://zenodo.org/record/8343107/files/figure.png" pageId="65" pageNumber="416">23A–C</figureCitation>
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</paragraph>
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<paragraph id="8BE59B56FF98B14151BA9DDAF68FFE1F" blockId="65.[640,1157,458,556]" box="[640,760,458,484]" pageId="65" pageNumber="416">Etymology</paragraph>
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<paragraph id="8BE59B56FF98B14151BC9DFFF772FDD7" blockId="65.[640,1157,458,556]" pageId="65" pageNumber="416">
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For Dee Range, the
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<typeStatus id="54E125F4FF98B14150459DFFF7C7FDF2" box="[895,944,495,521]" pageId="65" pageNumber="416">type</typeStatus>
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locality; noun in apposition.
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</paragraph>
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<paragraph id="8BE59B56FF98B14151BA9E4AF723FD8F" blockId="65.[640,1157,602,880]" box="[640,852,602,628]" pageId="65" pageNumber="416">Material examined</paragraph>
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<paragraph id="8BE59B56FF98B14151BC9E6FF736FCFE" blockId="65.[640,1157,602,880]" pageId="65" pageNumber="416">
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<emphasis id="B92E4744FF98B14151BC9E6FF6BCFD63" box="[646,715,639,664]" italics="true" pageId="65" pageNumber="416">Types:</emphasis>
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<materialsCitation id="3B32910BFF98B14151ED9E6EF736FCFE" collectingDate="1989-06-25" collectionCode="QM" collectorName="J. Stanisic & D. Potter & J. Chaseling" latitude="-23.633333" location="Dee Ra" longLatPrecision="1254" longitude="150.45" municipality="Northern Spur" pageId="65" pageNumber="416" specimenCode="MO85834" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54E125F4FF98B14151ED9E6EF735FD63" box="[727,834,638,664]" pageId="65" pageNumber="416">Holotype</typeStatus>
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:
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<collectionCode id="ED4B0393FF98B14150689E6EF70AFD63" box="[850,893,638,664]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="65" pageNumber="416" type="Museum">QM</collectionCode>
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<specimenCode id="DBFC332DFF98B14150B39E6EF783FD62" box="[905,1012,638,665]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="65" pageNumber="416" type="Museum">MO85834</specimenCode>
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(
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<location id="8E85CD8DFF98B14157329E6EF015FD62" LSID="urn:lsid:plazi:treatment:03F32A40FF98B14451BB9D4FF4AEFC4C:8E85CD8DFF98B14157329E6EF015FD62" box="[1032,1122,638,665]" latitude="-23.633333" longLatPrecision="1254" longitude="150.45" municipality="Northern Spur" name="Dee Ra" pageId="65" pageNumber="416">Dee Ra</location>
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, c.
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<locationDeviation id="9A578274FF98B14151BC9EB2F71CFD46" box="[646,875,674,701]" location="Mt Morgan" pageId="65" pageNumber="416">
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<quantity id="4CA236B3FF98B14151BC9EB2F6A7FD47" box="[646,720,674,701]" metricMagnitude="3" metricUnit="m" metricValue="5.2" pageId="65" pageNumber="416" unit="km" value="5.2">5.2 km</quantity>
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E Mt Morgan
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</locationDeviation>
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,
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<collectingMunicipality id="6B81012CFF98B141504C9EB2F06FFD47" box="[886,1048,674,700]" pageId="65" pageNumber="416">Northern Spur</collectingMunicipality>
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,
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<geoCoordinate id="EE6EFD91FF98B14157189EB2F0F7FD47" box="[1058,1152,674,700]" degrees="23" direction="south" minutes="38" orientation="latitude" pageId="65" pageNumber="416" precision="925" value="-23.633333">23° 38' S</geoCoordinate>
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,
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<geoCoordinate id="EE6EFD91FF98B14151BD9ED6F685FD1B" box="[647,754,710,737]" degrees="150" direction="east" minutes="27" orientation="longitude" pageId="65" pageNumber="416" precision="925" value="150.45">150° 27' E</geoCoordinate>
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, coll.
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<date id="FFE4BD96FF98B141500E9ED6F7C2FD1B" box="[820,949,710,737]" pageId="65" pageNumber="416" value="1989-06-25">
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<collectingDate id="EFA0447EFF98B141500E9ED6F7C2FD1B" box="[820,949,710,737]" pageId="65" pageNumber="416" value="1989-06-25">25 Jun 1989</collectingDate>
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</date>
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,
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<collectorName id="26AFFE80FF98B14150FA9ED6F059FD1B" box="[960,1070,710,736]" pageId="65" pageNumber="416">J. Stanisic</collectorName>
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,
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<collectorName id="26AFFE80FF98B14157039ED6F6D2FCFF" pageId="65" pageNumber="416">D. Potter</collectorName>
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,
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<collectorName id="26AFFE80FF98B14151959EFAF742FCFE" box="[687,821,746,773]" pageId="65" pageNumber="416">J. Chaseling</collectorName>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BE59B56FF98B141519F9F1EF68DFCB6" blockId="65.[640,1157,602,880]" pageId="65" pageNumber="416">
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<materialsCitation id="3B32910BFF98B141519F9F1EF681FCB6" collectingDate="1989-06-25" collectionCode="QM" collectorName="J. Stanisic & D. Potter & J. Chaseling" latitude="-23.633333" location="Dee Ra" longLatPrecision="1254" longitude="150.45" municipality="Northern Spur" pageId="65" pageNumber="416" specimenCode="MO 21578" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54E125F4FF98B141519F9F1EF763FCD3" box="[677,788,782,808]" pageId="65" pageNumber="416">Paratypes</typeStatus>
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:
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<collectionCode id="ED4B0393FF98B141501E9F1EF738FCD3" box="[804,847,782,808]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="65" pageNumber="416" type="Museum">QM</collectionCode>
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<specimenCode id="DBFC332DFF98B14150659F1EF7A6FCD3" box="[863,977,781,808]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="65" pageNumber="416" type="Museum">MO 21578</specimenCode>
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(same data as holotype)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BE59B56FF98B141519F9F46F798FC8B" blockId="65.[640,1157,602,880]" box="[677,1007,853,880]" pageId="65" pageNumber="416">
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<emphasis id="B92E4744FF98B141519F9F46F707FC94" box="[677,880,853,879]" italics="true" pageId="65" pageNumber="416">Non-type material:</emphasis>
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See
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<tableCitation id="C6D8AEEDFF98B14150989F45F79CFC8B" box="[930,1003,853,880]" captionStart="TABLE 1" captionStartId="3.[56,116,402,428]" captionTargetBox="[56,1638,477,1528]" captionTargetPageId="3" captionText="TABLE 1 Material examined. Abbreviations: AM, Australian Museum, Sydney; H, holotype; NMV,Museums Victoria, Melbourne; P, Paratype; QM, Queensland Museum, Brisbane; S, syntype; NHRM, Naturhistoriska Riksmuseet, Stockholm, Sweden" httpUri="http://table.plazi.org/id/DF25CBDEFFDAB10353029D82F71DFE37" pageId="65" pageNumber="416" tableUuid="DF25CBDEFFDAB10353029D82F71DFE37">table 1</tableCitation>
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.
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</paragraph>
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<paragraph id="8BE59B56FF98B14151BA9F8DF69AFC4C" blockId="65.[640,1219,925,1810]" box="[640,749,925,951]" pageId="65" pageNumber="416">Diagnosis</paragraph>
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<paragraph id="8BE59B56FF98B14151BC9FD1F02FFAE5" blockId="65.[640,1219,925,1810]" pageId="65" pageNumber="416">
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<emphasis id="B92E4744FF98B14151BC9FD1F702FC20" box="[646,885,961,987]" italics="true" pageId="65" pageNumber="416">External morphology</emphasis>
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: Shell (fig. 19B) medium-sized (
|
||
<quantity id="4CA236B3FF98B14151C79FF5F7E3FBFB" box="[765,916,997,1024]" metricMagnitude="-2" metricUnit="m" metricValue="1.225" metricValueMax="1.28" metricValueMin="1.17" pageId="65" pageNumber="416" unit="mm" value="12.25" valueMax="12.8" valueMin="11.7">11.7–12.8 mm</quantity>
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||
), golden amber, very transparent, 4.0 whorls, subglobose with a low spire. Body
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<quantity id="4CA236B3FF98B1415037983DF7F1FBBC" box="[781,902,1069,1096]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="65" pageNumber="416" unit="mm" value="24.5" valueMax="25.0" valueMin="24.0">24–25 mm</quantity>
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long, colour (in ethanol) beige, darker grey on tail and head. Mantle lobes large, finely pustulose, shell lappets moderately large, finely pustulose, each with several rows of larger unpigmented warts; some smaller pustules have brown pigmentation. Tail not keeled, slime wrinkles weak.
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</paragraph>
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<paragraph id="8BE59B56FF98B144519F9938F582FF12" blockId="65.[640,1219,925,1810]" lastBlockId="68.[129,639,171,234]" lastPageId="68" lastPageNumber="419" pageId="65" pageNumber="416">
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<emphasis id="B92E4744FF98B141519F9938F71DFAB9" box="[677,874,1320,1346]" italics="true" pageId="65" pageNumber="416">Genital anatomy:</emphasis>
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Genitalia (figs. 22C–D, 23A–C) with moderately short vagina; bursa copulatrix moderate length; duct broad, long, bursa small, oval shaped. Penis moderate size, cylindrical, inner penial wall sculptured with coarse pustules arranged in irregular transverse rows, penis interior with one proximal penial stimulator; 55% of penis contained in penial tunica. Epiphallus approx. same length as penis, entering through simple pore; epiphallus 2 longer than epiphallus 1; epiphallic caecum short, hooked; flagellum moderately long, slender, coiled. Spermatophore with a single, finely toothed, spiraling ridge only on terminal half of tail-pipe.
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</paragraph>
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<caption id="DF25CBDEFF9BB14253BB9A3EF69DF985" ID-DOI="http://doi.org/10.5281/zenodo.8343097" ID-Zenodo-Dep="8343097" httpUri="https://zenodo.org/record/8343097/files/figure.png" pageId="66" pageNumber="417" startId="66.[129,202,1582,1603]" targetBox="[129,1187,188,1550]" targetPageId="66" targetType="figure">
|
||
<paragraph id="8BE59B56FF9BB14253BB9A3EF69DF985" blockId="66.[129,1173,1581,1667]" pageId="66" pageNumber="417">
|
||
FIGURE 18 Distribution of Clade B species. █ =
|
||
<taxonomicName id="4C5AE0D5FF9BB142517D9A3EF6E6F9B8" box="[583,657,1582,1603]" class="Gastropoda" family="Helicarionidae" genus="Dimidarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="alyssa">
|
||
<emphasis id="B92E4744FF9BB142517D9A3EF6E6F9B8" box="[583,657,1582,1603]" italics="true" pageId="66" pageNumber="417">F. alyssa</emphasis>
|
||
</taxonomicName>
|
||
, ▼ =
|
||
<taxonomicName id="4C5AE0D5FF9BB14251F09A3EF769F9B8" authorityName="Hyman & Köhler" authorityYear="2019" box="[714,798,1582,1603]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="deensis">
|
||
<emphasis id="B92E4744FF9BB14251F09A3EF769F9B8" box="[714,798,1582,1603]" italics="true" pageId="66" pageNumber="417">F.deensis</emphasis>
|
||
</taxonomicName>
|
||
, × =
|
||
<taxonomicName id="4C5AE0D5FF9BB14250739A3EF7EDF9B8" authorityName="Hyman & Köhler" authorityYear="2019" box="[841,922,1582,1603]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="ephelis">
|
||
<emphasis id="B92E4744FF9BB14250739A3EF7EDF9B8" box="[841,922,1582,1603]" italics="true" pageId="66" pageNumber="417">F.ephelis</emphasis>
|
||
</taxonomicName>
|
||
, ♢ =
|
||
<taxonomicName id="4C5AE0D5FF9BB14250F39A3EF04AF9B8" authorityName="Stanisic" authorityYear="2010" box="[969,1085,1582,1603]" class="Gastropoda" family="Helicarionidae" genus="Hymanarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="hannianus">
|
||
<emphasis id="B92E4744FF9BB14250F39A3EF04AF9B8" box="[969,1085,1582,1603]" italics="true" pageId="66" pageNumber="417">F.hannianus</emphasis>
|
||
</taxonomicName>
|
||
, ▲ =
|
||
<taxonomicName id="4C5AE0D5FF9BB14257489A3EF530F99B" authorityName="Hyman & Köhler" authorityYear="2019" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="insularis">
|
||
<emphasis id="B92E4744FF9BB14257489A3EF530F99B" italics="true" pageId="66" pageNumber="417">F. insularis</emphasis>
|
||
</taxonomicName>
|
||
, ◻ =
|
||
<taxonomicName id="4C5AE0D5FF9BB14252429A5CF594F99B" authorityName="Hyman & Köhler" authorityYear="2019" box="[376,483,1611,1633]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="katatonos">
|
||
<emphasis id="B92E4744FF9BB14252429A5CF594F99B" box="[376,483,1611,1633]" italics="true" pageId="66" pageNumber="417">F.katatonos</emphasis>
|
||
</taxonomicName>
|
||
, ∆ =
|
||
<taxonomicName id="4C5AE0D5FF9BB14251359A5CF6EDF99B" authorityName="Hyman & Köhler" authorityYear="2019" box="[527,666,1611,1633]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="longimentula">
|
||
<emphasis id="B92E4744FF9BB14251359A5CF6EDF99B" box="[527,666,1611,1633]" italics="true" pageId="66" pageNumber="417">F.longimentula</emphasis>
|
||
</taxonomicName>
|
||
, ● =
|
||
<taxonomicName id="4C5AE0D5FF9BB14251F39A5CF74BF99B" authorityName="Stanisic" authorityYear="1993" box="[713,828,1611,1633]" class="Gastropoda" family="Helicarionidae" genus="Eungarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="mcdonaldi">
|
||
<emphasis id="B92E4744FF9BB14251F39A5CF74BF99B" box="[713,828,1611,1633]" italics="true" pageId="66" pageNumber="417">F. mcdonaldi</emphasis>
|
||
</taxonomicName>
|
||
, ♠ =
|
||
<taxonomicName id="4C5AE0D5FF9BB14250519A5CF7BFF99A" box="[875,968,1612,1633]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="minerva">
|
||
<emphasis id="B92E4744FF9BB14250519A5CF7BFF99A" box="[875,968,1612,1633]" italics="true" pageId="66" pageNumber="417">F. minerva</emphasis>
|
||
</taxonomicName>
|
||
, ◆ =
|
||
<taxonomicName id="4C5AE0D5FF9BB142573A9A5CF019F99B" box="[1024,1134,1611,1633]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="papillosus">
|
||
<emphasis id="B92E4744FF9BB142573A9A5CF019F99B" box="[1024,1134,1611,1633]" italics="true" pageId="66" pageNumber="417">F.papillosus</emphasis>
|
||
</taxonomicName>
|
||
, ♣ =
|
||
<taxonomicName id="4C5AE0D5FF9BB14253C09A79F51BF986" box="[250,364,1640,1662]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="pustulosus">
|
||
<emphasis id="B92E4744FF9BB14253C09A79F51BF986" box="[250,364,1640,1662]" italics="true" pageId="66" pageNumber="417">F.pustulosus</emphasis>
|
||
</taxonomicName>
|
||
, ♥ =
|
||
<taxonomicName id="4C5AE0D5FF9BB14252A79A79F599F985" authorityName="Hyman & Köhler" authorityYear="2019" box="[413,494,1641,1662]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="rowani">
|
||
<emphasis id="B92E4744FF9BB14252A79A79F599F985" box="[413,494,1641,1662]" italics="true" pageId="66" pageNumber="417">F.rowani</emphasis>
|
||
</taxonomicName>
|
||
, Ǫ =
|
||
<taxonomicName id="4C5AE0D5FF9BB14251269A79F61FF985" box="[540,616,1641,1662]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="sarina">
|
||
<emphasis id="B92E4744FF9BB14251269A79F61FF985" box="[540,616,1641,1662]" italics="true" pageId="66" pageNumber="417">F.sarina</emphasis>
|
||
</taxonomicName>
|
||
, ▽ =
|
||
<taxonomicName id="4C5AE0D5FF9BB14251A79A79F692F986" authorityName="Hyman & Köhler" authorityYear="2019" box="[669,741,1640,1662]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="417" phylum="Mollusca" rank="species" species="tuljun">
|
||
<emphasis id="B92E4744FF9BB14251A79A79F692F986" box="[669,741,1640,1662]" italics="true" pageId="66" pageNumber="417">F. tuljun</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF25CBDEFF9AB143535899D8F60CF951" ID-DOI="http://doi.org/10.5281/zenodo.8343099" ID-Zenodo-Dep="8343099" httpUri="https://zenodo.org/record/8343099/files/figure.png" pageId="67" pageNumber="418" startId="67.[98,171,1480,1501]" targetBox="[98,1104,171,1448]" targetPageId="67" targetType="figure">
|
||
<paragraph id="8BE59B56FF9AB143535899D8F60CF951" blockId="67.[98,1145,1479,1707]" pageId="67" pageNumber="418">
|
||
FIGURE 19 Shells of Clade B species. A.
|
||
<taxonomicName id="4C5AE0D5FF9AB14352E699D8F608FA27" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[476,639,1479,1501]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="alyssa">
|
||
<emphasis id="B92E4744FF9AB14352E699D8F608FA27" box="[476,639,1479,1501]" italics="true" pageId="67" pageNumber="418">Fastosarion alyssa</emphasis>
|
||
</taxonomicName>
|
||
(type locality of
|
||
<taxonomicName id="4C5AE0D5FF9AB143502399D8F7E3FA27" box="[793,916,1479,1501]" class="Gastropoda" family="Helicarionidae" genus="Dimidarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="peterbrocki">
|
||
<emphasis id="B92E4744FF9AB143502399D8F7E3FA27" box="[793,916,1479,1501]" italics="true" pageId="67" pageNumber="418">D. peterbrocki</emphasis>
|
||
</taxonomicName>
|
||
), NMV F230748. B.
|
||
<taxonomicName id="4C5AE0D5FF9AB14353E199F5F5C3FA00" authority=", MO" authorityName="MO" box="[219,436,1509,1531]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="deensis">
|
||
<emphasis id="B92E4744FF9AB14353E199F5F5FFFA01" box="[219,392,1509,1530]" italics="true" pageId="67" pageNumber="418">Fastosarion deensis</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
85834 (holotype). C.
|
||
<taxonomicName id="4C5AE0D5FF9AB143514B99F5F730FA00" authority=", MO" authorityName="MO" box="[625,839,1509,1531]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="ephelis">
|
||
<emphasis id="B92E4744FF9AB143514B99F5F76CFA01" box="[625,795,1509,1530]" italics="true" pageId="67" pageNumber="418">Fastosarion ephelis</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
85835 (holotype). D.
|
||
<taxonomicName id="4C5AE0D5FF9AB143573E99F5F51EF9E3" authority=", MO" authorityName="MO" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="hannianus">
|
||
<emphasis id="B92E4744FF9AB143573E99F5F54BF9EC" italics="true" pageId="67" pageNumber="418">Fastosarion hannianus</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
56684. E.
|
||
<taxonomicName id="4C5AE0D5FF9AB14352849A13F600F9EC" authority="Hyman & Köhler, 2019" authorityName="Hyman & Köhler" authorityYear="2019" box="[446,631,1538,1560]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="insularis" status="sp. nov.">
|
||
<emphasis id="B92E4744FF9AB14352849A13F600F9EC" box="[446,631,1538,1560]" italics="true" pageId="67" pageNumber="418">Fastosarion insularis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DFA3FFF9AB14351479A13F6B5F9E3" box="[637,706,1539,1560]" pageId="67" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, MO76354. F.
|
||
<taxonomicName id="4C5AE0D5FF9AB14350049A13F059F9E3" authority=", MO" authorityName="MO" box="[830,1070,1538,1560]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="katatonos">
|
||
<emphasis id="B92E4744FF9AB14350049A13F075F9EC" box="[830,1026,1538,1560]" italics="true" pageId="67" pageNumber="418">Fastosarion katatonos</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
85837 (holotype). G.
|
||
<taxonomicName id="4C5AE0D5FF9AB14352649A30F619F9CE" authority=", MO" authorityName="MO" box="[350,622,1567,1589]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="longimentula">
|
||
<emphasis id="B92E4744FF9AB14352649A30F634F9CF" box="[350,579,1567,1589]" italics="true" pageId="67" pageNumber="418">Fastosarion longimentula</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
65313. H.
|
||
<taxonomicName id="4C5AE0D5FF9AB14351F99A30F7CCF9CE" authority=", MO" authorityName="MO" box="[707,955,1567,1589]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="mcdonaldi">
|
||
<emphasis id="B92E4744FF9AB14351F99A30F7F8F9CF" box="[707,911,1567,1589]" italics="true" pageId="67" pageNumber="418">Fastosarion mcdonaldi</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
50434. I.
|
||
<taxonomicName id="4C5AE0D5FF9AB14357319A30F50CF9A8" authority=", MO" authorityName="MO" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="minerva">
|
||
<emphasis id="B92E4744FF9AB14357319A30F539F9A9" italics="true" pageId="67" pageNumber="418">Fastosarion minerva</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
78358. J.
|
||
<taxonomicName id="4C5AE0D5FF9AB14352FD9A2DF6CCF9A8" authority=", MO" authorityName="MO" box="[455,699,1597,1619]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="papillosus">
|
||
<emphasis id="B92E4744FF9AB14352FD9A2DF6F8F9A9" box="[455,655,1597,1618]" italics="true" pageId="67" pageNumber="418">Fastosarion papillosus</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
78896 (
|
||
<taxonomicName id="4C5AE0D5FF9AB14351C69A2DF701F9A8" box="[764,886,1597,1619]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="superbus">Mt Superbus</taxonomicName>
|
||
). K.
|
||
<taxonomicName id="4C5AE0D5FF9AB14350A69A2DF48AF98B" authority=", MO" authorityName="MO" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="papillosus">
|
||
<emphasis id="B92E4744FF9AB14350A69A2DF014F9A9" box="[924,1123,1597,1618]" italics="true" pageId="67" pageNumber="418">Fastosarion papillosus</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
62397 (Kroombit Tops). L.
|
||
<taxonomicName id="4C5AE0D5FF9AB14352D49A4BF692F994" authority=", MO" authorityName="MO" box="[494,741,1626,1648]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="pustulosus">
|
||
<emphasis id="B92E4744FF9AB14352D49A4BF6CEF994" box="[494,697,1626,1648]" italics="true" pageId="67" pageNumber="418">Fastosarion pustulosus</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
78894 (Kalpowar SF). M.
|
||
<taxonomicName id="4C5AE0D5FF9AB14350F09A4BF489F976" authority=", MO" authorityName="MO" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="rowani">
|
||
<emphasis id="B92E4744FF9AB14350F09A4BF003F98B" box="[970,1140,1627,1648]" italics="true" pageId="67" pageNumber="418">Fastosarion rowani</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
85839 (holotype). N.
|
||
<taxonomicName id="4C5AE0D5FF9AB14352819A68F6FAF976" authority=", MO" authorityName="MO" box="[443,653,1655,1677]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="sarina">
|
||
<emphasis id="B92E4744FF9AB14352819A68F617F976" box="[443,608,1656,1677]" italics="true" pageId="67" pageNumber="418">Fastosarion sarina</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
85840 (holotype). O.
|
||
<taxonomicName id="4C5AE0D5FF9AB14350719A68F56AF951" authority="(Denham Range)" baseAuthorityName="Denham Range" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="67" pageNumber="418" phylum="Mollusca" rank="species" species="tujun">
|
||
<emphasis id="B92E4744FF9AB14350719A68F790F976" box="[843,999,1656,1677]" italics="true" pageId="67" pageNumber="418">Fastosarion tujun</emphasis>
|
||
(Denham Range)
|
||
</taxonomicName>
|
||
, MO76081 (holotype). Scale bar 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE59B56FF9DB14453419D07F4AAFECA" blockId="68.[123,639,279,951]" box="[123,221,279,305]" pageId="68" pageNumber="419">Remarks</paragraph>
|
||
<paragraph id="8BE59B56FF9DB14453BB9D2CF4AEFC4C" blockId="68.[123,639,279,951]" pageId="68" pageNumber="419">
|
||
<taxonomicName id="4C5AE0D5FF9DB14453BB9D2CF521FEAE" authorityName="Hyman & Köhler" authorityYear="2019" box="[129,342,315,341]" class="Gastropoda" family="Helicarionidae" genus="Fastosation" kingdom="Animalia" order="Stylommatophora" pageId="68" pageNumber="419" phylum="Mollusca" rank="species" species="deensis">
|
||
<emphasis id="B92E4744FF9DB14453BB9D2CF521FEAE" box="[129,342,315,341]" italics="true" pageId="68" pageNumber="419">Fastosation deensis</emphasis>
|
||
</taxonomicName>
|
||
(referred to above as ‘
|
||
<taxonomicName id="4C5AE0D5FF9DB144516F9D2BF570FE82" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="68" pageNumber="419" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
|
||
sp. Mt Morgan’) is known from a single population
|
||
<quantity id="4CA236B3FF9DB14452489D92F5D8FE66" box="[370,431,386,413]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="68" pageNumber="419" unit="km" value="5.0">5 km</quantity>
|
||
east of Mt Morgan (south of Rockhampton), living under logs in semi-evergreen vine thicket (fig. 18). This population was previously included in
|
||
<taxonomicName id="4C5AE0D5FF9DB14453BB9E03F565FDD7" box="[129,274,530,556]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="68" pageNumber="419" phylum="Mollusca" rank="species" species="pustulosus">
|
||
<emphasis id="B92E4744FF9DB14453BB9E03F565FDD7" box="[129,274,530,556]" italics="true" pageId="68" pageNumber="419">F. pustulosus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFCBE6A7FF9DB14452799E02F647FDD7" author="Stanisic, J. & Shea, M. & Potter, D. & Griffiths, O." box="[323,560,530,557]" pageId="68" pageNumber="419" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. & Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al. (2010)</bibRefCitation>
|
||
but is both genetically and anatomically distinct from this species, having a flatter shell and pustulose shell lappets with more numerous rows of warts as well as a distinct genital anatomy. Several other congeners live in close proximity, including
|
||
<taxonomicName id="4C5AE0D5FF9DB14452869EFBF4CBFCDC" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="68" pageNumber="419" phylum="Mollusca" rank="species" species="katatonos">
|
||
<emphasis id="B92E4744FF9DB14452869EFBF4CBFCDC" italics="true" pageId="68" pageNumber="419">Fastosarion katatonos</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5AE0D5FF9DB14453C79F1EF512FCDC" box="[253,357,782,807]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="68" pageNumber="419" phylum="Mollusca" rank="species" species="rowani">
|
||
<emphasis id="B92E4744FF9DB14453C79F1EF512FCDC" box="[253,357,782,807]" italics="true" pageId="68" pageNumber="419">F. rowani</emphasis>
|
||
</taxonomicName>
|
||
, but none are recorded in sympatry.
|
||
<taxonomicName id="4C5AE0D5FF9DB14452239F22F585FCB7" box="[281,498,818,844]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="68" pageNumber="419" phylum="Mollusca" rank="species" species="deensis">
|
||
<emphasis id="B92E4744FF9DB14452239F22F585FCB7" box="[281,498,818,844]" italics="true" pageId="68" pageNumber="419">Fastosarion deensis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from the sympatric
|
||
<taxonomicName id="4C5AE0D5FF9DB14452C09F46F608FC8B" baseAuthorityName="Ferussac" baseAuthorityYear="1821" box="[506,639,854,880]" class="Gastropoda" family="Helicarionidae" genus="Stanisicarion" kingdom="Animalia" order="Stylommatophora" pageId="68" pageNumber="419" phylum="Mollusca" rank="species" species="freycineti">
|
||
<emphasis id="B92E4744FF9DB14452C09F46F608FC8B" box="[506,639,854,880]" italics="true" pageId="68" pageNumber="419">S. freycineti</emphasis>
|
||
</taxonomicName>
|
||
by its smaller size and the warts on its shell lappets.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |